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<mods:title id="6006BAA8AD490C10D1B52EAE66896860">Two Eocene species of peacock mites (Acari: Tetranychoidea: Tuckerellidae)</mods:title>
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<treatment id="03832F724A1D4932FE17FB178C81FAC6" ID-DOI="http://doi.org/10.5281/zenodo.4501403" ID-GBIF-Taxon="177209876" ID-Zenodo-Dep="4501403" LSID="urn:lsid:plazi:treatment:03832F724A1D4932FE17FB178C81FAC6" httpUri="http://treatment.plazi.org/id/03832F724A1D4932FE17FB178C81FAC6" lastPageId="12" lastPageNumber="110" pageId="8" pageNumber="106">
<subSubSection id="C330CDEF4A1D4936FE17FB178C1FFB06" box="[491,1359,1198,1226]" pageId="8" pageNumber="106" type="nomenclature">
<paragraph id="8B959E644A1D4936FE17FB178C1FFB06" blockId="8.[491,1359,1198,1226]" box="[491,1359,1198,1226]" pageId="8" pageNumber="106">
<heading id="D0DD29084A1D4936FE17FB178C1FFB06" bold="true" box="[491,1359,1198,1226]" fontSize="12" level="2" pageId="8" pageNumber="106" reason="6">
<emphasis id="B95E42764A1D4936FE17FB178C1FFB06" bold="true" box="[491,1359,1198,1226]" pageId="8" pageNumber="106">
<taxonomicName id="4C2AE5E74A1D4936FE17FB178C1BFB06" authority="Khaustov, Sergeyenko &amp; Perkovsky, 2014" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityYear="2014" box="[491,1355,1198,1226]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="106" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A1D4936FE17FB178A57FB06" bold="true" box="[491,775,1198,1226]" italics="true" pageId="8" pageNumber="106">Tuckerella fossilibus</emphasis>
<bibRefCitation id="EFBBE3954A1D4936FCF2FB178C1BFB06" author="Khaustov A. A. &amp; Sergeyenko A. L. &amp; Perkovsky E. E." box="[782,1355,1198,1226]" pageId="8" pageNumber="106" pagination="367 - 369" refId="ref8354" refString="Khaustov A. A., Sergeyenko A. L., Perkovsky E. E. 2014 - First fossil record of mites of the family Tuckerellidae (Acari: Tetranychoidea) from Rovno amber with description of a new species - Int. J. Acarology, 40 (5): 367 - 369. doi: 10.1080 / 01647954.2014.933253" type="journal article" year="2014">Khaustov, Sergeyenko &amp; Perkovsky, 2014</bibRefCitation>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C330CDEF4A1D4936FE17FB6A8B29FB23" box="[491,633,1235,1263]" pageId="8" pageNumber="106" type="description">
<paragraph id="8B959E644A1D4936FE17FB6A8B29FB23" blockId="8.[491,633,1235,1263]" box="[491,633,1235,1263]" pageId="8" pageNumber="106">
<emphasis id="B95E42764A1D4936FE17FB6A8B29FB23" bold="true" box="[491,633,1235,1263]" pageId="8" pageNumber="106">
(
<figureCitation id="131182E14A1D4936FE08FB6A8B20FB23" box="[500,624,1235,1263]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-2="Figure 7" captionStartId-0="9.[119,190,1903,1924]" captionStartId-1="10.[515,586,1440,1461]" captionStartId-2="11.[515,586,1189,1210]" captionTargetBox-0="[163,1433,233,1850]" captionTargetBox-1="[583,1390,221,1384]" captionTargetBox-2="[504,1482,220,1135]" captionTargetId-0="figure-2@9.[163,1433,233,1850]" captionTargetId-1="figure-2@10.[503,1487,219,1386]" captionTargetId-2="figure-2@11.[503,1487,220,1135]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." captionText-1="Figure 6 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph dorsally. Abbreviation: sty, stylophore." captionText-2="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi-0="http://doi.org/10.5281/zenodo.4487591" figureDoi-1="http://doi.org/10.5281/zenodo.4487593" figureDoi-2="http://doi.org/10.5281/zenodo.4487595" httpUri-0="https://zenodo.org/record/4487591/files/figure.png" httpUri-1="https://zenodo.org/record/4487593/files/figure.png" httpUri-2="https://zenodo.org/record/4487595/files/figure.png" pageId="8" pageNumber="106">Figs 57</figureCitation>
)
</emphasis>
</paragraph>
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<subSubSection id="C330CDEF4A1D4936FE17FAB08C20FAED" box="[491,1392,1289,1313]" pageId="8" pageNumber="106" type="reference_group">
<paragraph id="8B959E644A1D4936FE17FAB08C20FAED" blockId="8.[491,1536,1289,2009]" box="[491,1392,1289,1313]" pageId="8" pageNumber="106">
<taxonomicName id="4C2AE5E74A1D4936FE17FAB08D8DFAED" authority="Khaustov, Sergeyenko &amp; Perkovsky, 2014, p. 367" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityPageNumber="367" authorityYear="2014" box="[491,1245,1289,1313]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="106" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A1D4936FE17FAB08B95FAED" box="[491,709,1289,1313]" italics="true" pageId="8" pageNumber="106">Tuckerella fossilibus</emphasis>
<bibRefCitation id="EFBBE3954A1D4936FD30FAB08DDEFAED" author="Khaustov A. A. &amp; Sergeyenko A. L. &amp; Perkovsky E. E." box="[716,1166,1289,1313]" pageId="8" pageNumber="106" pagination="367 - 369" refId="ref8354" refString="Khaustov A. A., Sergeyenko A. L., Perkovsky E. E. 2014 - First fossil record of mites of the family Tuckerellidae (Acari: Tetranychoidea) from Rovno amber with description of a new species - Int. J. Acarology, 40 (5): 367 - 369. doi: 10.1080 / 01647954.2014.933253" type="journal article" year="2014">Khaustov, Sergeyenko &amp; Perkovsky, 2014</bibRefCitation>
, p. 367
</taxonomicName>
,
<figureCitation id="131182E14A1D4936FB1BFAB08C3DFAED" box="[1255,1389,1289,1313]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="4.[119,190,1834,1855]" captionStartId-1="5.[515,586,945,966]" captionStartId-2="6.[515,586,1371,1392]" captionStartId-3="7.[119,190,1487,1508]" captionStartId-4="9.[119,190,1903,1924]" captionTargetBox-0="[166,1433,279,1779]" captionTargetBox-1="[504,1488,222,892]" captionTargetBox-2="[608,1425,223,1316]" captionTargetBox-3="[307,1290,221,1431]" captionTargetBox-4="[163,1433,233,1850]" captionTargetId-0="figure-2@4.[166,1433,279,1779]" captionTargetId-1="figure-2@5.[503,1489,221,892]" captionTargetId-2="figure-2@6.[503,1487,220,1318]" captionTargetId-3="figure-2@7.[307,1291,219,1434]" captionTargetId-4="figure-2@9.[163,1433,233,1850]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="9" captionText-0="Figure 1 Tuckerella weiterschani n. sp. holotype. Layered DIC micrographs: A dorsal view, inserts show two aspects of amber piece containing holotype and its package, not to scale; B ventral; C posterior; D, E, details of idiosomal setae, ventral view; F, eyes; G, posterior idiosomal setae laterally; H, detail of prodorsal setav1 and peritreme; J, lateral aspect. Rectangles in A, B, J show positions of Figs D-G. Abbreviations: an, anal area; ge, genital area; oc, eyes, per, peritreme; plp, palp; sub, subcapitulum; IIV, legs." captionText-1="Figure 2 Tuckerella weiterschani n. sp. holotype. Layered POL micrograph. Gnathosoma laterally, arrow points to adoral seta. Abbreviations: palp segments: Fe, femur; Ge, genu; Ta, tarsus; Ti, tibia; ti.cl, palptibial claw." captionText-2="Figure 3 Tuckerella weiterschani n. sp. holotype dorsally." captionText-3="Figure 4 Tuckerella weiterschani n. sp. holotype details. A, right tarsus I adaxially; A, left leg I abaxially, gnathosoma and anterior idiosoma dorsally; B, leg II abaxially; C, leg III abaxially; D, leg IV abaxially; D, tarsus and tibia IV dorsally. Abbreviations: oc, eyes; per, peritreme; plp, left (poorly visible) palp; sub, subcapitulum; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." captionText-4="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi-0="http://doi.org/10.5281/zenodo.4487581" figureDoi-1="http://doi.org/10.5281/zenodo.4487585" figureDoi-2="http://doi.org/10.5281/zenodo.4487587" figureDoi-3="http://doi.org/10.5281/zenodo.4487589" figureDoi-4="http://doi.org/10.5281/zenodo.4487591" httpUri-0="https://zenodo.org/record/4487581/files/figure.png" httpUri-1="https://zenodo.org/record/4487585/files/figure.png" httpUri-2="https://zenodo.org/record/4487587/files/figure.png" httpUri-3="https://zenodo.org/record/4487589/files/figure.png" httpUri-4="https://zenodo.org/record/4487591/files/figure.png" pageId="8" pageNumber="106">Figures 15</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330CDEF4A1D4932FDEFFA958C81FAC6" lastPageId="12" lastPageNumber="110" pageId="8" pageNumber="106" type="description">
