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<document id="DCAD6AD6BB36F67973ED76C5FD83688A" ID-DOI="10.11646/zootaxa.5512.4.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849236" ID-ZooBank="631FD956-27A5-4D98-9968-0D2CB66889D6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727440606981" checkinUser="plazi" docAuthor="Liu, Dongxia, Yang, Ding &amp; Li, Weihai" docDate="2024" docId="0384878EFFDDFFAA7499F9B832BDFF05" docLanguage="en" docName="zootaxa.5512.4.8.pdf" docOrigin="Zootaxa 5512 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5512.4.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Indonemoura geminus Zwick 1977" docType="treatment" docVersion="2" lastPageNumber="591" masterDocId="FFBDFFF6FFDCFFA8740EFFC13066FFD0" masterDocTitle="New species and new records of Indonemoura (Plecoptera: Nemouridae) from Xizang of southwestern China" masterLastPageNumber="596" masterPageNumber="590" pageNumber="591" updateTime="1727441779009" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="95C3B07529C016F03620E1A916071477">New species and new records of Indonemoura (Plecoptera: Nemouridae) from Xizang of southwestern China</mods:title>
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<mods:namePart id="110987387003F29381C11AF95B764980">Yang, Ding</mods:namePart>
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<taxonomicName id="4C2D4D1BFFDDFFA97499F9B8322DF944" ID-CoL="3PNF5" authority="Zwick, 1977" authorityName="Zwick" authorityYear="1977" box="[151,587,1657,1684]" class="Insecta" family="Nemouridae" genus="Indonemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B959EA8AFFDDFFA97499F9B8322DF944" bold="true" box="[151,587,1657,1684]" pageId="1" pageNumber="591">
<emphasis id="B959EA8AFFDDFFA97499F9B831CAF944" bold="true" box="[151,428,1657,1684]" italics="true" pageId="1" pageNumber="591">Indonemoura geminus</emphasis>
<bibRefCitation id="EFBC4B69FFDDFFA975BDF9B8322DF944" author="Zwick, P." box="[435,587,1657,1684]" pageId="1" pageNumber="591" pagination="85 - 134" refId="ref2988" refString="Zwick, P. (1977) Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Plecoptera. Entomologica Basiliensia, 2, 85 - 134." type="journal article" year="1977">Zwick, 1977</bibRefCitation>
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<figureCitation id="13162A1DFFDDFFA97499F95F30BFF968" box="[151,217,1694,1720]" captionStart="FIGURE 4" captionStartId="4.[151,250,1981,2005]" captionTargetBox="[269,1319,1376,1952]" captionTargetId="figure-61@4.[269,1319,1376,1958]" captionTargetPageId="4" captionText="FIGURE 4. Indonemoura geminus Zwick, 1977 male. a: terminalia, dorsal view; b: cleared epiproct, dorsal view; c: cleared paraproct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849248" httpUri="https://zenodo.org/record/13849248/files/figure.png" pageId="1" pageNumber="591">Fig. 4</figureCitation>
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<taxonomicName id="4C2D4D1BFFDDFFA97499F9233251F92A" ID-CoL="3PNF5" authority="Zwick, 1977: 104" authorityName="Zwick" authorityPageNumber="104" authorityYear="1977" box="[151,567,1762,1786]" class="Insecta" family="Nemouridae" genus="Indonemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B959EA8AFFDDFFA97499F923311CF92A" box="[151,378,1762,1786]" italics="true" pageId="1" pageNumber="591">Indonemoura geminus</emphasis>
<bibRefCitation id="EFBC4B69FFDDFFA9758FF9233251F92A" author="Zwick, P." box="[385,567,1762,1786]" pageId="1" pageNumber="591" pagination="85 - 134" refId="ref2988" refString="Zwick, P. (1977) Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Plecoptera. Entomologica Basiliensia, 2, 85 - 134." type="journal article" year="1977">Zwick, 1977: 104</bibRefCitation>
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;
<treatmentCitation id="0A8C1089FFDDFFA9764CF9233323F92A" author="Muranyi, D. &amp; Li, W. H." box="[578,837,1762,1786]" page="173" pageId="1" pageNumber="591" year="2013">
<bibRefCitation id="EFBC4B69FFDDFFA9764CF9233323F92A" author="Muranyi, D. &amp; Li, W. H." box="[578,837,1762,1786]" pageId="1" pageNumber="591" pagination="167 - 177" refId="ref2765" refString="Muranyi, D. &amp; Li, W. H. (2013) Two new species of stoneflies (Plecoptera: Nemouridae) from Northeastern India, with a checklist of the family in the Indian Subcontinent. Zootaxa, 3694 (2), 167 - 177. https: // doi. org / 10.11646 / zootaxa. 3694.2.6" type="journal article" year="2013">Murányi &amp; Li, 2013: 173</bibRefCitation>
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.
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<emphasis id="B959EA8AFFDDFFA97499F8E9311DF892" bold="true" box="[151,379,1832,1858]" pageId="1" pageNumber="591">Examined material</emphasis>
.
