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treatments-xml/data/03/EF/62/03EF6217BF7AFF870AC0FD919F22AE23.xml
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<document id="CE234702C3075DF0CB0172ECD04E83B4" ID-CLB-Dataset="20762" ID-DOI="10.11646/zootaxa.3365.1.1" ID-GBIF-Dataset="1db9bc74-46dd-4263-9784-88a59bd05d59" ID-ISSN="1175­5334" ID-Zenodo-Dep="6038900" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629787146833" checkinUser="felipe" docAuthor="Garcia, Francisco Hita &amp; Fisher, Brian L." docDate="2012" docId="03EF6217BF7AFF870AC0FD919F22AE23" docLanguage="en" docName="zt03365p123.pdf" docOrigin="Zootaxa 3365" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Tetramorium macki Garcia &amp; Fisher 2012, sp. n." docType="treatment" docVersion="9" lastPageNumber="70" masterDocId="FFD61A6FBF39FFC20A57FF889C5BAB77" masterDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups" masterLastPageNumber="123" masterPageNumber="1" pageNumber="68" updateTime="1699142484625" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="07687A55DD571B3EBDDBE9F39C6F07BB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="837CA7C5CE0EE51218538A719F06C47D">
<mods:title id="9ED848A3DE33D1128F2B18E915F9A93D">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups</mods:title>
</mods:titleInfo>
<mods:name id="4F71B924A4F7F0AEE220CB055FAA6347" type="personal">
<mods:role id="03C4601C33A32E0C7F2DB575A9D87255">
<mods:roleTerm id="8870E35009440AADDFB533019B66237C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13D7D21B21C53463EDB3489DAC55DE41">Garcia, Francisco Hita</mods:namePart>
</mods:name>
<mods:name id="3E98E6628D3A25689A5FE4BB4C4F33D4" type="personal">
<mods:role id="FEE93A31FD880CA7D2F0BA21CD966C9F">
<mods:roleTerm id="B4199660A1B40B8F2B6460FC474AE499">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EED14D2177866EF4EA7F5B06472F6314">Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E9C4E969F543E8E19D6391E03890BEF3">text</mods:typeOfResource>
<mods:relatedItem id="95F75A7A8EB2F964FC69E90ADAD08C7B" type="host">
<mods:titleInfo id="B4D3290B21A8A4A2B6358714FFC44638">
<mods:title id="B993B1DDC7D84CCDA9178BF00F005E97">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="12E157B0E49B06D12E3A047E7894EC40">
<mods:date id="0C9C1CE286F108B75D3C0DBD56F22F46">2012</mods:date>
<mods:detail id="B11D1B3A60EDE12C9A1E4443FCFB01BB" type="pubDate">
<mods:number id="F07BDE9FA74EFFA152B6CED01268623F">2012-07-04</mods:number>
</mods:detail>
<mods:detail id="DBA7BDBD24FAB4EF8C0D8D372E0DAB90" type="volume">
<mods:number id="F05BE9DBAF933C82C35836E192ADB849">3365</mods:number>
</mods:detail>
<mods:extent id="4A2A1562F36D8E5EB1637EACD05C2637" unit="page">
<mods:start id="0C433EDC3E87B297B2B6A3E691A52903">1</mods:start>
<mods:end id="7EA564678B5F0F0BF1C532A9C8F2E021">123</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="28D6A1E08DBCB06BF3DD17D8C7EDF029">journal article</mods:classification>
<mods:identifier id="AB68824CECC5DFE9D4BC978087D93420" type="CLB-Dataset">20762</mods:identifier>
<mods:identifier id="38CC1B3627046F264FFC265F8F12B757" type="DOI">10.11646/zootaxa.3365.1.1</mods:identifier>
<mods:identifier id="5549DF19D62F041CB2B5A1CF6D3DCCCE" type="GBIF-Dataset">1db9bc74-46dd-4263-9784-88a59bd05d59</mods:identifier>
<mods:identifier id="F51DF1FA29C35729F30AE361BCD7055E" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="270931D478E4C58E768237DDF620FA02" type="Zenodo-Dep">6038900</mods:identifier>
</mods:mods>
<treatment id="03EF6217BF7AFF870AC0FD919F22AE23" ID-DOI="http://doi.org/10.5281/zenodo.5253660" ID-GBIF-Taxon="193281035" ID-Zenodo-Dep="5253660" LSID="urn:lsid:plazi:treatment:03EF6217BF7AFF870AC0FD919F22AE23" httpUri="http://treatment.plazi.org/id/03EF6217BF7AFF870AC0FD919F22AE23" lastPageId="69" lastPageNumber="70" pageId="67" pageNumber="68">
<subSubSection id="C35C808ABF7AFF810AC0FD919D94A945" box="[151,463,536,563]" pageId="67" pageNumber="68" type="nomenclature">
<paragraph id="8BF9D301BF7AFF810AC0FD919D94A945" blockId="67.[151,463,536,596]" box="[151,463,536,563]" pageId="67" pageNumber="68">
<heading id="D0B1646DBF7AFF810AC0FD919D94A945" bold="true" box="[151,463,536,563]" fontSize="11" level="1" pageId="67" pageNumber="68" reason="1">
<emphasis id="B9320F13BF7AFF810AC0FD919D94A945" bold="true" box="[151,463,536,563]" pageId="67" pageNumber="68">
<taxonomicName id="4C46A882BF7AFF810AC0FD919DDDA944" authority="Garcia &amp; Fisher, 2012" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[151,390,537,563]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="macki" status="sp. nov.">
