495 lines
70 KiB
XML
495 lines
70 KiB
XML
<document id="25DEB968D9302A6B529BCFC509B5E274" ID-CLB-Dataset="45698" ID-DOI="10.5281/zenodo.279609" ID-GBIF-Dataset="2774854a-4f4f-4bf8-a2df-2bcc57ec574a" ID-ISSN="1175-5326" ID-Zenodo-Dep="279609" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460424672434" checkinUser="plazi" docAuthor="Alvarez, Fernando, Iliffe, Thomas M., Gonzalez, Brett & Villalobos, José Luis" docDate="2012" docId="03C33854FFDA710AFF18FF2B324BFDDC" docLanguage="en" docName="zt03154p068.pdf" docOrigin="Zootaxa 3154" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Triacanthoneus akumalensis Alvarez, Iliffe, Gonzalez & Villalobos, 2012, sp. nov." docType="treatment" docVersion="7" lastPageNumber="68" masterDocId="FFFA402CFFDB710DFF8FFFBC362FFFE3" masterDocTitle="Triacanthoneus akumalensis, a new species of alpheid shrimp (Crustacea: Caridea: Alpheidae) from an anchialine cave in Quintana Roo, Mexico" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698330418051" updateUser="ExternalLinkService">
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<mods:titleInfo id="BF6902E0F96129A90042BD5A33106952">
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<mods:title id="F9D56B9C74124086A7E4DE9187872953">Triacanthoneus akumalensis, a new species of alpheid shrimp (Crustacea: Caridea: Alpheidae) from an anchialine cave in Quintana Roo, Mexico</mods:title>
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<mods:name id="9C3C4E1C02EF03AA4CE9E1830A6881EB" type="personal">
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<mods:namePart id="6240FD557E4BF3D2BDD914F54A93631B">Alvarez, Fernando</mods:namePart>
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<mods:namePart id="3F8F86A161D1E1CDB22A7825C00807D6">Iliffe, Thomas M.</mods:namePart>
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<mods:namePart id="1AD9F02B30C05811478F6DD2606B9A0F">Gonzalez, Brett</mods:namePart>
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</mods:role>
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<mods:namePart id="7CFDAFFF3C3ED6561C00B17A7AE23D56">Villalobos, José Luis</mods:namePart>
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<mods:typeOfResource id="4DF8F6B57530991E57A7C24306B3E717">text</mods:typeOfResource>
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<mods:title id="827C99573CE87F9660290F573A41D785">Zootaxa</mods:title>
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<mods:part id="CCAA82846E3C57A504FB106BD1286443">
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<mods:date id="851C1AED35FF08CAA11FFAD39CC8C95E">2012</mods:date>
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<mods:number id="6454FD72F83EC432D3EB8053F716FB00">3154</mods:number>
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<treatment id="03C33854FFDA710AFF18FF2B324BFDDC" ID-DOI="http://doi.org/10.5281/zenodo.6178317" ID-GBIF-Taxon="119428211" ID-Zenodo-Dep="6178317" LSID="urn:lsid:plazi:treatment:03C33854FFDA710AFF18FF2B324BFDDC" httpUri="http://treatment.plazi.org/id/03C33854FFDA710AFF18FF2B324BFDDC" lastPageId="7" lastPageNumber="68" pageId="1" pageNumber="62">
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<subSubSection id="C370DAC9FFDA710CFF18FF2B373DFF31" pageId="1" pageNumber="62" type="nomenclature">
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<paragraph id="8BD58942FFDA710CFF18FF2B3476FF52" blockId="1.[151,601,151,210]" box="[151,601,151,177]" pageId="1" pageNumber="62">
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<heading id="D09D3E2EFFDA710CFF18FF2B3476FF52" bold="true" box="[151,601,151,177]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
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<emphasis id="B91E5550FFDA710CFF18FF2B3476FF52" bold="true" box="[151,601,151,177]" pageId="1" pageNumber="62">
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<taxonomicName id="4C6AF2C1FFDA710CFF18FF2B37DBFF52" ID-CoL="85ZFQ" box="[151,500,151,177]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
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<emphasis id="B91E5550FFDA710CFF18FF2B37DBFF52" bold="true" box="[151,500,151,177]" italics="true" pageId="1" pageNumber="62">Triacanthoneus akumalensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22DE82BFFDA710CFE73FF2B3476FF52" box="[508,601,151,177]" pageId="1" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD58942FFDA710CFF18FF06373DFF31" blockId="1.[151,601,151,210]" box="[151,274,186,210]" pageId="1" pageNumber="62">
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(
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<figureCitation id="135195C7FFDA710CFF10FF063726FF31" box="[159,265,186,210]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1962,1985]" captionStartId-1="3.[151,250,1953,1976]" captionStartId-2="4.[151,250,1915,1938]" captionTargetBox-0="[209,1335,212,1911]" captionTargetBox-1="[224,1351,236,1911]" captionTargetBox-2="[260,1294,232,1842]" captionTargetId-0="figure@2.[197,1350,195,1934]" captionTargetId-1="figure@3.[218,1368,193,1932]" captionTargetId-2="figure@4.[249,1319,193,1865]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm." captionText-1="FIGURE 2. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bars = 1 mm." captionText-2="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri-0="https://zenodo.org/record/279610/files/figure.png" httpUri-1="https://zenodo.org/record/279611/files/figure.png" httpUri-2="https://zenodo.org/record/279612/files/figure.png" pageId="1" pageNumber="62">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C370DAC9FFDA710CFF18FEBF33B9FE6A" pageId="1" pageNumber="62" type="materials_examined">
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<paragraph id="8BD58942FFDA710CFF18FEBF33B9FE6A" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
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<emphasis id="B91E5550FFDA710CFF18FEBF376BFEF8" bold="true" box="[151,324,258,283]" pageId="1" pageNumber="62">
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<typeStatus id="54D137E0FFDA710CFF18FEBF36FFFEF8" box="[151,208,259,283]" pageId="1" pageNumber="62">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54D137E0FFDA710CFEC0FEBE3793FEF9" box="[335,444,258,282]" pageId="1" pageNumber="62" type="holotype">Holotype</typeStatus>
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: ovigerous female, cl
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<quantity id="4C9224A7FFDA710CFD37FEBE353DFEF9" box="[696,786,258,282]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="1" pageNumber="62" unit="mm" value="8.9">8.9 mm</quantity>
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, tl
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<quantity id="4C9224A7FFDA710CFCB8FEBE35A5FEF9" box="[823,906,258,282]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="62" unit="mm" value="26.0">26 mm</quantity>
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,
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<collectingCountry id="F37DC9D2FFDA710CFC18FEBE35DFFEF9" box="[919,1008,258,282]" name="Mexico" pageId="1" pageNumber="62">Mexico</collectingCountry>
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, Quintana Roo, Akumal, Cenote Aak Kimin, underwater cave (
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<geoCoordinate id="EE5EEF85FFDA710CFE35FE953401FEA2" box="[442,558,294,321]" direction="north" orientation="latitude" pageId="1" pageNumber="62" precision="925" value="20.4">20o 24’ N</geoCoordinate>
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,
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<geoCoordinate id="EE5EEF85FFDA710CFDB4FE95349BFEA2" box="[571,692,294,321]" direction="west" orientation="longitude" pageId="1" pageNumber="62" precision="925" value="-87.3">87o 18’ W</geoCoordinate>
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), depth
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<quantity id="4C9224A7FFDA710CFC9DFE953556FEA2" box="[786,889,297,321]" metricMagnitude="1" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="1" pageNumber="62" unit="m" value="32.5" valueMax="40.0" valueMin="25.0">25–40 m</quantity>
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,
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<date id="FFD4AF82FFDA710CFC08FE953204FEA2" box="[903,1067,297,321]" pageId="1" pageNumber="62" value="2008-04-15">15 April 2008</date>
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, coll. T.M. Iliffe,
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<collectionCode id="ED7B1187FFDA710CFB75FE953369FEA2" box="[1274,1350,297,321]" pageId="1" pageNumber="62">CNCR</collectionCode>
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26315.
