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<document id="F5C976885C87292DD01D2FC41F9F79A8" ID-DOI="10.5281/zenodo.293434" ID-GBIF-Dataset="90a8372c-5dc2-4af5-abc6-d6974ab05a11" ID-ISSN="1175-5326" ID-Zenodo-Dep="293434" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460479489793" checkinUser="plazi" docAuthor="Theron, Elizabeth Omukunda Pieter D &amp; Ueckermann, E A" docDate="2012" docId="03FCB14BFFB4FFEDE2D2FB8EFAB0FC1C" docLanguage="en" docName="zt03304p024.pdf" docOrigin="Zootaxa 3304" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Spinibdella namibiensis Theron &amp; Ueckermann, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="FFC5C933FFBAFFFCE245FFFFFF98FFB0" masterDocTitle="Spinibdella Thor (Acari: Bdellidae) from southern Africa: descriptions of five new species and the redescription of S. thori (Meyer &amp; Ryke)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="15" updateTime="1698307322454" updateUser="plazi">
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<mods:title id="373D78036DFDDF784BF58ED28F6261EA">Spinibdella Thor (Acari: Bdellidae) from southern Africa: descriptions of five new species and the redescription of S. thori (Meyer &amp; Ryke)</mods:title>
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<mods:namePart id="F7FEEE60EA765350EC56B1FFE02F9006">Theron, Elizabeth Omukunda Pieter D</mods:namePart>
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<mods:namePart id="115AD07BE4E687E445AC63D394104231">Ueckermann, E A</mods:namePart>
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<mods:title id="0CD5623F6D3C31B50F11CF52B0A299DB">Zootaxa</mods:title>
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<mods:date id="39BC003F5F40BB9C30D21C819E6B2DDB">2012</mods:date>
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<treatment id="03FCB14BFFB4FFEDE2D2FB8EFAB0FC1C" ID-DOI="http://doi.org/10.5281/zenodo.6168967" ID-GBIF-Taxon="119516920" ID-Zenodo-Dep="6168967" LSID="urn:lsid:plazi:treatment:03FCB14BFFB4FFEDE2D2FB8EFAB0FC1C" httpUri="http://treatment.plazi.org/id/03FCB14BFFB4FFEDE2D2FB8EFAB0FC1C" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="C34F53D6FFB4FFF2E2D2FB8EFE8CFB1C" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BEA005DFFB4FFF2E2D2FB8EFC7EFB3B" blockId="14.[151,998,1137,1196]" box="[151,998,1137,1163]" pageId="14" pageNumber="15">
<heading id="D0A2B731FFB4FFF2E2D2FB8EFC7EFB3B" bold="true" box="[151,998,1137,1163]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B921DC4FFFB4FFF2E2D2FB8EFC7EFB3B" bold="true" box="[151,998,1137,1163]" pageId="14" pageNumber="15">
<taxonomicName id="4C557BDEFFB4FFF2E2D2FB8EFE20FB3B" box="[151,440,1137,1163]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov.">
<emphasis id="B921DC4FFFB4FFF2E2D2FB8EFE20FB3B" bold="true" box="[151,440,1137,1163]" italics="true" pageId="14" pageNumber="15">Spinibdella namibiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2126134FFB4FFF2E3FAFB8EFD85FB3B" box="[447,541,1137,1163]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
Omukunda, Theron &amp; Ueckermann
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA005DFFB4FFF2E2D2FB6BFE8CFB1C" blockId="14.[151,998,1137,1196]" box="[151,276,1172,1196]" pageId="14" pageNumber="15">
<figureCitation id="136E1CD8FFB4FFF2E2D2FB6BFE8CFB1C" box="[151,276,1172,1196]" captionStart-0="FIGURES 37 42" captionStart-1="FIGURES 43 44" captionStartId-0="15.[151,264,1629,1652]" captionStartId-1="16.[151,264,1663,1686]" captionTargetBox-0="[189,1380,304,1582]" captionTargetBox-1="[153,1366,291,1662]" captionTargetId-0="figure@15.[151,1436,266,1609]" captionTargetId-1="figure@16.[151,1436,259,1671]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 37 42. Spinibdella namibiensis sp. n.: 37. Female chelicera; 38. Female palp; 39. Female hypostome; 40. Female hypostome; 41. Female dorsal view; 42. Female ventral view." captionText-1="FIGURES 43 44. Spinibdella namibiensis sp. n.: 43. Female leg I; 44. Female leg II." httpUri-0="https://zenodo.org/record/293443/files/figure.png" httpUri-1="https://zenodo.org/record/294177/files/figure.png" pageId="14" pageNumber="15">Figs 3744</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F53D6FFB4FFEDE2D2FB23FA04FDF4" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="description">
<paragraph id="8BEA005DFFB4FFF2E2D2FB23FB75FA1B" blockId="14.[151,1437,1244,2003]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E2D2FB23FE11FB45" bold="true" box="[151,393,1244,1269]" pageId="14" pageNumber="15">Description. Female</emphasis>
(n = 9). Dimensions: length including gnathosoma 8651060 (890); width 311426 (422); length of gnathosoma 223233 (233); length of chelicerae 204210 (206);
<emphasis id="B921DC4FFFB4FFF2E1B1FAFEFBB2FAA8" box="[1012,1066,1281,1304]" italics="true" pageId="14" pageNumber="15">DHS</emphasis>