<emphasis id="B9320F13BF7AFF810AC0FD919DDDA944" bold="true" box="[151,390,537,563]" italics="true" pageId="67" pageNumber="68">Tetramorium macki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201B268BF7AFF810BDAFD909D94A945" box="[397,463,536,562]" pageId="67" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7AFF810AC0FDB49DC5A923" box="[151,414,572,596]" pageId="67" pageNumber="68" type="description">
<paragraph id="8BF9D301BF7AFF810AC0FDB49DC5A923" blockId="67.[151,463,536,596]" box="[151,414,572,596]" pageId="67" pageNumber="68">(figs 76, 101, 102, 103)</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7AFF810AC0FDF59F74A852" pageId="67" pageNumber="68" type="materials_examined">
<paragraph id="8BF9D301BF7AFF810AC0FDF59F74A852" blockId="67.[151,1436,637,805]" pageId="67" pageNumber="68">
<materialsCitation id="3B2ED95CBF7AFF810AC0FDF59D28A9AA" ID-GBIF-Occurrence="3462706286" collectingDate="1999-01-21" collectingDateMax="1999-01-24" collectingDateMin="1999-01-21" collectionCode="CASC" collectorName="F. C. Sandranantitra &amp; H. J. Ratsirarson" country="Madagascar" elevation="450" latitude="-18.04833" location="Toamasina" longLatPrecision="1" longitude="49.09167" pageId="67" pageNumber="68" specimenCode="CASENT0189093" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="54FD6DA3BF7AFF810AC0FDF59CA4A9E2" box="[151,255,637,661]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
worker,
<collectingCountry id="F3519391BF7AFF810B32FDF59E74A9E2" box="[357,559,637,661]" name="Madagascar" pageId="67" pageNumber="68">MADAGASCAR</collectingCountry>
,
<collectingRegion id="49821DE3BF7AFF81086BFDF59EE3A9E2" box="[572,696,637,661]" country="Madagascar" name="Toamasina" pageId="67" pageNumber="68">Toamasina</collectingRegion>
,
<collectorName id="26B3B6D7BF7AFF810892FDF59FF9A9E2" box="[709,930,637,661]" pageId="67" pageNumber="68">F.C. Sandranantitra</collectorName>
,
<geoCoordinate id="EE72B5C6BF7AFF8109F8FDF5986AA9E2" box="[943,1073,637,661]" degrees="18.04833" direction="south" orientation="latitude" pageId="67" pageNumber="68" precision="1" value="-18.04833">18.04833 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF7AFF810E68FDF5989FA9E2" box="[1087,1220,637,661]" degrees="49.09167" direction="east" orientation="longitude" pageId="67" pageNumber="68" precision="1" value="49.09167">49.09167 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF7AFF810E86FDF59947A9E2" box="[1233,1308,637,661]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="67" pageNumber="68" unit="m" value="450.0">
<elevation id="006B3432BF7AFF810E86FDF59947A9E2" box="[1233,1308,637,661]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="67" pageNumber="68" unit="m" value="450.0">450 m</elevation>
</quantity>
, rainforest, sifted litter (leaf mold, rotten wood), collection code HJR102,
<date id="FFF8F5C1BF7AFF8109FAFD299812A9CE" box="[941,1097,673,697]" pageId="67" pageNumber="68" value="1999-01-21" valueMax="1999-01-24" valueMin="1999-01-21">
<collectingDate id="EFBC0C29BF7AFF8109FAFD299812A9CE" box="[941,1097,673,697]" pageId="67" pageNumber="68" value="1999-01-21" valueMax="1999-01-24" valueMin="1999-01-21">21.–24.I.1999</collectingDate>
</date>
(
<emphasis id="B9320F13BF7AFF810E33FD2A997CA9CE" box="[1124,1319,674,697]" italics="true" pageId="67" pageNumber="68">
<collectorName id="26B3B6D7BF7AFF810E33FD2A997CA9CE" box="[1124,1319,674,697]" pageId="67" pageNumber="68">H.J. Ratsirarson</collectorName>
</emphasis>
) (
<collectionCode id="ED574BC4BF7AFF810F1DFD2999C2A9CE" box="[1354,1433,673,697]" pageId="67" pageNumber="68">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF7AFF810AC0FD4D9D30A9AA" box="[151,363,709,733]" collectionCode="CASC" pageId="67" pageNumber="68">CASENT0189093</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B2ED95CBF7AFF810B2AFD4E99C7A876" ID-GBIF-Occurrence="3462705424" collectingDate="1998-12-04" collectingDateMax="1998-12-07" collectingDateMin="1998-12-04" collectionCode="CASC" collectorName="F. C. Andriantantely &amp; H. J. Ratsirarson" country="Madagascar" elevation="530" latitude="-18.695" location="Toamasina" longLatPrecision="55" longitude="48.81333" pageId="67" pageNumber="68" specimenCode="CASENT0217710, CASENT0217711, CASENT0218032, CASENT0247151" specimenCount="1" specimenCount-worker="1" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus id="54FD6DA3BF7AFF810B2AFD4E9DB6A9A9" box="[381,493,710,734]" pageId="67" pageNumber="68" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D401888BF7AFF810BAFFD4E9EC2A9AA" box="[504,665,709,733]" pageId="67" pageNumber="68" type="worker">seven workers</specimenCount>
from
<collectingRegion id="49821DE3BF7AFF81088AFD4D9F02A9AA" box="[733,857,709,733]" country="Madagascar" name="Toamasina" pageId="67" pageNumber="68">Toamasina</collectingRegion>
,
<collectorName id="26B3B6D7BF7AFF810933FD4D9819A9AA" box="[868,1090,709,733]" pageId="67" pageNumber="68">F.C. Andriantantely</collectorName>