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||
<typeStatus id="54D137E0FFDA710CFF18FEF23727FE85" box="[151,264,334,358]" pageId="1" pageNumber="62" type="paratype">Paratypes</typeStatus>
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||
: 1 non-ovigerous specimen (male?), cl
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||
<quantity id="4C9224A7FFDA710CFD4DFEF13536FE86" box="[706,793,333,357]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="62" unit="mm" value="9.1">9.1 mm</quantity>
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||
, tl
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||
<quantity id="4C9224A7FFDA710CFCB5FEF135B0FE86" box="[826,927,333,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="1" pageNumber="62" unit="mm" value="25.9">25.9 mm</quantity>
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||
, same data as for the
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<typeStatus id="54D137E0FFDA710CFB02FEF132DDFE86" box="[1165,1266,333,357]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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||
,
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||
<collectionCode id="ED7B1187FFDA710CFB73FEF13367FE86" box="[1276,1352,333,357]" pageId="1" pageNumber="62">CNCR</collectionCode>
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||
26316; 1 ovigerous specimen with appendix masculina, cl
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||
<quantity id="4C9224A7FFDA710CFD5FFECD353DFE6A" box="[720,786,369,393]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="62" unit="mm" value="8.0">8 mm</quantity>
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||
, tl
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||
<quantity id="4C9224A7FFDA710CFCBCFECD35B6FE6A" box="[819,921,369,393]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="1" pageNumber="62" unit="mm" value="23.7">23.7 mm</quantity>
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||
, same data as for the
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||
<typeStatus id="54D137E0FFDA710CFB09FECD32C4FE6A" box="[1158,1259,369,393]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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||
,
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||
<collectionCode id="ED7B1187FFDA710CFB79FECD336DFE6A" box="[1270,1346,369,393]" pageId="1" pageNumber="62">CNCR</collectionCode>
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26486.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C370DAC9FFDA7108FF48FE2932E4FCD1" lastPageId="5" lastPageNumber="66" pageId="1" pageNumber="62" type="description">
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<paragraph id="8BD58942FFDA710CFF48FE2932AFFDFA" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
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<emphasis id="B91E5550FFDA710CFF48FE293776FE4D" bold="true" box="[199,345,405,430]" pageId="1" pageNumber="62">Description.</emphasis>
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Carapace broadly rounded laterally, with faint anterolateral suture and two strong, acute postorbital teeth; dorsal midline with prominent sharp tooth in epigastric position, low carina present between epigastric tooth and orbit; posterior margin with deep cardiac notch (
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<figureCitation id="135195C7FFDA710CFCAEFE613543FE16" box="[801,876,477,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1962,1985]" captionTargetBox="[209,1335,212,1911]" captionTargetId="figure@2.[197,1350,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm." httpUri="https://zenodo.org/record/279610/files/figure.png" pageId="1" pageNumber="62">Figs. 1</figureCitation>
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a, b). Rostrum reaching mid-length of third antennular article, tip slightly pointing upwards, ventral margin with small tooth (
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<figureCitation id="135195C7FFDA710CFC64FDBD3219FDFA" box="[1003,1078,513,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1962,1985]" captionTargetBox="[209,1335,212,1911]" captionTargetId="figure@2.[197,1350,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm." httpUri="https://zenodo.org/record/279610/files/figure.png" pageId="1" pageNumber="62">Figs. 1</figureCitation>
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b, 3a).
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</paragraph>
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<paragraph id="8BD58942FFDA710CFF48FD9A357EFD82" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
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Eyes completely covered by carapace, not visible in dorsal view; lateral region pigmented, cornea not faceted; dorsal margin with small acute projection present (
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<figureCitation id="135195C7FFDA710CFD40FDF53533FD82" box="[719,796,585,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1938]" captionTargetBox="[260,1294,232,1842]" captionTargetId="figure@4.[249,1319,193,1865]" captionTargetPageId="4" captionText="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="1" pageNumber="62">Figs. 3</figureCitation>
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a, f).
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</paragraph>
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<paragraph id="8BD58942FFDA710CFF48FDD134A5FD2E" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">First three abdominal somites with lateroventral margins of pleura rounded; fourth somite with lateroventral margin subtriangular; fifth somite with lateroventral margin produced in a small acute tooth; sixth somite longer than other somites, without articulated plate.</paragraph>
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<paragraph id="8BD58942FFDA710CFF48FD6532D5FCBE" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
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Telson approximately 2.6 times as long as its anterior width; posterior margin with deep “V” shaped notch, each posterolateral angle with a pair of long spines, lateral longest; posterior half of dorsal surface with shallow median longitudinal groove and two pairs of short spines (
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<figureCitation id="135195C7FFDA710CFCB3FC9D35AEFCDA" box="[828,897,801,825]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1938]" captionTargetBox="[260,1294,232,1842]" captionTargetId="figure@4.[249,1319,193,1865]" captionTargetPageId="4" captionText="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="1" pageNumber="62">Fig. 3</figureCitation>
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b). Uropods with both rami longer than telson, rounded distally; exopod longer than endopod, with complete diaeresis and strong distolateral spine.
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</paragraph>
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<paragraph id="8BD58942FFDA710CFF48FCD53518FBF2" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
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||
Antennule with prominent stylocerite, its acute tip reaching beyond mid-length of second article (
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<figureCitation id="135195C7FFDA710CFA87FCD53365FC62" box="[1288,1354,873,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1938]" captionTargetBox="[260,1294,232,1842]" captionTargetId="figure@4.[249,1319,193,1865]" captionTargetPageId="4" captionText="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="1" pageNumber="62">Fig. 3</figureCitation>
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a); second article longer than wide, about as long as third article; external flagellum with four basal articles, accessory branch with about eight articles bearing long aesthetascs (
|
||
<figureCitation id="135195C7FFDA710CFC91FC0D354FFC2A" box="[798,864,945,969]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1938]" captionTargetBox="[260,1294,232,1842]" captionTargetId="figure@4.[249,1319,193,1865]" captionTargetPageId="4" captionText="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="1" pageNumber="62">Fig. 3</figureCitation>
|
||
c). Antenna with basicerite armed with stout, sharp distal tooth; carpocerite short, not reaching beyond half-length of scaphocerite; scaphocerite ovate, with rounded anterior margin reaching beyond distolateral tooth (
|
||
<figureCitation id="135195C7FFDA710CFD57FC453535FBF2" box="[728,794,1017,1041]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1938]" captionTargetBox="[260,1294,232,1842]" captionTargetId="figure@4.[249,1319,193,1865]" captionTargetPageId="4" captionText="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="1" pageNumber="62">Fig. 3</figureCitation>
|
||
a).
