treatments-xml/data/03/84/87/038487AF1F03FFDB133FC44F7392FAFC.xml

<document id="4789BD35429BC07C72E2A5FB8AE200F5" ID-DOI="10.5281/zenodo.5401510" ID-ISSN="1638-9387" ID-Zenodo-Dep="5401510" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630334724433" checkinUser="marcus" docAuthor="Cumberlidge, Neil &amp; Sternberg, Richard V." docDate="2002" docId="038487AF1F03FFDB133FC44F7392FAFC" docLanguage="en" docName="Zoosystema.2002.24.1.41-79.pdf" docOrigin="Zoosystema 24 (1)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Boreas uglowi Cumberlidge &amp; Sternberg 2002, n. sp." docType="treatment" docVersion="4" lastPageNumber="77" masterDocId="FFBDFFD71F21FFFF120AC105702FFFF9" masterDocTitle="The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea)" masterLastPageNumber="79" masterPageNumber="41" pageNumber="75" updateTime="1699232587643" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="700F1A673DFD94D83198321191532B2C">The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea)</mods:title>
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<mods:name id="AFE770A1DC954D88C0EC859A007D9C5D" type="personal">
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<mods:namePart id="3CD91555BC2F0285E0ECD3282AF9784C">Cumberlidge, Neil</mods:namePart>
<mods:affiliation id="F7436EB4C12C78FB4282754ADFBB7477">Department of Biology, Northern Michigan University, Marquette, MI, 49855 (USA) ncumberl @ nmu. edu</mods:affiliation>
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<mods:namePart id="6DBD2C96CD1DB318473B7EC54FFB56AE">Sternberg, Richard V.</mods:namePart>
<mods:affiliation id="E816DA1309871BA647952A0F7CF4CDF0">NHB- 163, Department of Invertebrate Zoology, Smithsonian Institution, Washington D. C., 20560 (USA) Sternberg. Richard @ nmnh. si. edu.</mods:affiliation>
<mods:nameIdentifier id="0E0FAA356416C5CDF454B68B92138AA3" type="email">ichard@nmnh.si.edu</mods:nameIdentifier>
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<mods:title id="73B51C2A1BA339B8C5E39E98CFC0186B">Zoosystema</mods:title>
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<mods:date id="01B4E10AB415335C66395693DAA0622F">2002</mods:date>
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<heading id="D0DA81D51F03FFDD133FC44F7221FA9D" box="[309,526,1354,1381]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="75" reason="2">
<taxonomicName id="4C2D4D3A1F03FFDD133FC44F71E4FA9C" authority="Cumberlidge &amp; Sternberg, 2002" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[309,459,1354,1381]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="75" phylum="Arthropoda" rank="species" species="uglowi" status="sp. nov.">
<emphasis id="B959EAAB1F03FFDD133FC44F71E4FA9C" bold="true" box="[309,459,1354,1381]" italics="true" pageId="34" pageNumber="75">Boreas uglowi</emphasis>
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<taxonomicNameLabel id="A26A57D01F03FFDD13D9C44F7221FA9D" box="[467,526,1354,1380]" pageId="34" pageNumber="75" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C33765321F03FFDD129DC46E724DFA2C" pageId="34" pageNumber="75" type="description">
<paragraph id="8B9236B91F03FFDD129DC46E7283FA7C" blockId="34.[151,684,1354,1413]" box="[151,684,1386,1413]" pageId="34" pageNumber="75">
(
<figureCitation id="13162A3C1F03FFDD1294C46E70C3FA7C" box="[158,236,1387,1413]" captionStart="FIG" captionStartId="11.[151,162,1354,1371]" captionTargetBox="[192,1245,168,1311]" captionTargetId="figure-26@11.[187,1212,162,1321]" captionTargetPageId="11" captionText="FIG. 2. — Dorsal view of the carapace of adult males of Madagascan freshwater crabs (A-C based on Ng &amp; Takeda 1994); A, Skeleosophusa gollhardi (Bott, 1965), holotype (MNHN B-5165); B, S. prolixa Ng &amp; Takeda, 1994, holotype, (MRSN); C, S. eumeces Ng &amp; Takeda, 1994, holotype, (MRSN); D, Boreas uglowi n. gen., n. sp., holotype (MNHN B- 11292); E, Madagapotamon humberti Bott, 1965; F, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002, holotype, (AMNH 17833). Scale bar: A, 7.2 mm; B, 7.1 mm; C, 6.5 mm; D, 9 mm; E, 9.4 mm; F, 6 mm." figureDoi="http://doi.org/10.5281/zenodo.5401514" httpUri="https://zenodo.org/record/5401514/files/figure.png" pageId="34" pageNumber="75">Fig 2D</figureCitation>
; 
<figureCitation id="13162A3C1F03FFDD12FDC46E7134FA7C" box="[247,283,1387,1413]" captionStart="FIG" captionStartId="14.[151,162,1356,1373]" captionTargetBox="[156,1239,168,1322]" captionTargetId="figure-22@14.[146,1253,158,1327]" captionTargetPageId="14" captionText="FIG. 3. — Inferior view of the sternum of adult males of Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb., G, Madagapotamon humberti Bott, 1965; H, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; I, Boreas uglowi n. gen., n. sp. Scale bar: A, 11.7 mm; B, 17.2 mm; C, 15.9 mm; D, 14.5 mm; E, 10.2 mm; F, 7 mm; G, 9.6 mm; H, 6.4 mm; I, 9.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5401516" httpUri="https://zenodo.org/record/5401516/files/figure.png" pageId="34" pageNumber="75">3G</figureCitation>
; 
<figureCitation id="13162A3C1F03FFDD132CC46E7169FA7C" box="[294,326,1387,1413]" captionStart="FIG" captionStartId="16.[151,162,1071,1088]" captionTargetBox="[155,1241,171,1034]" captionTargetId="figure-282@16.[138,1257,160,1041]" captionTargetPageId="16" captionText="FIG. 4. — Left mandible of Madagascan freshwater crabs, frontal view showing the palp (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2000) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2000; H, Boreas uglowi n. gen., n. sp.; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Madagapotamon humberti Bott, 1965. Scale bar: A-I, K, L, 1.0 mm; J, 1.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5401518" httpUri="https://zenodo.org/record/5401518/files/figure.png" pageId="34" pageNumber="75">4L</figureCitation>
; 