,
<geoCoordinate id="EE72B5C6BF7AFF810E1AFD4D98E9A9AA" box="[1101,1202,709,733]" degrees="18.695" direction="south" orientation="latitude" pageId="67" pageNumber="68" precision="55" value="-18.695">18.695 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF7AFF810EE8FD4D9919A9AA" box="[1215,1346,709,733]" degrees="48.81333" direction="east" orientation="longitude" pageId="67" pageNumber="68" precision="1" value="48.81333">48.81333 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF7AFF810F19FD4D99CCA9AA" box="[1358,1431,709,733]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="67" pageNumber="68" unit="m" value="530.0">
<elevation id="006B3432BF7AFF810F19FD4D99CCA9AA" box="[1358,1431,709,733]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="67" pageNumber="68" unit="m" value="530.0">530 m</elevation>
</quantity>
, rainforest, sifted litter, collection code HJR121,
<date id="FFF8F5C1BF7AFF810882FD619F28A876" box="[725,883,745,769]" pageId="67" pageNumber="68" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">
<collectingDate id="EFBC0C29BF7AFF810882FD619F28A876" box="[725,883,745,769]" pageId="67" pageNumber="68" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">4.–7.XII.1998</collectingDate>
</date>
(
<emphasis id="B9320F13BF7AFF8109DEFD629813A876" box="[905,1096,746,769]" italics="true" pageId="67" pageNumber="68">
<collectorName id="26B3B6D7BF7AFF8109DEFD629813A876" box="[905,1096,746,769]" pageId="67" pageNumber="68">H.J. Ratsirarson</collectorName>
</emphasis>
) (
<collectionCode id="ED574BC4BF7AFF810E30FD6198EDA876" box="[1127,1206,745,769]" pageId="67" pageNumber="68">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF7AFF810E91FD6199C3A876" box="[1222,1432,745,769]" collectionCode="CASC" pageId="67" pageNumber="68">CASENT0217710</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2ED95CBF7AFF810AC0FC859D30A852" ID-GBIF-Occurrence="3462705475" box="[151,363,781,805]" collectingDate="1998-12-04" collectingDateMax="1998-12-07" collectingDateMin="1998-12-04" collectionCode="CASC" collectorName="F. C. Andriantantely &amp; H. J. Ratsirarson" country="Madagascar" elevation="530" latitude="-18.695" location="Toamasina" longLatPrecision="55" longitude="48.81333" pageId="67" pageNumber="68" specimenCode="CASENT0217710, CASENT0217711, CASENT0218032, CASENT0247151" specimenCount="1" specimenCount-worker="1" stateProvince="Toamasina" typeStatus="paratype">
<specimenCode id="DBE07B7ABF7AFF810AC0FC859D3CA852" box="[151,359,781,805]" collectionCode="CASC" pageId="67" pageNumber="68">CASENT0217711</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2ED95CBF7AFF810B24FC859E13A852" ID-GBIF-Occurrence="3462705603" box="[371,584,781,805]" collectingDate="1998-12-04" collectingDateMax="1998-12-07" collectingDateMin="1998-12-04" collectionCode="CASC" collectorName="F. C. Andriantantely &amp; H. J. Ratsirarson" country="Madagascar" elevation="530" latitude="-18.695" location="Toamasina" longLatPrecision="55" longitude="48.81333" pageId="67" pageNumber="68" specimenCode="CASENT0217710, CASENT0217711, CASENT0218032, CASENT0247151" specimenCount="1" specimenCount-worker="1" stateProvince="Toamasina" typeStatus="paratype">
<specimenCode id="DBE07B7ABF7AFF810B24FC859E1EA852" box="[371,581,781,805]" collectionCode="CASC" pageId="67" pageNumber="68">CASENT0218032</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2ED95CBF7AFF810818FC859F77A852" ID-GBIF-Occurrence="3462706280" box="[591,812,781,805]" collectingDate="1998-12-04" collectingDateMax="1998-12-07" collectingDateMin="1998-12-04" collectionCode="CASC" collectorName="F. C. Andriantantely &amp; H. J. Ratsirarson" country="Madagascar" elevation="530" latitude="-18.695" location="Toamasina" longLatPrecision="55" longitude="48.81333" pageId="67" pageNumber="68" specimenCode="CASENT0217710, CASENT0217711, CASENT0218032, CASENT0247151" specimenCount="1" specimenCount-worker="1" stateProvince="Toamasina" typeStatus="paratype">
<specimenCode id="DBE07B7ABF7AFF810818FC859F7FA852" box="[591,804,781,805]" collectionCode="CASC" pageId="67" pageNumber="68">CASENT0247151</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7AFF810AC0FCDD9EB8AF56" pageId="67" pageNumber="68" type="diagnosis">
<paragraph id="8BF9D301BF7AFF810AC0FCDD9D51A819" blockId="67.[151,1436,853,1058]" box="[151,266,853,878]" pageId="67" pageNumber="68">
<heading id="D0B1646DBF7AFF810AC0FCDD9D51A819" bold="true" box="[151,266,853,878]" fontSize="10" level="2" pageId="67" pageNumber="68" reason="2">
<emphasis id="B9320F13BF7AFF810AC0FCDD9D51A819" bold="true" box="[151,266,853,878]" pageId="67" pageNumber="68">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7AFF810AC0FCF29EB8AF56" blockId="67.[151,1436,853,1058]" pageId="67" pageNumber="68">
