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<document id="2872649BC00103563415FC3A841807FA" ID-DOI="10.11646/phytotaxa.218.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13635877" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725333068986" checkinUser="felipe" docAuthor="Karremans, Adam P., Bogarín, Diego, Pupulin, Franco, Luer, Carlyle A. &amp; Gravendeel, Barbara" docDate="2015" docId="038487AAE471FFB9FF4446ADFF37FEF6" docLanguage="en" docName="phytotaxa.218.2.1.pdf" docOrigin="Phytotaxa 218 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.218.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Specklinia gersonii Bogarin &amp; Karremans Karremans 2015, sp. nov." docType="treatment" docVersion="2" lastPageNumber="114" masterDocId="FFBDFFD2E47AFFB4FFCC451BFFEDFFD3" masterDocTitle="The glandulous Specklinia: morphological convergence versus phylogenetic divergence" masterLastPageNumber="127" masterPageNumber="101" pageNumber="112" updateTime="1725334363440" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="1FE372FFB503DE541AACE176372202D8">
<mods:title id="ED0EE4FEFEB4AEAF96BD07059055C994">The glandulous Specklinia: morphological convergence versus phylogenetic divergence</mods:title>
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<mods:role id="228055097293A58D6694CFF4994F98BB">
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<mods:namePart id="CA89B8D918E0F3472EF4AC1085AA4C5D">Karremans, Adam P.</mods:namePart>
<mods:affiliation id="8E7983A2C5B1A65861CC81C4F8078EE0">Naturalis Biodiversity Center-Leiden University, The Netherlands.</mods:affiliation>
</mods:name>
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<mods:namePart id="431CB9690C6A4C0D9188E8DD84C6D2D4">Bogarín, Diego</mods:namePart>
<mods:affiliation id="27C446DA763ADE51C50788C486EAC1B9">Herbario UCH, Universidad Autónoma de Chiriquí, 0427, David, Chiriquí, Panama.</mods:affiliation>
</mods:name>
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<mods:namePart id="4100F6CDDC1E24AEC9845893AFCF350A">Pupulin, Franco</mods:namePart>
<mods:affiliation id="8CC25872407CE6EC66BDCF9EEFB77D07">Harvard University Herbaria, Cambridge, MA, U. S. A.; Marie Selby Botanical Gardens, Sarasota, FL, U. S. A.</mods:affiliation>
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<mods:namePart id="C45A86246EB2D2B34759442D2747FF8F">Luer, Carlyle A.</mods:namePart>
<mods:affiliation id="857F61033BD89E5FB983133D2DCB8015">Curator emeritus: Missouri Botanical Garden, St. Louis, Missouri, U. S. A.</mods:affiliation>
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<mods:namePart id="9AD8BF729FAA11F1A9BCB1CD537238C3">Gravendeel, Barbara</mods:namePart>
<mods:affiliation id="AF042BCC66A8FB484D28E6358453A588">Naturalis Biodiversity Center-Leiden University, The Netherlands.</mods:affiliation>
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<mods:title id="C013B210A64C64CE24B2490D883A9D91">Phytotaxa</mods:title>
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<mods:date id="ACD97828FF20C004DF941A41DBC6EECA">2015</mods:date>
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<mods:number id="42EDB5C7731111B7511DF5D888B30200">2015-07-02</mods:number>
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<mods:number id="5C22FDCECEFE40D17349BF5E723B6843">218</mods:number>
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<mods:classification id="946F97D77B02AF661B11167DE43FA224">journal article</mods:classification>
<mods:identifier id="611006CCD890769A8CFEFC8CF6232DBD" type="DOI">10.11646/phytotaxa.218.2.1</mods:identifier>
<mods:identifier id="4D2735133531E82906EDDA9E84698BE9" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="356D0CF1A607BD05020CF958F203ACCC" type="Zenodo-Dep">13635877</mods:identifier>
