treatments-xml/data/03/EF/87/03EF879CFFAFFFBDFF37FA6191A71384.xml
2024-06-21 12:22:17 +02:00

653 lines
97 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="F6413EC6CC47B04201A40E638D9FBB1C" ID-CLB-Dataset="46619" ID-DOI="10.11646/zootaxa.3755.1.3" ID-GBIF-Dataset="9b22d33f-c1f8-49e7-b123-3877690f282b" ID-ISSN="1175-5326" ID-Zenodo-Dep="252659" ID-ZooBank="0486D86E-EEFF-453E-BD77-395094BED088" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460372518285" checkinUser="plazi" docAuthor="Chani-Posse, Mariana" docDate="2014" docId="03EF879CFFAFFFBDFF37FA6191A71384" docLanguage="en" docName="zt03755p086.pdf" docOrigin="Zootaxa 3755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philonthina" docType="key" docVersion="8" masterDocId="FFD6FFE4FFA3FFB3FFA0FFFA943C1748" masterDocTitle="An illustrated key to the New World genera of Philonthina Kirby (Coleoptera: Staphylinidae), with morphological, taxonomical and distributional notes" updateTime="1698295903052" updateUser="ExternalLinkService">
<mods:mods id="BC8102537236B4A73A3231B5D1C1099B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DDD9B2763B62040B55BC280EC450EC57">
<mods:title id="9A88EB2B3E759E470CA84C8BAC2E365A">An illustrated key to the New World genera of Philonthina Kirby (Coleoptera: Staphylinidae), with morphological, taxonomical and distributional notes</mods:title>
</mods:titleInfo>
<mods:name id="A772E4A42CD58C9DEE21D9ACA13CCFD3" type="personal">
<mods:role id="4B2A45CCC2E116E9040EF45C2BCAE6AD">
<mods:roleTerm id="24330F355AC7D81B2D65D86D8FA506EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5DEF81AE17CB228EAFADAD196D45B398">Chani-Posse, Mariana</mods:namePart>
</mods:name>
<mods:typeOfResource id="AAB73C2B3335D728E61124216AF4AF3C">text</mods:typeOfResource>
<mods:relatedItem id="D316ECBD2F468C5FC15FEA8546E0412C" type="host">
<mods:titleInfo id="0B27D8C117E6A48901BC1E150A796B23">
<mods:title id="887D307D83F2BD18B00D6F30169B30F8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="042E9AD9BD51B2A46238527A674DCA63">
<mods:date id="EDD9A5F3513E4FE142CFAA7BBD1FD522">2014</mods:date>
<mods:detail id="15BED1A64D739FECA9D59D43517663EF" type="volume">
<mods:number id="7D49B9589A18C6845191DEA25A4E151E">3755</mods:number>
</mods:detail>
<mods:detail id="CEA57A9E33A23326D7B79392E93631D0" type="issue">
<mods:number id="B533D6898317FE0D485517C8A098350E">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="F1FCB3AEB649409EE5415AA95194003D">journal volume</mods:classification>
<mods:identifier id="9080973CC51444C73A073D03D3F470C3" type="CLB-Dataset">46619</mods:identifier>
<mods:identifier id="116C63B1C3A07938D18E5554511100C7" type="DOI">10.11646/zootaxa.3755.1.3</mods:identifier>
<mods:identifier id="BA262BFD3D3B32A58FE7C472C9999B63" type="GBIF-Dataset">9b22d33f-c1f8-49e7-b123-3877690f282b</mods:identifier>
<mods:identifier id="7939322F96536464C80A50E6D59FD2B8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F90338EDDF8FA50CA27ED3537D6434B6" type="Zenodo-Dep">252659</mods:identifier>
<mods:identifier id="8B36275D172405FF4F121AAD90BA918C" type="ZooBank">0486D86E-EEFF-453E-BD77-395094BED088</mods:identifier>
</mods:mods>
<treatment id="03EF879CFFAFFFBDFF37FA6191A71384" ID-DOI="http://doi.org/10.5281/zenodo.6140180" ID-GBIF-Taxon="119412363" ID-Zenodo-Dep="6140180" LSID="urn:lsid:plazi:treatment:03EF879CFFAFFFBDFF37FA6191A71384" httpUri="http://treatment.plazi.org/id/03EF879CFFAFFFBDFF37FA6191A71384" lastPageId="14" pageId="12">
<subSubSection id="C35C6501FFAFFFBFFF37FA6196FF12FD" box="[151,707,1435,1461]" pageId="12" type="nomenclature">
<paragraph id="8BF9368AFFAFFFBFFF37FA6196FF12FD" blockId="12.[151,707,1435,1461]" box="[151,707,1435,1461]" pageId="12">
<heading id="D0B181E6FFAFFFBFFF37FA6196FF12FD" bold="true" box="[151,707,1435,1461]" fontSize="11" level="1" pageId="12" reason="1">
<emphasis id="B932EA98FFAFFFBFFF37FA6196FF12FD" bold="true" box="[151,707,1435,1461]" pageId="12">
Key to the New World genera of
<taxonomicName id="4C464D09FFAFFFBFFD90FA6196FF12FD" ID-CoL="8VRDC" box="[560,707,1435,1461]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C6501FFAFFFBDFF37FA1A91A71384" lastPageId="14" pageId="12" type="key">
<keyStep id="30B22E2FFFAFFFBFFF37FA1A91A711C1" pageId="12">
<paragraph id="8BF9368AFFAFFFBFFF37FA1A91A71179" blockId="12.[151,1436,1504,2025]" pageId="12">
<keyLead id="30B795BFFFAFFFBFFF37FA1A91A71179" pageId="12">
1 Pronotum with anterolateral margins forming a distinct ridge above the superior marginal line of hypomeron (
<figureCitation id="137D2A0FFFAFFFBFFABDFA1A916412BE" box="[1309,1368,1504,1526]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="12">Fig. 4</figureCitation>
K) and hypomeron with additional oblique line connecting the superior and inferior lines (
<figureCitation id="137D2A0FFFAFFFBFFC5FFA049004115C" box="[1023,1080,1534,1556]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="12">Fig. 4</figureCitation>
D); tarsi with segment 3 enclosing a rather minute segment 4 (
<figureCitation id="137D2A0FFFAFFFBFFE76F9E196331179" box="[470,527,1563,1585]" captionStart="FIGURE 6" captionStartId="12.[151,250,941,963]" captionTargetBox="[272,1315,193,919]" captionTargetId="figure@12.[272,1315,193,920]" captionTargetPageId="12" captionText="FIGURE 6. (A D) Front tarsus: (A) Delgadobius amazonensis, (B) Proxenobius borgmeieri, (C) Belonuchus mordens, (D) Pescolinus schmidti." httpUri="https://zenodo.org/record/252665/files/figure.png" pageId="12">Fig. 6</figureCitation>
A)...............................................................
<emphasis id="B932EA98FFAFFFBFFABCF9E691A71179" box="[1308,1435,1564,1585]" italics="true" pageId="12">Delgadobius</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAFFFBFFF37F9C391A711C1" blockId="12.[151,1436,1504,2025]" pageId="12">
<keyLead id="30B795BFFFAFFFBFFF37F9C391A711C1" pageId="12">
- Pronotum with anterolateral margins not forming a ridge above the superior marginal line of hypomeron (
<figureCitation id="137D2A0FFFAFFFBFFAA0F9C291791106" box="[1280,1349,1592,1614]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="12">Figs. 4</figureCitation>
EJ) and hypomeron with or without additional oblique line connecting the superior and inferior lines (
<figureCitation id="137D2A0FFFAFFFBFFBC7F9AC90961124" box="[1127,1194,1622,1644]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="12">Figs. 4</figureCitation>
A, C); tarsi with segment 3 and 4 not distinctly different (
<figureCitation id="137D2A0FFFAFFFBFFDB0F989966E11C1" box="[528,594,1651,1673]" captionStart="FIGURE 6" captionStartId="12.[151,250,941,963]" captionTargetBox="[272,1315,193,919]" captionTargetId="figure@12.[272,1315,193,920]" captionTargetPageId="12" captionText="FIGURE 6. (A D) Front tarsus: (A) Delgadobius amazonensis, (B) Proxenobius borgmeieri, (C) Belonuchus mordens, (D) Pescolinus schmidti." httpUri="https://zenodo.org/record/252665/files/figure.png" pageId="12">Figs. 6</figureCitation>
BD, 7EH)........................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAFFFBFFF37F96A91A011B6" pageId="12">
<paragraph id="8BF9368AFFAFFFBFFF37F96A91A7118C" blockId="12.[151,1436,1504,2025]" pageId="12">
<keyLead id="30B795BFFFAFFFBFFF37F96A91A7118C" pageId="12">
2 Head with ventral basal ridge strongly (to moderately) projecting anteriad (
<figureCitation id="137D2A0FFFAFFFBFFC79F96A902411EE" box="[985,1048,1680,1702]" captionStart="FIGURE 2" captionStartId="8.[151,250,1603,1625]" captionTargetBox="[156,1430,193,1579]" captionTargetId="figure@8.[156,1430,193,1581]" captionTargetPageId="8" captionText="FIGURE 2. (A K) Head, ventral view: (A) Hesperus fasciatus (Sharp), (B) Belonuchus mordens Erichson, (C) Neobisnius fraternus Bernhauer, (D) Paederomimus contractus Sharp, (E) Ecitophytes coniceps Wasmann, (F) Delgadobius amazonensis Chani-Posse &amp; Couturier, (G, K) Flohria subcoerulea, (H) Laetulonthus laetulus (Say), (I) Holisus humilis Erichson, (J) Phileciton badariottii Wasmann. Abbreviations: ioR, infraorbital ridge; pgR, postgenal ridge; pmR, postmandibular ridge; poS, postoccipital suture; PtC, proesternal transverse carina; vbR, ventral basal ridge." httpUri="https://zenodo.org/record/252661/files/figure.png" pageId="12">
Fig.
<date id="FFF8104AFFAFFFBFFBAAF96A902411EE" box="[1034,1048,1680,1702]" pageId="12">2</date>
</figureCitation>
I); pronotal hypomeron with additional oblique line connecting the superior and inferior lines (
<figureCitation id="137D2A0FFFAFFFBFFD50F9549715118C" box="[752,809,1710,1732]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="12">Fig. 4</figureCitation>
C)...............................