1519 (15); length of palp segments:
<date id="FFEB269DFFB4FFF2E2ACFADAFE80FA8C" box="[233,280,1316,1340]" pageId="14" pageNumber="15" value="1913-01">I 13</date>
; II + III 191214 (195);
<date id="FFEB269DFFB4FFF2E069FADAFD0FFA8C" box="[556,663,1316,1340]" pageId="14" pageNumber="15" value="1919-04-25">IV 1925</date>
(20);
<date id="FFEB269DFFB4FFF2E099FADAFC8BFA8C" box="[732,787,1316,1340]" pageId="14" pageNumber="15" value="1934-05">V 34</date>
36 (36);
<emphasis id="B921DC4FFFB4FFF2E1C7FADAFC05FA8D" box="[898,925,1317,1341]" italics="true" pageId="14" pageNumber="15">de</emphasis>
254271 (263);
<emphasis id="B921DC4FFFB4FFF2E61DFAD9FBE4FA8D" box="[1112,1148,1318,1341]" italics="true" pageId="14" pageNumber="15">ves</emphasis>
172187 (172); length of legs: I 304344 (331); II 319365 (323); III 397441 (389); IV 511564 (562); length of setae:
<emphasis id="B921DC4FFFB4FFF2E697FAB5FB7EFAD1" box="[1234,1254,1354,1377]" italics="true" pageId="14" pageNumber="15">vi</emphasis>
134143 (134);
<emphasis id="B921DC4FFFB4FFF2E2D2FA91FF28FA35" box="[151,176,1390,1413]" italics="true" pageId="14" pageNumber="15">ve</emphasis>
5254 (51);
<emphasis id="B921DC4FFFB4FFF2E307FA91FEF9FA35" box="[322,353,1390,1413]" italics="true" pageId="14" pageNumber="15">sci</emphasis>
99132 (100);
<emphasis id="B921DC4FFFB4FFF2E04AFA91FDABFA35" box="[527,563,1390,1413]" italics="true" pageId="14" pageNumber="15">sce</emphasis>
5559 (59);
<emphasis id="B921DC4FFFB4FFF2E080FA91FD42FA39" box="[709,730,1390,1417]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
54;
<emphasis id="B921DC4FFFB4FFF2E149FA91FCB8FA39" box="[780,800,1390,1417]" italics="true" pageId="14" pageNumber="15">c2</emphasis>
7686 (79);
<emphasis id="B921DC4FFFB4FFF2E1F7FA92FC58FA35" box="[946,960,1389,1413]" italics="true" pageId="14" pageNumber="15">d</emphasis>
56;
<emphasis id="B921DC4FFFB4FFF2E1B7FA91FC66FA35" box="[1010,1022,1390,1413]" italics="true" pageId="14" pageNumber="15">e</emphasis>
6162 (63);
<emphasis id="B921DC4FFFB4FFF2E6D4FA92FB3EFA35" box="[1169,1190,1389,1413]" italics="true" pageId="14" pageNumber="15">f1</emphasis>
7374 (72);
<emphasis id="B921DC4FFFB4FFF2E77CFA92FAD6FA35" box="[1337,1358,1389,1413]" italics="true" pageId="14" pageNumber="15">f2</emphasis>
7077 (77);
<emphasis id="B921DC4FFFB4FFF2E291FA6CFF68FA1B" box="[212,240,1427,1451]" italics="true" pageId="14" pageNumber="15">h1</emphasis>
8893 (81);
<emphasis id="B921DC4FFFB4FFF2E3C7FA6CFE06FA1B" box="[386,414,1427,1451]" italics="true" pageId="14" pageNumber="15">h2</emphasis>
8688 (83); distance:
<emphasis id="B921DC4FFFB4FFF2E0DFFA6BFD48FA1B" box="[666,720,1428,1451]" italics="true" pageId="14" pageNumber="15">
vi
<date id="FFEB269DFFB4FFF2E0F9FA6BFD48FA1B" box="[700,720,1428,1451]" pageId="14" pageNumber="15" value="1944-06">vi</date>
</emphasis>
4448 (44); first interspace (
<emphasis id="B921DC4FFFB4FFF2E652FA6BFBB4FA1F" box="[1047,1068,1428,1455]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
to
<emphasis id="B921DC4FFFB4FFF2E609FA6CFBC2FA1B" box="[1100,1114,1427,1451]" italics="true" pageId="14" pageNumber="15">d</emphasis>
) 7897 (81).
</paragraph>
<paragraph id="8BEA005DFFB4FFF2E28CFA45FC74F936" blockId="14.[151,1437,1244,2003]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E28CFA45FEFAFA63" bold="true" box="[201,354,1466,1491]" pageId="14" pageNumber="15">Gnathosoma</emphasis>
(
<figureCitation id="136E1CD8FFB4FFF2E337FA45FE6EFA62" box="[370,502,1466,1490]" captionStart="FIGURES 37 42" captionStartId="15.[151,264,1629,1652]" captionTargetBox="[189,1380,304,1582]" captionTargetId="figure@15.[151,1436,266,1609]" captionTargetPageId="15" captionText="FIGURES 37 42. Spinibdella namibiensis sp. n.: 37. Female chelicera; 38. Female palp; 39. Female hypostome; 40. Female hypostome; 41. Female dorsal view; 42. Female ventral view." httpUri="https://zenodo.org/record/293443/files/figure.png" pageId="14" pageNumber="15">Figs 3739</figureCitation>
). Chelicerae have rough longitudinal striations and a pair of strong setae dorsally. Both movable and fixed digits straight with former longest. Setae inserted as shown in figure 37. Palp extends beyond hypostome. Palp tibiotarsus has a long blunt solenidion ventrally. Distal setae comparatively long as shown in figure 38. Palp chaetotaxy from trochanter to tibiotarsus as follows: 0-9-1-4-4 sts, 1ω, 1
<emphasis id="B921DC4FFFB4FFF2E6DBF9D9FB5BF98E" box="[1182,1219,1574,1598]" italics="true" pageId="14" pageNumber="15">des</emphasis>
, 1
<emphasis id="B921DC4FFFB4FFF2E6ADF9D8FA93F98E" box="[1256,1291,1575,1598]" italics="true" pageId="14" pageNumber="15">ves</emphasis>
. Hypostome has weak longitudinal striations while base has fine strong transverse striations. Two pairs of strong hypostomal setae inserted as shown in figure 39. Two pairs of short adoral setae present.