<figureCitation id="13162A3C1F03FFDD1345C46E715AFA7C" box="[335,373,1387,1413]" captionStart="FIG" captionStartId="21.[151,162,1262,1279]" captionTargetBox="[248,1159,165,1192]" captionTargetId="figure-109@21.[228,1169,158,1197]" captionTargetPageId="21" captionText="FIG. 5. — Left third maxilliped of Madagascan freshwater crabs, frontal view (J, K based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.0 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401520" httpUri="https://zenodo.org/record/5401520/files/figure.png" pageId="34" pageNumber="75">5H</figureCitation>
; 
<figureCitation id="13162A3C1F03FFDD138AC46E71B0FA7C" box="[384,415,1387,1413]" captionStart="FIG" captionStartId="23.[151,162,768,785]" captionTargetBox="[258,1160,169,728]" captionTargetId="figure-337@23.[233,1163,157,745]" captionTargetPageId="23" captionText="FIG. 6. — Superior view of the carpus of the cheliped (pereiopod 1) of adult males of Madagascan freshwater crabs (J, K based on Ng &amp; Takeda 1994); A Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Malagasya antongilensis (Rathbun, 1905) n. comb.; F, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; G, Madagapotamon humberti Bott, 1965; H, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; I, Skeleosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp. Scale bar: A-D, 5.5 mm; E-L, 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5401522" httpUri="https://zenodo.org/record/5401522/files/figure.png" pageId="34" pageNumber="75">6L</figureCitation>
; 
<figureCitation id="13162A3C1F03FFDD13A3C46E71C1FA7D" box="[425,494,1386,1413]" captionStart="FIG" captionStartId="25.[151,162,1567,1584]" captionTargetBox="[244,1126,230,1497]" captionTargetId="figure-22@25.[232,1187,219,1521]" captionTargetPageId="25" captionText="FIG. 7. — Superior and inferior views of the merus of the cheliped (pereiopod 1) of adult males of Madagascan freshwater crabs; A, B, Hydrothelphusa agilis A. Milne-Edwards, 1872; C, D, H. madagascariensis (A. Milne-Edwards, 1872); E, F, H. bombetokensis (Rathbun, 1904); G, H, H. goudoti (H. Milne Edwards, 1853) n. comb.; I, J, Malagasya antongilensis (Rathbun, 1905) n. comb.; K, L, Madagapotamon humberti Bott, 1965; M, N, Skelosophusa gollhardi (Bott, 1965); O, P, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; Q, R, Boreas uglowi n. gen., n. sp. Scale bar: A-H, 5.0 mm; I-T, 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5401524" httpUri="https://zenodo.org/record/5401524/files/figure.png" pageId="34" pageNumber="75">7Q, R</figureCitation>
; 
<figureCitation id="13162A3C1F03FFDD13F2C46E7237FA7C" box="[504,536,1387,1413]" captionStart="FIG" captionStartId="29.[151,162,1545,1562]" captionTargetBox="[151,1247,251,1506]" captionTargetId="figure-43@29.[153,1248,238,1515]" captionTargetPageId="29" captionText="FIG. 8. — Last walking leg (pereiopod 5) of adult males of Madagascan freshwater crabs (H based on Ng &amp; Takeda 1994); A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853) n. comb.; E, Boreas uglowi n. gen., n. sp.; F, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; G, Madagapotamon humberti Bott, 1965; H, Skelosophusa prolixa Ng &amp; Takeda, 1994; I, Malagasya antongilensis (Rathbun, 1905) n. comb.; J, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb. Scale bar: A-D, 5.0 mm; E-L, 2.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5401526" httpUri="https://zenodo.org/record/5401526/files/figure.png" pageId="34" pageNumber="75">8E</figureCitation>
; 
<figureCitation id="13162A3C1F03FFDD1028C46E7241FA7D" box="[546,622,1386,1413]" captionStart="FIG" captionStartId="32.[151,162,1363,1380]" captionTargetBox="[149,1247,169,1337]" captionTargetId="figure-94@32.[141,1252,160,1342]" captionTargetPageId="32" captionText="FIG. 10. — Right first gonopod (dorsal and ventral views) of adult males of Madagascan freshwater crabs; A, B, Boreas uglowi n. gen., n. sp.; C, D, Malagasya antongilensis (Rathbun, 1905) n. comb.; E, F, Skeleosophusa gollhardi (Bott, 1965); G, H, Madagapotamon humberti Bott, 1965; I, J, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002. Scale bar: 4.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5401530" httpUri="https://zenodo.org/record/5401530/files/figure.png" pageId="34" pageNumber="75">10A, B</figureCitation>
; 
<figureCitation id="13162A3C1F03FFDD1073C46E728CFA7C" box="[633,675,1387,1413]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="34" pageNumber="75">11L</figureCitation>
)
</paragraph>
<paragraph id="8B9236B91F03FFDD129DC4A1724DFA2C" blockId="34.[151,684,1443,1493]" pageId="34" pageNumber="75">
<taxonomicName id="4C2D4D3A1F03FFDD129DC4A1719CFA45" baseAuthorityName="Nobili" baseAuthorityYear="1906" box="[151,435,1444,1468]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="75" phylum="Arthropoda" rank="species" species="ankaraharae">
<emphasis id="B959EAAB1F03FFDD129DC4A1719CFA45" box="[151,435,1444,1468]" italics="true" pageId="34" pageNumber="75">Madagapotamon ankaraharae</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFBC4B481F03FFDD13C4C4A6721BFA45" author="BOTT R." box="[462,564,1443,1468]" pageId="34" pageNumber="75" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott 1965</bibRefCitation>
(part): 347, 348, fig. 9d, e, pl. 5, figs 23-25 (non fig. 9a-c).
</paragraph>
</subSubSection>
<subSubSection id="C33765321F03FFDD129DC4E8744AFE7A" pageId="34" pageNumber="75" type="materials_examined">
<paragraph id="8B9236B91F03FFDD129DC4E87457FEA3" blockId="34.[151,684,1516,1668]" lastBlockId="34.[714,1248,169,584]" pageId="34" pageNumber="75">
MATERIAL EXAMINED. — 
<emphasis id="B959EAAB1F03FFDD13BBC4E97213F9FD" bold="true" box="[433,572,1516,1540]" pageId="34" pageNumber="75">
<collectingCountry id="F33A76291F03FFDD13BBC4E97217F9FD" box="[433,568,1516,1540]" name="Madagascar" pageId="34" pageNumber="75">Madagascar</collectingCountry>
.