<taxonomicName id="4C46A882BF7AFF810AC0FCF29D21A8E6" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[151,378,889,913]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7AFF810AC0FCF29D21A8E6" box="[151,378,889,913]" italics="true" pageId="67" pageNumber="68">Tetramorium macki</emphasis>
</taxonomicName>
can be recognised easily within the
<taxonomicName id="4C46A882BF7AFF810912FCF29F94A8E6" authorityName="Forel" authorityYear="1892" box="[837,975,889,913]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="steinheili">
<emphasis id="B9320F13BF7AFF810912FCF29F94A8E6" box="[837,975,889,913]" italics="true" pageId="67" pageNumber="68">T. steinheili</emphasis>
</taxonomicName>
group due to the following character combination: short antennal scapes (SI 70–74); comparatively short propodeal spines (PSLI 21–24); weakly cuneiform petiolar node shape with the anterodorsal angle situated higher than the posterodorsal, so that the dorsum tapers backwards posteriorly; mesosomal dorsum with longitudinally rugose sculpture; completely unsculptured waist segments; yellowish colouration.
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7AFF810AC0FBD99D6EAC9A" pageId="67" pageNumber="68" type="description">
<paragraph id="8BF9D301BF7AFF810AC0FBD99D7AAF1D" blockId="67.[151,1436,1105,2029]" box="[151,289,1105,1130]" pageId="67" pageNumber="68">
<heading id="D0B1646DBF7AFF810AC0FBD99D7AAF1D" bold="true" box="[151,289,1105,1130]" fontSize="10" level="2" pageId="67" pageNumber="68" reason="2">
<emphasis id="B9320F13BF7AFF810AC0FBD99D7AAF1D" bold="true" box="[151,289,1105,1130]" pageId="67" pageNumber="68">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7AFF810AC0FBFE9907AE6A" blockId="67.[151,1436,1105,2029]" pageId="67" pageNumber="68">
HL 0.49–0.58 (0.53); HW 0.45–0.54 (0.48); SL 0.32–0.39 (0.35); EL 0.09–0.12 (0.10);
<collectingCountry id="F3519391BF7AFF810E2CFBFE98C4AFFA" box="[1147,1183,1142,1165]" name="Philippines" pageId="67" pageNumber="68">PH</collectingCountry>
0.25–0.30 (0.27); PW 0.34–0.43 (0.38); WL 0.57–0.71 (0.62); PSL 0.11–0.14 (0.12); PTL 0.11–0.13 (0.12); PTH 0.20–0.23 (0.21); PTW 0.16–0.19 (0.17); PPL 0.14–0.16 (0.15); PPH 0.19–0.21 (0.20); PPW 0.21–0.24 (0.22); CI 90–94 (92); SI 70–74 (72); OI 20–23 (21); DMI 58–63 (61); LMI 42–45 (43); PSLI 21–24 (23); PeNI 43–49 (46); LPeI 51–59 (55); DPeI 142–156 (150); PpNI 52–62 (58); LPpI 71–76 (73); DPpI 147–157 (151); PPI 122–132 (127) (13 measured).
</paragraph>
<paragraph id="8BF9D301BF7AFF810A90FAA19D6EAC9A" blockId="67.[151,1436,1105,2029]" pageId="67" pageNumber="68">Head distinctly longer than wide (CI 90–94). Anterior clypeal margin medially impressed. Frontal carinae moderately developed and fine, ending between posterior eye margin and posterior head margin. Antennal scrobes faint. Antennal scapes short, ending between posterior eye margin and posterior head margin (SI 70–74). Eyes small to moderate (OI 20–23). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma, promesonotal suture and metanotal groove absent; mesosoma in profile comparatively stout and compact (LMI 42–45). Propodeal spines short to moderate, elongate-triangular and acute (PSLI 21–24). Propodeal lobes short and triangular. Petiolar node in profile high nodiform to cuneiform, approximately 1.7 to 2 times higher than long (LPeI 51–59), anterior and posterior faces not parallel, anterodorsal margin situated much higher than posterodorsal margin, dorsum distinctly tapering backwards posteriorly; node in dorsal view approximately 1.5 times wider than long (DPeI 142–156). Postpetiole in profile rounded, approximately 1.3 to 1.4 times higher than long (LPpI 71–76), in dorsal view approximately 1.5 times wider than long (DPpI 147–157). Postpetiole in profile approximately as voluminous as petiolar node, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 122–132). Mandibles finely striate but generally fairly shiny; clypeus with three to five longitudinal rugae or rugulae; cephalic dorsum between frontal carinae with six to eight longitudinal rugae, rugae running almost unbroken to posterior head margin, cross-meshes rare; lateral head weakly sculptured, anteriorly with irregular longitudinal rugulae mostly. Ground sculpture generally faint. Lateropronotum with weak sculpture only, remainder of lateral mesosoma with mainly longitudinally arranged, irregular rugae or rugulae. Both waist segments and gaster unsculptured, smooth and shining. All dorsal surfaces of body with long, fine, erect to suberect pilosity. Head, mesosoma, waist segments, and gaster of yellowish to light brownish colour; appendages of weakly lighter colour.</paragraph>
</subSubSection>