<taxonomicName id="4C464D09FFAFFFBFFB69F95491A7118C" ID-CoL="8X7GJ" authority="Hyptiomina" authorityName="Hyptiomina" box="[1225,1435,1710,1732]" class="Insecta" family="Staphylinidae" genus="Holisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAFFFBFFB69F954912E118B" box="[1225,1298,1710,1731]" italics="true" pageId="12">Holisus</emphasis>
(Hyptiomina)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAFFFBFFF37F93691A011B6" blockId="12.[151,1436,1504,2025]" pageId="12">
<keyLead id="30B795BFFFAFFFBFFF37F93691A011B6" pageId="12">
- Head with ventral basal ridge extending more or less parallel to ventral portion of postoccipital suture (
<figureCitation id="137D2A0FFFAFFFBFFB5FF931917911A9" box="[1279,1349,1739,1761]" captionStart="FIGURE 2" captionStartId="8.[151,250,1603,1625]" captionTargetBox="[156,1430,193,1579]" captionTargetId="figure@8.[156,1430,193,1581]" captionTargetPageId="8" captionText="FIGURE 2. (A K) Head, ventral view: (A) Hesperus fasciatus (Sharp), (B) Belonuchus mordens Erichson, (C) Neobisnius fraternus Bernhauer, (D) Paederomimus contractus Sharp, (E) Ecitophytes coniceps Wasmann, (F) Delgadobius amazonensis Chani-Posse &amp; Couturier, (G, K) Flohria subcoerulea, (H) Laetulonthus laetulus (Say), (I) Holisus humilis Erichson, (J) Phileciton badariottii Wasmann. Abbreviations: ioR, infraorbital ridge; pgR, postgenal ridge; pmR, postmandibular ridge; poS, postoccipital suture; PtC, proesternal transverse carina; vbR, ventral basal ridge." httpUri="https://zenodo.org/record/252661/files/figure.png" pageId="12">Figs. 2</figureCitation>
AH, J); hypomeron without additional oblique line connecting the superior and inferior lines (
<figureCitation id="137D2A0FFFAFFFBFFBB9F912906E11B6" box="[1049,1106,1768,1790]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="12">Fig. 4</figureCitation>
A)........................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAFFFBFFF37F8FC91A010D9" pageId="12">
<paragraph id="8BF9368AFFAFFFBFFF37F8FC9567103C" blockId="12.[151,1436,1504,2025]" pageId="12">
<keyLead id="30B795BFFFAFFFBFFF37F8FC9567103C" pageId="12">
3 Front femur with medio-ventral process (
<figureCitation id="137D2A0FFFAFFFBFFDCCF8FC96991054" box="[620,677,1798,1820]" captionStart="FIGURE 7" captionStartId="15.[151,250,1400,1422]" captionTargetBox="[173,1433,193,1355]" captionTargetId="figure@15.[154,1433,193,1379]" captionTargetPageId="15" captionText="FIGURE 7. (A H) (A, B, H) Atopocentrum mirabile Bernhauer: (A) front femur, (B) front tibia, (H) hind tibiae. (C) Pescolinus schmidti: front coxa and femur. (D, E) Proxenobius borgmeieri: (D) front leg, (E) front tarsus. (F, G) Front tibia and tarsus: (F) Laetulonthus laetulus, (G) Hesperus baltimorensis (Gravenhorst)." httpUri="https://zenodo.org/record/252666/files/figure.png" pageId="12">Fig. 7</figureCitation>
A); front tibia abruptly dilated apically (
<figureCitation id="137D2A0FFFAFFFBFFB85F8FC90631054" box="[1061,1119,1798,1820]" captionStart="FIGURE 7" captionStartId="15.[151,250,1400,1422]" captionTargetBox="[173,1433,193,1355]" captionTargetId="figure@15.[154,1433,193,1379]" captionTargetPageId="15" captionText="FIGURE 7. (A H) (A, B, H) Atopocentrum mirabile Bernhauer: (A) front femur, (B) front tibia, (H) hind tibiae. (C) Pescolinus schmidti: front coxa and femur. (D, E) Proxenobius borgmeieri: (D) front leg, (E) front tarsus. (F, G) Front tibia and tarsus: (F) Laetulonthus laetulus, (G) Hesperus baltimorensis (Gravenhorst)." httpUri="https://zenodo.org/record/252666/files/figure.png" pageId="12">Fig. 7</figureCitation>
B); hind tibia distinctly projected apically (
<figureCitation id="137D2A0FFFAFFFBFFE9AF8D9954F1071" box="[314,371,1827,1849]" captionStart="FIGURE 7" captionStartId="15.[151,250,1400,1422]" captionTargetBox="[173,1433,193,1355]" captionTargetId="figure@15.[154,1433,193,1379]" captionTargetPageId="15" captionText="FIGURE 7. (A H) (A, B, H) Atopocentrum mirabile Bernhauer: (A) front femur, (B) front tibia, (H) hind tibiae. (C) Pescolinus schmidti: front coxa and femur. (D, E) Proxenobius borgmeieri: (D) front leg, (E) front tarsus. (F, G) Front tibia and tarsus: (F) Laetulonthus laetulus, (G) Hesperus baltimorensis (Gravenhorst)." httpUri="https://zenodo.org/record/252666/files/figure.png" pageId="12">Fig. 7</figureCitation>
H).........................................................................
<emphasis id="B932EA98FFAFFFBFFAA8F8DE91A71071" box="[1288,1435,1828,1849]" italics="true" pageId="12">
<taxonomicName id="4C464D09FFAFFFBFFAA8F8DE91AF1071" ID-CoL="8XJSD" box="[1288,1427,1828,1849]" class="Insecta" family="Staphylinidae" genus="Atopocentrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">Atopocentrum</taxonomicName>
*
</emphasis>
* based on two male specimens from the
<typeStatus id="54FD8828FFAFFFBFFDD3F8BB96A0101E" box="[627,668,1857,1878]" pageId="12">type</typeStatus>
series of
<taxonomicName id="4C464D09FFAFFFBFFD5BF8BB97E7101E" ID-CoL="JHHF" box="[763,987,1857,1878]" class="Insecta" family="Staphylinidae" genus="Atopocentrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="mirabile">
<emphasis id="B932EA98FFAFFFBFFD5BF8BB97E7101E" box="[763,987,1857,1878]" italics="true" pageId="12">Atopocentrum mirabile</emphasis>
</taxonomicName>
(
<figureCitation id="137D2A0FFFAFFFBFFC49F8BA9010101E" box="[1001,1068,1856,1878]" captionStart="FIGURE 11" captionStartId="18.[151,250,1908,1930]" captionTargetBox="[266,1307,200,1856]" captionTargetId="figure@18.[265,1311,195,1870]" captionTargetPageId="18" captionText="FIGURE 11. (A J) Habitus: (A) Atopocentrum mirabile, (B) Belonuchus aluticollis (Bernhauer), (C) Cafius xantholoma, (D) Chroaptomus mexicanus Chani-Posse &amp; Navarrete-Heredia, (E) Delgadobius amazonensis Chani-Posse &amp; Couturier, (F) Ecitophytes coniceps Wasmann, (G) Flohria subcoerulea (LeConte), (H) Gabrius nidicola (Bernhauer), (I) Gabronthus maritimus (Motschulsky), (J), Hesperus fasciatus (Sharp)." httpUri="https://zenodo.org/record/252670/files/figure.png" pageId="12">Fig. 11</figureCitation>
A) and the original description (
<bibRefCitation id="EFD74B7BFFAFFFBFFAC4F8BA956C103C" author="Bernhauer" pageId="12" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der k. k. zoologisch-Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article" year="1906">Bernhauer 1906</bibRefCitation>
).
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAFFFBFFF37F88691A010D9" blockId="12.[151,1436,1504,2025]" box="[151,1436,1915,1937]" pageId="12">
<keyLead id="30B795BFFFAFFFBFFF37F88691A010D9" box="[151,1436,1915,1937]" pageId="12">
<emphasis id="B932EA98FFAFFFBFFF37F88694A310D9" box="[151,159,1916,1937]" italics="true" pageId="12">-</emphasis>
Front femur, front and hind tibiae not as above..............................................................4
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BF9368AFFAFFFBFFF37F86291A71084" blockId="12.[151,1436,1504,2025]" pageId="12">
<keyStep id="30B22E2FFFAFFFBFFF37F86291A71084" pageId="12">
<keyLead id="30B795BFFFAFFFBFFF37F86291A71084" pageId="12">
4 Antennae inserted at or below the anterior margin of frontoclypeus (
<figureCitation id="137D2A0FFFAFFFBFFCCAF862979E10E6" box="[874,930,1944,1966]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="12">Fig. 3</figureCitation>
E); abdomen conical (“petiolate” after
<bibRefCitation id="EFD74B7BFFAFFFBFFAB4F86291A810E6" author="Seevers" box="[1300,1428,1944,1966]" pageId="12" refString="Seevers, C. H. (1965) The systematics, evolution and zoogeography of staphylinid beetles associated with army ants (Coleoptera, Staphylinidae). Fieldiana, Zoology, 47 (2), 139 - 351." type="journal article" year="1965">Seevers 1965</bibRefCitation>
) (
<figureCitation id="137D2A0FFFAFFFBFFF48F84C951D1084" box="[232,289,1974,1996]" captionStart="FIGURE 9" captionStartId="16.[151,250,1973,1995]" captionTargetBox="[352,1232,1477,1962]" captionTargetId="figure@16.[339,1248,1477,1964]" captionTargetPageId="16" captionText="FIGURE 9. Abdominal shape, lateral view: (A) Ecitophytes coniceps, (B) Proxenobius borgmeieri, (C) Flohria subcoerulea, (D) Philonthus bonariensis." httpUri="https://zenodo.org/record/252668/files/figure.png" pageId="12">Fig. 9</figureCitation>
A) or subconical (
<figureCitation id="137D2A0FFFAFFFBFFE6DF84C963A1084" box="[461,518,1974,1996]" captionStart="FIGURE 9" captionStartId="16.[151,250,1973,1995]" captionTargetBox="[352,1232,1477,1962]" captionTargetId="figure@16.[339,1248,1477,1964]" captionTargetPageId="16" captionText="FIGURE 9. Abdominal shape, lateral view: (A) Ecitophytes coniceps, (B) Proxenobius borgmeieri, (C) Flohria subcoerulea, (D) Philonthus bonariensis." httpUri="https://zenodo.org/record/252668/files/figure.png" pageId="12">Fig. 9</figureCitation>
B)......................................................................... 5
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BF9368AFFAFFFBFFF37F82E91A010A1" blockId="12.[151,1436,1504,2025]" box="[151,1436,2003,2025]" pageId="12">
<keyLead id="30B795BFFFAFFFBFFF37F82E91A010A1" box="[151,1436,2003,2025]" pageId="12">
- Antennae inserted behind the anterior margin of frontoclypeus (
<figureCitation id="137D2A0FFFAFFFBFFCF0F82997AF10A1" box="[848,915,2003,2025]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="12">Figs. 3</figureCitation>
AD); abdomen dorsoventrally flattened (
<figureCitation id="137D2A0FFFAFFFBFFA8DF829915B10A1" box="[1325,1383,2003,2025]" captionStart="FIGURE 9" captionStartId="16.[151,250,1973,1995]" captionTargetBox="[352,1232,1477,1962]" captionTargetId="figure@16.[339,1248,1477,1964]" captionTargetPageId="16" captionText="FIGURE 9. Abdominal shape, lateral view: (A) Ecitophytes coniceps, (B) Proxenobius borgmeieri, (C) Flohria subcoerulea, (D) Philonthus bonariensis." httpUri="https://zenodo.org/record/252668/files/figure.png" pageId="12">Fig. 9</figureCitation>
D) or
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF41FF6D91A017AF" blockId="13.[151,1436,151,2021]" pageId="13">
subcylindrical (