<paragraph id="8B959E644A1D4936FDEFFA958C7EFA88" blockId="8.[491,1536,1289,2009]" box="[531,1326,1324,1348]" pageId="8" pageNumber="106">Redescription — Deutonymph. Length of idiosoma 245, width 165.</paragraph>
<paragraph id="8B959E644A1D4936FDEFFAF68DFAF9B7" blockId="8.[491,1536,1289,2009]" pageId="8" pageNumber="106">
Gnathosoma (
<figureCitation id="131182E14A1D4936FD51FAF68A59FAAB" box="[685,777,1359,1383]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="8" pageNumber="106">Figs 5D</figureCitation>
,
<figureCitation id="131182E14A1D4936FCEEFAF68A02FAAB" box="[786,850,1359,1383]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="8" pageNumber="106">7B, C</figureCitation>
) — Left palp covers main part of right palp dorsally, both palps preserved in a way to cover infracapitulum dorsally. Palpgenu dorsally with seta
<emphasis id="B95E42764A1D4936FAB6FAC88C0AFA45" box="[1354,1370,1393,1417]" italics="true" pageId="8" pageNumber="106">d</emphasis>
slightly not reaching base of tibial claw. Palptibia with large, hooked tibial claw and three pairs of setae:
<emphasis id="B95E42764A1D4936FA37FA2D8C8BFA60" box="[1483,1499,1428,1452]" italics="true" pageId="8" pageNumber="106">d</emphasis>
slightly longer than length of palpgenu; short
<emphasis id="B95E42764A1D4936FC31FA0F8AB3FA02" box="[973,995,1462,1486]" italics="true" pageId="8" pageNumber="106">l”</emphasis>
, reaching to tip of tibial claw; medium-sized
<emphasis id="B95E42764A1D4936FA3EFA0F8C82FA02" box="[1474,1490,1462,1486]" italics="true" pageId="8" pageNumber="106">l</emphasis>
, about length of palptibia+tibial claw. Inner margin of tibial claw transversely striate. Palptarsus with 5 visible phaneres: 3 terminal, blunt-ended (probably palpal eupathidia), one short dorsal and one short ventral. Subcapitulum long and narrow with visible stylet protruding apically (
<figureCitation id="131182E14A1D4936FE08F9F98B02F994" box="[500,594,1600,1624]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="8" pageNumber="106">Figs 5D</figureCitation>
,
<figureCitation id="131182E14A1D4936FDA1F9F88B2FF994" box="[605,639,1601,1624]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="8" pageNumber="106">7B</figureCitation>
, st); ventrally with one pair of long subcapitular setae
<emphasis id="B95E42764A1D4936FB40F9F88D9FF994" box="[1212,1231,1601,1624]" italics="true" pageId="8" pageNumber="106">m</emphasis>
. Peritremes not evident. Oval stylophore, ca 100x40, visible inside body (
<figureCitation id="131182E14A1D4936FC00F9DA8D0AF9B7" box="[1020,1114,1635,1659]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="8" pageNumber="106">Figs 5A</figureCitation>
,
<figureCitation id="131182E14A1D4936FB99F9DA8D24F9B7" box="[1125,1140,1635,1659]" captionStart="Figure 6" captionStartId="10.[515,586,1440,1461]" captionTargetBox="[583,1390,221,1384]" captionTargetId="figure-2@10.[503,1487,219,1386]" captionTargetPageId="10" captionText="Figure 6 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph dorsally. Abbreviation: sty, stylophore." figureDoi="http://doi.org/10.5281/zenodo.4487593" httpUri="https://zenodo.org/record/4487593/files/figure.png" pageId="8" pageNumber="106">6</figureCitation>
, sty).
</paragraph>
<paragraph id="8B959E644A1D4934FDEFF93C8CC1F9B1" blockId="8.[491,1536,1289,2009]" lastBlockId="10.[491,1548,1602,2041]" lastPageId="10" lastPageNumber="108" pageId="8" pageNumber="106">
Idiosomal dorsum (
<figureCitation id="131182E14A1D4936FD10F93C8A65F951" box="[748,821,1669,1693]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="8" pageNumber="106">Figs 5</figureCitation>
,
<figureCitation id="131182E14A1D4936FCBCF93C8A1FF951" box="[832,847,1669,1693]" captionStart="Figure 6" captionStartId="10.[515,586,1440,1461]" captionTargetBox="[583,1390,221,1384]" captionTargetId="figure-2@10.[503,1487,219,1386]" captionTargetPageId="10" captionText="Figure 6 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph dorsally. Abbreviation: sty, stylophore." figureDoi="http://doi.org/10.5281/zenodo.4487593" httpUri="https://zenodo.org/record/4487593/files/figure.png" pageId="8" pageNumber="106">6</figureCitation>
,
<figureCitation id="131182E14A1D4936FCA6F93C8A2CF951" box="[858,892,1669,1693]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="8" pageNumber="106">7B</figureCitation>
) — Cuticular ornamentation, where visible anterodorsally, irregularly reticulate (
<figureCitation id="131182E14A1D4936FD2BF9118A69F90C" box="[727,825,1704,1728]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="8" pageNumber="106">Fig. 5A</figureCitation>
). Number of dorsal idiosomal setae as in
<taxonomicName id="4C2AE5E74A1D4936FB06F9108CF0F90C" authority="Sidorchuk &amp; Khaustov, 2018" authorityName="Sidorchuk &amp; Khaustov" authorityYear="2018" box="[1274,1440,1704,1728]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="106" phylum="Arthropoda" rank="species" species="weiterschani" status="n.">
<emphasis id="B95E42764A1D4936FB06F9108C59F90C" box="[1274,1289,1705,1728]" italics="true" pageId="8" pageNumber="106">T</emphasis>
.
<emphasis id="B95E42764A1D4936FAE4F9118CF0F90C" box="[1304,1440,1704,1728]" italics="true" pageId="8" pageNumber="106">weiterschani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DFF0D4A1D4936FA54F9108CEDF90C" box="[1448,1469,1705,1728]" pageId="8" pageNumber="106">n.</taxonomicNameLabel>
sp. Setae
<emphasis id="B95E42764A1D4936FDA9F9728B31F92E" box="[597,609,1739,1762]" italics="true" pageId="8" pageNumber="106">v</emphasis>
<subScript id="17AE9C214A1D4936FD9DF96E8B3AF924" attach="left" box="[609,618,1751,1768]" fontSize="7" pageId="8" pageNumber="106">2</subScript>
,
<emphasis id="B95E42764A1D4936FD86F9728BC0F92E" box="[634,656,1739,1762]" italics="true" pageId="8" pageNumber="106">sc</emphasis>
<subScript id="17AE9C214A1D4936FD6CF96E8BC9F924" attach="left" box="[656,665,1751,1768]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
and
<emphasis id="B95E42764A1D4936FD33F9728BB5F92E" box="[719,741,1739,1762]" italics="true" pageId="8" pageNumber="106">sc</emphasis>
<subScript id="17AE9C214A1D4936FD19F96E8BBEF924" attach="left" box="[741,750,1751,1768]" fontSize="7" pageId="8" pageNumber="106">2</subScript>
fan-like (
<figureCitation id="131182E14A1D4936FCABF9738A85F92E" box="[855,981,1738,1762]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="8" pageNumber="106">Fig. 5C, D</figureCitation>