</paragraph>
<paragraph id="8BEA005DFFB4FFF2E28CF96CFA8BF81C" blockId="14.[151,1437,1244,2003]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E28CF96CFEB2F91B" bold="true" box="[201,298,1683,1707]" pageId="14" pageNumber="15">Dorsum</emphasis>
(
<figureCitation id="136E1CD8FFB4FFF2E37DF96DFE24F91A" box="[312,444,1682,1706]" captionStart="FIGURES 37 42" captionStartId="15.[151,264,1629,1652]" captionTargetBox="[189,1380,304,1582]" captionTargetId="figure@15.[151,1436,266,1609]" captionTargetPageId="15" captionText="FIGURES 37 42. Spinibdella namibiensis sp. n.: 37. Female chelicera; 38. Female palp; 39. Female hypostome; 40. Female hypostome; 41. Female dorsal view; 42. Female ventral view." httpUri="https://zenodo.org/record/293443/files/figure.png" pageId="14" pageNumber="15">Figs 4041</figureCitation>
). Naso simple, median eye detectable. Prodorsal setae simple. Setae
<emphasis id="B921DC4FFFB4FFF2E6F9F96CFB48F91A" box="[1212,1232,1683,1706]" italics="true" pageId="14" pageNumber="15">vi</emphasis>
and
<emphasis id="B921DC4FFFB4FFF2E742F96CFAB3F91A" box="[1287,1323,1683,1706]" italics="true" pageId="14" pageNumber="15">sce</emphasis>
relatively long. Setae
<emphasis id="B921DC4FFFB4FFF2E35FF948FEABF97E" box="[282,307,1719,1742]" italics="true" pageId="14" pageNumber="15">ve</emphasis>
, straight, long and occur in anterior half of prodorsum as shown in figure 40. Setae
<emphasis id="B921DC4FFFB4FFF2E698F948FA99F97E" box="[1245,1281,1719,1742]" italics="true" pageId="14" pageNumber="15">sce</emphasis>
straight, long and slightly thicker than
<emphasis id="B921DC4FFFB4FFF2E3F6F924FE5FF942" box="[435,455,1755,1778]" italics="true" pageId="14" pageNumber="15">vi</emphasis>
. Middle prodorsum has roughly broken striations as shown in figure 41. Internal apodemes halfway between
<emphasis id="B921DC4FFFB4FFF2E3EEF900FE27F8A6" box="[427,447,1791,1814]" italics="true" pageId="14" pageNumber="15">vi</emphasis>
and
<emphasis id="B921DC4FFFB4FFF2E3B3F900FD8DF8A6" box="[502,533,1791,1814]" italics="true" pageId="14" pageNumber="15">sci</emphasis>
. Two pairs of detectable eyes separated by distance equal to diameter of smaller posterior pair. Longitudinal striations between anterior and posterior pairs of eyes. Longitudinal striations on dorsolateral prodorsum. Sejugal groove conspicuous. Setae
<emphasis id="B921DC4FFFB4FFF2E149F8B8FCB8F8D2" box="[780,800,1863,1890]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
about 2/3 first interspace. Setae
<emphasis id="B921DC4FFFB4FFF2E6CFF8B8FB0EF8EE" box="[1162,1174,1863,1886]" italics="true" pageId="14" pageNumber="15">c</emphasis>
<date id="FFEB269DFFB4FFF2E6D3F8AAFB4CF8EF" box="[1174,1236,1863,1890]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E6D3F8AAFB06F8D2" box="[1174,1182,1877,1890]" italics="true" pageId="14" pageNumber="15">2</emphasis>
may
</date>
be pilose in some individuals and long but not as in
<emphasis id="B921DC4FFFB4FFF2E052F892FCB3F835" box="[535,811,1901,1925]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C557BDEFFB4FFF2E052F892FCB0F835" box="[535,808,1901,1925]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pongolensis">Spinibdella pongolensis</taxonomicName>
.
</emphasis>
Setae
<emphasis id="B921DC4FFFB4FFF2E133F892FC1EF839" box="[886,902,1901,1929]" italics="true" pageId="14" pageNumber="15">f2</emphasis>
occur on horizontal line with
<emphasis id="B921DC4FFFB4FFF2E69FF892FB68F839" box="[1242,1264,1901,1929]" italics="true" pageId="14" pageNumber="15">h1</emphasis>
strikingly long in some individuals. In some individuals, anal cleft not dorsally visible. Setae
<emphasis id="B921DC4FFFB4FFF2E647F86AFBBBF800" box="[1026,1059,1941,1968]" italics="true" pageId="14" pageNumber="15">ps1</emphasis>
and
<emphasis id="B921DC4FFFB4FFF2E61CF86AFBE2F800" box="[1113,1146,1941,1968]" italics="true" pageId="14" pageNumber="15">ps2</emphasis>
finely pilose.