</emphasis>
Nosy-Bé, Madagascar, 
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, 
<typeStatus id="5496881B1F03FFDD1367C70271ECF9E6" box="[365,451,1543,1567]" pageId="34" pageNumber="75" type="holotype">holotype</typeStatus>
cw 29.3, cl 21.3, ch 10, fw 
<quantity id="4CD59B5C1F03FFDD12B0C7257121F9CF" box="[186,270,1567,1592]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="34" pageNumber="75" unit="mm" value="8.5">8.5 mm</quantity>
(
<collectionCode id="ED3CAE7C1F03FFDD1315C72571BEF9C1" box="[287,401,1568,1592]" pageId="34" pageNumber="75">MNHN-B</collectionCode>
12292). — Diego-Suarez (now 
<collectingRegion id="49E9F85B1F03FFDD12D9C73C7160F9A8" box="[211,335,1593,1617]" country="Madagascar" name="Antsiranana" pageId="34" pageNumber="75">Antsiranana</collectingRegion>
), coll. J. Millot, 
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cw 24.3, cl 18.8, ch 8.6, fw 
<quantity id="4CD59B5C1F03FFDD136FC7567196F990" box="[357,441,1618,1643]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="34" pageNumber="75" unit="mm" value="7.6">7.6 mm</quantity>
(
<collectionCode id="ED3CAE7C1F03FFDD13C3C7567215F992" box="[457,570,1619,1643]" pageId="34" pageNumber="75">MNHN-B</collectionCode>
5028). — Nosy-Bé, Autumn 1927, coll. J. Millot, 
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, cw 
<quantity id="4CD59B5C1F03FFDD10E1C1AF7365FF39" box="[747,842,169,194]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="34" pageNumber="75" unit="mm" value="19.8">19.8 mm</quantity>
, broken in pieces (
<collectionCode id="ED3CAE7C1F03FFDD1602C1AF745BFF3B" box="[1032,1140,170,194]" pageId="34" pageNumber="75">MNHN-B</collectionCode>
5029). — Nosy-Bé, 
<date id="FF9310791F03FFDD1127C1C67356FF22" box="[813,889,195,219]" pageId="34" pageNumber="75" value="1922-10">X.1922</date>
, coll. J. Millot, 
<specimenCount id="9D2BFD301F03FFDD1612C1C67465FF22" box="[1048,1098,195,219]" pageId="34" pageNumber="75" type="generic">1 ad.</specimenCount>
cw 
<quantity id="4CD59B5C1F03FFDD169AC1C674F4FF23" box="[1168,1243,195,219]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="75" unit="mm" value="25.0">25 mm</quantity>
, carapace damaged (
<collectionCode id="ED3CAE7C1F03FFDD11AEC1D87436FF0C" box="[932,1049,221,245]" pageId="34" pageNumber="75">MNHN-B</collectionCode>
26147). — Nosy Komba, 
<quantity id="4CD59B5C1F03FFDD112FC1F37346FEF5" box="[805,873,245,270]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="75" unit="m" value="500.0">500 m</quantity>
altitude, on a path, 
<date id="FF9310791F03FFDD164EC1F37489FEF7" box="[1092,1190,246,270]" pageId="34" pageNumber="75" value="1922-12">XII.1922</date>
, coll. J. Millot, 
<specimenCount id="9D2BFD301F03FFDD1120C00A7383FEDE" box="[810,940,271,295]" pageId="34" pageNumber="75" type="subadult">1 subadult</specimenCount>
cw 22, cl 16, ch 8, fw 
<quantity id="4CD59B5C1F03FFDD1681C00A74F5FEDF" box="[1163,1242,271,295]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="34" pageNumber="75" unit="mm" value="6.5">6.5 mm</quantity>
, 1, broken cw 28.5, cl 
<quantity id="4CD59B5C1F03FFDD11D7C02C746CFEC6" box="[989,1091,296,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="34" pageNumber="75" unit="mm" value="19.5">19.5 mm</quantity>
, 
<specimenCount id="9D2BFD301F03FFDD165AC02C74A0FEB8" box="[1104,1167,297,321]" pageId="34" pageNumber="75" type="generic">1 juv.</specimenCount>
cw 18, cl 13.5, ch 6, fw 
<quantity id="4CD59B5C1F03FFDD1165C0477385FEA1" box="[879,938,321,346]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="75" unit="mm" value="5.0">5 mm</quantity>
(
<collectionCode id="ED3CAE7C1F03FFDD11BDC047740CFEA3" box="[951,1059,322,346]" pageId="34" pageNumber="75">MNHN-B</collectionCode>
12293).
</paragraph>
<paragraph id="8B9236B91F03FFDD10C0C06F744AFE7A" blockId="34.[714,1248,169,584]" box="[714,1125,362,387]" pageId="34" pageNumber="75">
TYPELOCALITY. — Nosy-Bé, 
<collectingCountry id="F33A76291F03FFDD11E0C06E744FFE7A" box="[1002,1120,363,387]" name="Madagascar" pageId="34" pageNumber="75">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F03FFDD10C0C097748CFE0E" pageId="34" pageNumber="75" type="etymology">
<paragraph id="8B9236B91F03FFDD10C0C097748CFE0E" blockId="34.[714,1248,169,584]" pageId="34" pageNumber="75">
ETYMOLOGY. — The species is named for Dr. Roger Frederick Uglow, Department of Applied Biology, University of Hull, 
<collectingCountry id="F33A76291F03FFDD1186C0C07390FE25" box="[908,959,453,476]" name="United Kingdom" pageId="34" pageNumber="75">U.K.</collectingCountry>
, in recognition of his prolific contributions to decapod crustacean physiology.