);
<emphasis id="B95E42764A1D4936FC1FF9728ABFF92E" box="[995,1007,1739,1762]" italics="true" pageId="8" pageNumber="106">v</emphasis>
<subScript id="17AE9C214A1D4936FC13F96E8AA8F924" attach="left" box="[1007,1016,1751,1768]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
considerably different from other prodorsal setae, with smooth margins and weakly tapered tip (
<figureCitation id="131182E14A1D4936FBCAF9548DF0F8C9" box="[1078,1184,1773,1797]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="8" pageNumber="106">Figs. 5D</figureCitation>
,
<figureCitation id="131182E14A1D4936FB50F9548DECF8C9" box="[1196,1212,1773,1797]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="8" pageNumber="106">7</figureCitation>
). Two pairs of eyes located anterolaterally to bases of setae
<emphasis id="B95E42764A1D4936FCBEF8A88A08F8E4" box="[834,856,1809,1832]" italics="true" pageId="8" pageNumber="106">sc</emphasis>
<subScript id="17AE9C214A1D4936FCA4F8A58A31F8E1" attach="left" box="[856,865,1820,1837]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
; posterior eye distinctly larger than anterior (
<figureCitation id="131182E14A1D4936FAADF8A98CFEF8EB" box="[1361,1454,1808,1832]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="8" pageNumber="106">Fig. 7B</figureCitation>
, oc). All setae of C, D, E and F-rows broad, flat, ovate to obovate in shape:
<emphasis id="B95E42764A1D4936FB2FF88A8D8FF886" box="[1235,1247,1843,1866]" italics="true" pageId="8" pageNumber="106">c</emphasis>
<subScript id="17AE9C214A1D4936FB23F8878DB8F883" attach="left" box="[1247,1256,1854,1871]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
-
<emphasis id="B95E42764A1D4936FB0EF88A8DAEF886" box="[1266,1278,1843,1866]" italics="true" pageId="8" pageNumber="106">c</emphasis>
<subScript id="17AE9C214A1D4936FB02F8878C54F883" attach="left" box="[1278,1284,1854,1871]" fontSize="7" pageId="8" pageNumber="106">3</subScript>
,
<emphasis id="B95E42764A1D4936FAE9F88B8C73F886" box="[1301,1315,1842,1866]" italics="true" pageId="8" pageNumber="106">d</emphasis>
<subScript id="17AE9C214A1D4936FAD8F8878C7DF883" attach="left" box="[1316,1325,1854,1871]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
-
<emphasis id="B95E42764A1D4936FACBF88B8C15F886" box="[1335,1349,1842,1866]" italics="true" pageId="8" pageNumber="106">d</emphasis>
<subScript id="17AE9C214A1D4936FABAF8878C1CF883" attach="left" box="[1350,1356,1854,1871]" fontSize="7" pageId="8" pageNumber="106">3</subScript>
and
<emphasis id="B95E42764A1D4936FA7FF88A8CDFF886" box="[1411,1423,1843,1866]" italics="true" pageId="8" pageNumber="106">e</emphasis>
<subScript id="17AE9C214A1D4936FA73F8878CC8F883" attach="left" box="[1423,1432,1854,1871]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
,
<emphasis id="B95E42764A1D4936FA5AF88A8CE2F886" box="[1446,1458,1843,1866]" italics="true" pageId="8" pageNumber="106">e</emphasis>
<subScript id="17AE9C214A1D4936FA4EF8878CEBF883" attach="left" box="[1458,1467,1854,1871]" fontSize="7" pageId="8" pageNumber="106">2</subScript>
of medium size, subequal (16 18 x 24 28);
<emphasis id="B95E42764A1D4936FC44F8EC8AEFF8A1" box="[952,959,1877,1901]" italics="true" pageId="8" pageNumber="106">f</emphasis>
<subScript id="17AE9C214A1D4936FC39F8D88A9EF8BE" attach="none" box="[965,974,1889,1906]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
and
<emphasis id="B95E42764A1D4936FBFEF8EC8D59F8A1" box="[1026,1033,1877,1901]" italics="true" pageId="8" pageNumber="106">f</emphasis>
<subScript id="17AE9C214A1D4936FBF2F8D88D47F8BE" attach="left" box="[1038,1047,1889,1906]" fontSize="7" pageId="8" pageNumber="106">2</subScript>
slightly smaller than
<emphasis id="B95E42764A1D4936FB00F8EF8C58F8A1" box="[1276,1288,1878,1901]" italics="true" pageId="8" pageNumber="106">e</emphasis>
<subScript id="17AE9C214A1D4936FAF4F8D88C41F8BE" attach="left" box="[1288,1297,1889,1906]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
and
<emphasis id="B95E42764A1D4936FAB9F8EF8C01F8A1" box="[1349,1361,1878,1901]" italics="true" pageId="8" pageNumber="106">e</emphasis>
<subScript id="17AE9C214A1D4936FAADF8D88C0AF8BE" attach="left" box="[1361,1370,1889,1906]" fontSize="7" pageId="8" pageNumber="106">2</subScript>
;
<emphasis id="B95E42764A1D4936FA95F8EC8C20F8A1" box="[1385,1392,1877,1901]" italics="true" pageId="8" pageNumber="106">f</emphasis>
<subScript id="17AE9C214A1D4936FA89F8D88C2EF8BE" attach="left" box="[1397,1406,1889,1906]" fontSize="7" pageId="8" pageNumber="106">2</subScript>
situated on same transverse level as
<emphasis id="B95E42764A1D4936FCE9F8CE8A4CF843" box="[789,796,1911,1935]" italics="true" pageId="8" pageNumber="106">f</emphasis>
<subScript id="17AE9C214A1D4936FCDEF83A8A7BF858" attach="right" box="[802,811,1923,1940]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
. Lateral setae
<emphasis id="B95E42764A1D4936FC37F8C18A87F843" box="[971,983,1912,1935]" italics="true" pageId="8" pageNumber="106">c</emphasis>
<subScript id="17AE9C214A1D4936FC2BF83A8AB0F858" attach="left" box="[983,992,1923,1940]" fontSize="7" pageId="8" pageNumber="106">5</subScript>
-
<emphasis id="B95E42764A1D4936FC17F8C18AA7F843" box="[1003,1015,1912,1935]" italics="true" pageId="8" pageNumber="106">c</emphasis>
<subScript id="17AE9C214A1D4936FC0AF83A8D51F858" attach="left" box="[1014,1025,1923,1940]" fontSize="7" pageId="8" pageNumber="106">7</subScript>
,
<emphasis id="B95E42764A1D4936FBF3F8CE8D4DF843" box="[1039,1053,1911,1935]" italics="true" pageId="8" pageNumber="106">d</emphasis>
<subScript id="17AE9C214A1D4936FBE1F83A8D73F858" attach="left" box="[1053,1059,1923,1940]" fontSize="7" pageId="8" pageNumber="106">4</subScript>
-
<emphasis id="B95E42764A1D4936FBCDF8CE8D1EF843" box="[1073,1102,1911,1935]" italics="true" pageId="8" pageNumber="106">d6</emphasis>
and
<emphasis id="B95E42764A1D4936FB7CF8CE8DCAF843" box="[1152,1178,1911,1935]" italics="true" pageId="8" pageNumber="106">e3</emphasis>
distinctly larger than central, more elongate, with rounded tips, largest
<emphasis id="B95E42764A1D4936FC56F8238AE8F87E" box="[938,952,1946,1970]" italics="true" pageId="8" pageNumber="106">d</emphasis>
<subScript id="17AE9C214A1D4936FC45F81F8A92F87B" attach="left" box="[953,962,1958,1975]" fontSize="7" pageId="8" pageNumber="106">6</subScript>
28 x 60. Setae
<emphasis id="B95E42764A1D4936FB8CF8238D2DF87E" box="[1136,1149,1946,1970]" italics="true" pageId="8" pageNumber="106">h</emphasis>
<subScript id="17AE9C214A1D4936FB81F81F8DD6F87B" attach="left" box="[1149,1158,1958,1975]" fontSize="7" pageId="8" pageNumber="106">1</subScript>
large, flat, obovate (27 x 55), setae
<emphasis id="B95E42764A1D4936FDD4F8058B65F818" box="[552,565,1980,2004]" italics="true" pageId="8" pageNumber="106">h</emphasis>
<subScript id="17AE9C214A1D4936FDC9F8718B6EF815" attach="left" box="[565,574,1992,2009]" fontSize="7" pageId="8" pageNumber="106">2</subScript>
<emphasis id="B95E42764A1D4936FDAEF8058B0FF818" box="[594,607,1980,2004]" italics="true" pageId="8" pageNumber="106">h</emphasis>
<subScript id="17AE9C214A1D4936FDA3F8718B34F815" attach="left" box="[607,612,1992,2009]" fontSize="7" pageId="8" pageNumber="106">8</subScript>
flagellate:
<emphasis id="B95E42764A1D4936FD1AF8058BA3F818" box="[742,755,1980,2004]" italics="true" pageId="8" pageNumber="106">h</emphasis>
<subScript id="17AE9C214A1D4936FD0FF8718BACF815" attach="left" box="[755,764,1992,2009]" fontSize="7" pageId="8" pageNumber="106">2</subScript>
,
<emphasis id="B95E42764A1D4936FCF0F8058A49F818" box="[780,793,1980,2004]" italics="true" pageId="8" pageNumber="106">h</emphasis>
<subScript id="17AE9C214A1D4936FCE5F8718A4FF815" attach="left" box="[793,799,1992,2009]" fontSize="7" pageId="8" pageNumber="106">4</subScript>
,
<emphasis id="B95E42764A1D4936FCCEF8058A6FF818" box="[818,831,1980,2004]" italics="true" pageId="8" pageNumber="106">h</emphasis>
<subScript id="17AE9C214A1D4936FCBCF8718A19F815" attach="left" box="[832,841,1992,2009]" fontSize="7" pageId="8" pageNumber="106">5</subScript>
,
<emphasis id="B95E42764A1D4936FCA5F8058A36F818" box="[857,870,1980,2004]" italics="true" pageId="8" pageNumber="106">h</emphasis>
<subScript id="17AE9C214A1D4936FC9AF8718A21F815" attach="left" box="[870,881,1992,2009]" fontSize="7" pageId="8" pageNumber="106">7</subScript>
long, barbed, tapered (180, 270, 180, 220 respectively),
<emphasis id="B95E42764A1F4934FE17F9FB88A8F996" box="[491,504,1602,1626]" italics="true" pageId="10" pageNumber="108">h</emphasis>
<subScript id="17AE9C214A1F4934FE05F9F78B52F993" attach="left" box="[505,514,1614,1631]" fontSize="7" pageId="10" pageNumber="108">6</subScript>
short, thick, heavily barbed, blunt (40);
<emphasis id="B95E42764A1F4934FC4FF9FB8A90F996" box="[947,960,1602,1626]" italics="true" pageId="10" pageNumber="108">h</emphasis>
<subScript id="17AE9C214A1F4934FC3CF9F78A96F993" attach="left" box="[960,966,1614,1631]" fontSize="7" pageId="10" pageNumber="108">3</subScript>
,
<emphasis id="B95E42764A1F4934FC24F9FB8AB5F996" box="[984,997,1602,1626]" italics="true" pageId="10" pageNumber="108">h</emphasis>
<subScript id="17AE9C214A1F4934FC1AF9F78ABBF993" attach="left" box="[998,1003,1614,1631]" fontSize="7" pageId="10" pageNumber="108">8</subScript>
intermediate in length and thickness, barbed, tapered (80). Only anterior margins of dorsal shields C and D clearly visible (
<figureCitation id="131182E14A1F4934FAC7F9DC8CD4F9B1" box="[1339,1412,1637,1661]" captionStart="Figure 6" captionStartId="10.[515,586,1440,1461]" captionTargetBox="[583,1390,221,1384]" captionTargetId="figure-2@10.[503,1487,219,1386]" captionTargetPageId="10" captionText="Figure 6 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph dorsally. Abbreviation: sty, stylophore." figureDoi="http://doi.org/10.5281/zenodo.4487593" httpUri="https://zenodo.org/record/4487593/files/figure.png" pageId="10" pageNumber="108">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF55CEEC4A1C4937FF8BF8D68DB6F87E" ID-DOI="http://doi.org/10.5281/zenodo.4487591" ID-Zenodo-Dep="4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="9" pageNumber="107" startId="9.[119,190,1903,1924]" targetBox="[163,1433,233,1850]" targetPageId="9">
<paragraph id="8B959E644A1C4937FF8BF8D68DB6F87E" blockId="9.[119,1525,1903,1970]" pageId="9" pageNumber="107">
<emphasis id="B95E42764A1C4937FF8BF8D68980F848" bold="true" box="[119,208,1903,1924]" pageId="9" pageNumber="107">Figure 5</emphasis>
<taxonomicName id="4C2AE5E74A1C4937FF25F8D68A66F849" authority="Khaustov, Sergeyenko &amp; Perkovsky, 2014" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityYear="2014" box="[217,822,1903,1925]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="107" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A1C4937FF25F8D688CDF849" box="[217,413,1903,1925]" italics="true" pageId="9" pageNumber="107">Tuckerella fossilibus</emphasis>
<bibRefCitation id="EFBBE3954A1C4937FE5FF8D68A66F849" author="Khaustov A. A. &amp; Sergeyenko A. L. &amp; Perkovsky E. E." box="[419,822,1903,1925]" pageId="9" pageNumber="107" pagination="367 - 369" refId="ref8354" refString="Khaustov A. A., Sergeyenko A. L., Perkovsky E. E. 2014 - First fossil record of mites of the family Tuckerellidae (Acari: Tetranychoidea) from Rovno amber with description of a new species - Int. J. Acarology, 40 (5): 367 - 369. doi: 10.1080 / 01647954.2014.933253" type="journal article" year="2014">Khaustov, Sergeyenko &amp; Perkovsky, 2014</bibRefCitation>
</taxonomicName>
holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw.
</paragraph>
</caption>
<caption id="DF55CEEC4A1F4934FDFFFA198A08FA00" ID-DOI="http://doi.org/10.5281/zenodo.4487593" ID-Zenodo-Dep="4487593" httpUri="https://zenodo.org/record/4487593/files/figure.png" pageId="10" pageNumber="108" startId="10.[515,586,1440,1461]" targetBox="[583,1390,221,1384]" targetPageId="10">
<paragraph id="8B959E644A1F4934FDFFFA198A08FA00" blockId="10.[515,1482,1440,1484]" pageId="10" pageNumber="108">
<emphasis id="B95E42764A1F4934FDFFFA198B0CFA79" bold="true" box="[515,604,1440,1461]" pageId="10" pageNumber="108">Figure 6</emphasis>
<taxonomicName id="4C2AE5E74A1F4934FD9FFA198DECFA7A" authority="Khaustov, Sergeyenko &amp; Perkovsky, 2014" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityYear="2014" box="[611,1212,1440,1462]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="108" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A1F4934FD9FFA198A76FA7A" box="[611,806,1440,1462]" italics="true" pageId="10" pageNumber="108">Tuckerella fossilibus</emphasis>
<bibRefCitation id="EFBBE3954A1F4934FCD0FA198DECFA7A" author="Khaustov A. A. &amp; Sergeyenko A. L. &amp; Perkovsky E. E." box="[812,1212,1440,1462]" pageId="10" pageNumber="108" pagination="367 - 369" refId="ref8354" refString="Khaustov A. A., Sergeyenko A. L., Perkovsky E. E. 2014 - First fossil record of mites of the family Tuckerellidae (Acari: Tetranychoidea) from Rovno amber with description of a new species - Int. J. Acarology, 40 (5): 367 - 369. doi: 10.1080 / 01647954.2014.933253" type="journal article" year="2014">Khaustov, Sergeyenko &amp; Perkovsky, 2014</bibRefCitation>
</taxonomicName>
holotype deutonymph dorsally. Abbreviation: sty, stylophore.
</paragraph>
</caption>
<paragraph id="8B959E644A1F4934FDEFF93E8D5BF928" blockId="10.[491,1548,1602,2041]" pageId="10" pageNumber="108">
Idiosomal venter (
<figureCitation id="131182E14A1F4934FD22F93E8A6AF953" box="[734,826,1671,1695]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="10" pageNumber="108">Figs 5B</figureCitation>
,
<figureCitation id="131182E14A1F4934FCBBF93E8A98F953" box="[839,968,1671,1695]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="10" pageNumber="108">7A, D, E, F</figureCitation>
) — Dark and collapsed, obscured by legs. Only coxal setae are partly discernible:
<emphasis id="B95E42764A1F4934FCA4F9138AD8F90E" box="[856,904,1706,1730]" italics="true" pageId="10" pageNumber="108">1a-c</emphasis>
and
<emphasis id="B95E42764A1F4934FC47F9138A85F90E" box="[955,981,1706,1730]" italics="true" pageId="10" pageNumber="108">2b</emphasis>
smooth, fine,
<emphasis id="B95E42764A1F4934FB93F9138DD9F90E" box="[1135,1161,1706,1730]" italics="true" pageId="10" pageNumber="108">1a</emphasis>
longest;
<emphasis id="B95E42764A1F4934FB17F9138C54F90E" box="[1259,1284,1706,1730]" italics="true" pageId="10" pageNumber="108">2c</emphasis>
long, thick, heavily barbed. Genital and anal openings indiscernible.