</paragraph>
<paragraph id="8BEA005DFFB4FFF3E28CF844FB4DFF47" blockId="14.[151,1437,1244,2003]" lastBlockId="15.[151,1436,151,252]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E28CF844FE81F863" bold="true" box="[201,281,1979,2003]" pageId="14" pageNumber="15">Venter</emphasis>
(
<figureCitation id="136E1CD8FFB4FFF2E369F845FE1CF862" box="[300,388,1978,2002]" captionStart="FIGURES 37 42" captionStartId="15.[151,264,1629,1652]" captionTargetBox="[189,1380,304,1582]" captionTargetId="figure@15.[151,1436,266,1609]" captionTargetPageId="15" captionText="FIGURES 37 42. Spinibdella namibiensis sp. n.: 37. Female chelicera; 38. Female palp; 39. Female hypostome; 40. Female hypostome; 41. Female dorsal view; 42. Female ventral view." httpUri="https://zenodo.org/record/293443/files/figure.png" pageId="14" pageNumber="15">Fig. 42</figureCitation>
). All ventral setae pilose. Forty long blunt pilose aggenitals. First setae unpaired between coxae III as shown in figure 42. Ten pairs of linearly arranged blunt pilose genital setae equal in length on genital plates. Three pairs of small genital discs, first pair in anterior half and rest in posterior half. Eighteen setae occur on ovipositor. Three pairs of anal setae present (
<emphasis id="B921DC4FFFB5FFF3E0CBFF1FFC8EFF4C" box="[654,790,224,252]" italics="true" pageId="15" pageNumber="16">ad1, ad2, an1</emphasis>
). Anal cleft closed in some individuals.
</paragraph>
<caption id="DF2A50D5FFB5FFF3E2D2F9A2FC9FF922" httpUri="https://zenodo.org/record/293443/files/figure.png" pageId="15" pageNumber="16" targetBox="[189,1380,304,1582]" targetPageId="15">
<paragraph id="8BEA005DFFB5FFF3E2D2F9A2FC9FF922" blockId="15.[151,1436,1629,1682]" pageId="15" pageNumber="16">
<emphasis id="B921DC4FFFB5FFF3E2D2F9A2FECDF9C3" bold="true" box="[151,341,1629,1652]" pageId="15" pageNumber="16">FIGURES 3742.</emphasis>
<taxonomicName id="4C557BDEFFB5FFF3E319F9A1FDC8F9C4" box="[348,592,1630,1652]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov.">
<emphasis id="B921DC4FFFB5FFF3E319F9A1FDC8F9C4" box="[348,592,1630,1652]" italics="true" pageId="15" pageNumber="16">Spinibdella namibiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2126134FFB5FFF3E012F9A0FD09F9C4" box="[599,657,1631,1652]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
: 37. Female chelicera; 38. Female palp; 39. Female hypostome; 40. Female hypostome; 41. Female dorsal view; 42. Female ventral view.
</paragraph>
</caption>
<paragraph id="8BEA005DFFB5FFECE28CF945FD74FF47" blockId="15.[151,1437,1722,2035]" lastBlockId="16.[151,1437,151,248]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B921DC4FFFB5FFF3E28CF945FE99F962" bold="true" box="[201,257,1722,1746]" pageId="15" pageNumber="16">Legs</emphasis>
(
<figureCitation id="136E1CD8FFB5FFF3E354F945FE0AF962" box="[273,402,1722,1746]" captionStart="FIGURES 43 44" captionStartId="16.[151,264,1663,1686]" captionTargetBox="[153,1366,291,1662]" captionTargetId="figure@16.[151,1436,259,1671]" captionTargetPageId="16" captionText="FIGURES 43 44. Spinibdella namibiensis sp. n.: 43. Female leg I; 44. Female leg II." httpUri="https://zenodo.org/record/294177/files/figure.png" pageId="15" pageNumber="16">Figs 4344</figureCitation>
). Legs shorter than idiosoma. Some individuals have leg II shorter than leg I. Rest increase progressively. All coxal setae pilose. Other leg segments may have either both pilose and simple or only simple setae. Basifemora and telofemora III and IV have a macroseta each. Genua I and II have duplex setae each in distal half while III have a reduced attenuate solenidion. Tibiae I have two attenuate solenidia, one shorter than other, a solid attenuate famulus and a trichobothrium distally as shown in figure 43. Tibiae II have a reduced blunt solenidion each as seen in figure 44. Tibiae III have an attenuate solenidion, tibiae IV have a trichobothrium. Setae on tarsi smooth. Tarsi I have one attenuate solenidion proximally and two blunt solenidia distally. Pretarsi longer than claws which lack rays. Tarsus II has a simple solid famulus proximally and two blunt solenidia distally. Tarsus III has a proximal trichobothrium. Tarsus IV has a proximal trichobothrium. Leg chaetotaxy as follows: coxae
<date id="FFEB269DFFB5FFECE722F824FF7AFF1F" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" value="2010-01-08" valueMax="2010-04-06" valueMin="2010-01-08">IIV 10-8-6</date>
-6sts; trochanters
<date id="FFEB269DFFAAFFECE3ECFF67FD9FFF1F" box="[425,519,151,175]" pageId="16" pageNumber="17">IIV 1-1</date>
-2-1 sts; basifemora
<date id="FFEB269DFFAAFFECE0B4FF67FCE1FF1F" box="[753,889,151,175]" pageId="16" pageNumber="17" value="2010-01-09" valueMax="2010-04-08" valueMin="2010-01-09">IIV 10-9-8</date>
-3 sts; telofemora
<date id="FFEB269DFFAAFFECE606FF67FB3DFF1F" box="[1091,1189,151,175]" pageId="16" pageNumber="17">IIV 7-7</date>
-4-5 sts; genua
<date id="FFEB269DFFAAFFECE714FF67FA03FF1F" box="[1361,1435,151,175]" pageId="16" pageNumber="17">IIV 6</date>
sts, 1duplex -6 sts, 1duplex -6 sts, 1σ -7 sts; tibiae
<date id="FFEB269DFFAAFFECE093FF42FCB7FF64" box="[726,815,188,212]" pageId="16" pageNumber="17" value="1914-01" valueMax="1914-04-31" valueMin="1914-01-14">IIV 14</date>
sts, 2φ, 1ĸ, 1tr -14 sts, 1φ -14 sts, 1φ -14 sts, 1tr; tarsi
<date id="FFEB269DFFAAFFECE2D2FF1FFF77FF47" box="[151,239,223,247]" pageId="16" pageNumber="17" value="1930-01" valueMax="1930-04-31" valueMin="1930-01-30">IIV 30</date>
sts, 3ω-30 sts, 2ω, 1 ĸ -30 sts, 1tr -27 sts, 1tr.