</paragraph>
</subSubSection>
<subSubSection id="C33765321F03FFDD10C0C30274E6FDE6" box="[714,1225,519,543]" pageId="34" pageNumber="75" type="distribution">
<paragraph id="8B9236B91F03FFDD10C0C30274E6FDE6" blockId="34.[714,1248,169,584]" box="[714,1225,519,543]" pageId="34" pageNumber="75">
DISTRIBUTION. — 
<collectingCountry id="F33A76291F03FFDD118EC30273D5FDE6" box="[900,1018,519,543]" name="Madagascar" pageId="34" pageNumber="75">Madagascar</collectingCountry>
, Nosy-Bé (
<figureCitation id="13162A3C1F03FFDD166CC3027492FDE6" box="[1126,1213,519,543]" captionStart="FIG" captionStartId="35.[151,162,1408,1425]" captionTargetBox="[170,1219,175,1376]" captionTargetId="figure-28@35.[157,1238,162,1394]" captionTargetPageId="35" captionText="FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; A, Hydrothelphusa agilis A. Milne-Edwards, 1872; B, H. madagascariensis (A. Milne-Edwards, 1872); C, H. bombetokensis (Rathbun, 1904); D, H. goudoti (H. Milne Edwards, 1853); E, Marojejy longimerus Cumberlidge, Boyko &amp; Harvey, 2002; F, Malagasya antongilensis (Rathbun, 1905) n. comb.; G, M. goodmani (Cumberlidge, Boyko &amp; Harvey, 2002) n. comb.; H, Madagapotamon humberti Bott, 1965; I, Skelosophusa gollhardi (Bott, 1965); J, S. prolixa Ng &amp; Takeda, 1994; K, S. eumeces Ng &amp; Takeda, 1994; L, Boreas uglowi n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="34" pageNumber="75">Fig. 11L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33765321F03FFDD10C0C32A73CEFDBE" box="[714,993,559,584]" pageId="34" pageNumber="75" type="diagnosis">
<paragraph id="8B9236B91F03FFDD10C0C32A73CEFDBE" blockId="34.[714,1248,169,584]" box="[714,993,559,584]" pageId="34" pageNumber="75">DIAGNOSIS. — As for genus.</paragraph>
</subSubSection>
<subSubSection id="C33765321F03FFDB10C0C38F7257FB9C" lastPageId="36" lastPageNumber="77" pageId="34" pageNumber="75" type="description">
<paragraph id="8B9236B91F03FFDD10C0C38F7374FD5B" blockId="34.[714,1248,650,1669]" box="[714,859,650,676]" pageId="34" pageNumber="75">DESCRIPTION</paragraph>
<paragraph id="8B9236B91F03FFDD10C0C3AE748AFABC" blockId="34.[714,1248,650,1669]" pageId="34" pageNumber="75">
Carapace outline transversely oval (wider than long, cw/fw = 3.2, cl/fw = 2.3), moderately arched (ch/fw = 1.1); epigastric, postorbital crests faint, not fused; postorbital crests not meeting anterolateral margins; exorbital, epibranchial teeth similar, both low, blunt, continuous with anterolateral margin; anterior regions of carapace surface smooth; anterolateral, posterolateral regions with short carinae; semi-circular, urogastric grooves deep; cervical grooves very short. Suborbital margin raised, faintly granulated. Front highly deflexed; frontal margin smooth, straight; frontal, posterior margins of carapace wide (cw 
<quantity id="4CD59B5C1F03FFDD1148C54E7387FB9D" box="[834,936,1098,1125]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="75" unit="mm" value="0.3">0.3 mm</quantity>
). Suborbital, subhepatic, regions of carapace sidewall smooth or with faint granules; pterygostomial region smooth with sparse short setae or a few small granules near third maxilliped. Carapace sidewall in three parts; distinct vertical sulcus on sidewall running from epibranchial tooth to longitudinal (epimeral) groove, dividing suborbital, hepatic regions.
</paragraph>
<paragraph id="8B9236B91F03FFDC10C0C44E716BF97C" blockId="34.[714,1248,650,1669]" lastBlockId="35.[151,684,1579,1669]" lastPageId="35" lastPageNumber="76" pageId="34" pageNumber="75">Mandibular palp two-segmented, terminal segment with flat basal thickening at junction between segments; ischium of third maxilliped smooth, vertical sulcus either faint or completely absent; exopod of third maxilliped medium length, reaching only proximal part of merus, exopod with long flagellum. Thoracic sternal sulcus s1/s2 not visible; sternal sulcus s2/s3 deep, horizontal; sternal sulcus s3/s4 reduced to two short side notches; anterior sternum smooth; episternal sulci s4/e4, s5/e5, s6/e6, s7/e7 smooth, no groove visible. Male abdomen slim, triangular.</paragraph>
<caption id="DF5266311F02FFDC129DC485746FFA1E" ID-DOI="http://doi.org/10.5281/zenodo.5401532" ID-Zenodo-Dep="5401532" httpUri="https://zenodo.org/record/5401532/files/figure.png" pageId="35" pageNumber="76" startId="35.[151,162,1408,1425]" targetBox="[170,1219,175,1376]" targetPageId="35">
<paragraph id="8B9236B91F02FFDC129DC485746FFA1E" blockId="35.[151,1249,1408,1511]" pageId="35" pageNumber="76">
FIG. 11. — Summaries of the known distribution of the Madagascan freshwater crabs; 
<emphasis id="B959EAAB1F02FFDC1179C48573AFFA68" bold="true" box="[883,896,1408,1425]" pageId="35" pageNumber="76">A</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC1181C48570ECFA5E" authority="A. Milne-Edwards, 1872" authorityName="A. Milne-Edwards" authorityYear="1872" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="B959EAAB1F02FFDC1181C4857411FA68" box="[907,1086,1408,1425]" italics="true" pageId="35" pageNumber="76">Hydrothelphusa agilis</emphasis>
A. Milne-Edwards, 1872
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC12C6C49370F6FA5E" bold="true" box="[204,217,1430,1447]" pageId="35" pageNumber="76">B</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC12EFC493725EFA5E" authority="(A. Milne-Edwards, 1872)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1872" box="[229,625,1430,1447]" class="Malacostraca" family="Potamidae" genus="Thelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B959EAAB1F02FFDC12EFC49371BCFA5E" box="[229,403,1430,1447]" italics="true" pageId="35" pageNumber="76">H. madagascariensis</emphasis>
(A. Milne-Edwards, 1872)
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC1070C49372A7FA5E" bold="true" box="[634,648,1430,1447]" pageId="35" pageNumber="76">C</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC109EC4937393FA5E" authority="(Rathbun, 1904)" baseAuthorityName="Rathbun" baseAuthorityYear="1904" box="[660,956,1430,1447]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="bombetokensis">
<emphasis id="B959EAAB1F02FFDC109EC4937303FA5E" box="[660,812,1430,1447]" italics="true" pageId="35" pageNumber="76">H. bombetokensis</emphasis>
(
<bibRefCitation id="EFBC4B481F02FFDC113DC4937398FA5E" author="RATHBUN M. J." box="[823,951,1430,1447]" pageId="35" pageNumber="76" pagination="255 - 312" refId="ref26930" refString="RATHBUN M. J. 1904. - Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle 6 (4): 255 - 312." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
)
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC11CFC49373FCFA5E" bold="true" box="[965,979,1430,1447]" pageId="35" pageNumber="76">D</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC11D5C49370E4FA45" authority="(H. Milne Edwards, 1853)" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Potamidae" genus="Thelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="goudoti">
<emphasis id="B959EAAB1F02FFDC11D5C4937417FA5E" box="[991,1080,1430,1447]" italics="true" pageId="35" pageNumber="76">H. goudoti</emphasis>
(H. 