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<treatment id="038487AAE471FFB9FF4446ADFF37FEF6" ID-DOI="http://doi.org/10.5281/zenodo.13635897" ID-Zenodo-Dep="13635897" LSID="urn:lsid:plazi:treatment:038487AAE471FFB9FF4446ADFF37FEF6" httpUri="http://treatment.plazi.org/id/038487AAE471FFB9FF4446ADFF37FEF6" lastPageId="13" lastPageNumber="114" pageId="11" pageNumber="112">
<subSubSection id="C3376537E471FFBFFF4446ADFD24FC02" box="[136,713,950,977]" pageId="11" pageNumber="112" type="nomenclature">
<paragraph id="8B9236BCE471FFBFFF4446ADFD24FC02" blockId="11.[136,713,950,977]" box="[136,713,950,977]" pageId="11" pageNumber="112">
<heading id="D0DA81D0E471FFBFFF4446ADFD24FC02" box="[136,713,950,977]" fontSize="11" level="2" pageId="11" pageNumber="112" reason="3">
<taxonomicName id="4C2D4D3FE471FFBFFF4446ADFD84FC02" ID-CoL="4YGWQ" authority="Bogarin &amp; Karremans" authorityName="Bogarin &amp; Karremans Karremans" authorityYear="2015" box="[136,617,950,977]" class="Liliopsida" family="Orchidaceae" genus="Specklinia" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="112" phylum="Tracheophyta" rank="species" species="gersonii" status="sp. nov.">
<emphasis id="B959EAAEE471FFBFFF4446ADFE88FC03" bold="true" box="[136,357,950,977]" italics="true" pageId="11" pageNumber="112">Specklinia gersonii</emphasis>
Bogarín &amp; Karremans
</taxonomicName>
,
<emphasis id="B959EAAEE471FFBFFDB846A3FD24FC02" box="[628,713,952,977]" italics="true" pageId="11" pageNumber="112">
<taxonomicNameLabel id="A26A57D5E471FFBFFDB846A3FD24FC02" box="[628,713,952,977]" pageId="11" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3376537E471FFBFFF444119FBD2F7F2" pageId="11" pageNumber="112" type="materials_examined">
<paragraph id="8B9236BCE471FFBFFF444119FD31FB8C" blockId="11.[136,1452,1026,1119]" pageId="11" pageNumber="112">
<materialsCitation id="3B453CE1E471FFBFFF444119FD35FB8C" collectingDate="2012-04-12" collectorName="La Argentina &amp; Gerson Villalobos &amp; Daniel Matamoros &amp; el Jardin Botanico Lankester &amp; D. Bogarin" country="Costa Rica" county="Type" elevation="600" location="Pocora" municipality="Guacimo" pageId="11" pageNumber="112" specimenCount="2" stateProvince="Limon" typeStatus="holotype">
<collectingCounty id="62F34E30E471FFBFFF444119FF55FBC4" box="[136,184,1026,1047]" pageId="11" pageNumber="112">Type</collectingCounty>
:—
<collectingCountry id="F33A762CE471FFBFFF184119FE8BFBC4" box="[212,358,1026,1047]" name="Costa Rica" pageId="11" pageNumber="112">COSTA RICA</collectingCountry>
.
<collectingRegion id="49E9F85EE471FFBFFEBD4119FE58FBC4" box="[369,437,1026,1047]" country="Costa Rica" name="Limon" pageId="11" pageNumber="112">Limón</collectingRegion>
:
<collectingMunicipality id="6BF6ACC6E471FFBFFE0C4119FDF7FBC4" box="[448,538,1026,1047]" pageId="11" pageNumber="112">Guácimo</collectingMunicipality>
,
<location id="8EF26067E471FFBFFDEA4119FD86FBC4" LSID="urn:lsid:plazi:treatment:038487AAE471FFB9FF4446ADFF37FEF6:8EF26067E471FFBFFDEA4119FD86FBC4" box="[550,619,1026,1047]" country="Costa Rica" county="Type" municipality="Guacimo" name="Pocora" pageId="11" pageNumber="112" stateProvince="Limon">Pocora</location>
,
<collectorName id="26D8536AE471FFBFFDBB4119FD14FBC4" box="[631,761,1026,1047]" pageId="11" pageNumber="112">
<location id="8EF26067E471FFBFFDBB4119FD14FBC4" LSID="urn:lsid:plazi:treatment:038487AAE471FFB9FF4446ADFF37FEF6:8EF26067E471FFBFFDBB4119FD14FBC4" box="[631,761,1026,1047]" country="Costa Rica" county="Type" municipality="Guacimo" name="La Argentina" pageId="11" pageNumber="112" stateProvince="Limon">La Argentina</location>
</collectorName>
, camino a la catarata del
<location id="8EF26067E471FFBFFC344119FB72FBC4" LSID="urn:lsid:plazi:treatment:038487AAE471FFB9FF4446ADFF37FEF6:8EF26067E471FFBFFC344119FB72FBC4" box="[1016,1183,1026,1047]" country="Costa Rica" county="Type" municipality="Guacimo" name="rio Dos Novillos" pageId="11" pageNumber="112" stateProvince="Limon">río Dos Novillos</location>
, ca.