<materialsCitation id="3B453CC5FFDDFFA97589F8E8327AF8B7" collectingDate="2019-06-13" collectionCode="CAU" collectorName="Qicheng Yang." county="Chayu County" elevation="2394" latitude="28.6669" location="Shuidianzhan" longLatPrecision="6" longitude="97.475" pageId="1" pageNumber="591" specimenCount="1" specimenCount-male="1" stateProvince="Xizang">
<specimenCount id="9D2BFD11FFDDFFA97589F8E831B6F892" box="[391,464,1832,1859]" count="1" pageId="1" pageNumber="591" type="male">1 male</specimenCount>
(
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),
<collectingRegion id="49E9F87AFFDDFFA97621F8E932E5F892" box="[559,643,1832,1858]" country="China" name="Xizang" pageId="1" pageNumber="591">Xizang</collectingRegion>
,
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,
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,
<quantity id="4CD59B7DFFDDFFA977D5F8E83457F893" box="[987,1073,1833,1859]" metricMagnitude="3" metricUnit="m" metricValue="2.394" pageId="1" pageNumber="591" unit="m" value="2394.0">
<elevation id="0000D1ABFFDDFFA977D5F8E83457F893" box="[987,1073,1833,1859]" metricMagnitude="3" metricUnit="m" metricValue="2.394" pageId="1" pageNumber="591" unit="m" value="2394.0">2394 m</elevation>
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,
<date id="FF931058FFDDFFA97034F8E834DAF893" box="[1082,1212,1833,1859]" pageId="1" pageNumber="591" value="2019-06-13">
<collectingDate id="EFD7E9B0FFDDFFA97034F8E834DAF893" box="[1082,1212,1833,1859]" pageId="1" pageNumber="591" value="2019-06-13">2019.VI.13</collectingDate>
</date>
,
<collectingMethod id="526C4E8FFFDDFFA970C8F8E9354AF893" box="[1222,1324,1832,1859]" pageId="1" pageNumber="591">light trap</collectingMethod>
,
<geoCoordinate id="EE19505FFFDDFFA97138F8E930C8F8B7" degrees="28.6669" direction="north" orientation="latitude" pageId="1" pageNumber="591" precision="5" value="28.6669">28.6669° N</geoCoordinate>
,
<geoCoordinate id="EE19505FFFDDFFA974B7F88D315CF8B7" box="[185,314,1868,1895]" degrees="97.4750" direction="east" orientation="longitude" pageId="1" pageNumber="591" precision="5" value="97.475">97.4750° E</geoCoordinate>
, leg.
<collectorName id="26D8534EFFDDFFA97578F88D327AF8B7" box="[374,540,1868,1895]" pageId="1" pageNumber="591">Qicheng Yang.</collectorName>
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</paragraph>
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<subSubSection id="C3376513FFDDFFAA74C9F8B032BDFF05" lastPageId="2" lastPageNumber="592" pageId="1" pageNumber="591" type="discussion">
<paragraph id="8B923698FFDDFFAA74C9F8B032BDFF05" blockId="1.[151,1437,1832,2039]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="592" pageId="1" pageNumber="591">
<emphasis id="B959EA8AFFDDFFA974C9F8B03152F85B" bold="true" box="[199,308,1905,1931]" pageId="1" pageNumber="591">Remarks</emphasis>
. Tergum IX, epiproct, and paraproct characteristics (
<figureCitation id="13162A1DFFDDFFA97780F8B13463F85B" box="[910,1029,1904,1931]" captionStart="FIGURE 4" captionStartId="4.[151,250,1981,2005]" captionTargetBox="[269,1319,1376,1952]" captionTargetId="figure-61@4.[269,1319,1376,1958]" captionTargetPageId="4" captionText="FIGURE 4. Indonemoura geminus Zwick, 1977 male. a: terminalia, dorsal view; b: cleared epiproct, dorsal view; c: cleared paraproct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849248" httpUri="https://zenodo.org/record/13849248/files/figure.png" pageId="1" pageNumber="591">Figs. 4ac</figureCitation>
) match well with the description of the
<typeStatus id="5496883AFFDDFFA974CFF8553145F87E" box="[193,291,1940,1966]" pageId="1" pageNumber="591" type="holotype">holotype</typeStatus>
male, indicating that the specimen examined above belongs to
<taxonomicName id="4C2D4D1BFFDDFFA977FAF854340EF87E" authorityName="Zwick" authorityYear="1977" box="[1012,1128,1941,1966]" class="Insecta" family="Nemouridae" genus="Indonemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B959EA8AFFDDFFA977FAF854340EF87E" box="[1012,1128,1941,1966]" italics="true" pageId="1" pageNumber="591">I. geminus</emphasis>
</taxonomicName>
and not its close relative
<taxonomicName id="4C2D4D1BFFDDFFA97182F85431C9F802" authority="(Aubert 1967)" baseAuthorityName="Aubert" baseAuthorityYear="1967" class="Insecta" family="Nemouridae" genus="Indonemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="shergaoni">