<bibRefCitation id="EFBC4B481F02FFDC1657C49370E9FA45" author="MILNE EDWARDS H." pageId="35" pageNumber="76" pagination="20" refId="ref26442" refString="MILNE EDWARDS H. 1853. - Observations sur les affinites zoologiques et la classification naturelle des Crustaces. Annales des Sciences naturelles, Zoologie, Serie 3, 20: 163 - 28." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
)
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC12DFC4AE70CEFA45" bold="true" box="[213,225,1451,1468]" pageId="35" pageNumber="76">E</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC12E4C4AE72DCFA45" authority="Cumberlidge, Boyko &amp; Harvey, 2002" authorityName="Cumberlidge, Boyko &amp; Harvey" authorityYear="2002" box="[238,755,1451,1468]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="longimerus">
<emphasis id="B959EAAB1F02FFDC12E4C4AE718EFA45" box="[238,417,1451,1468]" italics="true" pageId="35" pageNumber="76">Marojejy longimerus</emphasis>
Cumberlidge, Boyko &amp; Harvey, 2002
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC10F7C4AE7327FA45" bold="true" box="[765,776,1451,1468]" pageId="35" pageNumber="76">F</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC111FC4AE74ACFA45" authority="(Rathbun, 1905)" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" baseAuthorityName="Rathbun" baseAuthorityYear="1905" box="[789,1155,1451,1468]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="antongilensis" status="comb. nov.">
<emphasis id="B959EAAB1F02FFDC111FC4AE73C2FA45" box="[789,1005,1451,1468]" italics="true" pageId="35" pageNumber="76">Malagasya antongilensis</emphasis>
(
<bibRefCitation id="EFBC4B481F02FFDC11F3C4AE7451FA45" author="RATHBUN M. J." box="[1017,1150,1451,1468]" pageId="35" pageNumber="76" pagination="159 - 322" refId="ref26961" refString="RATHBUN M. J. 1905. - Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle 7 (4): 159 - 322." type="journal article" year="1905">Rathbun, 1905</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F02FFDC1680C4AE74F1FA45" box="[1162,1246,1451,1468]" pageId="35" pageNumber="76" rank="species">n. comb.</taxonomicNameLabel>
; 
<emphasis id="B959EAAB1F02FFDC129DC4C5708AFA28" bold="true" box="[151,165,1472,1489]" pageId="35" pageNumber="76">G</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC12A5C4C57275FA28" authority="(Cumberlidge, Boyko &amp; Harvey, 2002)" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" baseAuthorityName="Cumberlidge, Boyko &amp; Harvey" baseAuthorityYear="2002" box="[175,602,1472,1489]" class="Malacostraca" family="Potamonautidae" genus="Malagasya" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="goodmani" status="comb. nov.">
<emphasis id="B959EAAB1F02FFDC12A5C4C57132FA28" box="[175,285,1472,1489]" italics="true" pageId="35" pageNumber="76">M. goodmani</emphasis>
(Cumberlidge, Boyko &amp; Harvey, 2002)
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F02FFDC106AC4C57281FA28" box="[608,686,1472,1489]" pageId="35" pageNumber="76" rank="species">n. comb.</taxonomicNameLabel>
; 
<emphasis id="B959EAAB1F02FFDC10BEC4C572EDFA28" bold="true" box="[692,706,1472,1489]" pageId="35" pageNumber="76">H</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC10C7C4C57428FA28" authority="Bott, 1965" authorityName="Bott" authorityYear="1965" box="[717,1031,1472,1489]" class="Malacostraca" family="Potamonautidae" genus="Madagapotamon" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="humberti">
<emphasis id="B959EAAB1F02FFDC10C7C4C57387FA28" box="[717,936,1472,1489]" italics="true" pageId="35" pageNumber="76">Madagapotamon humberti</emphasis>
<bibRefCitation id="EFBC4B481F02FFDC11A4C4C57428FA28" author="BOTT R." box="[942,1031,1472,1489]" pageId="35" pageNumber="76" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott, 1965</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC1605C4C5743BFA28" bold="true" box="[1039,1044,1472,1489]" pageId="35" pageNumber="76">I</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC162AC4C570D4FA1E" authority="(Bott, 1965)" baseAuthorityName="Bott" baseAuthorityYear="1965" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="gollhardi">
<emphasis id="B959EAAB1F02FFDC162AC4C574F1FA28" box="[1056,1246,1472,1489]" italics="true" pageId="35" pageNumber="76">Skelosophusa gollhardi</emphasis>
(
<bibRefCitation id="EFBC4B481F02FFDC1296C4D370DAFA1E" author="BOTT R." box="[156,245,1494,1511]" pageId="35" pageNumber="76" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott, 1965</bibRefCitation>
)
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC1308C4D37123FA1E" bold="true" box="[258,268,1494,1511]" pageId="35" pageNumber="76">J</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC131DC4D37224FA1E" authority="Ng &amp; Takeda, 1994" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[279,523,1494,1511]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="prolixa">
<emphasis id="B959EAAB1F02FFDC131DC4D3714BFA1E" box="[279,356,1494,1511]" italics="true" pageId="35" pageNumber="76">S. prolixa</emphasis>
<bibRefCitation id="EFBC4B481F02FFDC1363C4D37224FA1E" author="NG P. K. L. &amp; TAKEDA M." box="[361,523,1494,1511]" pageId="35" pageNumber="76" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda, 1994</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC1019C4D3720FFA1E" bold="true" box="[531,544,1494,1511]" pageId="35" pageNumber="76">K</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC1021C4D3731BFA1E" authority="Ng &amp; Takeda, 1994" authorityName="Ng &amp; Takeda" authorityYear="1994" box="[555,820,1494,1511]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="eumeces">
<emphasis id="B959EAAB1F02FFDC1021C4D372A2FA1E" box="[555,653,1494,1511]" italics="true" pageId="35" pageNumber="76">S. eumeces</emphasis>