<caption id="DF398389BF7DFF860AC0F9639DB4AC57" ID-DOI="http://doi.org/10.5281/zenodo.6039003" ID-Zenodo-Dep="6039003" httpUri="https://zenodo.org/record/6039003/files/figure.png" pageId="68" pageNumber="69" startId="68.[151,264,1771,1794]" targetBox="[151,1436,193,1750]" targetPageId="68">
<paragraph id="8BF9D301BF7DFF860AC0F9639DB4AC57" blockId="68.[151,1436,1771,1824]" pageId="68" pageNumber="69">
<emphasis id="B9320F13BF7DFF860AC0F9639D35AC75" bold="true" box="[151,366,1771,1794]" pageId="68" pageNumber="69">FIGURES 101–103.</emphasis>
<taxonomicName id="4C46A882BF7DFF860B23F9649D91AC75" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[372,458,1772,1794]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7DFF860B23F9649D91AC75" box="[372,458,1772,1794]" italics="true" pageId="68" pageNumber="69">T. macki</emphasis>
</taxonomicName>
, holotype—CASENT0189093 (Estella Ortega 2011). 101. Body in profile. 102. Body in dorsal view. 103. Head in full-face view.
</paragraph>
</caption>
<subSubSection id="C35C808ABF7DFF870AC0F8C599CBA89B" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="discussion">
<paragraph id="8BF9D301BF7DFF860AC0F8C59C81AC12" blockId="68.[151,1436,1869,2037]" box="[151,218,1869,1893]" pageId="68" pageNumber="69">
<heading id="D0B1646DBF7DFF860AC0F8C59C81AC12" bold="true" box="[151,218,1869,1893]" fontSize="10" level="2" pageId="68" pageNumber="69" reason="2">
<emphasis id="B9320F13BF7DFF860AC0F8C59C81AC12" bold="true" box="[151,218,1869,1893]" pageId="68" pageNumber="69">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7DFF860AC0F8F99C89AC82" blockId="68.[151,1436,1869,2037]" pageId="68" pageNumber="69">
At present, the new species is only known to occur in Andriantantely, Betampona, Didy, and Sandranantitra. All four localities are comparatively close to each other geographically, and represent lowland rainforests lying at altitudes between
<quantity id="4CBE7EE4BF7DFF860B37F8319DFDACA6" box="[352,422,1977,2001]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="68" pageNumber="69" unit="m" value="450.0">450 m</quantity>
to
<specimenCount id="9D401888BF7DFF860B9FF8319E4EACA6" box="[456,533,1977,2001]" pageId="68" pageNumber="69" type="generic">
<quantity id="4CBE7EE4BF7DFF860B9FF8319E4AACA6" box="[456,529,1977,2001]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="68" pageNumber="69" unit="m" value="960.0">960 m</quantity>
.
</specimenCount>
On the basis of the available collection label data,
<taxonomicName id="4C46A882BF7DFF860E1DF83298FCACA6" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[1098,1191,1977,2001]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7DFF860E1DF83298FCACA6" box="[1098,1191,1977,2001]" italics="true" pageId="68" pageNumber="69">T. macki</emphasis>
</taxonomicName>
appears to live in leaf litter.
</paragraph>
<paragraph id="8BF9D301BF7CFF870A90FF109F3CAAF0" blockId="69.[151,1438,151,1004]" pageId="69" pageNumber="70">
<taxonomicName id="4C46A882BF7CFF870A90FF109DF8ABC7" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[199,419,152,176]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870A90FF109DF8ABC7" box="[199,419,152,176]" italics="true" pageId="69" pageNumber="70">Tetramorium macki</emphasis>
</taxonomicName>
can be straightforwardly identified within the
<taxonomicName id="4C46A882BF7CFF8709E3FF119875ABC7" authorityName="Bolton" authorityYear="1979" box="[948,1070,152,176]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="dysalum">
<emphasis id="B9320F13BF7CFF8709E3FF119875ABC7" box="[948,1070,152,176]" italics="true" pageId="69" pageNumber="70">T. dysalum</emphasis>
</taxonomicName>
group due to its combination of short propodeal spines (PSLI 21–24), weakly cuneiform petiolar node, and yellow colouration. The other two species with short propodeal spines/teeth,
<taxonomicName id="4C46A882BF7CFF8708D1FF699E91AB8F" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[646,714,225,248]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="orc">
<emphasis id="B9320F13BF7CFF8708D1FF699E91AB8F" box="[646,714,225,248]" italics="true" pageId="69" pageNumber="70">T. orc</emphasis>
</taxonomicName>
and
<taxonomicName id="4C46A882BF7CFF87095DFF699FDAAB8F" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[778,897,224,248]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="robitika">
<emphasis id="B9320F13BF7CFF87095DFF699FDAAB8F" box="[778,897,224,248]" italics="true" pageId="69" pageNumber="70">T. robitika</emphasis>
</taxonomicName>
, are not likely to be confused with
<taxonomicName id="4C46A882BF7CFF870F65FF6999CEAB8F" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[1330,1429,224,248]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870F65FF6999CEAB8F" box="[1330,1429,224,248]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
.