</paragraph>
<paragraph id="8B959E644A1F4935FDEFF9568C1EF822" blockId="10.[491,1548,1602,2041]" lastBlockId="11.[491,1534,1419,2030]" lastPageId="11" lastPageNumber="109" pageId="10" pageNumber="108">
Legs (
<figureCitation id="131182E14A1F4934FDA1F9568BF6F8CB" box="[605,678,1775,1799]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="10" pageNumber="108">Fig. 7</figureCitation>
) — Dark and curled, mostly visible in ventral aspect, with some setae poorly discernible. Leg I (
<figureCitation id="131182E14A1F4934FD3FF8A88A6FF8E5" box="[707,831,1809,1833]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="10" pageNumber="108">Fig. 7A, B</figureCitation>
): trochanter without visible setae; femur with three large obovate setae (
<emphasis id="B95E42764A1F4934FDCCF88D8B6DF880" box="[560,573,1844,1868]" italics="true" pageId="10" pageNumber="108">d</emphasis>
,
<emphasis id="B95E42764A1F4934FDB7F88D8B0BF880" box="[587,603,1844,1868]" italics="true" pageId="10" pageNumber="108">l</emphasis>
,
<emphasis id="B95E42764A1F4934FD94F88D8B2EF880" box="[616,638,1844,1868]" italics="true" pageId="10" pageNumber="108">l”</emphasis>
) and setiform
<emphasis id="B95E42764A1F4934FCE6F88D8A12F880" box="[794,834,1844,1868]" italics="true" pageId="10" pageNumber="108">bv”</emphasis>
, other ventral setae not visible. Genu with three large obovate setae (
<emphasis id="B95E42764A1F4934FDCCF8EF8B6DF8A2" box="[560,573,1878,1902]" italics="true" pageId="10" pageNumber="108">d</emphasis>
,
<emphasis id="B95E42764A1F4934FDB6F8EF8B0AF8A2" box="[586,602,1878,1902]" italics="true" pageId="10" pageNumber="108">l</emphasis>
,
<emphasis id="B95E42764A1F4934FD9BF8EF8B2DF8A2" box="[615,637,1878,1902]" italics="true" pageId="10" pageNumber="108">l”</emphasis>
) and setiform
<emphasis id="B95E42764A1F4934FCEBF8EE8A62F8A2" box="[791,818,1879,1902]" italics="true" pageId="10" pageNumber="108">v”</emphasis>
, other ventral setae not visible. Tibia with large, obovate seta
<emphasis id="B95E42764A1F4934FA3FF8EF8C89F8A2" box="[1475,1497,1878,1902]" italics="true" pageId="10" pageNumber="108">l”</emphasis>
and narrower
<emphasis id="B95E42764A1F4934FD82F8C08BDBF85D" box="[638,651,1913,1937]" italics="true" pageId="10" pageNumber="108">d</emphasis>
,
<emphasis id="B95E42764A1F4934FD64F8C08BF8F85D" box="[664,680,1913,1937]" italics="true" pageId="10" pageNumber="108">l</emphasis>
; ventral setae not visible; solenidion φ baculiform, situated on short tubercle. Tarsus with two claws and empodium, each with clearly visible tenent hairs; two solenidia present, ω
<subScript id="17AE9C214A1F4934FDA4F8738B31F817" attach="left" box="[600,609,1994,2011]" fontSize="7" pageId="10" pageNumber="108">1</subScript>
about 2 times longer than ω
<subScript id="17AE9C214A1F4934FC6EF8738ACBF817" attach="left" box="[914,923,1994,2011]" fontSize="7" pageId="10" pageNumber="108">2</subScript>
; setae: (
<emphasis id="B95E42764A1F4934FC07F8068D58F81A" box="[1019,1032,1983,2006]" italics="true" pageId="10" pageNumber="108">u</emphasis>
) simple and short, abaxial
<emphasis id="B95E42764A1F4934FACFF8068C12F81A" box="[1331,1346,1983,2006]" italics="true" pageId="10" pageNumber="108"></emphasis>
() members of pairs (
<emphasis id="B95E42764A1F4934FDD3F8598B6EF834" box="[559,574,2016,2040]" italics="true" pageId="10" pageNumber="108">ft</emphasis>
) and (
<emphasis id="B95E42764A1F4934FD79F8598BCAF834" box="[645,666,2016,2040]" italics="true" pageId="10" pageNumber="108">pl</emphasis>
) long, distinctly longer than tarsus,
<emphasis id="B95E42764A1F4934FBE0F8598D6AF834" box="[1052,1082,2016,2040]" italics="true" pageId="10" pageNumber="108">pl</emphasis>
about half shorter on right leg, absent on left leg; (
<emphasis id="B95E42764A1E4935FD8FFA358BD6FA6F" box="[627,646,1420,1443]" italics="true" pageId="11" pageNumber="109">tc</emphasis>
) situated on tubercles, fine, tapered, (
<emphasis id="B95E42764A1E4935FBDFFA358D60FA6F" box="[1059,1072,1420,1443]" italics="true" pageId="11" pageNumber="109">p</emphasis>
) blunt, short, apparently eupathidial, difficult to discern where hidden behind ambulacrum;
<emphasis id="B95E42764A1E4935FBC2FA178D30FA09" box="[1086,1120,1454,1477]" italics="true" pageId="11" pageNumber="109">pv</emphasis>
setiform. Leg II (
<figureCitation id="131182E14A1E4935FACEFA148CE8FA09" box="[1330,1464,1453,1477]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="11" pageNumber="109">Fig. 7C, D</figureCitation>
): trochanter not discernible; femur with long, thick, heavily barbed seta
<emphasis id="B95E42764A1E4935FB24FA698C50FA24" box="[1240,1280,1488,1512]" italics="true" pageId="11" pageNumber="109">bv”</emphasis>
and 3 obovate large setae
<emphasis id="B95E42764A1E4935FDD4FA4B8B68F9C6" box="[552,568,1522,1546]" italics="true" pageId="11" pageNumber="109">d</emphasis>
and (
<emphasis id="B95E42764A1E4935FD89FA4B8B2CF9C6" box="[629,636,1522,1546]" italics="true" pageId="11" pageNumber="109">l</emphasis>
) (
<emphasis id="B95E42764A1E4935FD6AFA4B8BF6F9C6" box="[662,678,1522,1546]" italics="true" pageId="11" pageNumber="109">d</emphasis>
and
<emphasis id="B95E42764A1E4935FD25FA4B8BBFF9C6" box="[729,751,1522,1546]" italics="true" pageId="11" pageNumber="109">l”</emphasis>
artificially split on left leg); genu with 3 obovate setae visible; tibia with 3 obovate and one or two long simple seta visible ((
<emphasis id="B95E42764A1E4935FBBAF9AF8D02F9E1" box="[1094,1106,1558,1581]" italics="true" pageId="11" pageNumber="109">v</emphasis>
) on left, only
<emphasis id="B95E42764A1E4935FB15F9AF8C53F9E1" box="[1257,1283,1558,1581]" italics="true" pageId="11" pageNumber="109">v”</emphasis>
on right leg); tarsus with ambulacrum as in leg I (well visible on right leg), one solenidion, and 10 smooth setae: long
<emphasis id="B95E42764A1E4935FDDEF9E38B10F9BE" box="[546,576,1626,1650]" italics="true" pageId="11" pageNumber="109">ft”</emphasis>
, shorter
<emphasis id="B95E42764A1E4935FD5CF9E38BE8F9BE" box="[672,696,1626,1650]" italics="true" pageId="11" pageNumber="109">ft</emphasis>
, (
<emphasis id="B95E42764A1E4935FD33F9E38BB4F9BE" box="[719,740,1626,1650]" italics="true" pageId="11" pageNumber="109">pl</emphasis>
), (
<emphasis id="B95E42764A1E4935FCF8F9E28A47F9BE" box="[772,791,1627,1650]" italics="true" pageId="11" pageNumber="109">tc</emphasis>
), (
<emphasis id="B95E42764A1E4935FCCBF9E28A14F9BE" box="[823,836,1627,1650]" italics="true" pageId="11" pageNumber="109">u</emphasis>
) (one member of pair visible on each leg),
<emphasis id="B95E42764A1E4935FAE6F9E28C66F9BE" box="[1306,1334,1627,1650]" italics="true" pageId="11" pageNumber="109">p”</emphasis>
and
<emphasis id="B95E42764A1E4935FA90F9E28CDEF9BE" box="[1388,1422,1627,1650]" italics="true" pageId="11" pageNumber="109">pv</emphasis>
. Leg III (
<figureCitation id="131182E14A1E4935FDEAF9C48BC1F959" box="[534,657,1661,1685]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="11" pageNumber="109">Fig. 7E, F</figureCitation>
): trochanter with single visible seta
<emphasis id="B95E42764A1E4935FBF2F9C48D4EF959" box="[1038,1054,1661,1685]" italics="true" pageId="11" pageNumber="109">l</emphasis>
thick, long, heavily barbed; femur with small obovate seta
<emphasis id="B95E42764A1E4935FD46F9268B9AF97B" box="[698,714,1695,1719]" italics="true" pageId="11" pageNumber="109">d</emphasis>
and smooth simple
<emphasis id="B95E42764A1E4935FC58F9198AECF97B" box="[932,956,1696,1719]" italics="true" pageId="11" pageNumber="109">ev</emphasis>
; genu with single visible small obovate seta
<emphasis id="B95E42764A1E4935FA50F9268CECF97B" box="[1452,1468,1695,1719]" italics="true" pageId="11" pageNumber="109">l</emphasis>
; tibia with 2 clearly visible setae, simple
<emphasis id="B95E42764A1E4935FC67F97A8AF7F916" box="[923,935,1731,1754]" italics="true" pageId="11" pageNumber="109">v</emphasis>
and small obovate
<emphasis id="B95E42764A1E4935FB7CF97B8DC0F916" box="[1152,1168,1730,1754]" italics="true" pageId="11" pageNumber="109">l</emphasis>
(visible transversally on right leg), and one spot that may be alveolus of seta
<emphasis id="B95E42764A1E4935FC18F95D8AA2F930" box="[996,1010,1764,1788]" italics="true" pageId="11" pageNumber="109">d</emphasis>
; tarsus with ambulacrum like in leg II and 7 simple setae (only 4 clearly visible on right leg): (
<emphasis id="B95E42764A1E4935FBF4F8BE8D47F8D3" box="[1032,1047,1799,1823]" italics="true" pageId="11" pageNumber="109">ft</emphasis>
), (
<emphasis id="B95E42764A1E4935FBCBF8B18D1AF8D3" box="[1079,1098,1800,1823]" italics="true" pageId="11" pageNumber="109">tc</emphasis>
), (
<emphasis id="B95E42764A1E4935FB96F8B18D27F8D3" box="[1130,1143,1800,1823]" italics="true" pageId="11" pageNumber="109">u</emphasis>
) and
<emphasis id="B95E42764A1E4935FB4AF8BE8D88F8D2" box="[1206,1240,1799,1822]" italics="true" pageId="11" pageNumber="109">pv</emphasis>
. Leg IV (
<figureCitation id="131182E14A1E4935FAB0F8BE8C9CF8D3" box="[1356,1484,1799,1823]" captionStart="Figure 7" captionStartId="11.[515,586,1189,1210]" captionTargetBox="[504,1482,220,1135]" captionTargetId="figure-2@11.[503,1487,220,1135]" captionTargetPageId="11" captionText="Figure 7 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures." figureDoi="http://doi.org/10.5281/zenodo.4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="11" pageNumber="109">Fig. 7 H, J</figureCitation>
): poorly visible, dark, trochanter and setation of femur indiscernible; genu with one well visible narrowly obovate seta
<emphasis id="B95E42764A1E4935FD1CF8F58BA0F8A8" box="[736,752,1868,1892]" italics="true" pageId="11" pageNumber="109">d</emphasis>
on left leg, no visible setae on right; tibia with 2 ventral setae well visible, two discernible on right leg, one well visible seta and one discernible on left; tarsus with two claws discernible, 3 setae visible on left and one on right leg. See figure for possible setal homologies. Observed setation (trochanter to tarsus, solenidia in parentheses): leg I:?-5-3-4(1)-11(2); leg II:?-4-3-5-10(1); leg III: 1-2-1-2/3?-7; leg IV:?-?-1-2?/4?-?.