</paragraph>
<caption id="DF2A50D5FFAAFFECE2D2F980FC63F925" box="[151,1019,1663,1686]" httpUri="https://zenodo.org/record/294177/files/figure.png" pageId="16" pageNumber="17" targetBox="[153,1366,291,1662]" targetPageId="16">
<paragraph id="8BEA005DFFAAFFECE2D2F980FC63F925" blockId="16.[151,1019,1663,1686]" box="[151,1019,1663,1686]" pageId="16" pageNumber="17">
<emphasis id="B921DC4FFFAAFFECE2D2F980FECBF925" bold="true" box="[151,339,1663,1686]" pageId="16" pageNumber="17">FIGURES 4344.</emphasis>
<taxonomicName id="4C557BDEFFAAFFECE31FF980FDD4F925" box="[346,588,1663,1685]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="namibiensis" status="sp. nov.">
<emphasis id="B921DC4FFFAAFFECE31FF980FDD4F925" box="[346,588,1663,1685]" italics="true" pageId="16" pageNumber="17">Spinibdella namibiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2126134FFAAFFECE017F97FFD13F925" box="[594,651,1664,1685]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
: 43. Female leg I; 44. Female leg II.
</paragraph>
</caption>
<paragraph id="8BEA005DFFAAFFECE28CF942FD80F8D7" blockId="16.[151,1436,1725,2011]" pageId="16" pageNumber="17">
<emphasis id="B921DC4FFFAAFFECE28CF942FECFF966" bold="true" box="[201,343,1725,1750]" pageId="16" pageNumber="17">Tritonymph</emphasis>
(n = 6). Dimensions: length including gnathosoma 564725; width 290325; length of gnathosoma 180193; length of chelicerae 153178;
<emphasis id="B921DC4FFFAAFFECE0D9F91DFD4AF949" box="[668,722,1762,1785]" italics="true" pageId="16" pageNumber="17">DHS</emphasis>
13; length of palp segments
<date id="FFEB269DFFAAFFECE657F91DFBD9F949" box="[1042,1089,1761,1785]" pageId="16" pageNumber="17" value="2010-01">I 10</date>
; II + III 159160;
<date id="FFEB269DFFAAFFECE74AF91DFAE5F949" box="[1295,1405,1761,1785]" pageId="16" pageNumber="17" value="1919-04-21">IV 1921</date>
;
<date id="FFEB269DFFAAFFECE7CDF91DFF2EF8AD" pageId="16" pageNumber="17" value="1931-05">V 31</date>
;
<emphasis id="B921DC4FFFAAFFECE287F8FAFF70F8AD" box="[194,232,1797,1821]" italics="true" pageId="16" pageNumber="17">des</emphasis>
224;
<emphasis id="B921DC4FFFAAFFECE362F8F9FED3F8AD" box="[295,331,1798,1821]" italics="true" pageId="16" pageNumber="17">ves</emphasis>
138147; length of legs: I 250262; II 246264; III 351353; IV 411579; length of setae:
<emphasis id="B921DC4FFFAAFFECE715F8F9FAFCF8AD" box="[1360,1380,1798,1821]" italics="true" pageId="16" pageNumber="17">vi</emphasis>
115;
<emphasis id="B921DC4FFFAAFFECE2D2F8D5FF28F8F1" box="[151,176,1834,1857]" italics="true" pageId="16" pageNumber="17">ve</emphasis>
46;
<emphasis id="B921DC4FFFAAFFECE2A7F8D5FE99F8F1" box="[226,257,1834,1857]" italics="true" pageId="16" pageNumber="17">sci</emphasis>
115;
<emphasis id="B921DC4FFFAAFFECE304F8D5FEFDF8F1" box="[321,357,1834,1857]" italics="true" pageId="16" pageNumber="17">sce</emphasis>
4448;
<emphasis id="B921DC4FFFAAFFECE387F8D5FE4EF8F5" box="[450,470,1834,1861]" italics="true" pageId="16" pageNumber="17">c1</emphasis>
4446;
<emphasis id="B921DC4FFFAAFFECE076F8D5FDDFF8F5" box="[563,583,1834,1861]" italics="true" pageId="16" pageNumber="17">c2</emphasis>
7176;
<emphasis id="B921DC4FFFAAFFECE0E1F8D6FD2AF8F1" box="[676,690,1833,1857]" italics="true" pageId="16" pageNumber="17">d</emphasis>
45;
<emphasis id="B921DC4FFFAAFFECE0A1F8D5FD68F8F1" box="[740,752,1834,1857]" italics="true" pageId="16" pageNumber="17">e</emphasis>
4950;
<emphasis id="B921DC4FFFAAFFECE10BF8D6FCFBF8F1" box="[846,867,1833,1857]" italics="true" pageId="16" pageNumber="17">f1</emphasis>
57;
<emphasis id="B921DC4FFFAAFFECE1D3F8D6FC33F8F1" box="[918,939,1833,1857]" italics="true" pageId="16" pageNumber="17">f2</emphasis>
5357;
<emphasis id="B921DC4FFFAAFFECE64CF8D6FBBCF8F1" box="[1033,1060,1833,1857]" italics="true" pageId="16" pageNumber="17">h1</emphasis>
7576;
<emphasis id="B921DC4FFFAAFFECE6C7F8D6FB06F8F1" box="[1154,1182,1833,1857]" italics="true" pageId="16" pageNumber="17">h2</emphasis>
6065; distance:
<emphasis id="B921DC4FFFAAFFECE720F8D5FA04F8F1" box="[1381,1436,1834,1857]" italics="true" pageId="16" pageNumber="17">
vi
<date id="FFEB269DFFAAFFECE7CDF8D5FA04F8F1" box="[1416,1436,1834,1857]" pageId="16" pageNumber="17" value="1936-06">vi</date>
</emphasis>
36; first interspace (
<emphasis id="B921DC4FFFAAFFECE332F8AEFE14F8DC" box="[375,396,1873,1900]" italics="true" pageId="16" pageNumber="17">c1</emphasis>
to
<emphasis id="B921DC4FFFAAFFECE3E9F8AFFE22F8D8" box="[428,442,1872,1896]" italics="true" pageId="16" pageNumber="17">d</emphasis>
) 6582.