<taxonomicName id="4C46A882BF7CFF870AC0FE8D9DD3AA6B" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[151,392,260,284]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="robitika">
<emphasis id="B9320F13BF7CFF870AC0FE8D9DD3AA6B" box="[151,392,260,284]" italics="true" pageId="69" pageNumber="70">Tetramorium robitika</emphasis>
</taxonomicName>
has a high nodiform petiolar node with anterodorsal and posterodorsal margins at about the same height while
<taxonomicName id="4C46A882BF7CFF870B3FFEA19D9DAA37" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[360,454,296,320]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870B3FFEA19D9DAA37" box="[360,454,296,320]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
has a cuneiform node with the dorsum tapering backwards posteriorly. Also,
<taxonomicName id="4C46A882BF7CFF870F73FEA19938AA37" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[1316,1379,297,320]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="orc">
<emphasis id="B9320F13BF7CFF870F73FEA19938AA37" box="[1316,1379,297,320]" italics="true" pageId="69" pageNumber="70">T. orc</emphasis>
</taxonomicName>
with its shorter antennal scapes (SI 68–69) and dark to very dark brown colouration differs strikingly from
<taxonomicName id="4C46A882BF7CFF870F51FEC5993FAA13" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[1286,1380,332,356]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870F51FEC5993FAA13" box="[1286,1380,332,356]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
with its longer antennal scapes (SI 70–74) and yellowish colouration.
</paragraph>
<paragraph id="8BF9D301BF7CFF870A90FE1C99CBA89B" blockId="69.[151,1438,151,1004]" pageId="69" pageNumber="70">
As noted above, within the
<taxonomicName id="4C46A882BF7CFF870BAEFE1D9E2AAADB" authorityName="Bolton" authorityYear="1979" box="[505,625,404,428]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="dysalum">
<emphasis id="B9320F13BF7CFF870BAEFE1D9E2AAADB" box="[505,625,404,428]" italics="true" pageId="69" pageNumber="70">T. dysalum</emphasis>
</taxonomicName>
group
<taxonomicName id="4C46A882BF7CFF870897FE1D9F44AADB" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[704,799,404,428]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870897FE1D9F44AADB" box="[704,799,404,428]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
is fairly easy to differentiate from the remaining species. However,
<taxonomicName id="4C46A882BF7CFF870B5AFE319D34AAA7" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[269,367,440,464]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870B5AFE319D34AAA7" box="[269,367,440,464]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
has a general gestalt that is comparatively close to the one seen in some members of the
<taxonomicName id="4C46A882BF7CFF870FD0FE319CA2AA83" authorityName="Forel" authorityYear="1895" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="ranarum">
<emphasis id="B9320F13BF7CFF870FD0FE319CA2AA83" italics="true" pageId="69" pageNumber="70">T. ranarum</emphasis>
</taxonomicName>
group (
<taxonomicName id="4C46A882BF7CFF870B0DFE559E65AA83" authority="Santschi" authorityName="Santschi" authorityYear="1911" box="[346,574,476,500]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="degener">
<emphasis id="B9320F13BF7CFF870B0DFE559D8FAA83" box="[346,468,476,500]" italics="true" pageId="69" pageNumber="70">T. degener</emphasis>
Santschi
</taxonomicName>
and few morphologically similar undescribed species), and without close examination they could be confused with each other. All are relatively small, yellowish species with a petiolar node which is higher and wider than it is long, and comparatively small propodeal spines. Yet these
<taxonomicName id="4C46A882BF7CFF870E84FDAD9909A94B" authorityName="Forel" authorityYear="1895" box="[1235,1362,549,572]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="ranarum">
<emphasis id="B9320F13BF7CFF870E84FDAD9909A94B" box="[1235,1362,549,572]" italics="true" pageId="69" pageNumber="70">T. ranarum</emphasis>
</taxonomicName>
group members either have propodeal lobes of approximately the same length and shape as the propodeal spines or a petiolar node with distinct sculpture, whereas
<taxonomicName id="4C46A882BF7CFF8708CBFDE59EA0A9F3" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[668,763,620,644]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF8708CBFDE59EA0A9F3" box="[668,763,620,644]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
possesses propodeal spines of short to moderate length that are much longer than the propodeal lobes, and a completely unsculptured petiolar node. These ants also differ in gastral pilosity. In
<taxonomicName id="4C46A882BF7CFF870B3AFD3D9D95A9BB" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[365,462,692,716]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870B3AFD3D9D95A9BB" box="[365,462,692,716]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
the gastral pilosity is mainly suberect to erect with few subdecumbent to decumbent hairs while the gastral pilosity of