<figureCitation id="137D2A0FFFAEFFBEFED6FF6D958C17E5" box="[374,432,151,173]" captionStart="FIGURE 9" captionStartId="16.[151,250,1973,1995]" captionTargetBox="[352,1232,1477,1962]" captionTargetId="figure@16.[339,1248,1477,1964]" captionTargetPageId="16" captionText="FIGURE 9. Abdominal shape, lateral view: (A) Ecitophytes coniceps, (B) Proxenobius borgmeieri, (C) Flohria subcoerulea, (D) Philonthus bonariensis." httpUri="https://zenodo.org/record/252668/files/figure.png" pageId="13">Fig. 9</figureCitation>
C).................................................................................7
<keyStep id="30B22E2FFFAEFFBEFF37FF4E91A017AF" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FF4E91A017AF" pageId="13">
5
<collectingCountry id="F351761AFFAEFFBEFF41FF4E952D1782" box="[225,273,180,202]" name="India" pageId="13">Hind</collectingCountry>
angles of head rounded to subangulate (
<figureCitation id="137D2A0FFFAEFFBEFD3BFF4E96E81782" box="[667,724,180,202]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="13">Fig. 3</figureCitation>
E); front tarsi with tarsomeres IIV more or less dorsoventrally flattened, not widened distally (
<figureCitation id="137D2A0FFFAEFFBEFE10FF2B95CE17AF" box="[432,498,209,231]" captionStart="FIGURE 7" captionStartId="15.[151,250,1400,1422]" captionTargetBox="[173,1433,193,1355]" captionTargetId="figure@15.[154,1433,193,1379]" captionTargetPageId="15" captionText="FIGURE 7. (A H) (A, B, H) Atopocentrum mirabile Bernhauer: (A) front femur, (B) front tibia, (H) hind tibiae. (C) Pescolinus schmidti: front coxa and femur. (D, E) Proxenobius borgmeieri: (D) front leg, (E) front tarsus. (F, G) Front tibia and tarsus: (F) Laetulonthus laetulus, (G) Hesperus baltimorensis (Gravenhorst)." httpUri="https://zenodo.org/record/252666/files/figure.png" pageId="13">Figs. 7</figureCitation>
D, E); abdomen subconical (
<figureCitation id="137D2A0FFFAEFFBEFCA5FF2B970217AF" box="[773,830,209,231]" captionStart="FIGURE 9" captionStartId="16.[151,250,1973,1995]" captionTargetBox="[352,1232,1477,1962]" captionTargetId="figure@16.[339,1248,1477,1964]" captionTargetPageId="16" captionText="FIGURE 9. Abdominal shape, lateral view: (A) Ecitophytes coniceps, (B) Proxenobius borgmeieri, (C) Flohria subcoerulea, (D) Philonthus bonariensis." httpUri="https://zenodo.org/record/252668/files/figure.png" pageId="13">Fig. 9</figureCitation>
B); habitus not myrmecoid (
<figureCitation id="137D2A0FFFAEFFBEFBECFF2B90A617AF" box="[1100,1178,209,231]" captionStart="FIGURE 13" captionStartId="20.[151,250,1495,1517]" captionTargetBox="[422,1158,193,1447]" captionTargetId="figure@20.[419,1158,193,1459]" captionTargetPageId="20" captionText="FIGURE 13. (A G) Habitus: (A) Pescolinus schmidti Bierig, (B) Phileciton badariottii Wasmann, (C) Philonthus bonariensis Bernhauer, (D) Proxenobius borgmeieri Seevers, (E) Pterygolaetus williamsii Bierig, (F) Xanthodermus vestitus (Sahlberg), (G) Xenobius rotundiceps Borgmeier." httpUri="https://zenodo.org/record/252672/files/figure.png" pageId="13">Figs. 13</figureCitation>
D, G)............... 6
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FF1591A7166A" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FF1591A7166A" pageId="13">
-
<collectingCountry id="F351761AFFAEFFBEFF41FF15952D164D" box="[225,273,239,261]" name="India" pageId="13">Hind</collectingCountry>
angles of head obsolete; front tarsi with tarsomeres IIV not as above; abdomen conical (
<figureCitation id="137D2A0FFFAEFFBEFB21FF159087164D" box="[1153,1211,239,261]" captionStart="FIGURE 9" captionStartId="16.[151,250,1973,1995]" captionTargetBox="[352,1232,1477,1962]" captionTargetId="figure@16.[339,1248,1477,1964]" captionTargetPageId="16" captionText="FIGURE 9. Abdominal shape, lateral view: (A) Ecitophytes coniceps, (B) Proxenobius borgmeieri, (C) Flohria subcoerulea, (D) Philonthus bonariensis." httpUri="https://zenodo.org/record/252668/files/figure.png" pageId="13">Fig. 9</figureCitation>
A); habitus myrmecoid (
<figureCitation id="137D2A0FFFAEFFBEFF48FEF69517166A" box="[232,299,268,290]" captionStart="FIGURE 11" captionStartId="18.[151,250,1908,1930]" captionTargetBox="[266,1307,200,1856]" captionTargetId="figure@18.[265,1311,195,1870]" captionTargetPageId="18" captionText="FIGURE 11. (A J) Habitus: (A) Atopocentrum mirabile, (B) Belonuchus aluticollis (Bernhauer), (C) Cafius xantholoma, (D) Chroaptomus mexicanus Chani-Posse &amp; Navarrete-Heredia, (E) Delgadobius amazonensis Chani-Posse &amp; Couturier, (F) Ecitophytes coniceps Wasmann, (G) Flohria subcoerulea (LeConte), (H) Gabrius nidicola (Bernhauer), (I) Gabronthus maritimus (Motschulsky), (J), Hesperus fasciatus (Sharp)." httpUri="https://zenodo.org/record/252670/files/figure.png" pageId="13">Fig. 11</figureCitation>
F)..................................................................................
<taxonomicName id="4C464D09FFAEFFBEFA8BFEF791A7166A" ID-CoL="8XL4L" box="[1323,1435,269,290]" class="Insecta" family="Staphylinidae" genus="Ecitophytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFA8BFEF791A7166A" box="[1323,1435,269,290]" italics="true" pageId="13">Ecitophytes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<keyStep id="30B22E2FFFAEFFBEFF37FED391A716DF" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FED391A01614" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FED391A01614" pageId="13">
6 Head about as wide as long with hind angles rounded, not extended beyond anterior margin of neck (
<figureCitation id="137D2A0FFFAEFFBEFB3FFED390D81677" box="[1183,1252,297,319]" captionStart="FIGURE 13" captionStartId="20.[151,250,1495,1517]" captionTargetBox="[422,1158,193,1447]" captionTargetId="figure@20.[419,1158,193,1459]" captionTargetPageId="20" captionText="FIGURE 13. (A G) Habitus: (A) Pescolinus schmidti Bierig, (B) Phileciton badariottii Wasmann, (C) Philonthus bonariensis Bernhauer, (D) Proxenobius borgmeieri Seevers, (E) Pterygolaetus williamsii Bierig, (F) Xanthodermus vestitus (Sahlberg), (G) Xenobius rotundiceps Borgmeier." httpUri="https://zenodo.org/record/252672/files/figure.png" pageId="13">Fig. 13</figureCitation>
G); pronotum with anterior angles rounded and hypomeron slightly inflexed, visible in lateral view (
<figureCitation id="137D2A0FFFAEFFBEFC49FEBD901E1615" box="[1001,1058,327,349]" captionStart="FIGURE 5" captionStartId="11.[151,250,1977,1999]" captionTargetBox="[158,1428,1631,1965]" captionTargetId="figure@11.[158,1428,1631,1966]" captionTargetPageId="11" captionText="FIGURE 5. (A D) Pronotum, lateral view: (A) Proxenobius borgmeieri, (B) Xenobius rotundiceps Borgmeier. Prosternum, lateral view: (C) Leptopeltus carchiensis Chani-Posse &amp; Asenjo, (D) Leptopeltoides columbiensis Chani-Posse &amp; Asenjo." httpUri="https://zenodo.org/record/252664/files/figure.png" pageId="13">Fig. 5</figureCitation>
B).....................
<taxonomicName id="4C464D09FFAEFFBEFAE3FEBD91A01614" box="[1347,1436,327,348]" class="Insecta" family="Staphylinidae" genus="Xenobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFAE3FEBD91A01614" box="[1347,1436,327,348]" italics="true" pageId="13">Xenobius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FE9F91A716DF" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FE9F91A716DF" pageId="13">
- Head longer than wide with hind angles subangulate, extended beyond anterior margin of neck (
<figureCitation id="137D2A0FFFAEFFBEFBD5FE9E908A1632" box="[1141,1206,356,378]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="13">Figs. 3</figureCitation>
E, 13D); pronotum with anterior angles distinctly angulate and hypomeron fully inflexed, not visible in lateral view (
<figureCitation id="137D2A0FFFAEFFBEFBF5FE7B90B116DF" box="[1109,1165,385,407]" captionStart="FIGURE 5" captionStartId="11.[151,250,1977,1999]" captionTargetBox="[158,1428,1631,1965]" captionTargetId="figure@11.[158,1428,1631,1966]" captionTargetPageId="11" captionText="FIGURE 5. (A D) Pronotum, lateral view: (A) Proxenobius borgmeieri, (B) Xenobius rotundiceps Borgmeier. Prosternum, lateral view: (C) Leptopeltus carchiensis Chani-Posse &amp; Asenjo, (D) Leptopeltoides columbiensis Chani-Posse &amp; Asenjo." httpUri="https://zenodo.org/record/252664/files/figure.png" pageId="13">Fig. 5</figureCitation>
A)..........
<taxonomicName id="4C464D09FFAEFFBEFA83FE7891A716DF" ID-CoL="8X7RL" box="[1315,1435,386,407]" class="Insecta" family="Staphylinidae" genus="Proxenobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFA83FE7891A716DF" box="[1315,1435,386,407]" italics="true" pageId="13">Proxenobius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FE6591A715CA" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FE6591A91544" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FE6591A91544" pageId="13">
7 Antennae with pubescence starting on antennomere V, lacking on antennomeres IIV (with only longer sparse setae) and segment I distinctly more than 1.5 times as wide as antennal segment II (
<figureCitation id="137D2A0FFFAEFFBEFC25FE469782169A" box="[901,958,444,466]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="13">Fig. 3</figureCitation>
A); sternopleural (anapleural) suture more or less curved so that medial part of suture is more longitudinal and lateral part more transverse (
<figureCitation id="137D2A0FFFAEFFBEFB9EFE23904B16A7" box="[1086,1143,473,495]" captionStart="FIGURE 8" captionStartId="16.[151,250,1354,1376]" captionTargetBox="[155,1431,359,1332]" captionTargetId="figure@16.[155,1431,359,1332]" captionTargetPageId="16" captionText="FIGURE 8. (A E) Mesoventrite, sternopleural (anapleural) suture: (A) Flohria laticornis, (B), Belonuchus mordens, (C) Paederomimus contractus, (D) Cafius xantholoma (Gravenhorst), (E), Neobisnius fraternus. Abbreviation: stS, sternopleural suture." httpUri="https://zenodo.org/record/252667/files/figure.png" pageId="13">Fig. 8</figureCitation>
A); abdomen subcylindrical in lateral view (
<figureCitation id="137D2A0FFFAEFFBEFEFEFE0D95AB1545" box="[350,407,503,525]" captionStart="FIGURE 9" captionStartId="16.[151,250,1973,1995]" captionTargetBox="[352,1232,1477,1962]" captionTargetId="figure@16.[339,1248,1477,1964]" captionTargetPageId="16" captionText="FIGURE 9. Abdominal shape, lateral view: (A) Ecitophytes coniceps, (B) Proxenobius borgmeieri, (C) Flohria subcoerulea, (D) Philonthus bonariensis." httpUri="https://zenodo.org/record/252668/files/figure.png" pageId="13">Fig. 9</figureCitation>
C)............................................................................