</paragraph>
<caption id="DF55CEEC4A1E4935FDFFFB1C8C45FAD9" ID-DOI="http://doi.org/10.5281/zenodo.4487595" ID-Zenodo-Dep="4487595" httpUri="https://zenodo.org/record/4487595/files/figure.png" pageId="11" pageNumber="109" startId="11.[515,586,1189,1210]" targetBox="[504,1482,220,1135]" targetPageId="11">
<paragraph id="8B959E644A1E4935FDFFFB1C8C45FAD9" blockId="11.[515,1513,1189,1301]" pageId="11" pageNumber="109">
<emphasis id="B95E42764A1E4935FDFFFB1C8B0DFB76" bold="true" box="[515,605,1189,1210]" pageId="11" pageNumber="109">Figure 7</emphasis>
<taxonomicName id="4C2AE5E74A1E4935FD99FB1C8D92FB77" authority="Khaustov, Sergeyenko &amp; Perkovsky, 2014" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityYear="2014" box="[613,1218,1189,1211]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="109" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A1E4935FD99FB1C8A79FB77" box="[613,809,1189,1211]" italics="true" pageId="11" pageNumber="109">Tuckerella fossilibus</emphasis>
<bibRefCitation id="EFBBE3954A1E4935FCCCFB1C8D92FB77" author="Khaustov A. A. &amp; Sergeyenko A. L. &amp; Perkovsky E. E." box="[816,1218,1189,1211]" pageId="11" pageNumber="109" pagination="367 - 369" refId="ref8354" refString="Khaustov A. A., Sergeyenko A. L., Perkovsky E. E. 2014 - First fossil record of mites of the family Tuckerellidae (Acari: Tetranychoidea) from Rovno amber with description of a new species - Int. J. Acarology, 40 (5): 367 - 369. doi: 10.1080 / 01647954.2014.933253" type="journal article" year="2014">Khaustov, Sergeyenko &amp; Perkovsky, 2014</bibRefCitation>
</taxonomicName>
holotype deutonymph detail: A right tarsus I in ventral view of mite; B leg I, gnathosoma and anterior proterosoma, dorsal view; CJ, legs in ventral view of mite: C right leg II; D left leg II and seta 3a; E right leg III F left leg III; H right leg IV; J left leg IV. Abbreviations: oc, eyes; st, cheliceral stylets; ta, palptarsus; ti.cl, palptibial claw. Gray lines represent poorly discernible structures.
</paragraph>
</caption>
<paragraph id="8B959E644A1E4935FA15FB6B8CBDFB2B" box="[1513,1517,1234,1255]" pageId="11" pageNumber="109">;</paragraph>
<paragraph id="8B959E644A194932FDEFFEB18C53FDF8" blockId="12.[491,1526,264,565]" pageId="12" pageNumber="110">
Remarks —
<bibRefCitation id="EFBBE3954A194932FD66FEB18AD9FEEC" author="Khaustov A. A. &amp; Sergeyenko A. L. &amp; Perkovsky E. E." box="[666,905,264,288]" pageId="12" pageNumber="110" pagination="367 - 369" refId="ref8354" refString="Khaustov A. A., Sergeyenko A. L., Perkovsky E. E. 2014 - First fossil record of mites of the family Tuckerellidae (Acari: Tetranychoidea) from Rovno amber with description of a new species - Int. J. Acarology, 40 (5): 367 - 369. doi: 10.1080 / 01647954.2014.933253" type="journal article" year="2014">
Khaustov
<emphasis id="B95E42764A194932FCFBFEB08A6DFEEC" box="[775,829,264,288]" italics="true" pageId="12" pageNumber="110">et al.</emphasis>
(2014)
</bibRefCitation>
in original description of
<taxonomicName id="4C2AE5E74A194932FB5FFEB08C71FEEC" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityYear="2014" box="[1187,1313,264,288]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A194932FB5FFEB08C71FEEC" box="[1187,1313,264,288]" italics="true" pageId="12" pageNumber="110">T. fossilibus</emphasis>
</taxonomicName>
were not able to identify its life stage. Now, after more detailed description we can suggest that it is probably a deutonymph. Our opinion is based on the setation of leg I, which shows: (1) two tarsal solenidia, (2) absence of the tarsal seta
<emphasis id="B95E42764A194932FC65FEC98AF0FE44" box="[921,928,368,392]" italics="true" pageId="12" pageNumber="110">l</emphasis>
<subScript id="17AE9C214A194932FC5DFEC58AFAFE41" attach="left" box="[929,938,380,397]" fontSize="7" pageId="12" pageNumber="110">1</subScript>
<emphasis id="B95E42764A194932FC57FEC98AE4FE4B" box="[939,948,368,391]" italics="true" pageId="12" pageNumber="110"></emphasis>
, (3) three obovate setae on each genu and tibia. According to observations by
<bibRefCitation id="EFBBE3954A194932FCC9FE2B8DF2FE66" author="Quiros-Gonzalez M. J. &amp; Baker E. W." box="[821,1186,402,426]" pageId="12" pageNumber="110" pagination="166 - 173" refId="ref9065" refString="Quiros-Gonzalez M. J., Baker E. W. 1984 - Systematics and taxonomy of Acari. 5.5 Idiosomal and leg chaetotaxy in the Tuckerellidae Baker &amp; Pritchard; ontogeny and nomenclature - In: Griffiths D. A., Bowman C. E. (Eds) Acarology 6. Vol. 1., Chichester: Ellis Horwood Ltd. pp. 166 - 173." type="book chapter" year="1984">Quiros-Gonzalez and Baker (1984)</bibRefCitation>
,
<bibRefCitation id="EFBBE3954A194932FB50FE2B8C0BFE66" author="Lindquist E. E." box="[1196,1371,402,426]" pageId="12" pageNumber="110" pagination="3 - 28" refId="ref8609" refString="Lindquist E. E. 1985 - External anatomy - In: Helle W., Sabelis M. W. (Eds) Spider mites - Their Biology, Natural Enemies, and Control, Amsterdam: Elsevier Science BV. pp. 3 - 28." type="book chapter" year="1985">Lindquist (1985)</bibRefCitation>
,
<bibRefCitation id="EFBBE3954A194932FA99FE2B8BD0FE01" author="Beard J. J. &amp; Walter D. E." pageId="12" pageNumber="110" pagination="49 - 60" refId="ref7947" refString="Beard J. J., Walter D. E. 2005 - A new Tuckerella (Prostigmata: Tetranychoidea; Tuckerellidae) from Australia with description of all stages and a discussion of the tritonymphal stage - Acarologia, 45 (1): 49 - 60." type="journal article" year="2005">Beard and Walter (2005)</bibRefCitation>
and
<bibRefCitation id="EFBBE3954A194932FD45FE0C8A3BFE01" author="Beard J. J. &amp; Ochoa R. &amp; Childers C. C. &amp; Bauchan G. R. &amp; Shepard M." box="[697,875,437,461]" pageId="12" pageNumber="110" refId="ref7880" refString="Beard J. J., Ochoa R., Childers C. C., Bauchan G. R., Shepard M. 2013 - Travelling with tea Tuckerella: a ' s" type="book" year="2013">
Beard
<emphasis id="B95E42764A194932FD05FE0F8A7FFE01" box="[761,815,437,461]" italics="true" pageId="12" pageNumber="110">et al.</emphasis>
2013
</bibRefCitation>
, although there is some interspecific variation, the second tarsal solenidion is deutonymphal, tarsal seta
<emphasis id="B95E42764A194932FC29FE6E8A8CFE23" box="[981,988,471,495]" italics="true" pageId="12" pageNumber="110">l</emphasis>
<subScript id="17AE9C214A194932FC21FE5A8AB6FE38" attach="left" box="[989,998,483,500]" fontSize="7" pageId="12" pageNumber="110">1</subScript>
<emphasis id="B95E42764A194932FC14FE618AA1FE23" box="[1000,1009,472,495]" italics="true" pageId="12" pageNumber="110"></emphasis>
is tritonymphal, and three obovate setae on first genu and tibia are only present in deutonymphs: known protonymphs are found to have 12 obovate setae per segment, while tritonymphs and adults have five.