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDA710CFF48FBA135D0FAD2" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
|
||
Mouthparts typical for genus. Mandible with incisor process bearing nine teeth distally; molar process cylindrical, stout, with blunt distal surface; palp two-articulated, distal article oval-shaped (
|
||
<figureCitation id="135195C7FFDA710CFB0AFBFD32E2FBBA" box="[1157,1229,1089,1113]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1976]" captionTargetBox="[224,1351,236,1911]" captionTargetId="figure@3.[218,1368,193,1932]" captionTargetPageId="3" captionText="FIGURE 2. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bars = 1 mm." httpUri="https://zenodo.org/record/279611/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
|
||
a). Maxillule with bilobed palp, each bearing one long seta (
|
||
<figureCitation id="135195C7FFDA710CFDE7FBD93483FB9E" box="[616,684,1125,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1976]" captionTargetBox="[224,1351,236,1911]" captionTargetId="figure@3.[218,1368,193,1932]" captionTargetPageId="3" captionText="FIGURE 2. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bars = 1 mm." httpUri="https://zenodo.org/record/279611/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
|
||
b). Maxilla with simple elongate palp, devoid of setae; distal endite bilobed, lobes divided by a deep cleft; scaphognathite about 3.2 times as long as wide (
|
||
<figureCitation id="135195C7FFDA710CFBF0FB3532EAFB42" box="[1151,1221,1161,1185]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1976]" captionTargetBox="[224,1351,236,1911]" captionTargetId="figure@3.[218,1368,193,1932]" captionTargetPageId="3" captionText="FIGURE 2. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bars = 1 mm." httpUri="https://zenodo.org/record/279611/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
|
||
c). First maxilliped with stout palp bearing distally two long setae covered with short setules; distal endite broad, subtrapezoidal; exopod elongate, caridean lobe evident; epipod approximately semicircular (
|
||
<figureCitation id="135195C7FFDA710CFC49FB6D3225FB0A" box="[966,1034,1233,1257]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1976]" captionTargetBox="[224,1351,236,1911]" captionTargetId="figure@3.[218,1368,193,1932]" captionTargetPageId="3" captionText="FIGURE 2. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bars = 1 mm." httpUri="https://zenodo.org/record/279611/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
|
||
d). Second maxilliped with elongate exopod; endopod with ischium and dactylus of about the same length; epipod relatively broad, somewhat elongate, as long as ischium of endopod; podobranch-like gill well-developed (
|
||
<figureCitation id="135195C7FFDA710CFC2FFAA535CBFAD2" box="[928,996,1305,1329]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1976]" captionTargetBox="[224,1351,236,1911]" captionTargetId="figure@3.[218,1368,193,1932]" captionTargetPageId="3" captionText="FIGURE 2. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bars = 1 mm." httpUri="https://zenodo.org/record/279611/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDA710CFF4AFA81331DFA22" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
|
||
Third maxilliped pediform, slender; endopod three-articulated, antepenultimate article subequal in length to penultimate and ultimate articles combined; ultimate article longer than penultimate, mesial surface covered with rows of short setae, tip with tuft of long setae; exopod shorter than antepenultimate article of endopod; epipod present as mastigobranch emerging from base of rounded lateral plate; arthrobranch well-developed (
|
||
<figureCitation id="135195C7FFDA710CFB59FA153335FA22" box="[1238,1306,1449,1473]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1976]" captionTargetBox="[224,1351,236,1911]" captionTargetId="figure@3.[218,1368,193,1932]" captionTargetPageId="3" captionText="FIGURE 2. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bars = 1 mm." httpUri="https://zenodo.org/record/279611/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
|
||
f).
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDA710CFF4AFA713710F8E6" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
|
||
First pair of pereopods asymmetrical, major cheliped always on the right side (
|
||
<figureCitation id="135195C7FFDA710CFBC3FA7132A2FA06" box="[1100,1165,1485,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1962,1985]" captionTargetBox="[209,1335,212,1911]" captionTargetId="figure@2.[197,1350,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm." httpUri="https://zenodo.org/record/279610/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
|
||
a). Major cheliped 0.75 times of total body length; ischium about one third of merus length, with three ventral spines; merus with ventral margin depressed, dorsal margin straight, distally somewhat thickened; carpus shorter than merus, wider distally; propodus strongly arched; palm about 2.4 times as long as high, inflated, with low hump on dorsal surface, adjacent to articulation with dactylus; fingers about 1.3 times length of palm, strongly crossing distally; cutting edge of fingers serrated with minute teeth, and with four-five conspicuously larger teeth intercalated among smaller teeth (
|
||
<figureCitation id="135195C7FFDA710CFAE0F93D368DF95E" captionStart="FIGURE 1" captionStartId="2.[151,250,1962,1985]" captionTargetBox="[209,1335,212,1911]" captionTargetId="figure@2.[197,1350,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm." httpUri="https://zenodo.org/record/279610/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
|
||
h). Minor cheliped approximately half as long as major cheliped, much less stout (fig. 1a); ischium with three ventral spines; ischium, merus, carpus and propodus subequal in length; chela simple, dactylus slightly shorter than palm (
|
||
<figureCitation id="135195C7FFDA710CFF51F951370FF8E6" box="[222,288,1773,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1938]" captionTargetBox="[260,1294,232,1842]" captionTargetId="figure@4.[249,1319,193,1865]" captionTargetPageId="4" captionText="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="1" pageNumber="62">
|
||
Fig.
|
||
<quantity id="4C9224A7FFDA710CFE9DF951370FF8E6" box="[274,288,1773,1797]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="1" pageNumber="62" unit="g" value="3.0">3</quantity>
|
||
</figureCitation>
|
||
g).
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDA710CFF4AF8AD3513F83E" blockId="1.[151,1437,258,2013]" pageId="1" pageNumber="62">
|
||
Second pair of pereopods slender (
|
||
<figureCitation id="135195C7FFDA710CFDD8F8AD34B6F8CA" box="[599,665,1809,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1962,1985]" captionTargetBox="[209,1335,212,1911]" captionTargetId="figure@2.[197,1350,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm." httpUri="https://zenodo.org/record/279610/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
|
||
b,
|
||
<date id="FFD4AF82FFDA710CFD35F8AD34FDF8CA" box="[698,722,1809,1833]" pageId="1" pageNumber="62">3i</date>
|
||
); ischium and merus subequal in length; ischium with row of three spines on ventrolateral surface; carpus five-articulated, first carpal article longest, more than half-length of the entire carpus, remaining carpal articles of about the same length; chela simple, with fingers longer than palm. Third to fifth pairs of pereopods slender with simple dactyli, increasing in length posteriorly (
|
||
<figureCitation id="135195C7FFDA710CFB46F8C13324F876" box="[1225,1291,1917,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1962,1985]" captionTargetBox="[209,1335,212,1911]" captionTargetId="figure@2.[197,1350,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm." httpUri="https://zenodo.org/record/279610/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
|
||
b); third and fourth pereopods each with three spines on ventrolateral surface of ischium; carpus longest article in third and fourth pereopods; propodus longest article in fifth pereopod.