<quantity id="4CD59B59E471FFBFFB024119FAFDFBC4" box="[1230,1296,1026,1047]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="112" unit="m" value="600.0">
<elevation id="0000D18FE471FFBFFB024119FAFDFBC4" box="[1230,1296,1026,1047]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="112" unit="m" value="600.0">600 m</elevation>
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, invenit
<collectorName id="26D8536AE471FFBFFAAA4119FECDFBE8" pageId="11" pageNumber="112">Gerson Villalobos</collectorName>
et
<collectorName id="26D8536AE471FFBFFE8D413DFE14FBE8" box="[321,505,1062,1083]" pageId="11" pageNumber="112">Daniel Matamoros</collectorName>
, floreció en cultivo en
<collectorName id="26D8536AE471FFBFFD13413DFC14FBE8" box="[735,1017,1062,1083]" pageId="11" pageNumber="112">el Jardín Botánico Lankester</collectorName>
,
<date id="FF93107CE471FFBFFBC9413DFB7DFBE8" box="[1029,1168,1062,1083]" pageId="11" pageNumber="112" value="2012-04-12">
<collectingDate id="EFD7E994E471FFBFFBC9413DFB7DFBE8" box="[1029,1168,1062,1083]" pageId="11" pageNumber="112" value="2012-04-12">12 April 2012</collectingDate>
</date>
,
<emphasis id="B959EAAEE471FFBFFB50413DFAACFBE8" box="[1180,1345,1062,1083]" italics="true" pageId="11" pageNumber="112">
<collectorName id="26D8536AE471FFBFFB50413DFAE7FBE8" box="[1180,1290,1062,1083]" pageId="11" pageNumber="112">D. Bogarín</collectorName>
9565
</emphasis>
(
<typeStatus id="5496881EE471FFBFFA9C413DFA45FBE8" box="[1360,1448,1062,1083]" pageId="11" pageNumber="112" type="holotype">holotype</typeStatus>
, JBL-spirit, D5192!;
<typeStatus id="5496881EE471FFBFFE4D4151FE24FB8C" box="[385,457,1098,1119]" pageId="11" pageNumber="112" type="isotype">isotype</typeStatus>
, JBL-spirit, D5197!;
<figureCitation id="13162A39E471FFBFFD594151FD23FB8C" box="[661,718,1098,1119]" captionStart="FIGURE 8" captionStartId="12.[136,229,1868,1889]" captionTargetBox="[151,1436,190,1823]" captionTargetId="figure-14@12.[151,1436,190,1843]" captionTargetPageId="12" captionText="FIGURE 8. Specklinia gersonii Bogarín &amp; Karremans. A. habit. B. flower. C. dissected perianth. D. column and lip, lateral view. E. column ventral view. F. lip. G. anther and pollinaria. Drawn by D. Bogarín &amp; L. Oses from Bogarín 9565 (JBL-spirit)." figureDoi="http://doi.org/10.5281/zenodo.13635899" httpUri="https://zenodo.org/record/13635899/files/figure.png" pageId="11" pageNumber="112">Fig. 8</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9236BCE471FFBFFF444196FAC4FB37" blockId="11.[136,1452,1165,1252]" pageId="11" pageNumber="112">
<emphasis id="B959EAAEE471FFBFFF444196FAC4FB37" italics="true" pageId="11" pageNumber="112">
The species is similar to
<taxonomicName id="4C2D4D3FE471FFBFFE424196FC3FFB77" authority="(Ames) Pridgeon &amp; M. W. Chase" authorityName="Pridgeon &amp; M. W. Chase. A." authorityYear="2001" baseAuthorityName="Ames" baseAuthorityYear="1923" box="[398,978,1165,1188]" class="Liliopsida" family="Orchidaceae" genus="Specklinia" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="112" phylum="Tracheophyta" rank="species" species="glandulosa">Specklinia glandulosa (Ames) Pridgeon &amp; M.W.Chase</taxonomicName>
, but can be easily distinguished by the wider and sub-orbicular (vs. linear) leaves, the lack of a bract on the peduncle, the trialate ovary (vs. terete), the wider dorsal sepal (2.5 vs
<quantity id="4CD59B59E471FFBFFF1A41D6FEC5FB37" box="[214,296,1229,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="112" unit="mm" value="1.5">1.5 mm</quantity>
), and the widely rounded apical half of the sepals in natural position (vs. narrow and elongate).