</paragraph>
<paragraph id="8B959E644A194932FDEFFDD88A8AFBB3" blockId="12.[491,1517,609,1151]" pageId="12" pageNumber="110">
Addition to the differential diagnosis — Certain caution is necessary when using this comparison, for the immature specimen is being compared to adults. Based on the available descriptions (
<bibRefCitation id="EFBBE3954A194932FD83FD1F8A8CFD72" author="Quiros-Gonzalez M. J. &amp; Baker E. W." box="[639,988,678,702]" pageId="12" pageNumber="110" pagination="166 - 173" refId="ref9065" refString="Quiros-Gonzalez M. J., Baker E. W. 1984 - Systematics and taxonomy of Acari. 5.5 Idiosomal and leg chaetotaxy in the Tuckerellidae Baker &amp; Pritchard; ontogeny and nomenclature - In: Griffiths D. A., Bowman C. E. (Eds) Acarology 6. Vol. 1., Chichester: Ellis Horwood Ltd. pp. 166 - 173." type="book chapter" year="1984">Quiros-Gonzalez and Baker 1984</bibRefCitation>
,
<bibRefCitation id="EFBBE3954A194932FC1AFD1F8DD3FD72" author="Lindquist E. E." box="[998,1155,678,702]" pageId="12" pageNumber="110" pagination="3 - 28" refId="ref8609" refString="Lindquist E. E. 1985 - External anatomy - In: Helle W., Sabelis M. W. (Eds) Spider mites - Their Biology, Natural Enemies, and Control, Amsterdam: Elsevier Science BV. pp. 3 - 28." type="book chapter" year="1985">Lindquist 1985</bibRefCitation>
,
<bibRefCitation id="EFBBE3954A194932FB71FD1F8CDFFD72" author="Beard J. J. &amp; Walter D. E." box="[1165,1423,678,702]" pageId="12" pageNumber="110" pagination="49 - 60" refId="ref7947" refString="Beard J. J., Walter D. E. 2005 - A new Tuckerella (Prostigmata: Tetranychoidea; Tuckerellidae) from Australia with description of all stages and a discussion of the tritonymphal stage - Acarologia, 45 (1): 49 - 60." type="journal article" year="2005">Beard and Walter 2005</bibRefCitation>
,
<bibRefCitation id="EFBBE3954A194932FA65FD1F8C8EFD72" author="Beard J. J. &amp; Ochoa R. &amp; Childers C. C. &amp; Bauchan G. R. &amp; Shepard M." box="[1433,1502,678,702]" pageId="12" pageNumber="110" refId="ref7880" refString="Beard J. J., Ochoa R., Childers C. C., Bauchan G. R., Shepard M. 2013 - Travelling with tea Tuckerella: a ' s" type="book" year="2013">Beard</bibRefCitation>
<emphasis id="B95E42764A194932FE17FD738B71FD2D" box="[491,545,713,737]" italics="true" pageId="12" pageNumber="110">et al.</emphasis>
2013), ontogenetic changes of the opisthonotal setation may affect setae
<emphasis id="B95E42764A194932FAD5FD738C65FD2D" box="[1321,1333,714,737]" italics="true" pageId="12" pageNumber="110">v</emphasis>
<subScript id="17AE9C214A194932FAC8FD6C8C6DFD2A" attach="left" box="[1332,1341,725,742]" fontSize="7" pageId="12" pageNumber="110">1</subScript>
(in
<taxonomicName id="4C2AE5E74A194932FA95FD738C89FD2D" authorityName="Beard and Walter" authorityYear="2005" box="[1385,1497,713,737]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="revelata">
<emphasis id="B95E42764A194932FA95FD738C89FD2D" box="[1385,1497,713,737]" italics="true" pageId="12" pageNumber="110">T. revelata</emphasis>
</taxonomicName>
they are obovate, with rough edge, in larva, with outline smoothing out in the subsequent stages), leaf-shaped C-H-row setae (relatively narrower and more tapered in the immatures) and flagellate H-row setae (becoming relatively shorter during ontogeny).
<taxonomicName id="4C2AE5E74A194932FAECFC8B8CDFFC85" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityYear="2014" box="[1296,1423,817,841]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A194932FAECFC8B8CDFFC85" box="[1296,1423,817,841]" italics="true" pageId="12" pageNumber="110">T. fossilibus</emphasis>
</taxonomicName>
is so far unique tuckerellid species known to have obovate setae
<emphasis id="B95E42764A194932FB8FFCEA8DD0FCA7" box="[1139,1152,851,875]" italics="true" pageId="12" pageNumber="110">h</emphasis>
<subScript id="17AE9C214A194932FB7DFCE68DDAFCBC" attach="left" box="[1153,1162,863,880]" fontSize="7" pageId="12" pageNumber="110">1</subScript>
similar in size to the lateral setae of C-, D- and E rows. By the shape of proterosomal setae
<emphasis id="B95E42764A194932FB62FCCE8DFAFC42" box="[1182,1194,887,910]" italics="true" pageId="12" pageNumber="110">v</emphasis>
<subScript id="17AE9C214A194932FB55FC3B8DE2FC5F" attach="left" box="[1193,1202,898,915]" fontSize="7" pageId="12" pageNumber="110">1</subScript>
and the relative shortness of setae
<emphasis id="B95E42764A194932FDB9FC218B02FC7C" box="[581,594,920,944]" italics="true" pageId="12" pageNumber="110">h</emphasis>
<subScript id="17AE9C214A194932FDAFFC1D8B0CFC79" attach="left" box="[595,604,932,949]" fontSize="7" pageId="12" pageNumber="110">1</subScript>
and
<emphasis id="B95E42764A194932FD6FFC218BF0FC7C" box="[659,672,920,944]" italics="true" pageId="12" pageNumber="110">h</emphasis>
<subScript id="17AE9C214A194932FD5CFC1D8BF9FC79" attach="left" box="[672,681,932,949]" fontSize="7" pageId="12" pageNumber="110">6</subScript>
, deutonymphal
<taxonomicName id="4C2AE5E74A194932FCA4FC208A87FC7C" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityYear="2014" box="[856,983,920,944]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A194932FCA4FC208A87FC7C" box="[856,983,920,944]" italics="true" pageId="12" pageNumber="110">T. fossilibus</emphasis>
</taxonomicName>
is similar to adults of
<taxonomicName id="4C2AE5E74A194932FB35FC208B77FC1F" authority="(Ewing, 1922)" baseAuthorityName="Ewing" baseAuthorityYear="1922" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="pavoniformis">
<emphasis id="B95E42764A194932FB35FC208C20FC7C" box="[1225,1392,920,944]" italics="true" pageId="12" pageNumber="110">T. pavoniformis</emphasis>
(
<bibRefCitation id="EFBBE3954A194932FA7CFC218B70FC1F" author="Ewing H. E." pageId="12" pageNumber="110" pagination="104 - 108" refId="ref8176" refString="Ewing H. E. 1922 - Three new species of peculiar and injurious spider mites - Proc. Entomol. Soc. Wash., 24 (4): 104 - 108." type="journal article" year="1922">Ewing, 1922</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C2AE5E74A194932FDC5FC058A79FC1F" authority="Prasad, 1973" authorityName="Prasad" authorityYear="1973" box="[569,809,955,979]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95E42764A194932FDC5FC058BC9FC1F" box="[569,665,955,979]" italics="true" pageId="12" pageNumber="110">T. indica</emphasis>
<bibRefCitation id="EFBBE3954A194932FD63FC028A79FC1F" author="Prasad V." box="[671,809,955,979]" pageId="12" pageNumber="110" pagination="237 - 239" refId="ref9036" refString="Prasad V. 1973 - A new species of Tuckerella (Acarina: Tuckerellidae) from India - Acarologia, 15 (2): 237 - 239." type="journal article" year="1973">Prasad, 1973</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2AE5E74A194932FC9CFC058C1FFC1F" authority="Malik and Kumar, 1992" authorityName="Mallik and Kumar" authorityYear="1992" box="[864,1359,955,979]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="channabasavannai">
<emphasis id="B95E42764A194932FC9CFC058D15FC1F" box="[864,1093,955,979]" italics="true" pageId="12" pageNumber="110">T. channabasavannai</emphasis>
Malik and Kumar, 1992
</taxonomicName>
. It differs from them by much larger setae
<emphasis id="B95E42764A194932FCBEFC648A1FFC39" box="[834,847,989,1013]" italics="true" pageId="12" pageNumber="110">h</emphasis>