|
||
</paragraph>
|
||
<caption id="DF15D9CAFFD9710FFF18F81634B9F803" httpUri="https://zenodo.org/record/279610/files/figure.png" pageId="2" pageNumber="63" targetBox="[209,1335,212,1911]" targetPageId="2">
|
||
<paragraph id="8BD58942FFD9710FFF18F81634B9F803" blockId="2.[151,1436,1962,2016]" pageId="2" pageNumber="63">
|
||
<emphasis id="B91E5550FFD9710FFF18F816373DF823" bold="true" box="[151,274,1962,1985]" pageId="2" pageNumber="63">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C6AF2C1FFD9710FFE97F8173416F822" box="[280,569,1963,1985]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFD9710FFE97F8173416F822" box="[280,569,1963,1985]" italics="true" pageId="2" pageNumber="63">Triacanthoneus akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFD9710FFDB0F81734A0F823" bold="true" box="[575,655,1963,1984]" pageId="2" pageNumber="63">
|
||
<taxonomicNameLabel id="A22DE82BFFD9710FFDB0F81734A0F823" box="[575,655,1963,1984]" pageId="2" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF15D9CAFFD8710EFF18F81D3558F835" httpUri="https://zenodo.org/record/279611/files/figure.png" pageId="3" pageNumber="64" targetBox="[224,1351,236,1911]" targetPageId="3">
|
||
<paragraph id="8BD58942FFD8710EFF18F81D3558F835" blockId="3.[151,1436,1953,2006]" pageId="3" pageNumber="64">
|
||
<emphasis id="B91E5550FFD8710EFF18F81D373BF854" bold="true" box="[151,276,1953,1976]" pageId="3" pageNumber="64">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6AF2C1FFD8710EFE92F81D346FF854" box="[285,576,1953,1975]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFD8710EFE92F81D346FF854" box="[285,576,1953,1975]" italics="true" pageId="3" pageNumber="64">Triacanthoneus akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFD8710EFDC7F81E34B5F854" bold="true" box="[584,666,1954,1975]" pageId="3" pageNumber="64">
|
||
<taxonomicNameLabel id="A22DE82BFFD8710EFDC7F81E34B5F854" box="[584,666,1954,1975]" pageId="3" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ovigerous female, holotype: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF15D9CAFFDF7109FF18F8C732B4F80E" httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="4" pageNumber="65" targetBox="[260,1294,232,1842]" targetPageId="4">
|
||
<paragraph id="8BD58942FFDF7109FF18F8C732B4F80E" blockId="4.[151,1436,1915,2029]" pageId="4" pageNumber="65">
|
||
<emphasis id="B91E5550FFDF7109FF18F8C7373DF873" bold="true" box="[151,274,1915,1938]" pageId="4" pageNumber="65">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C6AF2C1FFDF7109FE97F8C73417F872" box="[280,568,1915,1937]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDF7109FE97F8C73417F872" box="[280,568,1915,1937]" italics="true" pageId="4" pageNumber="65">Triacanthoneus akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDF7109FDB1F8C034A1F872" bold="true" box="[574,654,1916,1937]" pageId="4" pageNumber="65">
|
||
<taxonomicNameLabel id="A22DE82BFFDF7109FDB1F8C034A1F872" box="[574,654,1916,1937]" pageId="4" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD58942FFDE7108FF48FF2B36EDFF37" blockId="5.[151,1437,151,356]" pageId="5" pageNumber="66">
|
||
Pleural process, strong and anteriorly arched, arising from lateral body wall, above coxa of fifth pereopod (
|
||
<figureCitation id="135195C7FFDE7108FAE0FF2B368CFF37" captionStart="FIGURE 1" captionStartId="2.[151,250,1962,1985]" captionTargetBox="[209,1335,212,1911]" captionTargetId="figure@2.[197,1350,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Triacanthoneus akumalensis sp. nov., ovigerous female, holotype: a, total lateral view, in life; b, total lateral view; c, major chela. Scale bars: a, b = 4 mm; c = 1 mm." httpUri="https://zenodo.org/record/279610/files/figure.png" pageId="5" pageNumber="66">Fig. 1</figureCitation>
|
||
c).
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDE7108FF48FF633770FE87" blockId="5.[151,1437,151,356]" pageId="5" pageNumber="66">
|
||
Second pair of pleopods of male
|
||
<typeStatus id="54D137E0FFDE7108FDC6FF5C3486FF1B" box="[585,681,224,248]" pageId="5" pageNumber="66" type="paratype">paratype</typeStatus>
|
||
with protopod bearing long thick setae curling distally (
|
||
<figureCitation id="135195C7FFDE7108FAB7FF633352FF14" box="[1336,1405,223,247]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1938]" captionTargetBox="[260,1294,232,1842]" captionTargetId="figure@4.[249,1319,193,1865]" captionTargetPageId="4" captionText="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="5" pageNumber="66">Fig. 3</figureCitation>
|
||
d); exopod longer than endopod; endopod with appendix interna and appendix masculine, former elongate, slightly longer than appendix masculina; appendix masculina cylindrical, with nine strong spines along lateral and distal margins (
|
||
<figureCitation id="135195C7FFDE7108FE8FFEF0376DFE87" box="[256,322,332,356]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1938]" captionTargetBox="[260,1294,232,1842]" captionTargetId="figure@4.[249,1319,193,1865]" captionTargetPageId="4" captionText="FIGURE 3. Triacanthoneus akumalensis sp. nov., a – c, f – h, ovigerous female holotype; d, e, male paratype: a, dorsal view of anterior portion of carapace; b, dorsal view of telson and uropods; c, detail of lateral flagellum of antennule; d, second pleopod of male; e, same, appendix masculina; f, frontal view of eyes; g, minor first pereopod; h, position of pleural process above coxa of fifth pereopod; i, detail of second pereopod. Scale bars: a – d, f – i = 1 mm, e = 0.2 mm, h = 4 mm." httpUri="https://zenodo.org/record/279612/files/figure.png" pageId="5" pageNumber="66">Fig. 3</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
<caption id="DF15D9CAFFDE7108FF18FE2C3572FE45" box="[151,861,400,422]" pageId="5" pageNumber="66">
|
||
<paragraph id="8BD58942FFDE7108FF18FE2C3572FE45" blockId="5.[151,1337,400,723]" box="[151,861,400,422]" pageId="5" pageNumber="66">
|
||
<emphasis id="B91E5550FFDE7108FF18FE2C372BFE46" bold="true" box="[151,260,400,421]" pageId="5" pageNumber="66">TABLE 1.</emphasis>
|
||
Branchial formula of
|
||
<taxonomicName id="4C6AF2C1FFDE7108FE6AFE2C3529FE45" box="[485,774,400,422]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDE7108FE6AFE2C3529FE45" box="[485,774,400,422]" italics="true" pageId="5" pageNumber="66">Triacanthoneus akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22DE82BFFDE7108FC83FE2D3572FE45" box="[780,861,401,422]" pageId="5" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD58942FFDE7108FE1DFE7E35ACFE3B" blockId="5.[151,1337,400,723]" box="[402,899,450,472]" pageId="5" pageNumber="66">Maxillipeds Pereopods</paragraph>
|
||
<paragraph id="8BD58942FFDE7108FE1DFE5332E4FCD1" blockId="5.[151,1337,400,723]" lastBlockId="5.[151,1437,794,1970]" pageId="5" pageNumber="66">
|
||
1rst 2nd 3rd 1rst 2nd 3rd 4rth 5th Pleurobranch - - - P P P P - Arthrobranch - - P - - - - - Podobranch - P - - - - - - Mastigobranch - P P P P P P - Exopod P P P - - - - -
|
||
<emphasis id="B91E5550FFDE7108FF48FCA73761FCD0" bold="true" box="[199,334,795,819]" pageId="5" pageNumber="66">Etymology.</emphasis>
|
||
The specific epithet refers to the
|
||
<typeStatus id="54D137E0FFDE7108FD4AFCA734DAFCD0" box="[709,757,795,819]" pageId="5" pageNumber="66">type</typeStatus>
|
||
locality, Akumal, Quintana Roo,
|
||
<collectingCountry id="F37DC9D2FFDE7108FBE1FCA632E8FCD1" box="[1134,1223,794,818]" name="Mexico" pageId="5" pageNumber="66">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C370DAC9FFDE7108FF4AFC8232BAF851" pageId="5" pageNumber="66" type="distribution">
|
||
<paragraph id="8BD58942FFDE7108FF4AFC82341EFC99" blockId="5.[151,1437,794,1970]" pageId="5" pageNumber="66">
|
||
<emphasis id="B91E5550FFDE7108FF4AFC823771FCB4" bold="true" box="[197,350,830,855]" pageId="5" pageNumber="66">Distribution.</emphasis>
|
||
Presently known only from the
|
||
<typeStatus id="54D137E0FFDE7108FD42FC8334D2FCB4" box="[717,765,831,855]" pageId="5" pageNumber="66">type</typeStatus>
|
||
locality, Cenote Aak Kimin, Akumal, Quintana Roo, Mexico’s Yucatan Peninsula (see below).