<taxonomicName id="4C46A882BF7CFF870843FD519ED1A987" authorityName="Santschi" authorityYear="1911" box="[532,650,728,752]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="degener">
<emphasis id="B9320F13BF7CFF870843FD519ED1A987" box="[532,650,728,752]" italics="true" pageId="69" pageNumber="70">T. degener</emphasis>
</taxonomicName>
and allies is mainly appressed to subdecumbent with few erect hairs. This character, in combination with the unsculptured, higher and wider petiolar node, justifies the placement of
<taxonomicName id="4C46A882BF7CFF870FD0FD759C81A84F" authorityName="Garcia &amp; Fisher" authorityYear="2012" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870FD0FD759C81A84F" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C46A882BF7CFF870B37FCA99D80A84F" authorityName="Bolton" authorityYear="1979" box="[352,475,800,824]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="dysalum">
<emphasis id="B9320F13BF7CFF870B37FCA99D80A84F" box="[352,475,800,824]" italics="true" pageId="69" pageNumber="70">T. dysalum</emphasis>
</taxonomicName>
group. Nevertheless, these ants represent a good example of seemingly convergent evolution. Despite belonging to different groups,
<taxonomicName id="4C46A882BF7CFF870898FCCD9F6BA82B" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[719,816,836,860]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870898FCCD9F6BA82B" box="[719,816,836,860]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
and
<taxonomicName id="4C46A882BF7CFF87093BFCCD9FBFA82B" authorityName="Santschi" authorityYear="1911" box="[876,996,836,860]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="degener">
<emphasis id="B9320F13BF7CFF87093BFCCD9FBFA82B" box="[876,996,836,860]" italics="true" pageId="69" pageNumber="70">T. degener</emphasis>
</taxonomicName>
and allies all are comparatively small species with reduced characters in comparison to their respective group members, and their general habitus might be misleading at first glance. Another difference is their preferred habitat, because
<taxonomicName id="4C46A882BF7CFF870E0BFC0598E5A8D3" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[1116,1214,908,932]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="macki">
<emphasis id="B9320F13BF7CFF870E0BFC0598E5A8D3" box="[1116,1214,908,932]" italics="true" pageId="69" pageNumber="70">T. macki</emphasis>
</taxonomicName>
, like most other
<taxonomicName id="4C46A882BF7CFF870FD0FC059CAFA8BF" authorityName="Bolton" authorityYear="1979" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="dysalum">
<emphasis id="B9320F13BF7CFF870FD0FC059CAFA8BF" italics="true" pageId="69" pageNumber="70">T. dysalum</emphasis>
</taxonomicName>
members, is only found in humid rainforests, whereas
<taxonomicName id="4C46A882BF7CFF8709DAFC399853A8BF" authorityName="Santschi" authorityYear="1911" box="[909,1032,944,968]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="degener">
<emphasis id="B9320F13BF7CFF8709DAFC399853A8BF" box="[909,1032,944,968]" italics="true" pageId="69" pageNumber="70">T. degener</emphasis>
</taxonomicName>
and allies prefer much more arid habitats, such as spiny forests or thickets, woodlands, tropical dry forests, or anthropogenically modified habitats.
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7CFF870AC0FB949912AF20" box="[151,1353,1052,1111]" pageId="69" pageNumber="70" type="etymology">
<paragraph id="8BF9D301BF7CFF870AC0FB949D43AF43" blockId="69.[151,1353,1052,1111]" box="[151,280,1052,1076]" pageId="69" pageNumber="70">
<heading id="D0B1646DBF7CFF870AC0FB949D43AF43" bold="true" box="[151,280,1052,1076]" fontSize="10" level="2" pageId="69" pageNumber="70" reason="2">
<emphasis id="B9320F13BF7CFF870AC0FB949D43AF43" bold="true" box="[151,280,1052,1076]" pageId="69" pageNumber="70">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7CFF870AC0FBB79912AF20" blockId="69.[151,1353,1052,1111]" box="[151,1353,1087,1111]" pageId="69" pageNumber="70">The new species is named in honor of Dawn M. Mack for her support to discover and identify life on earth.</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7CFF870AC0FB0F9F22AE23" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="8BF9D301BF7CFF870AC0FB0F9D22AFD7" blockId="69.[151,1436,1159,1364]" box="[151,377,1159,1184]" pageId="69" pageNumber="70">
<heading id="D0B1646DBF7CFF870AC0FB0F9D22AFD7" bold="true" box="[151,377,1159,1184]" fontSize="10" level="2" pageId="69" pageNumber="70" reason="2">
<emphasis id="B9320F13BF7CFF870AC0FB0F9D22AFD7" bold="true" box="[151,377,1159,1184]" pageId="69" pageNumber="70">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7CFF870AC0FB239F22AE23" blockId="69.[151,1436,1159,1364]" pageId="69" pageNumber="70">
<materialsCitation id="3B2ED95CBF7CFF870AC0FB239D77AF90" ID-GBIF-Occurrence="3462706024" pageId="69" pageNumber="70">
<emphasis id="B9320F13BF7CFF870AC0FB239D3FAFB3" bold="true" box="[151,356,1195,1220]" pageId="69" pageNumber="70">
<collectingCountry id="F3519391BF7CFF870AC0FB239D3FAFB3" box="[151,356,1195,1220]" name="Madagascar" pageId="69" pageNumber="70">MADAGASCAR</collectingCountry>
</emphasis>
:
<collectingRegion id="49821DE3BF7CFF870B23FB249DABAFB3" box="[372,496,1196,1220]" country="Madagascar" name="Toamasina" pageId="69" pageNumber="70">Toamasina</collectingRegion>