<bibRefCitation id="EFBC4B481F02FFDC1098C4D3731BFA1E" author="NG P. K. L. &amp; TAKEDA M." box="[658,820,1494,1511]" pageId="35" pageNumber="76" pagination="161 - 172" refId="ref26560" refString="NG P. K. L. &amp; TAKEDA M. 1994. - Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bulletin of the National Science Museum, Tokyo, Series A (Zoology) 20 (4): 161 - 172." type="journal article" year="1994">Ng &amp; Takeda, 1994</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B959EAAB1F02FFDC1136C4D37368FA1E" bold="true" box="[828,839,1494,1511]" pageId="35" pageNumber="76">L</emphasis>
, 
<taxonomicName id="4C2D4D3A1F02FFDC1158C4D373E8FA1E" authority="Cumberlidge &amp; Sternberg, 2002" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[850,967,1494,1511]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="76" phylum="Arthropoda" rank="species" species="uglowi" status="n. gen., n. sp.">
<emphasis id="B959EAAB1F02FFDC1158C4D373E8FA1E" box="[850,967,1494,1511]" italics="true" pageId="35" pageNumber="76">Boreas uglowi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57D01F02FFDC11C7C4D3746FFA1E" box="[973,1088,1494,1511]" pageId="35" pageNumber="76" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B9236B91F02FFDB10C0C72E710EFE3C" blockId="35.[714,1247,1579,1669]" lastBlockId="36.[151,684,170,1125]" lastPageId="36" lastPageNumber="77" pageId="35" pageNumber="76">Gonopod 1 terminal article straight, slim, short (ratio length terminal article to subterminal segment 0.25), tip directed weakly outward, longitudinal groove clearly visible on ventral, superior sides, not visible on dorsal side; terminal article/ subterminal segment junction distinct, with narrow dorsal membrane; lateral, medial folds of ventral side of terminal article equal in height, width. Gonopod 2 longer than gonopod 1; gonopod 2 terminal article medium flagellum, ratio length gonopod 2 terminal article to subterminal segment 0.6.</paragraph>
<paragraph id="8B9236B91F05FFDB129DC0CE7257FB9C" blockId="36.[151,684,170,1125]" pageId="36" pageNumber="77">Dactylus of right, left chelipeds relatively narrow (one-quarter height of palm), upper margin smooth, finger of dactylus curved; slim interspace between closed fingers; finger of propodus onethird height of palm lower margin of propodus of cheliped slightly indented. Dactylus with series of small teeth interspersed by larger teeth; finger of propodus of right cheliped with large fused molar tooth. Anterior, posterior inferior margins of ischium of p1 smooth. Inferior margins of merus faintly granulated, distal margin with single low distal meral tooth; superior margin of merus with row of granules. First carpal tooth of carpus of cheliped large, wide, with pointed tip; second carpal tooth small. With pointed tip, followed by small granules. Anterior, posterior margins of propodi of p5 slightly serrated. Walking legs (p2- p5) normal length, neither elongated nor shortened, ratio length merus p5 to cw 0.3. P5 propodus short, broad, margins serrated; dactylus of p5 short, half length of dactylus of p2-p4.</paragraph>
</subSubSection>
<subSubSection id="C33765321F05FFDB129DC58F7392FAFC" pageId="36" pageNumber="77" type="discussion">
<paragraph id="8B9236B91F05FFDB129DC58F70D4FB5B" blockId="36.[151,684,1162,1669]" box="[151,251,1162,1188]" pageId="36" pageNumber="77">REMARKS</paragraph>
<paragraph id="8B9236B91F05FFDB129DC5AE74BBFDBC" blockId="36.[151,684,1162,1669]" lastBlockId="36.[714,1248,171,1285]" pageId="36" pageNumber="77">
The generic assignment and validity of 
<taxonomicName id="4C2D4D3A1F05FFDB1044C5AF71E2FB1C" authorityName="sensu Nobili" authorityYear="1906" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae" subGenus="Geothelphusa">
<emphasis id="B959EAAB1F05FFDB1044C5AF7284FB3A" box="[590,683,1194,1219]" italics="true" pageId="36" pageNumber="77">Potamon</emphasis>
(
<emphasis id="B959EAAB1F05FFDB12AAC5CE711EFB1C" box="[160,305,1227,1253]" italics="true" pageId="36" pageNumber="77">Geothelphusa</emphasis>
) 
<emphasis id="B959EAAB1F05FFDB134EC5CE71E2FB1C" box="[324,461,1227,1253]" italics="true" pageId="36" pageNumber="77">ankaraharae</emphasis>
</taxonomicName>
remains uncertain. 
<bibRefCitation id="EFBC4B481F05FFDB129DC5EE7179FAFC" author="NOBILI G." box="[151,342,1259,1285]" pageId="36" pageNumber="77" pagination="1 - 2" refId="ref26777" refString="NOBILI G. 1906. - Spedizione al Ruwenzori di S. A. R. Luigi Amedeo di Savoia Duca degli Abruzzi. 9: Crostacei. Bollettino dei Musei di Zoologia ed Anatomia comparata della R. Universita di Torino 21 (544): 1 - 2." type="journal article" year="1906">Nobili’s (1906)</bibRefCitation>
<typeStatus id="5496881B1F05FFDB136BC5EF71BBFAFD" box="[353,404,1258,1284]" pageId="36" pageNumber="77">type</typeStatus>
is unavailable and the figures of the 
<typeStatus id="5496881B1F05FFDB1323C40F7178FADD" box="[297,343,1290,1316]" pageId="36" pageNumber="77">type</typeStatus>
in 
<bibRefCitation id="EFBC4B481F05FFDB1370C40F71DAFADC" author="BOTT R." box="[378,501,1290,1317]" pageId="36" pageNumber="77" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott (1965</bibRefCitation>
: fig. 9a-c) do not show the mandible or first gonopod. It should be noted that the male specimen cw 
<quantity id="4CD59B5C1F05FFDB1058C44E7283FA9D" box="[594,684,1354,1381]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="36" pageNumber="77" unit="mm" value="28.0">28 mm</quantity>
(MNHN Paris, no number provided) labeled 
<taxonomicName id="4C2D4D3A1F05FFDB129DC48F7168FA5C" authorityName="sensu Nobili" authorityYear="1906" box="[151,327,1418,1445]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae">
<emphasis id="B959EAAB1F05FFDB129DC48F7168FA5C" box="[151,327,1418,1445]" italics="true" pageId="36" pageNumber="77">M. ankaraharae</emphasis>
</taxonomicName>
in 
<bibRefCitation id="EFBC4B481F05FFDB137BC48F71DCFA5C" author="BOTT R." box="[369,499,1418,1445]" pageId="36" pageNumber="77" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott (1965</bibRefCitation>