</emphasis>
</paragraph>
<paragraph id="8B9236BCE471FFBFFF44400FFF06F866" blockId="11.[136,1452,1300,2081]" pageId="11" pageNumber="112">
Epiphytic, caespitose, ascending, erect herb to 4.0 cm tall. Roots fibrous, flexuous, glabrous, to
<quantity id="4CD59B59E471FFBFFB15400FFAF1FAFE" box="[1241,1308,1300,1325]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="112" unit="mm" value="1.0">1 mm</quantity>
in diameter. Stem abbreviated, terete, to
<quantity id="4CD59B59E471FFBFFE094023FDEBFA82" box="[453,518,1336,1361]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="112" unit="mm" value="5.0">5 mm</quantity>
long, concealed by a papyraceous, subancipitous, acute sheath to
<quantity id="4CD59B59E471FFBFFB234023FADCFA82" box="[1263,1329,1336,1361]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="112" unit="mm" value="4.0">4 mm</quantity>
long. Leaf elliptic to broadly elliptic, up to 20 ×
<quantity id="4CD59B59E471FFBFFDF64047FD61FAA6" box="[570,652,1372,1397]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="112" unit="mm" value="12.0">12 mm</quantity>
, minutely and irregularly emarginate at apex, the mid-vein protruding abaxially into a small apicule, gradually tapering toward the base into a deeply conduplicate petiole, subcoriaceous. Inflorescence borne laterally from the base of the leaf, without an annulus, successively single flowered, up to
<quantity id="4CD59B59E471FFBFFABF40BFFA46FA6E" box="[1395,1451,1444,1469]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="112" unit="cm" value="3.0">3 cm</quantity>
long, glandular; peduncle terete, to
<quantity id="4CD59B59E471FFBFFDD940D3FDA1FA32" box="[533,588,1480,1505]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="112" unit="cm" value="2.0">2 cm</quantity>
long, without bracts. Floral bracts infundibuliform, glandular, broadly ovate, acute to subacuminate,
<quantity id="4CD59B59E471FFBFFE5F40F7FE35F9D6" box="[403,472,1516,1541]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="112" unit="mm" value="4.0">4 mm</quantity>
. Pedicel terete, glandular,
<quantity id="4CD59B59E471FFBFFCC840F7FCB6F9D6" box="[772,859,1516,1541]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="112" unit="mm" value="3.5">3.5 mm</quantity>
long, persistent, appearing fascicled. Ovary trialate, subclavate,
<quantity id="4CD59B59E471FFBFFEC7430BFEA1F9FA" box="[267,332,1552,1577]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="112" unit="mm" value="2.0">2 mm</quantity>
long, greenish to orange. Flowers up to 4, only one developed at a time; with fruity fragrance around midday. Sepals fleshy, densely microscopically-glandulose on the outer surface; dorsal sepal narrowly-elliptic, 3- veined, acute, greenish orange, with bright orange-red, the margins microscopically glandulous, 7.0–8.0 ×
<quantity id="4CD59B59E471FFBFFA804343FA4AF9A2" box="[1356,1447,1624,1649]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="112" unit="mm" value="2.5">2.5 mm</quantity>