<taxonomicName id="4C464D09FFAEFFBEFAECFE0D91A91544" ID-CoL="8XLHH" box="[1356,1429,503,524]" class="Insecta" family="Staphylinidae" genus="Flohria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFAECFE0D91A91544" box="[1356,1429,503,524]" italics="true" pageId="13">Flohria</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FDEF91A715CA" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FDEF91A715CA" pageId="13">
- Antennae with pubescence starting on antennomere III or IV, lacking on antennomeres III or IIII (with only sparse longer setae) and segment I not more than 1.5 times as wide as antennal segment II (
<figureCitation id="137D2A0FFFAEFFBEFC42FDCB9020150F" box="[994,1052,561,583]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="13">Fig. 3</figureCitation>
BD); sternopleural (anapleural) suture transverse, or nearly transverse to distinctly oblique (medial end of suture anterior to its lateral end) (
<figureCitation id="137D2A0FFFAEFFBEFB1DFDB5913D152D" box="[1213,1281,591,613]" captionStart="FIGURE 8" captionStartId="16.[151,250,1354,1376]" captionTargetBox="[155,1431,359,1332]" captionTargetId="figure@16.[155,1431,359,1332]" captionTargetPageId="16" captionText="FIGURE 8. (A E) Mesoventrite, sternopleural (anapleural) suture: (A) Flohria laticornis, (B), Belonuchus mordens, (C) Paederomimus contractus, (D) Cafius xantholoma (Gravenhorst), (E), Neobisnius fraternus. Abbreviation: stS, sternopleural suture." httpUri="https://zenodo.org/record/252667/files/figure.png" pageId="13">Figs. 8</figureCitation>
B, C); abdomen flattened dorsoventrally in lateral view (
<figureCitation id="137D2A0FFFAEFFBEFDC1FD9696A615CA" box="[609,666,620,642]" captionStart="FIGURE 9" captionStartId="16.[151,250,1973,1995]" captionTargetBox="[352,1232,1477,1962]" captionTargetId="figure@16.[339,1248,1477,1964]" captionTargetPageId="16" captionText="FIGURE 9. Abdominal shape, lateral view: (A) Ecitophytes coniceps, (B) Proxenobius borgmeieri, (C) Flohria subcoerulea, (D) Philonthus bonariensis." httpUri="https://zenodo.org/record/252668/files/figure.png" pageId="13">Fig. 9</figureCitation>
D).............................................................8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FD7391A71592" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FD7391A015D7" blockId="13.[151,1436,151,2021]" box="[151,1436,649,671]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FD7391A015D7" box="[151,1436,649,671]" pageId="13">8 Front tarsi with segments IIV more or less cylindrical with only regular, unmodified marginal setae on ventral surface..... 9</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FD5D91A71592" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FD5D91A71592" pageId="13">
- Front tarsi with segments IIV more or less flattened dorsoventrally and widened distally, with modified pale (adhesive) setae on ventral surface (
<figureCitation id="137D2A0FFFAEFFBEFE3AFD3E95E01592" box="[410,476,708,730]" captionStart="FIGURE 6" captionStartId="12.[151,250,941,963]" captionTargetBox="[272,1315,193,919]" captionTargetId="figure@12.[272,1315,193,920]" captionTargetPageId="12" captionText="FIGURE 6. (A D) Front tarsus: (A) Delgadobius amazonensis, (B) Proxenobius borgmeieri, (C) Belonuchus mordens, (D) Pescolinus schmidti." httpUri="https://zenodo.org/record/252665/files/figure.png" pageId="13">Figs. 6</figureCitation>
D, 7F, G)...................................................................... 20
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FD1B91A714C2" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FD1B91A7147A" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FD1B91A7147A" pageId="13">
9 Superior marginal line of hypomeron not distinctly deflexed under anterior angle of pronotum; lateral puncture of pronotum bearing long seta situated close to superior line or at a distance no more than three times the diameter of puncture (
<figureCitation id="137D2A0FFFAEFFBEFA8CFD059159145D" box="[1324,1381,767,789]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="13">Fig. 4</figureCitation>
F)..................................................................................................... 10
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FCC091A714C2" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FCC091A714C2" pageId="13">
- Superior marginal line of hypomeron distinctly deflexed under anterior angle of pronotum; lateral puncture of pronotum bearing long seta situated away from superior line at a distance at least three times as large as diameter of puncture (
<figureCitation id="137D2A0FFFAEFFBEFAB1FCAD916F1425" box="[1297,1363,855,877]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="13">Figs. 4</figureCitation>
GI)..................................................................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FC6B91A71355" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FC6B91A714AA" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FC6B91A714AA" pageId="13">
10 Last segment of labial palpus distinctly narrower than preceding segment; sternum 9 of male genital segment with basal portion not reduced, more or less symmetrical (except in the New World species
<taxonomicName id="4C464D09FFAEFFBEFC6AFC559117148D" ID-CoL="6K7D2" authority="Horn (Smetana 1995)" authorityName="Horn (Smetana" authorityYear="1995" box="[970,1323,943,965]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B932EA98FFAEFFBEFC6AFC55906F148C" box="[970,1107,943,964]" italics="true" pageId="13">G. brevipennis</emphasis>
Horn (
<bibRefCitation id="EFD74B7BFFAEFFBEFB37FC559118148D" author="Smetana" box="[1175,1316,943,965]" pageId="13" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. http: // dx. doi. org / 10.2307 / 3495761" type="journal article" year="1995">Smetana 1995</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C464D09FFAEFFBEFA95FC55954214AA" ID-CoL="6K7BL" authority="Bernhauer" authorityName="Bernhauer" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="argentinus">
<emphasis id="B932EA98FFAEFFBEFA95FC55953E14AA" italics="true" pageId="13">G. argentinus</emphasis>
(Bernhauer)
</taxonomicName>
,
<taxonomicName id="4C464D09FFAEFFBEFE28FC37964214AA" ID-CoL="6JV4Z" authority="Bernhauer" authorityName="Bernhauer" box="[392,638,972,994]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="jujuyensis">
<emphasis id="B932EA98FFAEFFBEFE28FC37963E14AA" box="[392,514,973,994]" italics="true" pageId="13">G. jujuyensis</emphasis>
(Bernhauer)
</taxonomicName>
, and
<emphasis id="B932EA98FFAEFFBEFD10FC37972A14AA" box="[688,790,973,994]" italics="true" pageId="13">G. n i d i c o l a</emphasis>
(Bernhauer) (Chani-Posse de Maus 2009)).............
<taxonomicName id="4C464D09FFAEFFBEFAEDFC3791A714AA" ID-CoL="8VRP9" box="[1357,1435,973,994]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFAEDFC3791A714AA" box="[1357,1435,973,994]" italics="true" pageId="13">Gabrius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FC1091A71355" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FC1091A71355" pageId="13">- Last segment of labial palpus not distinctly narrower than preceding segment; sternum 9 of male genital segment with basal portion reduced, asymmetrical..........................................................................11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FBDE91A7131F" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FBDE91A71372" blockId="13.[151,1436,151,2021]" box="[151,1435,1060,1082]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FBDE91A71372" box="[151,1435,1060,1082]" pageId="13">
11 Elytra distinctly bicoloured, darker basally and testaceous distally (
<figureCitation id="137D2A0FFFAEFFBEFCC5FBDE97881372" box="[869,948,1060,1082]" captionStart="FIGURE 12" captionStartId="19.[151,250,1893,1915]" captionTargetBox="[291,1312,196,1862]" captionTargetId="figure@19.[283,1312,193,1868]" captionTargetPageId="19" captionText="FIGURE 12. (A J) Habitus: (A) Holisus humilis Erichson (Hyptiomina), (B) Laetulonthus laetulus (Say), (C) Leptopeltus weyrauchi Chani-Posse &amp; Asenjo, (D) Leptopeltoides ecuatoriensis Chani-Posse &amp; Asenjo, (E) Linoderus gracilipes Sharp, (F) Neobisnius fraternus Bernhauer, (G) Odontolinus fasciatus Sharp, (H) Ophionthus serpentinus Bernhauer, (I) Paederallus fragilis Sharp, (J) Paederomimus debilis Sharp." httpUri="https://zenodo.org/record/252671/files/figure.png" pageId="13">Figs. 12</figureCitation>
C, D).................................. 12
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FBB891A7131F" blockId="13.[151,1436,151,2021]" box="[151,1435,1089,1111]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FBB891A7131F" box="[151,1435,1089,1111]" pageId="13">- Elytra not bicoloured as above, usually unicolourous......................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FBA591A91384" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FBA591AA13DA" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FBA591AA13DA" pageId="13">
12 Labrum transverse; prosternum with distinct keel (
<figureCitation id="137D2A0FFFAEFFBEFD64FBA596C0133D" box="[708,764,1119,1141]" captionStart="FIGURE 5" captionStartId="11.[151,250,1977,1999]" captionTargetBox="[158,1428,1631,1965]" captionTargetId="figure@11.[158,1428,1631,1966]" captionTargetPageId="11" captionText="FIGURE 5. (A D) Pronotum, lateral view: (A) Proxenobius borgmeieri, (B) Xenobius rotundiceps Borgmeier. Prosternum, lateral view: (C) Leptopeltus carchiensis Chani-Posse &amp; Asenjo, (D) Leptopeltoides columbiensis Chani-Posse &amp; Asenjo." httpUri="https://zenodo.org/record/252664/files/figure.png" pageId="13">Fig. 5</figureCitation>
C); aedeagus with paramere completely fused to median lobe, reduced.......................................................................................... ..