</paragraph>
<paragraph id="8BEA005DFFAAFFEDE28CF887FB35FEAF" blockId="16.[151,1436,1725,2011]" lastBlockId="17.[151,1437,151,940]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Twenty eight blunt pilose aggenital setae, first unpaired between coxae IV. Five pairs of pilose genital setae linearly arranged on two genital plates. Three pairs of genital discs, proximal pair in anterior half while two other pairs in distal half. Setae
<emphasis id="B921DC4FFFAAFFECE3F9F83FFE44F86B" box="[444,476,1984,2011]" italics="true" pageId="16" pageNumber="17">ps1</emphasis>
on horizontal level with termination of anal cleft, last setae dorsally. Three pairs of anal setae, namely
<emphasis id="B921DC4FFFABFFEDE328FF67FE5CFF04" box="[365,452,152,180]" italics="true" pageId="17" pageNumber="18">ad2, an1</emphasis>
and
<emphasis id="B921DC4FFFABFFEDE3BEFF66FD87FF04" box="[507,543,153,180]" italics="true" pageId="17" pageNumber="18">an2</emphasis>
occur ventrally. Palp basifemora with seven setae. Leg chaetotaxy differs from that of female as follows: coxae
<date id="FFEB269DFFABFFEDE064FF3FFD1EFF67" box="[545,646,191,215]" pageId="17" pageNumber="18">IIV 8-6</date>
-5-4 sts; trochanters
<date id="FFEB269DFFABFFEDE13EFF3FFC46FF67" box="[891,990,191,215]" pageId="17" pageNumber="18">IIV 1-1</date>
-2-1 sts; basifemora
<date id="FFEB269DFFABFFEDE693FF3FFAA5FF67" box="[1238,1341,191,215]" pageId="17" pageNumber="18">IIV 6-8</date>
-7-3 sts; telofemora
<date id="FFEB269DFFABFFEDE352FF1CFEE1FF4A" box="[279,377,226,250]" pageId="17" pageNumber="18">IIV 5-6</date>
-4-4 sts; genua
<date id="FFEB269DFFABFFEDE062FF1CFDEAFF4A" box="[551,626,226,250]" pageId="17" pageNumber="18">IIV 4</date>
sts 1 duplex -4 sts, 1 duplex -4 sts, 1σ -5 sts; tibiae
<date id="FFEB269DFFABFFEDE685FF1CFA80FF4A" box="[1216,1304,226,250]" pageId="17" pageNumber="18" value="2010-01" valueMax="2010-04-31" valueMin="2010-01-10">IIV 10</date>
sts, 2φ, 1ĸ, 1tr -9 sts, 1φ -9 sts, 1φ -11 sts, 1tr; tarsi
<date id="FFEB269DFFABFFEDE012FEF7FD37FEAF" box="[599,687,263,287]" pageId="17" pageNumber="18" value="1925-01" valueMax="1925-04-31" valueMin="1925-01-25">IIV 25</date>
sts, 3ω-24 sts, 2 ω,1 ĸ -22 sts, 1tr -21 sts, 1tr.