<subScript id="17AE9C214A194932FCB3FC508A08FC36" attach="left" box="[847,856,1001,1018]" fontSize="7" pageId="12" pageNumber="110">1</subScript>
and by the differentiated shapes of the other H-row setae
<emphasis id="B95E42764A194932FA39FC678C8AFC39" box="[1477,1498,990,1013]" italics="true" pageId="12" pageNumber="110">T.</emphasis>
(
<taxonomicName id="4C2AE5E74A194932FE17FBB98B27FBD4" baseAuthorityName="Ewing" baseAuthorityYear="1922" box="[491,631,1024,1048]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="pavoniformis">
<emphasis id="B95E42764A194932FE17FBB98B27FBD4" box="[491,631,1024,1048]" italics="true" pageId="12" pageNumber="110">pavoniformis</emphasis>
</taxonomicName>
has setae
<emphasis id="B95E42764A194932FD1FFBB98BA0FBD4" box="[739,752,1024,1048]" italics="true" pageId="12" pageNumber="110">h</emphasis>
<subScript id="17AE9C214A194932FD0CFBB58BA9FBD1" attach="left" box="[752,761,1036,1053]" fontSize="7" pageId="12" pageNumber="110">2</subScript>
-
<emphasis id="B95E42764A194932FCF8FBB98A41FBD4" box="[772,785,1024,1048]" italics="true" pageId="12" pageNumber="110">h</emphasis>
<subScript id="17AE9C214A194932FCEDFBB58A4AFBD1" attach="left" box="[785,794,1036,1053]" fontSize="7" pageId="12" pageNumber="110">5</subScript>
,
<emphasis id="B95E42764A194932FCD5FBB98A66FBD4" box="[809,822,1024,1048]" italics="true" pageId="12" pageNumber="110">h</emphasis>
<subScript id="17AE9C214A194932FCCAFBB58A11FBD1" attach="left" box="[822,833,1036,1053]" fontSize="7" pageId="12" pageNumber="110">7</subScript>
and
<emphasis id="B95E42764A194932FC8FFBB98AD0FBD4" box="[883,896,1024,1048]" italics="true" pageId="12" pageNumber="110">h</emphasis>
<subScript id="17AE9C214A194932FC7DFBB58AD6FBD1" attach="left" box="[897,902,1036,1053]" fontSize="7" pageId="12" pageNumber="110">8</subScript>
all similar, whip-like (
<bibRefCitation id="EFBBE3954A194932FB7DFBB98CCFFBD4" author="Baker E. W. &amp; Pritchard A. E." box="[1153,1439,1024,1048]" pageId="12" pageNumber="110" refId="ref7775" refString="Baker E. W., Pritchard A. E. 1953 - The family categories of tetranychoid mites, with a review of" type="book" year="1953">Baker and Pritchard 1953</bibRefCitation>
), as do
<taxonomicName id="4C2AE5E74A194932FE17FB9A8B1BFBF6" authorityName="Prasad" authorityYear="1973" box="[491,587,1058,1082]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B95E42764A194932FE17FB9A8B1BFBF6" box="[491,587,1058,1082]" italics="true" pageId="12" pageNumber="110">T. indica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2AE5E74A194932FD81FB9A8A32FBF6" authorityName="Mallik and Kumar" authorityYear="1992" box="[637,866,1058,1082]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="channabasavannai">
<emphasis id="B95E42764A194932FD81FB9A8BDCFBF6" box="[637,652,1059,1082]" italics="true" pageId="12" pageNumber="110">T</emphasis>
.
<emphasis id="B95E42764A194932FD65FB9B8A32FBF6" box="[665,866,1058,1082]" italics="true" pageId="12" pageNumber="110">channabasavannai</emphasis>
</taxonomicName>
). By the shape of setae
<emphasis id="B95E42764A194932FB99FB9A8D21FBF6" box="[1125,1137,1059,1082]" italics="true" pageId="12" pageNumber="110">v</emphasis>
<subScript id="17AE9C214A194932FB8DFB978D2AFBF3" attach="left" box="[1137,1146,1070,1087]" fontSize="7" pageId="12" pageNumber="110">1</subScript>
<taxonomicName id="4C2AE5E74A194932FB7FFB9A8C51FBF6" authorityName="Khaustov, Sergeyenko &amp; Perkovsky" authorityYear="2014" box="[1155,1281,1058,1082]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="fossilibus">
<emphasis id="B95E42764A194932FB7FFB9A8C51FBF6" box="[1155,1281,1058,1082]" italics="true" pageId="12" pageNumber="110">T. fossilibus</emphasis>
</taxonomicName>
is also similar to
<emphasis id="B95E42764A194932FA3CFB9A8B18FB91" italics="true" pageId="12" pageNumber="110">
<taxonomicName id="4C2AE5E74A194932FA3CFB9A8C84FBF6" authority="Ehara, 1975" authorityName="Ehara" authorityYear="1975" box="[1472,1492,1059,1082]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="101" phylum="Arthropoda" rank="species" species="japonica">T.</taxonomicName>
japonica
</emphasis>
<bibRefCitation id="EFBBE3954A194932FDB3FBFC8B82FB91" author="Ehara S." box="[591,722,1093,1117]" pageId="12" pageNumber="110" pagination="1 - 5" refId="ref8136" refString="Ehara S. 1975 - Description of a new species of Tuckerella from Japan (Acarina: Tuckerellidae) - Int. J. Acarology, 1 (2): 1 - 5. doi: 10.1080 / 01647957508683741" type="journal article" year="1975">Ehara, 1975</bibRefCitation>
and to
<taxonomicName id="4C2AE5E74A194932FCE3FBFF8D17FB91" authority="Gupta, 1979" authorityName="Gupta" authorityYear="1979" box="[799,1095,1093,1117]" class="Arachnida" family="Tuckerellidae" genus="Tuckerella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="110" phylum="Arthropoda" rank="species" species="kumaonensis">
<emphasis id="B95E42764A194932FCE3FBFF8A92FB91" box="[799,962,1093,1117]" italics="true" pageId="12" pageNumber="110">T. kumaonensis</emphasis>
<bibRefCitation id="EFBBE3954A194932FC35FBFC8D17FB91" author="Gupta S. K." box="[969,1095,1093,1117]" pageId="12" pageNumber="110" refId="ref8254" refString="Gupta S. K. 1979 - Tuckerella kumaonensis, sp. nov. (Acarina: Tuckerellidae) from India - Orient." type="book" year="1979">Gupta, 1979</bibRefCitation>
</taxonomicName>
, but these species have three pairs of comparatively small leaf-like setae in
<emphasis id="B95E42764A194932FC7CFBDE8ADDFBB3" box="[896,909,1127,1151]" italics="true" pageId="12" pageNumber="110">h</emphasis>
-series.
</paragraph>
<paragraph id="8B959E644A194932FDEFFB158C81FAC6" blockId="12.[491,1526,1196,1290]" pageId="12" pageNumber="110">
<typeStatus id="549120C64A194932FDEFFB158B21FB08" box="[531,625,1196,1220]" pageId="12" pageNumber="110" type="holotype">Holotype</typeStatus>
container — Amber fragment with
<typeStatus id="549120C64A194932FBFEFB158D33FB08" box="[1026,1123,1196,1220]" pageId="12" pageNumber="110" type="holotype">holotype</typeStatus>
, sized 2x1x
<quantity id="4CD233814A194932FB18FB158C6AFB08" box="[1252,1338,1196,1220]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="110" unit="mm" value="0.5">0.5 mm</quantity>
, is mounted in the Buehler EpoThin epoxy resin between two
<quantity id="4CD233814A194932FC12FB768D72FB2B" box="[1006,1058,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="110" unit="cm" value="1.0">1 cm</quantity>
coverslips. The preparation is glued with a low-temperature hot-melt adhesive to a 1.5 mm-thick transparent plastic frame (
<figureCitation id="131182E14A194932FA94FB4B8C95FAC6" box="[1384,1477,1266,1290]" captionStart="Figure 5" captionStartId="9.[119,190,1903,1924]" captionTargetBox="[163,1433,233,1850]" captionTargetId="figure-2@9.[163,1433,233,1850]" captionTargetPageId="9" captionText="Figure 5 Tuckerella fossilibus Khaustov, Sergeyenko &amp; Perkovsky, 2014 holotype deutonymph. Layered DIC micrographs: A dorsal view, rectangles show positions of figures C, D, insert shows container with holotype; B ventral; C detail of idiosomal setae; D left leg I and anterior proterosoma dorsally. Abbreviations: st, cheliceral stylets; sty, stylophore; ta, palptarsus; ti.cl, palptibial claw." figureDoi="http://doi.org/10.5281/zenodo.4487591" httpUri="https://zenodo.org/record/4487591/files/figure.png" pageId="12" pageNumber="110">Fig. 5A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>