<taxonomicName id="4C464D09FFAEFFBEFA86FB8791AA13DA" ID-CoL="8XMBF" box="[1318,1430,1149,1170]" class="Insecta" family="Staphylinidae" genus="Leptopeltus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFA86FB8791AA13DA" box="[1318,1430,1149,1170]" italics="true" pageId="13">Leptopeltus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FB6091A91384" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FB6091A91384" pageId="13">
- Labrum subconical; prosternum without keel (
<figureCitation id="137D2A0FFFAEFFBEFD3DFB6396EA13E7" box="[669,726,1177,1199]" captionStart="FIGURE 5" captionStartId="11.[151,250,1977,1999]" captionTargetBox="[158,1428,1631,1965]" captionTargetId="figure@11.[158,1428,1631,1966]" captionTargetPageId="11" captionText="FIGURE 5. (A D) Pronotum, lateral view: (A) Proxenobius borgmeieri, (B) Xenobius rotundiceps Borgmeier. Prosternum, lateral view: (C) Leptopeltus carchiensis Chani-Posse &amp; Asenjo, (D) Leptopeltoides columbiensis Chani-Posse &amp; Asenjo." httpUri="https://zenodo.org/record/252664/files/figure.png" pageId="13">Fig. 5</figureCitation>
D); aedeagus with paramere fused to median lobe only at base, elongate..........................................................................................
<emphasis id="B932EA98FFAEFFBEFAA8FB4D91A91384" box="[1288,1429,1207,1228]" italics="true" pageId="13">Leptopeltoides</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FB2E91A7120A" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FB2E951C126D" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FB2E951C126D" pageId="13">
13 Both neck and basisternum transversely carinate.................................................
<taxonomicName id="4C464D09FFAEFFBEFAABFB2F91AC13A2" ID-CoL="8XPHM" box="[1291,1424,1237,1258]" class="Insecta" family="Staphylinidae" genus="Pterygolaetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFAABFB2F91AC13A2" box="[1291,1424,1237,1258]" italics="true" pageId="13">Pterygolaetus</emphasis>
</taxonomicName>
* * based on a single specimen of
<taxonomicName id="4C464D09FFAEFFBEFD8EFB089720124F" box="[558,796,1266,1287]" class="Insecta" family="Staphylinidae" genus="Pterygolaetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="williamsii">
<emphasis id="B932EA98FFAEFFBEFD8EFB089720124F" box="[558,796,1266,1287]" italics="true" pageId="13">Pterygolaetus williamsii</emphasis>
</taxonomicName>
(
<typeStatus id="54FD8828FFAEFFBEFC8BFB0B97BE124F" box="[811,898,1265,1287]" pageId="13" type="holotype">holotype</typeStatus>
, male) (
<figureCitation id="137D2A0FFFAEFFBEFC76FB0B9021124F" box="[982,1053,1265,1287]" captionStart="FIGURE 13" captionStartId="20.[151,250,1495,1517]" captionTargetBox="[422,1158,193,1447]" captionTargetId="figure@20.[419,1158,193,1459]" captionTargetPageId="20" captionText="FIGURE 13. (A G) Habitus: (A) Pescolinus schmidti Bierig, (B) Phileciton badariottii Wasmann, (C) Philonthus bonariensis Bernhauer, (D) Proxenobius borgmeieri Seevers, (E) Pterygolaetus williamsii Bierig, (F) Xanthodermus vestitus (Sahlberg), (G) Xenobius rotundiceps Borgmeier." httpUri="https://zenodo.org/record/252672/files/figure.png" pageId="13">Fig. 13</figureCitation>
E) and the original description (
<bibRefCitation id="EFD74B7BFFAEFFBEFAFEFB0B9529126D" author="Bierig" pageId="13" refString="Bierig, A. (1937) Nuevos Staphylinini neotropicales. Memorias de la Sociedad Cubana de Historia Natural &quot; Felipe Poey &quot;, 11 (3), 191 - 205." type="journal article" year="1937">Bierig 1937</bibRefCitation>
).
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FAD791A7120A" blockId="13.[151,1436,151,2021]" box="[151,1435,1324,1346]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FAD791A7120A" box="[151,1435,1324,1346]" pageId="13">
- Neck and basisternum not as above..................................................................
<taxonomicName id="4C464D09FFAEFFBEFAF5FAD791A7120A" ID-CoL="8VRJ5" box="[1365,1435,1325,1346]" class="Insecta" family="Staphylinidae" genus="Bisnius" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFAF5FAD791A7120A" box="[1365,1435,1325,1346]" italics="true" pageId="13">Bisnius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FAB391A71235" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FAB391A71217" blockId="13.[151,1436,151,2021]" box="[151,1435,1353,1375]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FAB391A71217" box="[151,1435,1353,1375]" pageId="13">14 Antennal insertions closer to eyes than to anterior margin of frontoclypeus or at equal distance at most................. 15</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FA9D91A71235" blockId="13.[151,1436,151,2021]" box="[151,1435,1383,1405]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FA9D91A71235" box="[151,1435,1383,1405]" pageId="13">- Antennal insertions closer to anterior margin of frontoclypeus than to eyes....................................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FA7E91A712BA" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FA7E91A712FF" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FA7E91A712FF" pageId="13">
15 Antennal insertions closer to eyes than to anterior margin of frontoclypeus, distance to frontoclypeus at least twice the distance to eye; labrum and frontoclypeus sexually dimorphic (
<figureCitation id="137D2A0FFFAEFFBEFD49FA5B971012FF" box="[745,812,1441,1463]" captionStart="FIGURE 11" captionStartId="18.[151,250,1908,1930]" captionTargetBox="[266,1307,200,1856]" captionTargetId="figure@18.[265,1311,195,1870]" captionTargetPageId="18" captionText="FIGURE 11. (A J) Habitus: (A) Atopocentrum mirabile, (B) Belonuchus aluticollis (Bernhauer), (C) Cafius xantholoma, (D) Chroaptomus mexicanus Chani-Posse &amp; Navarrete-Heredia, (E) Delgadobius amazonensis Chani-Posse &amp; Couturier, (F) Ecitophytes coniceps Wasmann, (G) Flohria subcoerulea (LeConte), (H) Gabrius nidicola (Bernhauer), (I) Gabronthus maritimus (Motschulsky), (J), Hesperus fasciatus (Sharp)." httpUri="https://zenodo.org/record/252670/files/figure.png" pageId="13">Fig. 11</figureCitation>
D).......................................
<taxonomicName id="4C464D09FFAEFFBEFAB9FA5891A712FF" ID-CoL="8X778" box="[1305,1435,1442,1463]" class="Insecta" family="Staphylinidae" genus="Chroaptomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFAB9FA5891A712FF" box="[1305,1435,1442,1463]" italics="true" pageId="13">Chroaptomus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37FA4591A712BA" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FA4591A712BA" pageId="13">
- Antennal insertions at equal distance to both anterior margin of frontoclypeus and eyes; microsculpture of head sexually dimorphic (
<figureCitation id="137D2A0FFFAEFFBEFEF0FA2695A912BA" box="[336,405,1500,1522]" captionStart="FIGURE 12" captionStartId="19.[151,250,1893,1915]" captionTargetBox="[291,1312,196,1862]" captionTargetId="figure@19.[283,1312,193,1868]" captionTargetPageId="19" captionText="FIGURE 12. (A J) Habitus: (A) Holisus humilis Erichson (Hyptiomina), (B) Laetulonthus laetulus (Say), (C) Leptopeltus weyrauchi Chani-Posse &amp; Asenjo, (D) Leptopeltoides ecuatoriensis Chani-Posse &amp; Asenjo, (E) Linoderus gracilipes Sharp, (F) Neobisnius fraternus Bernhauer, (G) Odontolinus fasciatus Sharp, (H) Ophionthus serpentinus Bernhauer, (I) Paederallus fragilis Sharp, (J) Paederomimus debilis Sharp." httpUri="https://zenodo.org/record/252671/files/figure.png" pageId="13">Fig. 12</figureCitation>
E)............................................................................
<taxonomicName id="4C464D09FFAEFFBEFA9AFA2791A712BA" ID-CoL="8X7JG" box="[1338,1435,1501,1522]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFA9AFA2791A712BA" box="[1338,1435,1501,1522]" italics="true" pageId="13">Linoderus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37FA0391A0112F" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37FA0391A01164" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37FA0391A01164" pageId="13">
16 Antennae with pubescence starting on antennomere III, lacking on antennomere I and II (with only longer sparse setae) and segments IVX distinctly constricted at middle; front tibia not spinose on lateral face (
<figureCitation id="137D2A0FFFAEFFBEFBBEF9ED905F1165" box="[1054,1123,1559,1581]" captionStart="FIGURE 12" captionStartId="19.[151,250,1893,1915]" captionTargetBox="[291,1312,196,1862]" captionTargetId="figure@19.[283,1312,193,1868]" captionTargetPageId="19" captionText="FIGURE 12. (A J) Habitus: (A) Holisus humilis Erichson (Hyptiomina), (B) Laetulonthus laetulus (Say), (C) Leptopeltus weyrauchi Chani-Posse &amp; Asenjo, (D) Leptopeltoides ecuatoriensis Chani-Posse &amp; Asenjo, (E) Linoderus gracilipes Sharp, (F) Neobisnius fraternus Bernhauer, (G) Odontolinus fasciatus Sharp, (H) Ophionthus serpentinus Bernhauer, (I) Paederallus fragilis Sharp, (J) Paederomimus debilis Sharp." httpUri="https://zenodo.org/record/252671/files/figure.png" pageId="13">
Fig.
<date id="FFF8104AFFAEFFBEFBEBF9ED905F1165" box="[1099,1123,1559,1581]" pageId="13" value="2012-01">12</date>
</figureCitation>
I)............. ..
<taxonomicName id="4C464D09FFAEFFBEFA88F9ED91A01164" ID-CoL="8XNHH" box="[1320,1436,1559,1580]" class="Insecta" family="Staphylinidae" genus="Paederallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFA88F9ED91A01164" box="[1320,1436,1559,1580]" italics="true" pageId="13">Paederallus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37F9CF91A0112F" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37F9CF91A0112F" pageId="13">- Antennae with pubescence starting on antennomere IV, lacking on antennomere I III (with only longer sparse setae) and segments IVX not constricted at middle; front tibia spinose on lateral face.........................................17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37F99591A711EA" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37F99591A711CC" blockId="13.[151,1436,151,2021]" box="[151,1435,1647,1669]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37F99591A711CC" box="[151,1435,1647,1669]" pageId="13">
17 Head and pronotum appearing dull due to distinct, dense and rugose sculpture (
<figureCitation id="137D2A0FFFAEFFBEFC79F995902011CD" box="[985,1052,1647,1669]" captionStart="FIGURE 13" captionStartId="20.[151,250,1495,1517]" captionTargetBox="[422,1158,193,1447]" captionTargetId="figure@20.[419,1158,193,1459]" captionTargetPageId="20" captionText="FIGURE 13. (A G) Habitus: (A) Pescolinus schmidti Bierig, (B) Phileciton badariottii Wasmann, (C) Philonthus bonariensis Bernhauer, (D) Proxenobius borgmeieri Seevers, (E) Pterygolaetus williamsii Bierig, (F) Xanthodermus vestitus (Sahlberg), (G) Xenobius rotundiceps Borgmeier." httpUri="https://zenodo.org/record/252672/files/figure.png" pageId="13">Fig. 13</figureCitation>
F)................. ..