</paragraph>
<paragraph id="8BEA005DFFABFFEDE28CFED4FC3CFE01" blockId="17.[151,1437,151,940]" pageId="17" pageNumber="18">
<emphasis id="B921DC4FFFABFFEDE28CFED4FEFDFEF3" bold="true" box="[201,357,299,323]" pageId="17" pageNumber="18">Deutonymph</emphasis>
(n = 1). Dimensions: length including gnathosoma 502; width 218; length of gnathosoma 168; length of chelicerae 143;
<emphasis id="B921DC4FFFABFFEDE3F7FEB0FE70FED6" box="[434,488,335,358]" italics="true" pageId="17" pageNumber="18">DHS</emphasis>
, 10; length of palp segments:
<date id="FFEB269DFFABFFEDE17DFEAFFCFFFED7" box="[824,871,335,359]" pageId="17" pageNumber="18" value="2010-01">I 10</date>
; II + III 134;
<date id="FFEB269DFFABFFEDE1B9FEAFFBD8FED7" box="[1020,1088,335,359]" pageId="17" pageNumber="18" value="1919-04">IV 19</date>
;
<date id="FFEB269DFFABFFEDE60EFEAFFB1DFED7" box="[1099,1157,335,359]" pageId="17" pageNumber="18" value="1927-05">V 27</date>
;
<emphasis id="B921DC4FFFABFFEDE6D5FEB0FB2EFED7" box="[1168,1206,335,359]" italics="true" pageId="17" pageNumber="18">des</emphasis>
185;
<emphasis id="B921DC4FFFABFFEDE6B1FEAFFA80FED7" box="[1268,1304,336,359]" italics="true" pageId="17" pageNumber="18">ves</emphasis>
117; length of legs: I 212; II 203; III 256; IV 309; length of setae:
<emphasis id="B921DC4FFFABFFEDE145FE8BFCA3FE3B" box="[768,827,371,395]" italics="true" pageId="17" pageNumber="18">vive</emphasis>
38;
<emphasis id="B921DC4FFFABFFEDE12BFE8BFC15FE3B" box="[878,909,372,395]" italics="true" pageId="17" pageNumber="18">sci</emphasis>
96;
<emphasis id="B921DC4FFFABFFEDE185FE8BFC7CFE3B" box="[960,996,372,395]" italics="true" pageId="17" pageNumber="18">sce</emphasis>
38;
<emphasis id="B921DC4FFFABFFEDE652FE8BFBB3FE3F" box="[1047,1067,372,399]" italics="true" pageId="17" pageNumber="18">c1</emphasis>
38;
<emphasis id="B921DC4FFFABFFEDE61BFE8BFBEAFE3F" box="[1118,1138,372,399]" italics="true" pageId="17" pageNumber="18">c2</emphasis>
65;
<emphasis id="B921DC4FFFABFFEDE6E0FE8CFB2BFE3B" box="[1189,1203,371,395]" italics="true" pageId="17" pageNumber="18">d</emphasis>
35;
<emphasis id="B921DC4FFFABFFEDE6A3FE8BFB6AFE3B" box="[1254,1266,372,395]" italics="true" pageId="17" pageNumber="18">e</emphasis>
, broken off;
<emphasis id="B921DC4FFFABFFEDE7C3FE8CFA03FE3B" box="[1414,1435,371,395]" italics="true" pageId="17" pageNumber="18">f1</emphasis>
51;
<emphasis id="B921DC4FFFABFFEDE287FE66FF4FFE01" box="[194,215,409,433]" italics="true" pageId="17" pageNumber="18">f2</emphasis>
45;
<emphasis id="B921DC4FFFABFFEDE34CFE66FEBCFE01" box="[265,292,409,433]" italics="true" pageId="17" pageNumber="18">h1</emphasis>
65;
<emphasis id="B921DC4FFFABFFEDE312FE66FEEAFE01" box="[343,370,409,433]" italics="true" pageId="17" pageNumber="18">h2</emphasis>
56; distance:
<emphasis id="B921DC4FFFABFFEDE04AFE65FDA5FE01" box="[527,573,410,433]" italics="true" pageId="17" pageNumber="18">viv</emphasis>
<date id="FFEB269DFFABFFEDE07BFE66FDF3FE01" box="[574,619,409,433]" pageId="17" pageNumber="18" value="1929-01">i 29</date>
; first interspace (
<emphasis id="B921DC4FFFABFFEDE168FE65FCDAFE06" box="[813,834,410,438]" italics="true" pageId="17" pageNumber="18">c1</emphasis>
to
<emphasis id="B921DC4FFFABFFEDE127FE66FCE8FE01" box="[866,880,409,433]" italics="true" pageId="17" pageNumber="18">d</emphasis>
) 61.
</paragraph>
<paragraph id="8BEA005DFFABFFEDE28CFE3FFA04FDF4" blockId="17.[151,1437,151,940]" pageId="17" pageNumber="18">
Sixteen long pilose aggenitals; first pair between coxae IV. One pair of pilose genital setae. Two pairs of genital discs. Venter with three pairs of anal setae as in tritonymph. Leg chaetotaxy differs from that of the female as follows: coxae
<date id="FFEB269DFFABFFEDE304FDF6FE3AFD90" box="[321,418,520,544]" pageId="17" pageNumber="18">IIV 6-4</date>
-4-2 sts; trochanters
<date id="FFEB269DFFABFFEDE0C3FDF6FD7BFD90" box="[646,739,520,544]" pageId="17" pageNumber="18">IIV 1-1</date>
-2-1 sts; basifemora
<date id="FFEB269DFFABFFEDE18EFDF6FBB5FD90" box="[971,1069,520,544]" pageId="17" pageNumber="18">IIV 5-5</date>
-4-1 sts; telofemora
<date id="FFEB269DFFABFFEDE749FDF6FAF6FD90" box="[1292,1390,520,544]" pageId="17" pageNumber="18">IIV 5-4</date>
-4-3 sts; genua
<date id="FFEB269DFFABFFEDE349FDD2FECDFDF4" box="[268,341,556,580]" pageId="17" pageNumber="18">IIV 4</date>
sts, 1 duplex -4 sts, 1 duplex -4, 1σ -4 sts; tarsi
<date id="FFEB269DFFABFFEDE123FDD2FC25FDF4" box="[870,957,556,580]" pageId="17" pageNumber="18" value="1920-01" valueMax="1920-04-31" valueMin="1920-01-20">IIV 20</date>
sts, 3φ -20sts, 2φ, 1ĸ -21sts, 1tr -19sts, 1tr.