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDE7108FF48FC3A36F8FA59" blockId="5.[151,1437,794,1970]" pageId="5" pageNumber="66">
|
||
<emphasis id="B91E5550FFDE7108FF48FC3A3705FC7C" bold="true" box="[199,298,902,927]" pageId="5" pageNumber="66">Habitat.</emphasis>
|
||
Cenote Aak Kimin is located
|
||
<quantity id="4C9224A7FFDE7108FDF5FC3A34EEFC7D" box="[634,705,902,926]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="66" unit="m" value="100.0">100 m</quantity>
|
||
inland from the Caribbean coast at Half Moon Bay, about
|
||
<quantity id="4C9224A7FFDE7108FAC1FC3A33B3FC7D" box="[1358,1436,902,926]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="66" unit="km" value="1.5">1.5 km</quantity>
|
||
northeast of Akumal and
|
||
<quantity id="4C9224A7FFDE7108FE39FC16342BFC21" box="[438,516,938,962]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="66" unit="km" value="0.5">0.5 km</quantity>
|
||
southwest of Yal-ku Lagoon. While most cave passages in the Yucatan Peninsula are primarily oriented perpendicular to the coast and act as subterranean rivers, this
|
||
<quantity id="4C9224A7FFDE7108FBCEFC723255FC05" box="[1089,1146,974,998]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="66" unit="km" value="2.0">2 km</quantity>
|
||
long cave runs parallel to and just inland from the shoreline under a mangrove swamp. The cave entrance is at a depth of
|
||
<quantity id="4C9224A7FFDE7108FB4CFC4E32C1FBE9" box="[1219,1262,1010,1034]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="66" unit="m" value="3.0">3 m</quantity>
|
||
in a small (
|
||
<quantity id="4C9224A7FFDE7108FAFEFC4E3682FBCD" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="5" pageNumber="66" unit="m" value="7.5" valueMax="8.0" valueMin="7.0">7–8 m</quantity>
|
||
diameter) brackish pool surrounded by red mangrove forest. The cave has two distinctly different sections with discrete modes of formation. The upper, shallow water (
|
||
<quantity id="4C9224A7FFDE7108FC8AFB86351EFBB1" box="[773,817,1082,1106]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="66" unit="m" value="6.0">6 m</quantity>
|
||
) passages were created by solutional processes beneath the mangrove swamp and form a network of low but wide tunnels extending parallel to the coast and connecting through to the Yal-ku Lagoon. The lower deep section of the cave runs beneath the shallow passages and consists of a narrow fissure with vertical walls. It was formed from slump faulting on the platform margin and enlarged by solutional processes. Small stalactites and stalagmites are present throughout the shallow section but are brittle to the touch due to mixing corrosion between fresh and salt water. Water in the shallow water section of the cave is regulated by tidal pumping and contains a mix of both stygobitic and ocean species. Tidal pumping does not appear to be a factor within the deeper section of the cave. This section is more biologically depauperate, with only stygobites observed, including the shrimps,
|
||
<taxonomicName id="4C6AF2C1FFDE7108FDC6FAE63498FA91" box="[585,695,1370,1394]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91E5550FFDE7108FDC6FAE63498FA91" box="[585,695,1370,1394]" italics="true" pageId="5" pageNumber="66">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
sp. and
|
||
<taxonomicName id="4C6AF2C1FFDE7108FC9AFAE73286FA91" authority="Kensley, 1988" authorityName="Kensley" authorityYear="1988" box="[789,1193,1370,1394]" class="Malacostraca" family="Alpheidae" genus="Yagerocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="cozumel">
|
||
<emphasis id="B91E5550FFDE7108FC9AFAE735D1FA91" box="[789,1022,1370,1394]" italics="true" pageId="5" pageNumber="66">Yagerocaris cozumel</emphasis>
|
||
<bibRefCitation id="EFFBF4B3FFDE7108FB8AFAE63286FA91" author="Kensley" box="[1029,1193,1370,1394]" pageId="5" pageNumber="66" refString="Kensley, B. (1988) New species and records of cave shrimps from the Yucatan Peninsula (Decapoda: Agostocarididae and Hippolytidae). Journal of Crustacean Biology, 8, 688 - 699." type="journal article" year="1988">Kensley, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
, and the isopod,
|
||
<taxonomicName id="4C6AF2C1FFDE7108FAE8FAE7372CFA75" class="Malacostraca" family="Cirolanidae" genus="Metacirolana" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91E5550FFDE7108FAE8FAE7372CFA75" italics="true" pageId="5" pageNumber="66">Metacirolana</emphasis>
|
||
</taxonomicName>
|
||
sp. The specimens of
|
||
<taxonomicName id="4C6AF2C1FFDE7108FD8BFAC2356AFA75" box="[516,837,1406,1430]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDE7108FD8BFAC2356AFA75" box="[516,837,1406,1430]" italics="true" pageId="5" pageNumber="66">Triacanthoneus akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDE7108FCC2FAC33587FA74" bold="true" box="[845,936,1407,1431]" pageId="5" pageNumber="66">
|
||
<taxonomicNameLabel id="A22DE82BFFDE7108FCC2FAC33587FA74" box="[845,936,1407,1431]" pageId="5" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were collected at depths between
|
||
<quantity id="4C9224A7FFDE7108FABEFAC23344FA75" box="[1329,1387,1406,1430]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="66" unit="m" value="25.0">25 m</quantity>
|
||
and
|
||
<specimenCount id="9D6C42CBFFDE7108FF18FA1E36F8FA59" box="[151,215,1442,1466]" pageId="5" pageNumber="66" type="generic">
|
||
<quantity id="4C9224A7FFDE7108FF18FA1E36FCFA59" box="[151,211,1442,1466]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="66" unit="m" value="40.0">40 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDE7108FF48FA7B32BAF851" blockId="5.[151,1437,794,1970]" pageId="5" pageNumber="66">
|
||
A water column profile in Cenote Aak Kimin was obtained in
|
||
<date id="FFD4AF82FFDE7108FC13FA7A3209FA3D" box="[924,1062,1478,1502]" pageId="5" pageNumber="66" value="2011-03">March 2011</date>
|
||
by cave divers using a YSI 600 XLM Multiparameter Water Quality Analyzer held forwards during the descent. Surface waters of the entrance pool were brackish (
|
||