; lateral sepals narrowly elliptic-oblanceolate, subfalcate, 3-veined, 6.5–7.5 × 4.0–5.0 mm, connate for about two thirds of their length, the midvein strongly carinate abaxially. Petals small, lanceolate-falcate, acute, 3.1–3.3 ×
<quantity id="4CD59B59E471FFBFFAD243BBFA4AF96A" box="[1310,1447,1696,1721]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="11" pageNumber="112" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
, 3-veined. Lip reddish-orange, small, longitudinally arched-convex in natural position, thinly articulate with the column foot by a hyaline claw, apically bifid-emarginate in natural position, sagittate to sub-triangular when expanded, obtuse, 3.5 ×
<quantity id="4CD59B59E471FFBFFF074217FEB9F8F6" box="[203,340,1804,1829]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="11" pageNumber="112" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
, provided with a pair of sharp, triangular lateral lobes at the base. Column dark-red, arched, terete and slender at the base,
<quantity id="4CD59B59E471FFBFFE51422BFE18F89A" box="[413,501,1840,1865]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="112" unit="mm" value="2.5">2.5 mm</quantity>
long without the foot, provided with membranous wings, the apex prolonged into a deeply cucullate, lacerate clinandrium; column foot, stout, fleshy,
<quantity id="4CD59B59E471FFBFFC43424FFC3FF8BE" box="[911,978,1876,1901]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="112" unit="mm" value="1.0">1 mm</quantity>
long. Anther cap deeply cucullate, ovate, 2-celled. Pollinia 2, obovate-complanate, minutely hooked at the base. *
<collectionCode id="ED3CAE79E471FFBFFC1C4263FBCFF842" box="[976,1058,1912,1937]" pageId="11" pageNumber="112">NOTE</collectionCode>
: Description based on
<emphasis id="B959EAAEE471FFBFFAE24263FA66F842" box="[1326,1419,1912,1937]" italics="true" pageId="11" pageNumber="112">DB9565</emphasis>
&amp;
<emphasis id="B959EAAEE471FFBFFF444287FF09F866" box="[136,228,1948,1973]" italics="true" pageId="11" pageNumber="112">
<collectionCode id="ED3CAE79E471FFBFFF444287FF43F866" box="[136,174,1948,1973]" country="New Zealand" lsid="urn:lsid:biocol.org:col:15426" name="Auckland War Memorial Museum" pageId="11" pageNumber="112" type="Herbarium">AK</collectionCode>
6025
</emphasis>
.
</paragraph>
<paragraph id="8B9236BCE471FFBFFF7142DBFBD2F7F2" blockId="11.[136,1452,1300,2081]" pageId="11" pageNumber="112">
<emphasis id="B959EAAEE471FFBFFF7142DBFE5BF80A" bold="true" box="[189,438,1984,2009]" pageId="11" pageNumber="112">Additional material:</emphasis>
—
<materialsCitation id="3B453CE1E471FFBFFE1E42DBFBD1F7F2" collectingDate="2013-11-25" collectorName="La Argentina &amp; Karremans &amp; Bogarin &amp; Villalobos" country="Costa Rica" county="Guacimo" elevation="591" latitude="10.102142" location="La Argentina" longLatPrecision="1" longitude="-83.65798" municipality="Pocora" pageId="11" pageNumber="112" specimenCount="1" stateProvince="Limon">
<collectingCountry id="F33A762CE471FFBFFE1E42DBFD91F80A" box="[466,636,1984,2009]" name="Costa Rica" pageId="11" pageNumber="112">COSTA RICA</collectingCountry>
.