</paragraph>
</subSubSection>
<subSubSection id="C34F53D6FFABFFEDE28CFDAEFD4AFD64" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BEA005DFFABFFEDE28CFDAEFD4AFD64" blockId="17.[151,1437,151,940]" pageId="17" pageNumber="18">
<emphasis id="B921DC4FFFABFFEDE28CFDAEFEEFFDD9" bold="true" box="[201,375,592,617]" pageId="17" pageNumber="18">
<typeStatus id="54EEBEFFFFABFFEDE28CFDAEFE9CFDD9" box="[201,260,593,617]" pageId="17" pageNumber="18">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="F34240CDFFABFFEDE33BFDAEFE65FDD9" box="[382,509,593,617]" name="Namibia" pageId="17" pageNumber="18">NAMIBIA</collectingCountry>
:
<typeStatus id="54EEBEFFFFABFFEDE042FDAFFDF7FDD8" box="[519,623,592,616]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
female,
<specimenCount id="9D53CBD4FFABFFEDE094FDAFFCAAFDD8" box="[721,818,592,616]" pageId="17" pageNumber="18" type="female">6 female</specimenCount>
<typeStatus id="54EEBEFFFFABFFEDE17FFDAEFC30FDD9" box="[826,936,593,617]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
, 3 tritonymph
<typeStatus id="54EEBEFFFFABFFEDE609FDAEFB37FDD9" box="[1100,1199,593,617]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
, 1 deutonymph
<typeStatus id="54EEBEFFFFABFFEDE727FDAEFF52FD3D" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
, Gobabeb P.D. Theron;
<collectingCountry id="F34240CDFFABFFEDE39AFD8BFD37FD3C" box="[479,687,628,652]" name="South Africa" pageId="17" pageNumber="18">SOUTH AFRICA</collectingCountry>
,
<emphasis id="B921DC4FFFABFFEDE0F9FD8AFC76FD3D" bold="true" box="[700,1006,628,653]" pageId="17" pageNumber="18">Northern Cape Province:</emphasis>
Springbok, Northern Cape from soil,
<date id="FFEB269DFFABFFEDE2D2FD67FEBCFD00" box="[151,292,664,688]" pageId="17" pageNumber="18" value="1980-03-11">11-III- 1980</date>
, 2
<typeStatus id="54EEBEFFFFABFFEDE302FD66FE3FFD01" box="[327,423,665,689]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
females and 3
<typeStatus id="54EEBEFFFFABFFEDE013FD66FD2EFD01" box="[598,694,665,689]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
tritonymphs, from
<taxonomicName id="4C557BDEFFABFFEDE1D1FD67FA00FD00" authority="Harv." authorityName="Harv." box="[916,1432,664,688]" class="Magnoliopsida" family="Menispermaceae" genus="Antizoma" kingdom="Plantae" order="Ranunculales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="miersiana">
<emphasis id="B921DC4FFFABFFEDE1D1FD67FBEFFD00" box="[916,1143,664,688]" italics="true" pageId="17" pageNumber="18">Antizoma miersiana</emphasis>
Harv. (Menispermaceae)
</taxonomicName>
,
<quantity id="4CADADB8FFABFFEDE2D2FD43FF46FD64" box="[151,222,700,724]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="17" pageNumber="18" unit="km" value="21.0">21 km</quantity>
south of Springbok, D.J. Botha,
<date id="FFEB269DFFABFFEDE00BFD43FD56FD64" box="[590,718,700,724]" pageId="17" pageNumber="18" value="1973-09-27">27-ix-1973</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F53D6FFABFFEDE28CFD1EFAB0FC1C" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BEA005DFFABFFEDE28CFD1EFCDEFD48" blockId="17.[151,1437,151,940]" box="[201,838,736,761]" pageId="17" pageNumber="18">
<emphasis id="B921DC4FFFABFFEDE28CFD1EFED0FD49" bold="true" box="[201,328,737,761]" pageId="17" pageNumber="18">Etymology</emphasis>
. This species is named after the
<typeStatus id="54EEBEFFFFABFFEDE0F0FD1EFD7DFD49" box="[693,741,737,761]" pageId="17" pageNumber="18">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BEA005DFFABFFEDE28CFCFBFAB0FC1C" blockId="17.[151,1437,151,940]" pageId="17" pageNumber="18">
<emphasis id="B921DC4FFFABFFEDE28CFCFBFE52FCAD" bold="true" box="[201,458,772,797]" pageId="17" pageNumber="18">Differential diagnosis</emphasis>
. This species closely resembles
<taxonomicName id="4C557BDEFFABFFEDE100FCFBFB9DFCAC" box="[837,1029,772,796]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="thori">
<emphasis id="B921DC4FFFABFFEDE100FCFBFB9DFCAC" box="[837,1029,772,796]" italics="true" pageId="17" pageNumber="18">Spinibdella thori</emphasis>
</taxonomicName>
but can be recognized by: roughly broken transverse striations medially on the prodorsum, unlike continuous in
<emphasis id="B921DC4FFFABFFEDE655FCD6FBFEFCF0" box="[1040,1126,808,832]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C557BDEFFABFFEDE655FCD6FBC7FCF0" box="[1040,1119,808,832]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="thori">S thori</taxonomicName>
;
</emphasis>
two attenuate solenidia on tibiae I, one which is about half the length of the longest one; tibia II has a reduced blunt solenidion; genua I and II have duplex setae while III have a reduced attenuate solenidion; leg IV strikingly long (over 500 µm). The South African material however has some variations in that the vertical setae are both simple and finely pilose.
</paragraph>
</subSubSection>
</treatment>
</document>