<taxonomicName id="4C464D09FFAEFFBEFAB0F99591A711CC" ID-CoL="8XQQ4" box="[1296,1435,1647,1668]" class="Insecta" family="Staphylinidae" genus="Xanthodermus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFAB0F99591A711CC" box="[1296,1435,1647,1668]" italics="true" pageId="13">Xanthodermus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37F97791A711EA" blockId="13.[151,1436,151,2021]" box="[151,1435,1676,1698]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37F97791A711EA" box="[151,1435,1676,1698]" pageId="13">- Head and pronotum glossy, with sculpture less dense........................................................ 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBEFF37F95391A7105F" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37F95391A01194" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37F95391A01194" pageId="13">
18 Head with interocular punctation extensive, with postmandibular ridge; last segment of both maxillary and labial palpi gradually narrowed to subacute apex (
<figureCitation id="137D2A0FFFAEFFBEFDB8F93D966E1195" box="[536,594,1735,1757]" captionStart="FIGURE 2" captionStartId="8.[151,250,1603,1625]" captionTargetBox="[156,1430,193,1579]" captionTargetId="figure@8.[156,1430,193,1581]" captionTargetPageId="8" captionText="FIGURE 2. (A K) Head, ventral view: (A) Hesperus fasciatus (Sharp), (B) Belonuchus mordens Erichson, (C) Neobisnius fraternus Bernhauer, (D) Paederomimus contractus Sharp, (E) Ecitophytes coniceps Wasmann, (F) Delgadobius amazonensis Chani-Posse &amp; Couturier, (G, K) Flohria subcoerulea, (H) Laetulonthus laetulus (Say), (I) Holisus humilis Erichson, (J) Phileciton badariottii Wasmann. Abbreviations: ioR, infraorbital ridge; pgR, postgenal ridge; pmR, postmandibular ridge; poS, postoccipital suture; PtC, proesternal transverse carina; vbR, ventral basal ridge." httpUri="https://zenodo.org/record/252661/files/figure.png" pageId="13">Fig. 2</figureCitation>
J); gular sutures not joined before neck and running close to the base.......
<taxonomicName id="4C464D09FFAEFFBEFA95F93D91A01194" box="[1333,1436,1735,1756]" class="Insecta" family="Staphylinidae" genus="Phileciton" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFA95F93D91A01194" box="[1333,1436,1735,1756]" italics="true" pageId="13">Phileciton</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37F91F91A7105F" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37F91F91A7105F" pageId="13">
- Head with few interocular punctures, usually not more than four, rarely with postmandibular ridge; last segment of both maxillary and labial palpi fusiform to subcylindrical (
<figureCitation id="137D2A0FFFAEFFBEFD0CF8FB96D3105F" box="[684,751,1793,1815]" captionStart="FIGURE 2" captionStartId="8.[151,250,1603,1625]" captionTargetBox="[156,1430,193,1579]" captionTargetId="figure@8.[156,1430,193,1581]" captionTargetPageId="8" captionText="FIGURE 2. (A K) Head, ventral view: (A) Hesperus fasciatus (Sharp), (B) Belonuchus mordens Erichson, (C) Neobisnius fraternus Bernhauer, (D) Paederomimus contractus Sharp, (E) Ecitophytes coniceps Wasmann, (F) Delgadobius amazonensis Chani-Posse &amp; Couturier, (G, K) Flohria subcoerulea, (H) Laetulonthus laetulus (Say), (I) Holisus humilis Erichson, (J) Phileciton badariottii Wasmann. Abbreviations: ioR, infraorbital ridge; pgR, postgenal ridge; pmR, postmandibular ridge; poS, postoccipital suture; PtC, proesternal transverse carina; vbR, ventral basal ridge." httpUri="https://zenodo.org/record/252661/files/figure.png" pageId="13">Figs. 2</figureCitation>
B, D); gular sutures joined before neck....................... 19
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFAEFFBDFF37F8E591A01677" lastPageId="14" pageId="13">
<paragraph id="8BF9368AFFAEFFBEFF37F8E591A910C4" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37F8E591A910C4" pageId="13">
19 Anterior angles of pronotum not or slightly produced beyond (anteriad of) anterior margin of prosternum (
<figureCitation id="137D2A0FFFAEFFBEFB41F8E59127107D" box="[1249,1307,1823,1845]" captionStart="FIGURE 2" captionStartId="8.[151,250,1603,1625]" captionTargetBox="[156,1430,193,1579]" captionTargetId="figure@8.[156,1430,193,1581]" captionTargetPageId="8" captionText="FIGURE 2. (A K) Head, ventral view: (A) Hesperus fasciatus (Sharp), (B) Belonuchus mordens Erichson, (C) Neobisnius fraternus Bernhauer, (D) Paederomimus contractus Sharp, (E) Ecitophytes coniceps Wasmann, (F) Delgadobius amazonensis Chani-Posse &amp; Couturier, (G, K) Flohria subcoerulea, (H) Laetulonthus laetulus (Say), (I) Holisus humilis Erichson, (J) Phileciton badariottii Wasmann. Abbreviations: ioR, infraorbital ridge; pgR, postgenal ridge; pmR, postmandibular ridge; poS, postoccipital suture; PtC, proesternal transverse carina; vbR, ventral basal ridge." httpUri="https://zenodo.org/record/252661/files/figure.png" pageId="13">Fig. 2</figureCitation>
B); neck usually without transverse carina (
<figureCitation id="137D2A0FFFAEFFBEFDABF8C69678101A" box="[523,580,1852,1874]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="13">Fig. 3</figureCitation>
D); abdominal tergites III to V with both anterior and posterior lines of transverse carina complete (
<figureCitation id="137D2A0FFFAEFFBEFEE6F8A395B01027" box="[326,396,1881,1903]" captionStart="FIGURE 10" captionStartId="17.[151,250,1655,1677]" captionTargetBox="[158,1428,193,1633]" captionTargetId="figure@17.[158,1428,193,1634]" captionTargetPageId="17" captionText="FIGURE 10. (A, B) Abdomen, tergite III V, basal transverse carinae: (A) Belonuchus haemorrhoidalis, (B) Odontolinus fasciatus. (C-F) Lateral tergal sclerites IX (styli): (C) Odontolinus fasciatus, (D) Pescolinus schmidti (male), (E) Pescolinus schmidti (female), (F), Philonthus bonariensis. Abbreviations: aL, anterior line; pL, posterior line." httpUri="https://zenodo.org/record/252669/files/figure.png" pageId="13">Fig. 10</figureCitation>
A); mesoventrite with sternopleural (anapleural) suture usually transverse, or nearly transverse (very slightly oblique) (
<figureCitation id="137D2A0FFFAEFFBEFE9EF88D954B10C5" box="[318,375,1911,1933]" captionStart="FIGURE 8" captionStartId="16.[151,250,1354,1376]" captionTargetBox="[155,1431,359,1332]" captionTargetId="figure@16.[155,1431,359,1332]" captionTargetPageId="16" captionText="FIGURE 8. (A E) Mesoventrite, sternopleural (anapleural) suture: (A) Flohria laticornis, (B), Belonuchus mordens, (C) Paederomimus contractus, (D) Cafius xantholoma (Gravenhorst), (E), Neobisnius fraternus. Abbreviation: stS, sternopleural suture." httpUri="https://zenodo.org/record/252667/files/figure.png" pageId="13">Fig. 8</figureCitation>
B)............................................................................
<taxonomicName id="4C464D09FFAEFFBEFA85F88D91A910C4" ID-CoL="8VRHV" box="[1317,1429,1911,1932]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFAEFFBEFA85F88D91A910C4" box="[1317,1429,1911,1932]" italics="true" pageId="13">Belonuchus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFAEFFBEFF37F86F91AB10AC" blockId="13.[151,1436,151,2021]" pageId="13">
<keyLead id="30B795BFFFAEFFBEFF37F86F91AB10AC" pageId="13">
- Anterior angles of pronotum distinctly produced beyond (anteriad of) anterior margin of prosternum (
<figureCitation id="137D2A0FFFAEFFBEFB77F86E912910E2" box="[1239,1301,1940,1962]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="13">Fig. 4</figureCitation>
B); neck with transverse carina (
<figureCitation id="137D2A0FFFAEFFBEFE2FF84895FB1080" box="[399,455,1970,1992]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="13">Fig. 3</figureCitation>
C); abdominal tergites III to V with only anterior line of transverse carina complete; mesoventrite with sternopleural (anapleural) usually distinctly oblique (medial end of suture anterior to its lateral end) (
<figureCitation id="137D2A0FFFAEFFBEFB08F83590E310AD" box="[1192,1247,1999,2021]" captionStart="FIGURE 8" captionStartId="16.[151,250,1354,1376]" captionTargetBox="[155,1431,359,1332]" captionTargetId="figure@16.[155,1431,359,1332]" captionTargetPageId="16" captionText="FIGURE 8. (A E) Mesoventrite, sternopleural (anapleural) suture: (A) Flohria laticornis, (B), Belonuchus mordens, (C) Paederomimus contractus, (D) Cafius xantholoma (Gravenhorst), (E), Neobisnius fraternus. Abbreviation: stS, sternopleural suture." httpUri="https://zenodo.org/record/252667/files/figure.png" pageId="13">Fig. 8</figureCitation>
C).............
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF43FF6D91A01677" blockId="14.[151,1437,151,1229]" pageId="14">
.........................................................................................