, F.C. Andriantantely,
<geoCoordinate id="EE72B5C6BF7CFF8708B0FB249F15AFB3" box="[743,846,1196,1220]" degrees="18.695" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="55" value="-18.695">18.695 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF7CFF87090CFB249F84AFB4" box="[859,991,1196,1220]" degrees="48.81333" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="48.81333">48.81333 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF7CFF8709BBFB24986CAFB4" box="[1004,1079,1196,1220]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="69" pageNumber="70" unit="m" value="530.0">530 m</quantity>
, rainforest,
<date id="FFF8F5C1BF7CFF870EE9FB249907AFB3" box="[1214,1372,1196,1220]" pageId="69" pageNumber="70" value="1998-12-04" valueMax="1998-12-07" valueMin="1998-12-04">4.–7.XII.1998</date>
(
<emphasis id="B9320F13BF7CFF870F3AFB259D40AF90" italics="true" pageId="69" pageNumber="70">H.J. Ratsirarson</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF7CFF870B69FB479D77AE7B" ID-GBIF-Occurrence="3462705498" pageId="69" pageNumber="70">
<collectingRegion id="49821DE3BF7CFF870B69FB479DE1AF90" box="[318,442,1231,1255]" country="Madagascar" name="Toamasina" pageId="69" pageNumber="70">Toamasina</collectingRegion>
, F.C. Didy,
<geoCoordinate id="EE72B5C6BF7CFF870837FB479EB7AF90" box="[608,748,1231,1255]" degrees="18.19833" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="1" value="-18.19833">18.19833 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF7CFF870954FB479FC9AF90" box="[771,914,1231,1255]" degrees="48.57833" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="48.57833">48.57833 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF7CFF8709FEFB479FA6AF90" box="[937,1021,1231,1255]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="69" pageNumber="70" unit="m" value="960.0">960 m</quantity>
, rainforest,
<date id="FFF8F5C1BF7CFF870ECFFB479909AF90" box="[1176,1362,1231,1255]" pageId="69" pageNumber="70" value="1998-12-16" valueMax="1998-12-23" valueMin="1998-12-16">16.–23.XII.1998</date>
(
<emphasis id="B9320F13BF7CFF870F3AFB599D40AE7B" italics="true" pageId="69" pageNumber="70">H.J. Ratsirarson</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF7CFF870B6FFB7C9D77AE58" ID-GBIF-Occurrence="3462705370" pageId="69" pageNumber="70">
<collectingRegion id="49821DE3BF7CFF870B6FFB7C9DEFAE7B" box="[312,436,1268,1292]" country="Madagascar" name="Toamasina" pageId="69" pageNumber="70">Toamasina</collectingRegion>
, F.C. Sandranantitra,
<geoCoordinate id="EE72B5C6BF7CFF8708E5FB7C9F63AE7B" box="[690,824,1268,1292]" degrees="18.04833" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="1" value="-18.04833">18.04833 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF7CFF87091EFB7C9F8BAE7C" box="[841,976,1268,1292]" degrees="49.09167" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="49.09167">49.09167 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF7CFF8709B6FB7C9875AE7C" box="[993,1070,1268,1292]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="69" pageNumber="70" unit="m" value="450.0">450 m</quantity>
, rainforest,
<date id="FFF8F5C1BF7CFF870EEBFB7C9903AE7B" box="[1212,1368,1268,1292]" pageId="69" pageNumber="70" value="1999-01-21" valueMax="1999-01-24" valueMin="1999-01-21">21.–24.I.1999</date>
(
<emphasis id="B9320F13BF7CFF870F3AFB7D9D40AE58" italics="true" pageId="69" pageNumber="70">H.J. Ratsirarson</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF7CFF870B63FA9F9F22AE23" ID-GBIF-Occurrence="3462705650" pageId="69" pageNumber="70">
<collectingRegion id="49821DE3BF7CFF870B63FA9F9DEBAE58" box="[308,432,1303,1327]" country="Madagascar" name="Toamasina" pageId="69" pageNumber="70">Toamasina</collectingRegion>
, Réserve Nationale Intégrale Betampona, Betampona
<quantity id="4CBE7EE4BF7CFF870E46FA9F9835AE58" box="[1041,1134,1303,1327]" metricMagnitude="4" metricUnit="m" metricValue="3.51" pageId="69" pageNumber="70" unit="km" value="35.1">35.1 km</quantity>
NW
<collectingRegion id="49821DE3BF7CFF870EFBFA9F9973AE58" box="[1196,1320,1303,1327]" country="Madagascar" name="Toamasina" pageId="69" pageNumber="70">Toamasina</collectingRegion>
,
<geoCoordinate id="EE72B5C6BF7CFF870F64FA9F9CF3AE23" degrees="17.91801" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="1" value="-17.91801">17.91801 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF7CFF870AE3FAB49D6CAE24" box="[180,311,1340,1364]" degrees="49.20074" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="49.20074">49.20074 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF7CFF870B14FAB49DD7AE24" box="[323,396,1340,1364]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="69" pageNumber="70" unit="m" value="500.0">500 m</quantity>
, rainforest,
<date id="FFF8F5C1BF7CFF870846FAB49EC1AE23" box="[529,666,1340,1364]" pageId="69" pageNumber="70" value="2007-12-16">16.XII.2007</date>
(
<emphasis id="B9320F13BF7CFF8708FCFAB59F32AE23" box="[683,873,1340,1364]" italics="true" pageId="69" pageNumber="70">B.L. Fisher et al.</emphasis>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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