: fig. 9d, e, pl. 5 figs 23-25) from Diego-Suarez (now 
<collectingRegion id="49E9F85B1F05FFDB1029C4AE728AFA3C" box="[547,677,1451,1477]" country="Madagascar" name="Antsiranana" pageId="36" pageNumber="77">Antsiranana</collectingRegion>
) was not part of the 
<typeStatus id="5496881B1F05FFDB1378C4CF718FFA1D" box="[370,416,1482,1508]" pageId="36" pageNumber="77">type</typeStatus>
series from Ankarahara, and probably does not represent this taxon. We consider it likely that this specimen was misidentified, and that it may not actually belong to the same species as 
<taxonomicName id="4C2D4D3A1F05FFDB134AC74F70FBF97C" authority="sensu Nobili, 1906" authorityName="sensu Nobili" authorityYear="1906" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae" subGenus="Geothelphusa">
<emphasis id="B959EAAB1F05FFDB134AC74F7178F99A" box="[320,343,1610,1635]" italics="true" pageId="36" pageNumber="77">P.</emphasis>
(
<emphasis id="B959EAAB1F05FFDB136DC74F71ADF99D" box="[359,386,1610,1636]" italics="true" pageId="36" pageNumber="77">G.</emphasis>
) 
<emphasis id="B959EAAB1F05FFDB1398C74E723AF99C" box="[402,533,1611,1637]" italics="true" pageId="36" pageNumber="77">ankaraharae</emphasis>
sensu 
<bibRefCitation id="EFBC4B481F05FFDB1054C74E70FBF97C" author="NOBILI G." pageId="36" pageNumber="77" pagination="1 - 2" refId="ref26777" refString="NOBILI G. 1906. - Spedizione al Ruwenzori di S. A. R. Luigi Amedeo di Savoia Duca degli Abruzzi. 9: Crostacei. Bollettino dei Musei di Zoologia ed Anatomia comparata della R. Universita di Torino 21 (544): 1 - 2." type="journal article" year="1906">Nobili, 1906</bibRefCitation>
</taxonomicName>
. Although 
<bibRefCitation id="EFBC4B481F05FFDB135EC76F71F4F97C" author="BOTT R." box="[340,475,1642,1669]" pageId="36" pageNumber="77" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott (1965)</bibRefCitation>
did not provide a catalogue number, it is likely that the specimen from Diego-Suarez (MNHN-B 28) is the same one that was used (in part) in the redescription of 
<taxonomicName id="4C2D4D3A1F05FFDB10C0C00F735BFEDC" authorityName="sensu Nobili" authorityYear="1906" box="[714,884,266,293]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae">
<emphasis id="B959EAAB1F05FFDB10C0C00F735BFEDC" box="[714,884,266,293]" italics="true" pageId="36" pageNumber="77">M. ankaraharae</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFBC4B481F05FFDB1194C00F7435FEDC" author="BOTT R." box="[926,1050,266,293]" pageId="36" pageNumber="77" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott (1965</bibRefCitation>
: pl. V, figs 23-25, fig. 9e, f). For this reason, the illustrations of gonopods 1 and 
<quantity id="4CD59B5C1F05FFDB118CC04E739CFE9C" box="[902,947,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="36" pageNumber="77" unit="in" value="2.0">2 in</quantity>
<bibRefCitation id="EFBC4B481F05FFDB11B6C04F7402FE9C" author="BOTT R." box="[956,1069,330,357]" pageId="36" pageNumber="77" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott 1965</bibRefCitation>
(fig. 9d, e), the photographs of the whole animal (pl. V, figs 23- 25), and the text description of the mandibular palp as simple, may describe these structures in 
<taxonomicName id="4C2D4D3A1F05FFDB10C0C0CF7301FE1C" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[714,814,458,485]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="uglowi">
<emphasis id="B959EAAB1F05FFDB10C0C0CF7301FE1C" box="[714,814,458,485]" italics="true" pageId="36" pageNumber="77">B. uglowi</emphasis>
</taxonomicName>
n. gen, n. sp., rather than 
<taxonomicName id="4C2D4D3A1F05FFDB165BC0CF73D9FDFC" authority="sensu Nobili, 1906" authorityName="sensu Nobili" authorityYear="1906" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae" subGenus="Geothelphusa">
<emphasis id="B959EAAB1F05FFDB165BC0CF7447FE1A" box="[1105,1128,458,483]" italics="true" pageId="36" pageNumber="77">P.</emphasis>
(
<emphasis id="B959EAAB1F05FFDB167DC0CF74BDFE1D" box="[1143,1170,458,484]" italics="true" pageId="36" pageNumber="77">G.</emphasis>
) 
<emphasis id="B959EAAB1F05FFDB16A9C0CE7336FDFC" italics="true" pageId="36" pageNumber="77">ankaraharae</emphasis>
sensu 
<bibRefCitation id="EFBC4B481F05FFDB116FC0EE73D9FDFC" author="NOBILI G." box="[869,1014,491,517]" pageId="36" pageNumber="77" pagination="1 - 2" refId="ref26777" refString="NOBILI G. 1906. - Spedizione al Ruwenzori di S. A. R. Luigi Amedeo di Savoia Duca degli Abruzzi. 9: Crostacei. Bollettino dei Musei di Zoologia ed Anatomia comparata della R. Universita di Torino 21 (544): 1 - 2." type="journal article" year="1906">Nobili, 1906</bibRefCitation>
</taxonomicName>
. This means that the gonopods and mandible of 
<taxonomicName id="4C2D4D3A1F05FFDB1608C30F73B4FDBC" authority="sensu Nobili, 1906" authorityName="sensu Nobili" authorityYear="1906" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae" subGenus="Geothelphusa">
<emphasis id="B959EAAB1F05FFDB1608C30F7436FDDA" box="[1026,1049,522,547]" italics="true" pageId="36" pageNumber="77">P.</emphasis>
(
<emphasis id="B959EAAB1F05FFDB1621C30F7468FDDD" box="[1067,1095,522,548]" italics="true" pageId="36" pageNumber="77">G.</emphasis>
) 
<emphasis id="B959EAAB1F05FFDB1653C30E74F0FDDC" box="[1113,1247,523,549]" italics="true" pageId="36" pageNumber="77">ankaraharae</emphasis>
sensu 
<bibRefCitation id="EFBC4B481F05FFDB1101C32E73B4FDBC" author="NOBILI G." box="[779,923,555,581]" pageId="36" pageNumber="77" pagination="1 - 2" refId="ref26777" refString="NOBILI G. 1906. - Spedizione al Ruwenzori di S. A. R. Luigi Amedeo di Savoia Duca degli Abruzzi. 9: Crostacei. Bollettino dei Musei di Zoologia ed Anatomia comparata della R. Universita di Torino 21 (544): 1 - 2." type="journal article" year="1906">Nobili, 1906</bibRefCitation>
</taxonomicName>
, still await description.