<taxonomicName id="4C464D09FFADFFBDFAACFF6D91A717E4" ID-CoL="8W232" box="[1292,1435,151,172]" class="Insecta" family="Staphylinidae" genus="Paederomimus" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFAACFF6D91A717E4" box="[1292,1435,151,172]" italics="true" pageId="14">Paederomimus</emphasis>
</taxonomicName>
20 Superior marginal line of hypomeron not distinctly deflexed under anterior angle of pronotum; lateral puncture of pronotum bearing long seta situated close to superior line or at a distance no more than three times the diameter of puncture (
<figureCitation id="137D2A0FFFADFFBDFA8CFF2B915917AF" box="[1324,1381,209,231]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="14">Fig. 4</figureCitation>
E).................................................................................................... 21
<keyLead id="30B795BFFFADFFBDFF37FEF791A01677" pageId="14">
- Superior marginal line of hypomeron distinctly deflexed under anterior angle of pronotum; lateral puncture of pronotum bearing long seta situated away from superior line at a distance at least three times as large as diameter of puncture (
<figureCitation id="137D2A0FFFADFFBDFAB1FED391771677" box="[1297,1355,297,319]" captionStart="FIGURE 4" captionStartId="11.[151,250,1446,1468]" captionTargetBox="[260,1326,397,1425]" captionTargetId="figure@11.[260,1326,397,1425]" captionTargetPageId="11" captionText="FIGURE 4. (A K) Proesternum and hypomeron: (A, F) Gabrius nidicola (Bernhauer), (B) Paederomimus contractus, (C) Holisus humilis, (D, K) Delgadobius amazonensis, (E) Philonthus bonariensis Bernhauer, (G) Belonuchus haemorrhoidalis, (H) Paederomimus contractus, (I) Phileciton badariottii, (J) Neobisnius fraternus. Abbreviations: aL, additional line of hypomeron; iL, inferior line of hypomeron; lp, lateral puncture of pronotum; PtC, proesternal transverse carina; sL, superior line of hypomeron." httpUri="https://zenodo.org/record/252663/files/figure.png" pageId="14">Fig. 4</figureCitation>
J)...23
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FEBD91A01632" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FEBD91A01614" blockId="14.[151,1437,151,1229]" box="[151,1436,327,349]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FEBD91A01614" box="[151,1436,327,349]" pageId="14">
21 Neck without dorsal basal ridge; last segment of labial palpus distinctly narrower than preceding segment.......
<taxonomicName id="4C464D09FFADFFBDFA8AFEBD91A01614" ID-CoL="8VRPD" box="[1322,1436,327,348]" class="Insecta" family="Staphylinidae" genus="Gabronthus" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFA8AFEBD91A01614" box="[1322,1436,327,348]" italics="true" pageId="14">Gabronthus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FE9F91A01632" blockId="14.[151,1437,151,1229]" box="[151,1436,356,378]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FE9F91A01632" box="[151,1436,356,378]" pageId="14">- Neck with dorsal basal ridge; last segment of labial palpus not distinctly narrower than preceding segment..............22</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FE7B91A0169A" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FE7B91A516DF" blockId="14.[151,1437,151,1229]" box="[151,1433,385,407]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FE7B91A516DF" box="[151,1433,385,407]" pageId="14">
22 Head with completely obliterated hind angles; last segment of both maxillary and labial palpi subulate……………….
<taxonomicName id="4C464D09FFADFFBDFAEBFE7891A516DF" ID-CoL="8VS2Q" box="[1355,1433,386,407]" class="Insecta" family="Staphylinidae" genus="Rabigus" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFAEBFE7891A516DF" box="[1355,1433,386,407]" italics="true" pageId="14">Rabigus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FE6591A0169A" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FE6591A0169A" pageId="14">
- Head with hind angles variably shaped; last segment of both maxillary and labial palpi fusiform to gradually narrowed into acute or subacute apex, never subulate............................................................ ..
<taxonomicName id="4C464D09FFADFFBDFA94FE4791A0169A" ID-CoL="8VRYR" box="[1332,1436,445,466]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFA94FE4791A0169A" box="[1332,1436,445,466]" italics="true" pageId="14">Philonthus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FE2391A71545" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FE2391A716A7" blockId="14.[151,1437,151,1229]" box="[151,1435,473,495]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FE2391A716A7" box="[151,1435,473,495]" pageId="14">23 Antennal segment 1 distinctly longer than half of head length.................................................. 24</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FE0D91A71545" blockId="14.[151,1437,151,1229]" box="[151,1435,503,525]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FE0D91A71545" box="[151,1435,503,525]" pageId="14">- Antennal segment 1 not longer than half of head length....................................................... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FDEE91A715CA" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FDEE91A7150F" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FDEE91A7150F" pageId="14">
24 Antennal segment 1 with apical spine-like seta (
<figureCitation id="137D2A0FFFADFFBDFD0AFDEE96DF1562" box="[682,739,532,554]" captionStart="FIGURE 3" captionStartId="10.[151,250,1683,1705]" captionTargetBox="[160,1427,735,1661]" captionTargetId="figure@10.[160,1427,735,1661]" captionTargetPageId="10" captionText="FIGURE 3. (A F) Head, dorsal view: (A) Flohria subcoerulea, (B, F) Odontolinus fasciatus Sharp, (C) Paederomimus contractus, (D) Belonuchus haemorrhoidalis (Fabricius), (E) Proxenobius borgmeieri Seevers. Abbreviations: alR, anterolateral-dorsal ridge; ioR, infraorbital ridge; NtC, transverse carina of neck; S, spine-like seta." httpUri="https://zenodo.org/record/252662/files/figure.png" pageId="14">Fig. 3</figureCitation>
F); elytra bicoloured, abdomen with distinct color pattern (
<figureCitation id="137D2A0FFFADFFBDFAB2FDEE916B1562" box="[1298,1367,532,554]" captionStart="FIGURE 12" captionStartId="19.[151,250,1893,1915]" captionTargetBox="[291,1312,196,1862]" captionTargetId="figure@19.[283,1312,193,1868]" captionTargetPageId="19" captionText="FIGURE 12. (A J) Habitus: (A) Holisus humilis Erichson (Hyptiomina), (B) Laetulonthus laetulus (Say), (C) Leptopeltus weyrauchi Chani-Posse &amp; Asenjo, (D) Leptopeltoides ecuatoriensis Chani-Posse &amp; Asenjo, (E) Linoderus gracilipes Sharp, (F) Neobisnius fraternus Bernhauer, (G) Odontolinus fasciatus Sharp, (H) Ophionthus serpentinus Bernhauer, (I) Paederallus fragilis Sharp, (J) Paederomimus debilis Sharp." httpUri="https://zenodo.org/record/252671/files/figure.png" pageId="14">Fig. 12</figureCitation>
G); lateral tergal sclerites IX (styli) dorsoventrally flattened, distinctly dilated (
<figureCitation id="137D2A0FFFADFFBDFC38FDCB97E0150F" box="[920,988,561,583]" captionStart="FIGURE 10" captionStartId="17.[151,250,1655,1677]" captionTargetBox="[158,1428,193,1633]" captionTargetId="figure@17.[158,1428,193,1634]" captionTargetPageId="17" captionText="FIGURE 10. (A, B) Abdomen, tergite III V, basal transverse carinae: (A) Belonuchus haemorrhoidalis, (B) Odontolinus fasciatus. (C-F) Lateral tergal sclerites IX (styli): (C) Odontolinus fasciatus, (D) Pescolinus schmidti (male), (E) Pescolinus schmidti (female), (F), Philonthus bonariensis. Abbreviations: aL, anterior line; pL, posterior line." httpUri="https://zenodo.org/record/252669/files/figure.png" pageId="14">Fig. 10</figureCitation>
C).........................
<taxonomicName id="4C464D09FFADFFBDFA85FDC891A7150F" ID-CoL="8X7MF" box="[1317,1435,562,583]" class="Insecta" family="Staphylinidae" genus="Odontolinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFA85FDC891A7150F" box="[1317,1435,562,583]" italics="true" pageId="14">Odontolinus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FDB591A715CA" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FDB591A715CA" pageId="14">
- Antennal segment 1 without apical spine-like seta; elytra unicolorous, abdomen with apical segments testaceous, distinctly lighter than the preceding segments (
<figureCitation id="137D2A0FFFADFFBDFDE3FD9696B515CA" box="[579,649,620,642]" captionStart="FIGURE 12" captionStartId="19.[151,250,1893,1915]" captionTargetBox="[291,1312,196,1862]" captionTargetId="figure@19.[283,1312,193,1868]" captionTargetPageId="19" captionText="FIGURE 12. (A J) Habitus: (A) Holisus humilis Erichson (Hyptiomina), (B) Laetulonthus laetulus (Say), (C) Leptopeltus weyrauchi Chani-Posse &amp; Asenjo, (D) Leptopeltoides ecuatoriensis Chani-Posse &amp; Asenjo, (E) Linoderus gracilipes Sharp, (F) Neobisnius fraternus Bernhauer, (G) Odontolinus fasciatus Sharp, (H) Ophionthus serpentinus Bernhauer, (I) Paederallus fragilis Sharp, (J) Paederomimus debilis Sharp." httpUri="https://zenodo.org/record/252671/files/figure.png" pageId="14">Fig. 12</figureCitation>
H); lateral tergal sclerites IX (styli) not as above..................
<taxonomicName id="4C464D09FFADFFBDFA88FD9791A715CA" ID-CoL="8XND3" box="[1320,1435,621,642]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFA88FD9791A715CA" box="[1320,1435,621,642]" italics="true" pageId="14">Ophionthus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FD7391A715BF" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FD7391A715F4" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FD7391A715F4" pageId="14">
25 Anterior angles of pronotum distinctly produced beyond (anteriad of) anterior margin of prosternum; lateral tergal sclerites IX (styli) dorsoventrally flattened, slightly to distinctly dilated (
<figureCitation id="137D2A0FFFADFFBDFC85FD5D974F15F5" box="[805,883,679,701]" captionStart="FIGURE 10" captionStartId="17.[151,250,1655,1677]" captionTargetBox="[158,1428,193,1633]" captionTargetId="figure@17.[158,1428,193,1634]" captionTargetPageId="17" captionText="FIGURE 10. (A, B) Abdomen, tergite III V, basal transverse carinae: (A) Belonuchus haemorrhoidalis, (B) Odontolinus fasciatus. (C-F) Lateral tergal sclerites IX (styli): (C) Odontolinus fasciatus, (D) Pescolinus schmidti (male), (E) Pescolinus schmidti (female), (F), Philonthus bonariensis. Abbreviations: aL, anterior line; pL, posterior line." httpUri="https://zenodo.org/record/252669/files/figure.png" pageId="14">Figs. 10</figureCitation>
D, E)................................
<taxonomicName id="4C464D09FFADFFBDFA93FD5D91A715F4" ID-CoL="8XNSR" box="[1331,1435,679,700]" class="Insecta" family="Staphylinidae" genus="Pescolinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFA93FD5D91A715F4" box="[1331,1435,679,700]" italics="true" pageId="14">Pescolinus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FD3F91A715BF" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FD3F91A715BF" pageId="14">
- Anterior angles of pronotum not or slightly produced beyond (anteriad of) anterior margin of prosternum; lateral tergal sclerites IX (styli) not as above (
<figureCitation id="137D2A0FFFADFFBDFE4BFD1B960C15BF" box="[491,560,737,759]" captionStart="FIGURE 10" captionStartId="17.[151,250,1655,1677]" captionTargetBox="[158,1428,193,1633]" captionTargetId="figure@17.[158,1428,193,1634]" captionTargetPageId="17" captionText="FIGURE 10. (A, B) Abdomen, tergite III V, basal transverse carinae: (A) Belonuchus haemorrhoidalis, (B) Odontolinus fasciatus. (C-F) Lateral tergal sclerites IX (styli): (C) Odontolinus fasciatus, (D) Pescolinus schmidti (male), (E) Pescolinus schmidti (female), (F), Philonthus bonariensis. Abbreviations: aL, anterior line; pL, posterior line." httpUri="https://zenodo.org/record/252669/files/figure.png" pageId="14">Fig. 10</figureCitation>
F).....................................................................26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FD0591A01425" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FD0591A7147A" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FD0591A7147A" pageId="14">
26 Disc of mesoventrite situated distinctly more ventrally than median part (mesoventral part of mesocoxal acetabula) and mid coxae contiguous (
<figureCitation id="137D2A0FFFADFFBDFE35FCE695F1147A" box="[405,461,796,818]" captionStart="FIGURE 8" captionStartId="16.[151,250,1354,1376]" captionTargetBox="[155,1431,359,1332]" captionTargetId="figure@16.[155,1431,359,1332]" captionTargetPageId="16" captionText="FIGURE 8. (A E) Mesoventrite, sternopleural (anapleural) suture: (A) Flohria laticornis, (B), Belonuchus mordens, (C) Paederomimus contractus, (D) Cafius xantholoma (Gravenhorst), (E), Neobisnius fraternus. Abbreviation: stS, sternopleural suture." httpUri="https://zenodo.org/record/252667/files/figure.png" pageId="14">Fig. 8</figureCitation>
E)......................................................................