<figureCitation id="135195C7FFDE7108FE0AF9B237E3F9C5" box="[389,460,1550,1574]" captionStart="FIGURE 4" captionStartId="6.[151,250,1310,1333]" captionTargetBox="[181,1409,210,1278]" captionTargetId="figure@6.[151,1436,193,1289]" captionTargetPageId="6" captionText="FIGURE 4. Vertical profiles in Cenote Aak Kimin: a, salinity; b, temperature; c, pH; d, dissolved oxygen. Individual measurements, indicated by w symbol, were taken at 2 sec. intervals between the surface and 47 m depth." httpUri="https://zenodo.org/record/279613/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
|
||
a). A halocline was present between 4.9 and
|
||
<quantity id="4C9224A7FFDE7108FC56F9B23233F9C5" box="[985,1052,1550,1574]" metricMagnitude="0" metricUnit="m" metricValue="9.3" pageId="5" pageNumber="66" unit="m" value="9.3">9.3 m</quantity>
|
||
marking the transition to marine waters (32.8 ppt) with salinity increasing to 36.26 ppt at depths below
|
||
<specimenCount id="9D6C42CBFFDE7108FC30F98E322EF9A9" box="[959,1025,1586,1610]" pageId="5" pageNumber="66" type="generic">
|
||
<quantity id="4C9224A7FFDE7108FC30F98E35D2F9A9" box="[959,1021,1586,1610]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="66" unit="m" value="30.0">30 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
While insolation warms the surface pool (25.83 °C), a small drop in temperature occurred at slightly deeper depths, followed by a gradual increase across the halocline (
|
||
<figureCitation id="135195C7FFDE7108FE06F9C637E0F971" box="[393,463,1658,1682]" captionStart="FIGURE 4" captionStartId="6.[151,250,1310,1333]" captionTargetBox="[181,1409,210,1278]" captionTargetId="figure@6.[151,1436,193,1289]" captionTargetPageId="6" captionText="FIGURE 4. Vertical profiles in Cenote Aak Kimin: a, salinity; b, temperature; c, pH; d, dissolved oxygen. Individual measurements, indicated by w symbol, were taken at 2 sec. intervals between the surface and 47 m depth." httpUri="https://zenodo.org/record/279613/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
|
||
b). A second noticeable temperature increase at
|
||
<quantity id="4C9224A7FFDE7108FC79F9C63269F971" box="[1014,1094,1658,1682]" metricMagnitude="1" metricUnit="m" metricValue="1.0699999999999998" pageId="5" pageNumber="66" unit="m" value="10.7">10.7 m</quantity>
|
||
is likely associated with tidal pumping from the nearby Yal-ku Lagoon. A temperature maximum (27.51 °C) at
|
||
<quantity id="4C9224A7FFDE7108FBC7F92232ACF955" box="[1096,1155,1694,1718]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="66" unit="m" value="25.0">25 m</quantity>
|
||
with was followed by a gradual decrease in temperature with depth, possibly associated with deep water circulation. A
|
||
<collectingCountry id="F37DC9D2FFDE7108FB33F97F32F1F939" box="[1212,1246,1731,1754]" name="Philippines" pageId="5" pageNumber="66">pH</collectingCountry>
|
||
minimum (7.08) occurred at the top of the halocline, followed by two maxima at 7 and
|
||
<quantity id="4C9224A7FFDE7108FC27F95A35CCF91D" box="[936,995,1766,1790]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="66" unit="m" value="15.0">15 m</quantity>
|
||
, with a gradual decrease then to 7.43 at maximum depth (
|
||
<figureCitation id="135195C7FFDE7108FEEAF8B63781F8C1" box="[357,430,1802,1826]" captionStart="FIGURE 4" captionStartId="6.[151,250,1310,1333]" captionTargetBox="[181,1409,210,1278]" captionTargetId="figure@6.[151,1436,193,1289]" captionTargetPageId="6" captionText="FIGURE 4. Vertical profiles in Cenote Aak Kimin: a, salinity; b, temperature; c, pH; d, dissolved oxygen. Individual measurements, indicated by w symbol, were taken at 2 sec. intervals between the surface and 47 m depth." httpUri="https://zenodo.org/record/279613/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
|
||
c). Dissolved oxygen at the surface of the entrance pool was
|
||
<quantity id="4C9224A7FFDE7108FB0AF8B632CBF8C1" box="[1157,1252,1802,1826]" metricMagnitude="-6" metricUnit="kg" metricValue="4.75" pageId="5" pageNumber="66" unit="mg" value="4.75">4.75 mg</quantity>
|
||
/L with a slight decrease at the halocline (
|
||
<figureCitation id="135195C7FFDE7108FE4FF8923429F8A5" box="[448,518,1838,1862]" captionStart="FIGURE 4" captionStartId="6.[151,250,1310,1333]" captionTargetBox="[181,1409,210,1278]" captionTargetId="figure@6.[151,1436,193,1289]" captionTargetPageId="6" captionText="FIGURE 4. Vertical profiles in Cenote Aak Kimin: a, salinity; b, temperature; c, pH; d, dissolved oxygen. Individual measurements, indicated by w symbol, were taken at 2 sec. intervals between the surface and 47 m depth." httpUri="https://zenodo.org/record/279613/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
|
||
d). A small rebound in dissolved oxygen was observed at
|
||
<quantity id="4C9224A7FFDE7108FB11F89232F4F8A5" box="[1182,1243,1838,1862]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="66" unit="m" value="15.0">15 m</quantity>
|
||
, probably due to horizontal water flow of tidally pumped water from nearby Yal-ku Lagoon. The dissolved oxygen concentration decreased to a low of
|
||
<quantity id="4C9224A7FFDE7108FE05F8CA37CBF86D" box="[394,484,1910,1934]" metricMagnitude="-7" metricUnit="kg" metricValue="4.3" pageId="5" pageNumber="66" unit="mg" value="0.43">0.43 mg</quantity>
|
||
/L at
|
||
<specimenCount id="9D6C42CBFFDE7108FD90F8CA3470F86D" box="[543,607,1910,1934]" pageId="5" pageNumber="66" type="generic">
|
||
<quantity id="4C9224A7FFDE7108FD90F8CA3474F86D" box="[543,603,1910,1934]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="5" pageNumber="66" unit="m" value="47.0">47 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Redox levels were distinctly bimodal (data not shown), suggesting significant reducing conditions below the halocline and throughout the deep sections of the cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF15D9CAFFDD710BFF18FAA2325EFAB0" httpUri="https://zenodo.org/record/279613/files/figure.png" pageId="6" pageNumber="67" targetBox="[181,1409,210,1278]" targetPageId="6">
|
||
<paragraph id="8BD58942FFDD710BFF18FAA2325EFAB0" blockId="6.[151,1435,1310,1363]" pageId="6" pageNumber="67">
|
||
<emphasis id="B91E5550FFDD710BFF18FAA2373DFAD0" bold="true" box="[151,274,1310,1333]" pageId="6" pageNumber="67">FIGURE 4.</emphasis>