<collectingRegion id="49E9F85EE471FFBFFD4642DBFD34F80A" box="[650,729,1984,2009]" country="Costa Rica" name="Limon" pageId="11" pageNumber="112">Limón</collectingRegion>
:
<collectingCounty id="62F34E30E471FFBFFD2442DBFCBFF80A" box="[744,850,1984,2009]" pageId="11" pageNumber="112">Guácimo</collectingCounty>
,
<collectingMunicipality id="6BF6ACC6E471FFBFFCAD42DBFC5FF80A" box="[865,946,1984,2009]" pageId="11" pageNumber="112">Pocora</collectingMunicipality>
,
<collectorName id="26D8536AE471FFBFFC0C42DBFBB4F80A" box="[960,1113,1984,2009]" pageId="11" pageNumber="112">
<location id="8EF26067E471FFBFFC0C42DBFBB4F80A" LSID="urn:lsid:plazi:treatment:038487AAE471FFB9FF4446ADFF37FEF6:8EF26067E471FFBFFC0C42DBFBB4F80A" box="[960,1113,1984,2009]" country="Costa Rica" county="Guacimo" latitude="10.102142" longLatPrecision="1" longitude="-83.65798" municipality="Pocora" name="La Argentina" pageId="11" pageNumber="112" stateProvince="Limon">La Argentina</location>
</collectorName>
, camino a la catarata del
<location id="8EF26067E471FFBFFA4042DBFECFF82E" LSID="urn:lsid:plazi:treatment:038487AAE471FFB9FF4446ADFF37FEF6:8EF26067E471FFBFFA4042DBFECFF82E" country="Costa Rica" county="Guacimo" latitude="10.102142" longLatPrecision="1" longitude="-83.65798" municipality="Pocora" name="rio Dos Novillos" pageId="11" pageNumber="112" stateProvince="Limon">río Dos Novillos</location>
,
<geoCoordinate id="EE19507BE471FFBFFEFC42FFFE0BF82E" box="[304,486,2020,2045]" degrees="10" direction="north" minutes="06" orientation="latitude" pageId="11" pageNumber="112" precision="1" seconds="07.71" value="10.102142">10°06’07.71” N</geoCoordinate>
<geoCoordinate id="EE19507BE471FFBFFE2342FFFD41F82E" box="[495,684,2020,2045]" degrees="83" direction="west" minutes="39" orientation="longitude" pageId="11" pageNumber="112" precision="1" seconds="28.74" value="-83.65798">83°39’28.74” W</geoCoordinate>
,
<quantity id="4CD59B59E471FFBFFD7542FFFCE8F82E" box="[697,773,2020,2045]" metricMagnitude="2" metricUnit="m" metricValue="5.91" pageId="11" pageNumber="112" unit="m" value="591.0">
<elevation id="0000D18FE471FFBFFD7542FFFCE8F82E" box="[697,773,2020,2045]" metricMagnitude="2" metricUnit="m" metricValue="5.91" pageId="11" pageNumber="112" unit="m" value="591.0">591 m</elevation>
</quantity>
, bosque muy húmedo tropical transición a premontano,
<date id="FF93107CE471FFBFFA4342FFFEADF7F2" pageId="11" pageNumber="112" value="2013-11-25">
<collectingDate id="EFD7E994E471FFBFFA4342FFFEADF7F2" pageId="11" pageNumber="112" value="2013-11-25">25 November 2013</collectingDate>
</date>
,
<emphasis id="B959EAAEE471FFBFFE874D13FCEBF7F2" box="[331,774,2056,2081]" italics="true" pageId="11" pageNumber="112">
<collectorName id="26D8536AE471FFBFFE874D13FE26F7F2" box="[331,459,2056,2081]" pageId="11" pageNumber="112">Karremans</collectorName>
,
<collectorName id="26D8536AE471FFBFFE1A4D13FDDFF7F2" box="[470,562,2056,2081]" pageId="11" pageNumber="112">Bogarín</collectorName>
&amp;
<collectorName id="26D8536AE471FFBFFD9A4D13FD2AF7F2" box="[598,711,2056,2081]" pageId="11" pageNumber="112">Villalobos</collectorName>
6025
</emphasis>
(JBL-spirit!;