<taxonomicName id="4C464D09FFADFFBDFA91FCE791A7147A" ID-CoL="8VRW8" box="[1329,1435,797,818]" class="Insecta" family="Staphylinidae" genus="Neobisnius" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFA91FCE791A7147A" box="[1329,1435,797,818]" italics="true" pageId="14">Neobisnius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FCC091A01425" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FCC091A01425" pageId="14">
- Disc of mesoventrite situated more or less in one plane with, or only slightly more ventrally than its median portion (mesoventral part of mesocoxal acetabula) and mid coxae more or less separated by elevated part of metaventrite (
<figureCitation id="137D2A0FFFADFFBDFB6CFCAD91381425" box="[1228,1284,855,877]" captionStart="FIGURE 8" captionStartId="16.[151,250,1354,1376]" captionTargetBox="[155,1431,359,1332]" captionTargetId="figure@16.[155,1431,359,1332]" captionTargetPageId="16" captionText="FIGURE 8. (A E) Mesoventrite, sternopleural (anapleural) suture: (A) Flohria laticornis, (B), Belonuchus mordens, (C) Paederomimus contractus, (D) Cafius xantholoma (Gravenhorst), (E), Neobisnius fraternus. Abbreviation: stS, sternopleural suture." httpUri="https://zenodo.org/record/252667/files/figure.png" pageId="14">Fig. 8</figureCitation>
D)........27
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FC8E91A714AA" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FC8E91A714EF" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FC8E91A714EF" pageId="14">
27 Head with infraorbital ridge moderately long, extending some distance in front of the lateral bend of the postgenal ridge (
<figureCitation id="137D2A0FFFADFFBDFAD5FC8E94D214EF" captionStart="FIGURE 2" captionStartId="8.[151,250,1603,1625]" captionTargetBox="[156,1430,193,1579]" captionTargetId="figure@8.[156,1430,193,1581]" captionTargetPageId="8" captionText="FIGURE 2. (A K) Head, ventral view: (A) Hesperus fasciatus (Sharp), (B) Belonuchus mordens Erichson, (C) Neobisnius fraternus Bernhauer, (D) Paederomimus contractus Sharp, (E) Ecitophytes coniceps Wasmann, (F) Delgadobius amazonensis Chani-Posse &amp; Couturier, (G, K) Flohria subcoerulea, (H) Laetulonthus laetulus (Say), (I) Holisus humilis Erichson, (J) Phileciton badariottii Wasmann. Abbreviations: ioR, infraorbital ridge; pgR, postgenal ridge; pmR, postmandibular ridge; poS, postoccipital suture; PtC, proesternal transverse carina; vbR, ventral basal ridge." httpUri="https://zenodo.org/record/252661/files/figure.png" pageId="14">Fig. 2</figureCitation>
H); front tibia not spinose on lateral face (
<figureCitation id="137D2A0FFFADFFBDFDDFFC6B968514EF" box="[639,697,913,935]" captionStart="FIGURE 7" captionStartId="15.[151,250,1400,1422]" captionTargetBox="[173,1433,193,1355]" captionTargetId="figure@15.[154,1433,193,1379]" captionTargetPageId="15" captionText="FIGURE 7. (A H) (A, B, H) Atopocentrum mirabile Bernhauer: (A) front femur, (B) front tibia, (H) hind tibiae. (C) Pescolinus schmidti: front coxa and femur. (D, E) Proxenobius borgmeieri: (D) front leg, (E) front tarsus. (F, G) Front tibia and tarsus: (F) Laetulonthus laetulus, (G) Hesperus baltimorensis (Gravenhorst)." httpUri="https://zenodo.org/record/252666/files/figure.png" pageId="14">Fig. 7</figureCitation>
F)..................................................
<taxonomicName id="4C464D09FFADFFBDFABEFC6891A714EF" ID-CoL="8VRS2" box="[1310,1435,914,935]" class="Insecta" family="Staphylinidae" genus="Laetulonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFABEFC6891A714EF" box="[1310,1435,914,935]" italics="true" pageId="14">Laetulonthus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FC5591A714AA" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FC5591A714AA" pageId="14">
- Head with infraorbital ridge very short, becoming obsolete just in front of the lateral portion of the postgenal ridge (
<figureCitation id="137D2A0FFFADFFBDFAE2FC559141148D" box="[1346,1405,943,965]" captionStart="FIGURE 2" captionStartId="8.[151,250,1603,1625]" captionTargetBox="[156,1430,193,1579]" captionTargetId="figure@8.[156,1430,193,1581]" captionTargetPageId="8" captionText="FIGURE 2. (A K) Head, ventral view: (A) Hesperus fasciatus (Sharp), (B) Belonuchus mordens Erichson, (C) Neobisnius fraternus Bernhauer, (D) Paederomimus contractus Sharp, (E) Ecitophytes coniceps Wasmann, (F) Delgadobius amazonensis Chani-Posse &amp; Couturier, (G, K) Flohria subcoerulea, (H) Laetulonthus laetulus (Say), (I) Holisus humilis Erichson, (J) Phileciton badariottii Wasmann. Abbreviations: ioR, infraorbital ridge; pgR, postgenal ridge; pmR, postmandibular ridge; poS, postoccipital suture; PtC, proesternal transverse carina; vbR, ventral basal ridge." httpUri="https://zenodo.org/record/252661/files/figure.png" pageId="14">Fig. 2</figureCitation>
A); front tibia spinose on lateral face (
<figureCitation id="137D2A0FFFADFFBDFD89FC36965D14AA" box="[553,609,972,994]" captionStart="FIGURE 7" captionStartId="15.[151,250,1400,1422]" captionTargetBox="[173,1433,193,1355]" captionTargetId="figure@15.[154,1433,193,1379]" captionTargetPageId="15" captionText="FIGURE 7. (A H) (A, B, H) Atopocentrum mirabile Bernhauer: (A) front femur, (B) front tibia, (H) hind tibiae. (C) Pescolinus schmidti: front coxa and femur. (D, E) Proxenobius borgmeieri: (D) front leg, (E) front tarsus. (F, G) Front tibia and tarsus: (F) Laetulonthus laetulus, (G) Hesperus baltimorensis (Gravenhorst)." httpUri="https://zenodo.org/record/252666/files/figure.png" pageId="14">Fig. 7</figureCitation>
G)................................................................ 28
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FC1391A7131F" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FC1391A71354" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FC1391A71354" pageId="14">
28 Last segment of both maxillary and labial palpi subcylindrical (
<figureCitation id="137D2A0FFFADFFBDFCE1FC13974714B7" box="[833,891,1001,1023]" captionStart="FIGURE 2" captionStartId="8.[151,250,1603,1625]" captionTargetBox="[156,1430,193,1579]" captionTargetId="figure@8.[156,1430,193,1581]" captionTargetPageId="8" captionText="FIGURE 2. (A K) Head, ventral view: (A) Hesperus fasciatus (Sharp), (B) Belonuchus mordens Erichson, (C) Neobisnius fraternus Bernhauer, (D) Paederomimus contractus Sharp, (E) Ecitophytes coniceps Wasmann, (F) Delgadobius amazonensis Chani-Posse &amp; Couturier, (G, K) Flohria subcoerulea, (H) Laetulonthus laetulus (Say), (I) Holisus humilis Erichson, (J) Phileciton badariottii Wasmann. Abbreviations: ioR, infraorbital ridge; pgR, postgenal ridge; pmR, postmandibular ridge; poS, postoccipital suture; PtC, proesternal transverse carina; vbR, ventral basal ridge." httpUri="https://zenodo.org/record/252661/files/figure.png" pageId="14">Fig. 2</figureCitation>
A); mesoventrite with intercoxal process truncate to subtruncate.......................................................................................
<taxonomicName id="4C464D09FFADFFBDFAE1FBFD91A71354" ID-CoL="8VRQP" box="[1345,1435,1031,1052]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFAE1FBFD91A71354" box="[1345,1435,1031,1052]" italics="true" pageId="14">Hesperus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FBDF91A7131F" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FBDF91A7131F" pageId="14">- Last segment of both maxillary and labial palpi rather fusiform, not subcylindrical; mesoventrite with intercoxal process either acute or rounded apically..............................................................................29</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B22E2FFFADFFBDFF37FBA591A71384" pageId="14">
<paragraph id="8BF9368AFFADFFBDFF37FBA591A713DA" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FBA591A713DA" pageId="14">
29 Mesoventrite with sternopleural (anapleural) suture distinctly oblique (medial end of suture anterior to its lateral end) and intercoxal process acute apically (
<figureCitation id="137D2A0FFFADFFBDFD86FB86966213DA" box="[550,606,1148,1170]" captionStart="FIGURE 8" captionStartId="16.[151,250,1354,1376]" captionTargetBox="[155,1431,359,1332]" captionTargetId="figure@16.[155,1431,359,1332]" captionTargetPageId="16" captionText="FIGURE 8. (A E) Mesoventrite, sternopleural (anapleural) suture: (A) Flohria laticornis, (B), Belonuchus mordens, (C) Paederomimus contractus, (D) Cafius xantholoma (Gravenhorst), (E), Neobisnius fraternus. Abbreviation: stS, sternopleural suture." httpUri="https://zenodo.org/record/252667/files/figure.png" pageId="14">Fig. 8</figureCitation>
D)..............................................................
<taxonomicName id="4C464D09FFADFFBDFAFCFB8791A713DA" ID-CoL="8VRJY" box="[1372,1435,1149,1170]" class="Insecta" family="Staphylinidae" genus="Cafius" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFAFCFB8791A713DA" box="[1372,1435,1149,1170]" italics="true" pageId="14">Cafius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF9368AFFADFFBDFF37FB6091A71384" blockId="14.[151,1437,151,1229]" pageId="14">
<keyLead id="30B795BFFFADFFBDFF37FB6091A71384" pageId="14">
- Mesoventrite with sternopleural (anapleural) suture transverse, or nearly transverse (very slightly oblique) and mesoventrite with intercoxal process rounded apically...............................................................
<taxonomicName id="4C464D09FFADFFBDFAF7FB4D91A71384" ID-CoL="8VS2V" box="[1367,1435,1207,1228]" class="Insecta" family="Staphylinidae" genus="Remus" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="B932EA98FFADFFBDFAF7FB4D91A71384" box="[1367,1435,1207,1228]" italics="true" pageId="14">Remus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>