<emphasis id="B959EA8AFFDDFFA97182F8543160F802" italics="true" pageId="1" pageNumber="591">I. shergaoni</emphasis>
(
<bibRefCitation id="EFBC4B69FFDDFFA97518F87931C1F802" author="Aubert, J." box="[278,423,1976,2002]" pageId="1" pageNumber="591" pagination="209 - 253" refId="ref2153" refString="Aubert, J. (1967) Les Nemouridae de l'Assam (Plecopteres). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 39, 209 - 253." type="journal article" year="1967">Aubert 1967</bibRefCitation>
)
</taxonomicName>
as discussed by
<bibRefCitation id="EFBC4B69FFDDFFA97661F879336AF803" author="Zwick, P." box="[623,780,1976,2003]" pageId="1" pageNumber="591" pagination="85 - 134" refId="ref2988" refString="Zwick, P. (1977) Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Plecoptera. Entomologica Basiliensia, 2, 85 - 134." type="journal article" year="1977">Zwick (1977)</bibRefCitation>
. Males of
<taxonomicName id="4C2D4D1BFFDDFFA9778BF878339CF802" authorityName="Zwick" authorityYear="1977" box="[901,1018,1977,2002]" class="Insecta" family="Nemouridae" genus="Indonemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B959EA8AFFDDFFA9778BF878339CF802" box="[901,1018,1977,2002]" italics="true" pageId="1" pageNumber="591">I. geminus</emphasis>
</taxonomicName>
are characterized by tergum IX with a symmetrical posterior margin (
<figureCitation id="13162A1DFFDDFFA97609F81D323CF827" box="[519,602,2012,2039]" captionStart="FIGURE 4" captionStartId="4.[151,250,1981,2005]" captionTargetBox="[269,1319,1376,1952]" captionTargetId="figure-61@4.[269,1319,1376,1958]" captionTargetPageId="4" captionText="FIGURE 4. Indonemoura geminus Zwick, 1977 male. a: terminalia, dorsal view; b: cleared epiproct, dorsal view; c: cleared paraproct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849248" httpUri="https://zenodo.org/record/13849248/files/figure.png" pageId="1" pageNumber="591">Fig. 4a</figureCitation>
). The epiproct has stout, curved flagellum (
<figureCitation id="13162A1DFFDDFFA97048F81D34FDF826" box="[1094,1179,2012,2038]" captionStart="FIGURE 4" captionStartId="4.[151,250,1981,2005]" captionTargetBox="[269,1319,1376,1952]" captionTargetId="figure-61@4.[269,1319,1376,1958]" captionTargetPageId="4" captionText="FIGURE 4. Indonemoura geminus Zwick, 1977 male. a: terminalia, dorsal view; b: cleared epiproct, dorsal view; c: cleared paraproct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849248" httpUri="https://zenodo.org/record/13849248/files/figure.png" pageId="1" pageNumber="591">Fig. 4b</figureCitation>
). The outer lobe of the paraproct is L-shaped with an enlarged rounded apex (
<figureCitation id="13162A1DFFDEFFAA7709FF56333DFF61" box="[775,859,151,177]" captionStart="FIGURE 4" captionStartId="4.[151,250,1981,2005]" captionTargetBox="[269,1319,1376,1952]" captionTargetId="figure-61@4.[269,1319,1376,1958]" captionTargetPageId="4" captionText="FIGURE 4. Indonemoura geminus Zwick, 1977 male. a: terminalia, dorsal view; b: cleared epiproct, dorsal view; c: cleared paraproct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849248" httpUri="https://zenodo.org/record/13849248/files/figure.png" pageId="2" pageNumber="592">Fig. 4c</figureCitation>
). This species was previously known from
<collectingCountry id="F33A7608FFDEFFAA7145FF5635FAFF61" box="[1355,1436,151,177]" name="Bhutan" pageId="2" pageNumber="592">Bhutan</collectingCountry>
(
<bibRefCitation id="EFBC4B69FFDEFFAA7491FF7A31EFFF05" author="Muranyi, D. &amp; Li, W. H." box="[159,393,187,213]" pageId="2" pageNumber="592" pagination="167 - 177" refId="ref2765" refString="Muranyi, D. &amp; Li, W. H. (2013) Two new species of stoneflies (Plecoptera: Nemouridae) from Northeastern India, with a checklist of the family in the Indian Subcontinent. Zootaxa, 3694 (2), 167 - 177. https: // doi. org / 10.11646 / zootaxa. 3694.2.6" type="journal article" year="2013">Murányi &amp; Li, 2013</bibRefCitation>
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<collectingCountry id="F33A7608FFDEFFAA769FFF7A32BEFF05" box="[657,728,187,213]" name="China" pageId="2" pageNumber="592">China</collectingCountry>
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</paragraph>
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</treatment>
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