|
||
Vertical profiles in Cenote Aak Kimin: a, salinity; b, temperature; c, pH; d, dissolved oxygen. Individual measurements, indicated by w symbol, were taken at 2 sec. intervals between the surface and 47 m depth.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C370DAC9FFDD710AFF48FA3C324BFDDC" lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67" type="discussion">
|
||
<paragraph id="8BD58942FFDD710BFF48FA3C3776F93F" blockId="6.[151,1436,1407,2008]" pageId="6" pageNumber="67">
|
||
<emphasis id="B91E5550FFDD710BFF48FA3C3714FA7B" bold="true" box="[199,315,1408,1432]" pageId="6" pageNumber="67">Remarks.</emphasis>
|
||
The new species differs from the other three species of
|
||
<taxonomicName id="4C6AF2C1FFDD710BFC26FAC33278FA74" box="[937,1111,1407,1431]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91E5550FFDD710BFC26FAC33278FA74" box="[937,1111,1407,1431]" italics="true" pageId="6" pageNumber="67">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
in several respects. The eyes of
|
||
<taxonomicName id="4C6AF2C1FFDD710BFF3AFA193773FA5F" box="[181,348,1444,1468]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDD710BFF3AFA193773FA5F" box="[181,348,1444,1468]" italics="true" pageId="6" pageNumber="67">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDD710BFEECFA183792FA5F" bold="true" box="[355,445,1444,1468]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A22DE82BFFDD710BFEECFA183792FA5F" box="[355,445,1444,1468]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are completely covered by the carapace and are not visible in dorsal view as they are in the other three species. The rostrum, as seen in the dorsal view, is elongate-triangular in the three previously known species, while it is more acuminate in the new species. In the three previously described species, the dentition of the major chela is formed by similar-sized teeth, alternating on both fingers, whilst in the new species, four or five large teeth on each finger are separated by regular spaces themselves bearing smaller teeth. The posterior margin of the telson has a deep notch in the new species; it is straight in the other three species. In addition, the rostra of
|
||
<taxonomicName id="4C6AF2C1FFDD710BFA08F9E436EAF977" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B91E5550FFDD710BFA08F9E436EAF977" italics="true" pageId="6" pageNumber="67">T. toro</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6AF2C1FFDD710BFF71F9C137AEF977" box="[254,385,1660,1684]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="pacificus">
|
||
<emphasis id="B91E5550FFDD710BFF71F9C137AEF977" box="[254,385,1660,1684]" italics="true" pageId="6" pageNumber="67">T. pacificus</emphasis>
|
||
</taxonomicName>
|
||
are shorter than in
|
||
<taxonomicName id="4C6AF2C1FFDD710BFDEDF9C13525F977" box="[610,778,1660,1684]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="akumalensis">
|
||
<emphasis id="B91E5550FFDD710BFDEDF9C13525F977" box="[610,778,1660,1684]" italics="true" pageId="6" pageNumber="67">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
, not reaching beyond the second article of the antennular peduncle, whilst
|
||
<taxonomicName id="4C6AF2C1FFDD710BFED4F91C37C7F954" box="[347,488,1695,1719]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B91E5550FFDD710BFED4F91C37C7F954" box="[347,488,1695,1719]" italics="true" pageId="6" pageNumber="67">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
has orbital teeth, which are absent from the other three species, including
|
||
<taxonomicName id="4C6AF2C1FFDD710BFAC3F91C36D6F93F" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDD710BFAC3F91C36D6F93F" italics="true" pageId="6" pageNumber="67">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDD710BFE8FF9783776F93F" bold="true" box="[256,345,1732,1756]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A22DE82BFFDD710BFE8FF9783776F93F" box="[256,345,1732,1756]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDD710BFF4AF95B340BF86C" blockId="6.[151,1436,1407,2008]" pageId="6" pageNumber="67">
|
||
The branchial formula reported for the previously described species of
|
||
<taxonomicName id="4C6AF2C1FFDD710BFC77F95B3365F91C" authority="Anker 2010" authorityName="Anker" authorityYear="2010" box="[1016,1354,1767,1791]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91E5550FFDD710BFC77F95B3288F91C" box="[1016,1191,1767,1791]" italics="true" pageId="6" pageNumber="67">Triacanthoneus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFBF4B3FFDD710BFB38F95B336CF91C" author="Anker" box="[1207,1347,1767,1791]" pageId="6" pageNumber="67" refString="Anker, A. (2010) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652, 47 - 63." type="journal article" year="2010">Anker 2010</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C6AF2C1FFDD710BFEACF8B137E6F8C7" box="[291,457,1804,1828]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDD710BFEACF8B137E6F8C7" box="[291,457,1804,1828]" italics="true" pageId="6" pageNumber="67">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDD710BFE5FF8B03406F8C7" bold="true" box="[464,553,1804,1828]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A22DE82BFFDD710BFE5FF8B03406F8C7" box="[464,553,1804,1828]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in lacking a pleurobranch on the fifth pereopod and having an arthrobranch on the second maxilliped. Furthermore,
|
||
<taxonomicName id="4C6AF2C1FFDD710BFDB9F88C34F3F8A4" box="[566,732,1839,1863]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDD710BFDB9F88C34F3F8A4" box="[566,732,1839,1863]" italics="true" pageId="6" pageNumber="67">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDD710BFD6BF88C3511F8AB" bold="true" box="[740,830,1840,1864]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A22DE82BFFDD710BFD6BF88C3511F8AB" box="[740,830,1840,1864]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a fairly strong process above the coxa of the fifth pereopod, in the area of insertion of a pleurobranch, perhaps a modified pleurobranch; this character is presumably absent from the other three species.