</paragraph>
<paragraph id="8B9236B91F05FFDB10C0C34E7392FAFC" blockId="36.[714,1248,171,1285]" pageId="36" pageNumber="77">
Characters of the carapace, thoracic sternum and walking legs of 
<taxonomicName id="4C2D4D3A1F05FFDB1163C36F73E6FD7C" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[873,969,618,645]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="uglowi">
<emphasis id="B959EAAB1F05FFDB1163C36F73E6FD7C" box="[873,969,618,645]" italics="true" pageId="36" pageNumber="77">B. uglowi</emphasis>
</taxonomicName>
n. gen, n. sp. are similar to those shown in the figures of the 
<typeStatus id="5496881B1F05FFDB1634C38F7441FD5D" box="[1086,1134,650,676]" pageId="36" pageNumber="77">type</typeStatus>
of 
<taxonomicName id="4C2D4D3A1F05FFDB1698C38F7365FD3C" authorityName="sensu Nobili" authorityYear="1906" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae" subGenus="Geothelphusa">
<emphasis id="B959EAAB1F05FFDB1698C38F7485FD5A" box="[1170,1194,650,675]" italics="true" pageId="36" pageNumber="77">P.</emphasis>
(
<emphasis id="B959EAAB1F05FFDB16B0C38F74FAFD5D" box="[1210,1237,650,676]" italics="true" pageId="36" pageNumber="77">G.</emphasis>
) 
<emphasis id="B959EAAB1F05FFDB10C0C3AE7365FD3C" box="[714,842,683,709]" italics="true" pageId="36" pageNumber="77">ankaraharae</emphasis>
</taxonomicName>
provided by 
<bibRefCitation id="EFBC4B481F05FFDB11DAC3AF7463FD3C" author="BOTT R." box="[976,1100,682,709]" pageId="36" pageNumber="77" pagination="335 - 350" refId="ref25493" refString="BOTT R. 1965. - Die Susswasserkrabben von Madagaskar. Bulletin du Museum national d'Histoire naturelle 37 (2): 335 - 350." type="journal article" year="1965">Bott (1965)</bibRefCitation>
. The two taxa have a similar carapace outline and texture and walking legs of similar proportions. The two taxa can be distinguished by examination of the ischium of the third maxilliped, which has a deep vertical groove in 
<taxonomicName id="4C2D4D3A1F05FFDB1138C24F742FFC9C" authorityName="sensu Nobili" authorityYear="1906" box="[818,1024,842,869]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae" subGenus="Geothelphusa">
<emphasis id="B959EAAB1F05FFDB1138C24F7366FC9A" box="[818,841,842,867]" italics="true" pageId="36" pageNumber="77">P.</emphasis>
(
<emphasis id="B959EAAB1F05FFDB115DC24F735EFC9D" box="[855,881,842,868]" italics="true" pageId="36" pageNumber="77">G.</emphasis>
) 
<emphasis id="B959EAAB1F05FFDB118AC24E742FFC9C" box="[896,1024,843,869]" italics="true" pageId="36" pageNumber="77">ankaraharae</emphasis>
</taxonomicName>
, but which is characteristically faint or lacking in 
<taxonomicName id="4C2D4D3A1F05FFDB11FCC26F7479FC7C" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[1014,1110,874,901]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="uglowi">
<emphasis id="B959EAAB1F05FFDB11FCC26F7479FC7C" box="[1014,1110,874,901]" italics="true" pageId="36" pageNumber="77">B. uglowi</emphasis>
</taxonomicName>
n. gen, n. sp. The specimens from northern 
<collectingCountry id="F33A76291F05FFDB162AC28E748FFC5C" box="[1056,1184,907,933]" name="Madagascar" pageId="36" pageNumber="77">Madagascar</collectingCountry>
from Diego-Suarez (MNHN-B 5028), Nosy-Bé (MNHN-B 5029) and (MNHN-B 26147), and Nosy Komba (MNHN-B 12293) that were formerly identified as 
<taxonomicName id="4C2D4D3A1F05FFDB11B7C50F749EFBDC" authorityName="sensu Nobili" authorityYear="1906" box="[957,1201,1034,1061]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="ankaraharae" subGenus="Geothelphusa">
<emphasis id="B959EAAB1F05FFDB11B7C50F73FAFBDA" box="[957,981,1034,1059]" italics="true" pageId="36" pageNumber="77">P.</emphasis>
(
<emphasis id="B959EAAB1F05FFDB11E1C50F7428FBDD" box="[1003,1031,1034,1060]" italics="true" pageId="36" pageNumber="77">G.</emphasis>
) 
<emphasis id="B959EAAB1F05FFDB1617C50E749EFBDC" box="[1053,1201,1035,1061]" italics="true" pageId="36" pageNumber="77">ankaraharae</emphasis>
</taxonomicName>
are included here in the new taxon, because they resemble 
<taxonomicName id="4C2D4D3A1F05FFDB1132C54F738CFB9C" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[824,931,1098,1125]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="uglowi">
<emphasis id="B959EAAB1F05FFDB1132C54F738CFB9C" box="[824,931,1098,1125]" italics="true" pageId="36" pageNumber="77">B. uglowi</emphasis>
</taxonomicName>
n. gen, n. sp. very closely. These specimens differ from 
<taxonomicName id="4C2D4D3A1F05FFDB1616C56F74AAFB7C" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[1052,1157,1130,1157]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="uglowi">
<emphasis id="B959EAAB1F05FFDB1616C56F74AAFB7C" box="[1052,1157,1130,1157]" italics="true" pageId="36" pageNumber="77">B. uglowi</emphasis>
</taxonomicName>
n. gen, n. sp. in the texture of the pterygostomial region of the carapace sidewall, which is smooth in 
<taxonomicName id="4C2D4D3A1F05FFDB10C0C5CF731DFB1C" authorityName="Cumberlidge &amp; Sternberg" authorityYear="2002" box="[714,818,1226,1253]" class="Malacostraca" family="Potamonautidae" genus="Boreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="77" phylum="Arthropoda" rank="species" species="uglowi">
<emphasis id="B959EAAB1F05FFDB10C0C5CF731DFB1C" box="[714,818,1226,1253]" italics="true" pageId="36" pageNumber="77">B. uglowi</emphasis>
</taxonomicName>
n. gen, n. sp. and distinctly setose in these other specimens.
</paragraph>
</subSubSection>
</treatment>
</document>