<figureCitation id="13162A39E471FFBFFC6C4D13FC02F7F2" box="[928,1007,2056,2081]" captionStart="FIGURE 1" captionStartId="3.[136,229,1989,2010]" captionTargetBox="[204,1384,190,1964]" captionTargetId="figure-14@3.[204,1384,190,1964]" captionTargetPageId="3" captionText="FIGURE 1. The glandulous Specklinia species. A. S. alajuelensis 1 (Karremans 3265). B. S. alajuelensis 2 (Bogarín 2895). C. S. chontalensis (Pupulin 6543). D. S. gersonii (Karremans 6025). E. S. glandulosa (Karremans 6306). F. S. vittariifolia (Karremans 2945). Photographs by A.P. Karremans." figureDoi="http://doi.org/10.5281/zenodo.13635879" httpUri="https://zenodo.org/record/13635879/files/figure.png" pageId="11" pageNumber="112">Fig. 1d</figureCitation>
&amp;
<figureCitation id="13162A39E471FFBFFBDF4D13FBDFF7F2" box="[1043,1074,2056,2081]" captionStart="FIGURE 2" captionStartId="4.[136,229,1062,1083]" captionTargetBox="[438,1149,281,1037]" captionTargetId="figure-54@4.[437,1149,281,1037]" captionTargetPageId="4" captionText="FIGURE 2. Comparison of the habits and flowers of species of the glandulous species of Specklinia. A. S. alajuelensis 1 (Karremans 3265). B. S. gersonii (Karremans 6025). C. S. glandulosa (Karremans 6306). D. S. vittariifolia (Karremans 2945). Scale bar = 1 cm. Photographs by A.P. Karremans." figureDoi="http://doi.org/10.5281/zenodo.13635881" httpUri="https://zenodo.org/record/13635881/files/figure.png" pageId="11" pageNumber="112">2b</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF526634E476FFB8FF444257FB12F856" ID-DOI="http://doi.org/10.5281/zenodo.13635899" ID-Zenodo-Dep="13635899" httpUri="https://zenodo.org/record/13635899/files/figure.png" pageId="12" pageNumber="113" startId="12.[136,229,1868,1889]" targetBox="[151,1436,190,1823]" targetPageId="12" targetType="figure">
<paragraph id="8B9236BCE476FFB8FF444257FB12F856" blockId="12.[136,1451,1868,1925]" pageId="12" pageNumber="113">
<emphasis id="B959EAAEE476FFB8FF444257FEEDF8B2" bold="true" box="[136,256,1868,1889]" pageId="12" pageNumber="113">FIGURE 8.</emphasis>
<taxonomicName id="4C2D4D3FE476FFB8FEC44257FD24F8B2" authority="Bogarin &amp; Karremans. A." authorityName="Bogarin &amp; Karremans Karremans" authorityYear="2015" box="[264,713,1868,1889]" class="Liliopsida" family="Orchidaceae" genus="Specklinia" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="gersonii">
<emphasis id="B959EAAEE476FFB8FEC44257FE2EF8B2" box="[264,451,1868,1889]" italics="true" pageId="12" pageNumber="113">Specklinia gersonii</emphasis>
Bogarín &amp; Karremans.
<emphasis id="B959EAAEE476FFB8FD7E4257FD2EF8B2" bold="true" box="[690,707,1868,1889]" pageId="12" pageNumber="113">A</emphasis>
.
</taxonomicName>
habit.
<emphasis id="B959EAAEE476FFB8FCC34257FCF2F8B2" bold="true" box="[783,799,1868,1889]" pageId="12" pageNumber="113">B</emphasis>
. flower.
<emphasis id="B959EAAEE476FFB8FCB54257FC67F8B2" bold="true" box="[889,906,1868,1889]" pageId="12" pageNumber="113">C</emphasis>
. dissected perianth.
<emphasis id="B959EAAEE476FFB8FB994257FB8BF8B2" bold="true" box="[1109,1126,1868,1889]" pageId="12" pageNumber="113">D</emphasis>
. column and lip, lateral view.
<emphasis id="B959EAAEE476FFB8FA594257FA48F8B2" bold="true" box="[1429,1445,1868,1889]" pageId="12" pageNumber="113">E</emphasis>
. column ventral view.
<emphasis id="B959EAAEE476FFB8FE94426BFE8AF856" bold="true" box="[344,359,1904,1925]" pageId="12" pageNumber="113">F</emphasis>
. lip.
<emphasis id="B959EAAEE476FFB8FE54426BFE46F856" bold="true" box="[408,427,1904,1925]" pageId="12" pageNumber="113">G</emphasis>
. anther and pollinaria. Drawn by D. Bogarín &amp; L. Oses from
<emphasis id="B959EAAEE476FFB8FC30426BFB6DF856" box="[1020,1152,1904,1925]" italics="true" pageId="12" pageNumber="113">Bogarín 9565</emphasis>
(JBL-spirit).
</paragraph>
</caption>
<subSubSection id="C3376537E476FFB8FF7142AEFEF3F821" pageId="12" pageNumber="113" type="etymology">
<paragraph id="8B9236BCE476FFB8FF7142AEFEF3F821" blockId="12.[136,1452,1973,2070]" pageId="12" pageNumber="113">
<emphasis id="B959EAAEE476FFB8FF7142AEFEA5F81D" bold="true" box="[189,328,1973,1998]" pageId="12" pageNumber="113">Etymology:</emphasis>
—The name honors Gerson Villalobos, a Costa Rican orchid enthusiast who brought this species to our attention.
</paragraph>
</subSubSection>
<subSubSection id="C3376537E476FFB9FF7142E6FF37FEF6" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113" type="distribution">
<paragraph id="8B9236BCE476FFB8FF7142E6FB2FF7C5" blockId="12.[136,1452,1973,2070]" box="[189,1218,2045,2070]" pageId="12" pageNumber="113">
<emphasis id="B959EAAEE476FFB8FF7142E6FEB4F7C5" bold="true" box="[189,345,2045,2070]" pageId="12" pageNumber="113">Distribution:</emphasis>
—Known only from
<collectingCountry id="F33A762CE476FFB8FD8942E6FD2CF7C5" box="[581,705,2045,2070]" name="Costa Rica" pageId="12" pageNumber="113">Costa Rica</collectingCountry>
, growing at elevations around
<quantity id="4CD59B59E476FFB8FBD642E6FB8CF7C5" box="[1050,1121,2045,2070]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="113" unit="m" value="600.0">600 m</quantity>
(
<figureCitation id="13162A39E476FFB8FBBC42E6FB59F7C5" box="[1136,1204,2045,2070]" captionStart="FIGURE 6" captionStartId="9.[136,229,918,939]" captionTargetBox="[151,1437,317,893]" captionTargetId="figure-56@9.[151,1437,317,893]" captionTargetPageId="9" captionText="FIGURE 6. General distribution map of the glandulous species of Specklinia. Edited by D. Bogarín." figureDoi="http://doi.org/10.5281/zenodo.13635893" httpUri="https://zenodo.org/record/13635893/files/figure.png" pageId="12" pageNumber="113">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8B9236BCE477FFB9FF7145BBFF37FEF6" blockId="13.[136,1452,160,293]" pageId="13" pageNumber="114">
<emphasis id="B959EAAEE477FFB9FF7145BBFEE4FF6A" bold="true" box="[189,265,160,185]" pageId="13" pageNumber="114">Notes:</emphasis>
—Despite its appealing bright orange flowers and easily recognizable broadly elliptic leaves it seems that
<taxonomicName id="4C2D4D3FE477FFB9FF4445DFFD8EFF0E" authority="Bogarin &amp; Karremans" authorityName="Bogarin &amp; Karremans Karremans" authorityYear="2015" box="[136,611,196,221]" class="Liliopsida" family="Orchidaceae" genus="Specklinia" kingdom="Plantae" order="Asparagales" pageId="13" pageNumber="114" phylum="Tracheophyta" rank="species" species="gersonii">
<emphasis id="B959EAAEE477FFB9FF4445DFFE8CFF0E" box="[136,353,196,221]" italics="true" pageId="13" pageNumber="114">Specklinia gersonii</emphasis>
Bogarín &amp; Karremans
</taxonomicName>
had eluded botanists so far. The species is as far as we know very rare; in fact we are aware only of a handful of plants, all from a single population on a trail along the Dos Novillos river in
<collectingRegion id="49E9F85EE477FFB9FF444417FF3BFEF6" box="[136,214,268,293]" country="Costa Rica" name="Limon" pageId="13" pageNumber="114">Limón</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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