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDD710AFF4AF827347BFE64" blockId="6.[151,1436,1407,2008]" lastBlockId="7.[151,1436,151,575]" lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67">
|
||
<bibRefCitation id="EFFBF4B3FFDD710BFF4AF8273771F850" author="Anker" box="[197,350,1947,1971]" pageId="6" pageNumber="67" refString="Anker, A. (2010) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652, 47 - 63." type="journal article" year="2010">Anker (2010)</bibRefCitation>
|
||
reported the presence of an appendix masculina in all individuals of
|
||
<taxonomicName id="4C6AF2C1FFDD710BFBFBF820330DF857" box="[1140,1314,1948,1972]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91E5550FFDD710BFBFBF820330DF857" box="[1140,1314,1948,1972]" italics="true" pageId="6" pageNumber="67">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
, including ovigerous females, suggesting some sort of hermaphroditism. In
|
||
<taxonomicName id="4C6AF2C1FFDD710BFC07F87D321DF83B" box="[904,1074,1984,2008]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDD710BFC07F87D321DF83B" box="[904,1074,1984,2008]" italics="true" pageId="6" pageNumber="67">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDD710BFBB3F87C32B6F83B" bold="true" box="[1084,1177,1984,2008]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A22DE82BFFDD710BFBB3F87C32B6F83B" box="[1084,1177,1984,2008]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the ovigerous female
|
||
<typeStatus id="54D137E0FFDC710AFF18FF2B36D6FF4C" box="[151,249,151,175]" pageId="7" pageNumber="68" type="holotype">holotype</typeStatus>
|
||
does not have an appendix masculina; the
|
||
<typeStatus id="54D137E0FFDC710AFD50FF24356FFF53" box="[735,832,152,176]" pageId="7" pageNumber="68" type="paratype">paratype</typeStatus>
|
||
specimen sexed as male does not show any evidence of egg incubation or maturation; however, the second
|
||
<typeStatus id="54D137E0FFDC710AFD78FF013578FF36" box="[759,855,189,213]" pageId="7" pageNumber="68" type="paratype">paratype</typeStatus>
|
||
has both eggs and a well-developed appendix masculina. At this stage, with the evidence coming from only three individuals, we could suggest the presence of another variation of hermaphroditism in
|
||
<taxonomicName id="4C6AF2C1FFDC710AFDF9FEB9350FFEFF" box="[630,800,260,284]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="akumalensis">
|
||
<emphasis id="B91E5550FFDC710AFDF9FEB9350FFEFF" box="[630,800,260,284]" italics="true" pageId="7" pageNumber="68">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
. Another difference between the previously described marine species of
|
||
<taxonomicName id="4C6AF2C1FFDC710AFEECFE94343DFEA3" box="[355,530,296,320]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91E5550FFDC710AFEECFE94343DFEA3" box="[355,530,296,320]" italics="true" pageId="7" pageNumber="68">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
and the cave-dwelling
|
||
<taxonomicName id="4C6AF2C1FFDC710AFC98FE953591FEA3" box="[791,958,296,320]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDC710AFC98FE953591FEA3" box="[791,958,296,320]" italics="true" pageId="7" pageNumber="68">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDC710AFC4AFE943230FEA3" bold="true" box="[965,1055,296,320]" pageId="7" pageNumber="68">
|
||
<taxonomicNameLabel id="A22DE82BFFDC710AFC4AFE943230FEA3" box="[965,1055,296,320]" pageId="7" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the size: the carapace length of
|
||
<taxonomicName id="4C6AF2C1FFDC710AFF18FEF13712FE87" box="[151,317,332,356]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDC710AFF18FEF13712FE87" box="[151,317,332,356]" italics="true" pageId="7" pageNumber="68">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDC710AFECAFEF0378FFE87" bold="true" box="[325,416,332,356]" pageId="7" pageNumber="68">
|
||
<taxonomicNameLabel id="A22DE82BFFDC710AFECAFEF0378FFE87" box="[325,416,332,356]" pageId="7" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(8.0, 8.9,
|
||
<quantity id="4C9224A7FFDC710AFD9BFEF03442FE87" box="[532,621,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="68" unit="mm" value="9.1">9.1 mm</quantity>
|
||
) is more than twice to three times that of the three previously described species (maximum cl 2.8, 3.7,
|
||
<quantity id="4C9224A7FFDC710AFE61FED33467FE64" box="[494,584,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="7" pageNumber="68" unit="mm" value="4.1">4.1 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD58942FFDC710AFF4AFE29324BFDDC" blockId="7.[151,1436,151,575]" pageId="7" pageNumber="68">
|
||
In respect to the anchialine caridean fauna of the Yucatan Peninsula,
|
||
<taxonomicName id="4C6AF2C1FFDC710AFC6FFE2832A1FE4F" box="[992,1166,404,428]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91E5550FFDC710AFC6FFE2832A1FE4F" box="[992,1166,404,428]" italics="true" pageId="7" pageNumber="68">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
represents the eleventh caridean genus and the second alpheid genus after
|
||
<taxonomicName id="4C6AF2C1FFDC710AFD5BFE05322DFE2C" authority="Kensley, 1988" authorityName="Kensley" authorityYear="1988" box="[724,1026,439,464]" class="Malacostraca" family="Alpheidae" genus="Yagerocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91E5550FFDC710AFD5BFE053576FE33" box="[724,857,441,464]" italics="true" pageId="7" pageNumber="68">Yagerocaris</emphasis>
|
||
<bibRefCitation id="EFFBF4B3FFDC710AFCEEFE0B322DFE2C" author="Kensley" box="[865,1026,439,463]" pageId="7" pageNumber="68" refString="Kensley, B. (1988) New species and records of cave shrimps from the Yucatan Peninsula (Decapoda: Agostocarididae and Hippolytidae). Journal of Crustacean Biology, 8, 688 - 699." type="journal article" year="1988">Kensley, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
to be recorded in this area (
|
||
<bibRefCitation id="EFFBF4B3FFDC710AFAB1FE0B36FDFE17" author="Kensley" pageId="7" pageNumber="68" refString="Kensley, B. (1988) New species and records of cave shrimps from the Yucatan Peninsula (Decapoda: Agostocarididae and Hippolytidae). Journal of Crustacean Biology, 8, 688 - 699." type="journal article" year="1988">Kensley 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFBF4B3FFDC710AFF51FE603747FE17" author="Anker" box="[222,360,476,500]" pageId="7" pageNumber="68" refString="Anker, A. (2008) A worldwide review of stygobiotic and stygophilic shrimps of the family Alpheidae (Crustacea, Decapoda, Caridea). Subterranean Biology, 6, 1 - 16." type="journal article" year="2008">Anker 2008</bibRefCitation>
|
||
). In addition,
|
||
<taxonomicName id="4C6AF2C1FFDC710AFD8CFE61348CFE17" box="[515,675,476,500]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="akumalensis">
|
||
<emphasis id="B91E5550FFDC710AFD8CFE61348CFE17" box="[515,675,476,500]" italics="true" pageId="7" pageNumber="68">T akumalensis</emphasis>
|
||
</taxonomicName>
|
||
becomes the 14th species of anchialine shrimp in Yucatan (
|
||
<bibRefCitation id="EFFBF4B3FFDC710AFACEFE60371BFDF9" author="Alvarez" pageId="7" pageNumber="68" refString="Alvarez, F. & Iliffe, T. M. (2008) Fauna anquihalina de Yucatan. In: Alvarez F. & Rodriguez-Almaraz, G. (Eds.), Crustaceos de Mexico: estado actual de su conocimiento, Universidad Autonoma de Nuevo Leon-Promep, pp. 379 - 418." type="book chapter" year="2008">Alvarez & Iliffe 2008</bibRefCitation>
|
||
). Within the family
|
||
<taxonomicName id="4C6AF2C1FFDC710AFD92FDBE34BCFDF9" box="[541,659,514,538]" class="Malacostraca" family="Alpheidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6AF2C1FFDC710AFD2FFDB83566FDF8" box="[672,841,515,539]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="akumalensis" status="sp. nov.">
|
||
<emphasis id="B91E5550FFDC710AFD2FFDB83566FDF8" box="[672,841,515,539]" italics="true" pageId="7" pageNumber="68">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E5550FFDC710AFCDDFDBF3581FDF8" bold="true" box="[850,942,515,539]" pageId="7" pageNumber="68">
|
||
<taxonomicNameLabel id="A22DE82BFFDC710AFCDDFDBF3581FDF8" box="[850,942,515,539]" pageId="7" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the 10th described species reported from cave and anchialine habitats, with several more awaiting description (see
|
||
<bibRefCitation id="EFFBF4B3FFDC710AFC41FD9B3277FDDC" author="Anker" box="[974,1112,551,575]" pageId="7" pageNumber="68" refString="Anker, A. (2008) A worldwide review of stygobiotic and stygophilic shrimps of the family Alpheidae (Crustacea, Decapoda, Caridea). Subterranean Biology, 6, 1 - 16." type="journal article" year="2008">Anker 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |