388 lines
44 KiB
XML
388 lines
44 KiB
XML
<document id="93C3698417B17421D064AD09C26F7DC4" ID-CLB-Dataset="28431" ID-DOI="10.11646/zootaxa.4564.1.2" ID-GBIF-Dataset="d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588621" ID-ZooBank="27F70B5F-24DD-4C9E-BBA7-2B87476BD49C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552162070752" checkinUser="plazi" docAuthor="Gunter, Nicole L. & Weir, Thomas A." docDate="2019" docId="03832713FFBD8D68A3DAFED950FC906E" docLanguage="en" docName="zootaxa.4564.1.2.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus gubara Gunter & Weir 2019, new species" docType="treatment" docUuid="4FE7F1BF-2BE6-41A5-8BE5-F15613BE95A8" docUuidSource="ZooBank" docVersion="9" lastPageNumber="67" masterDocId="FFBA5F6BFFAB8D72A34DFFB65569910A" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group" masterLastPageNumber="80" masterPageNumber="41" pageNumber="63" updateTime="1698700752634" updateUser="ExternalLinkService">
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<mods:title id="6746D2940AF0D5EEB321E2E08AECF506">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
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<mods:namePart id="09EE6361E6D6CA93E6F050651FBB4B33">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="20380537055D38617A25347655287CA1">Weir, Thomas A.</mods:namePart>
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<mods:title id="B3A5D5E75C8AEEF696EE7260317D6C30">Zootaxa</mods:title>
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<mods:date id="B961D4B4CEE0B3FF2D895044F39544F7">2019</mods:date>
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<treatment id="03832713FFBD8D68A3DAFED950FC906E" ID-DOI="http://doi.org/10.5281/zenodo.5922808" ID-GBIF-Taxon="154366349" ID-Zenodo-Dep="5922808" LSID="urn:lsid:zoobank.org:act:4FE7F1BF-2BE6-41A5-8BE5-F15613BE95A8" httpUri="http://treatment.plazi.org/id/03832713FFBD8D68A3DAFED950FC906E" lastPageId="26" lastPageNumber="67" pageId="22" pageNumber="63">
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<heading id="D0DD2169FFBD8D64A3DAFED957B19080" bold="true" box="[151,728,367,394]" fontSize="11" level="1" pageId="22" pageNumber="63" reason="1">
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<emphasis id="B95E4A17FFBD8D64A3DAFED957B19080" bold="true" box="[151,728,367,394]" pageId="22" pageNumber="63">
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<taxonomicName id="4C2AED86FFBD8D64A3DAFED9575D9080" ID-CoL="6P89W" authority="Gunter & Weir" authorityName="Gunter & Weir" authorityYear="2019" box="[151,564,367,394]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="gubara" status="sp. nov.">
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<emphasis id="B95E4A17FFBD8D64A3DAFED9540C9083" bold="true" box="[151,357,367,393]" italics="true" pageId="22" pageNumber="63">Lepanus gubara</emphasis>
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Gunter & Weir
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26DF76CFFBD8D64A10FFEC657B19080" box="[578,728,368,394]" pageId="22" pageNumber="63" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C330C58EFFBD8D64A3DAFE2454C690A1" box="[151,431,402,427]" pageId="22" pageNumber="63" type="description">
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<paragraph id="8B959605FFBD8D64A3DAFE2454C690A1" blockId="22.[151,728,367,427]" box="[151,431,402,427]" pageId="22" pageNumber="63">
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(
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<figureCitation id="13118A80FFBD8D64A3D2FE24546090A1" box="[159,265,402,427]" captionStart="FIGURE 5" captionStartId="15.[151,250,1945,1967]" captionTargetBox="[245,1336,223,1911]" captionTargetId="figure@15.[233,1355,193,1926]" captionTargetPageId="15" captionText="FIGURE 5. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588631/files/figure.png" pageId="22" pageNumber="63">Figs. 5D</figureCitation>
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,
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<figureCitation id="13118A80FFBD8D64A258FE25545290A1" box="[277,315,403,427]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="22" pageNumber="63">8D</figureCitation>
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,
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<figureCitation id="13118A80FFBD8D64A20AFE25541590A1" box="[327,380,403,427]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="22" pageNumber="63">10D</figureCitation>
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,
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<figureCitation id="13118A80FFBD8D64A2C5FE2554CF90A1" box="[392,422,403,427]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="22" pageNumber="63">14</figureCitation>
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)
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</paragraph>
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<emphasis id="B95E4A17FFBD8D64A3DAFE6C54C290F9" bold="true" box="[151,427,474,499]" pageId="22" pageNumber="63">
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Type series.
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<materialsCitation id="3B429C58FFBD8D64A27DFE6C54C290F9" ID-GBIF-Occurrence="2012745412" box="[304,427,474,499]" collectingDate="1993-12-14" collectingDateMax="1993-12-22" collectingDateMin="1993-12-14" collectionCode="AUST, NATURE, ANIC" county="Rainforest" latitude="-12.666667" location="Jabiru" longLatPrecision="21" longitude="133.13333" municipality="Gubarra Spring" pageId="22" pageNumber="63" specimenCode="CMNEN 00030233" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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:
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“AUST: NT: Kakadu NP, 25kmS
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,
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,
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</date>
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, FIT S&J Peck 93-127” [
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,
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] / “Canadian Museum of NATURE
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” / “ANIC Database 25-065549” / “
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<taxonomicName id="4C2AED86FFBD8D64A174FD9557F79331" authorityName=", Matthews" authorityYear="1974" box="[569,670,547,571]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
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NT8 det.
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<bibRefCitation id="EFBBEBF4FFBD8D64A07EFD9456859331" author="Gunter, N. L. & Monteith, G. B. & Cameron, S. L. & Weir, T." box="[819,1004,546,571]" pageId="22" pageNumber="63" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. & Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">T.A.Weir 2018</bibRefCitation>
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” in ANIC.
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:
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:
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,
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S Jabiru
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</locationDeviation>
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,
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<collectingMunicipality id="6BF10C7FFFBD8D64A00BFDF051609355" box="[838,1033,582,607]" pageId="22" pageNumber="63">Gubarra Spring</collectingMunicipality>
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, [
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,
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],
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<date id="FF94B0C5FFBD8D64A63DFDF154769389" pageId="22" pageNumber="63" value="1993-12-14" valueMax="1993-12-22" valueMin="1993-12-14">
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<collectingDate id="EFD0492DFFBD8D64A63DFDF154769389" pageId="22" pageNumber="63" value="1993-12-14" valueMax="1993-12-22" valueMin="1993-12-14">14– 22.xii.1993</collectingDate>
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</date>
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,
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<collectorName id="26DFF3D3FFBD8D64A263FDDD54899389" box="[302,480,618,643]" pageId="22" pageNumber="63">S.B. & J. Peck</collectorName>
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(
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<specimenCode id="DB8C3E7EFFBD8D64A2BCFDDD57B39389" box="[497,730,619,643]" collectionCode="CMNEN-" pageId="22" pageNumber="63">CMNEN-00030232</specimenCode>
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<specimenCount id="9D2C5D8CFFBD8D64A1AEFDDD57919388" box="[739,760,619,642]" pageId="22" pageNumber="63" type="female">♀</specimenCount>
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in
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<collectionCode id="ED3B0EC0FFBD8D64A06EFDDD56149389" box="[803,893,619,643]" pageId="22" pageNumber="63">CMNC</collectionCode>
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); The Amphitheatre,
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<location id="8EF5C0DEFFBD8D64A730FDDC50F19389" LSID="urn:lsid:plazi:treatment:03832713FFBD8D68A3DAFED950FC906E:8EF5C0DEFFBD8D64A730FDDC50F19389" box="[1149,1432,618,643]" county="Kakadu National Park" latitude="-14.15" longLatPrecision="21" longitude="132.38333" municipality="Gubarra Spring" name="Nitmiluk National Park" pageId="22" pageNumber="63" stateProvince="Northern Territory">Nitmiluk National Park</location>
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,
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<geoCoordinate id="EE1EF0C2FFBD8D64A3DAFD39547793AD" box="[151,286,655,679]" direction="south" orientation="latitude" pageId="22" pageNumber="63" precision="15" value="-14.15">14°09'00"S</geoCoordinate>
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,
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<geoCoordinate id="EE1EF0C2FFBD8D64A266FD3954AB93AD" box="[299,450,655,679]" direction="east" orientation="longitude" pageId="22" pageNumber="63" precision="15" value="132.38333">132°23'00"E</geoCoordinate>
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,
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<date id="FF94B0C5FFBD8D64A282FD38573893AD" box="[463,593,654,679]" pageId="22" pageNumber="63" value="1995-03-16">
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<collectingDate id="EFD0492DFFBD8D64A282FD38573893AD" box="[463,593,654,679]" pageId="22" pageNumber="63" value="1995-03-16">16.iii.1995</collectingDate>
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</date>
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,
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<collectorName id="26DFF3D3FFBD8D64A110FD39566D93AD" box="[605,772,654,679]" pageId="22" pageNumber="63">G.B. Monteith</collectorName>
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&
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||
<collectorName id="26DFF3D3FFBD8D64A064FD3956C493AD" box="[809,941,654,679]" pageId="22" pageNumber="63">H. Janetzki</collectorName>
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(T
|
||
<date id="FF94B0C5FFBD8D64A083FD38514F93AD" box="[974,1062,654,679]" pageId="22" pageNumber="63" value="52-11-19">
|
||
<collectingDate id="EFD0492DFFBD8D64A083FD38514F93AD" box="[974,1062,654,679]" pageId="22" pageNumber="63" value="52-11-19">111952</collectingDate>
|
||
</date>
|
||
<specimenCount id="9D2C5D8CFFBD8D64A763FD39512A93AC" box="[1070,1091,655,678]" pageId="22" pageNumber="63" type="male">♂</specimenCount>
|
||
in
|
||
<collectionCode id="ED3B0EC0FFBD8D64A724FD3951F393AD" box="[1129,1178,655,679]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="22" pageNumber="63" type="Museum">QM</collectionCode>
|
||
,
|
||
<specimenCode id="DB8C3E7EFFBD8D64A7EBFD38506793AD" box="[1190,1294,654,679]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="22" pageNumber="63" type="Museum">
|
||
T
|
||
<date id="FF94B0C5FFBD8D64A7F5FD38506793AD" box="[1208,1294,654,679]" pageId="22" pageNumber="63" value="50-11-19">
|
||
<quantity id="4CD23BE0FFBD8D64A7F5FD38506793AD" box="[1208,1294,654,679]" metricMagnitude="3" metricUnit="m" metricValue="2.8435299999999994" pageId="22" pageNumber="63" unit="in" value="111950.0">111950</quantity>
|
||
</date>
|
||
</specimenCode>
|
||
in
|
||
<collectionCode id="ED3B0EC0FFBD8D64A67EFD39500893AD" box="[1331,1377,655,679]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="22" pageNumber="63" type="Museum">QM</collectionCode>
|
||
<specimenCount id="9D2C5D8CFFBD8D64A624FD39501793AD" box="[1385,1406,655,679]" pageId="22" pageNumber="63" type="female">♀</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330C58EFFBD8D64A38AFD04579994B5" pageId="22" pageNumber="63" type="description">
|
||
<paragraph id="8B959605FFBD8D64A38AFD0456BE93C1" blockId="22.[151,1437,474,1759]" box="[199,983,690,715]" pageId="22" pageNumber="63">
|
||
<emphasis id="B95E4A17FFBD8D64A38AFD04543693C1" bold="true" box="[199,351,690,715]" pageId="22" pageNumber="63">Description.</emphasis>
|
||
Uniform dark brown in colour. Antennal clubs black.
|
||
</paragraph>
|
||
<paragraph id="8B959605FFBD8D64A38AFD60519893E4" blockId="22.[151,1437,474,1759]" box="[199,1265,726,751]" pageId="22" pageNumber="63">
|
||
Total length:
|
||
<quantity id="4CD23BE0FFBD8D64A229FD6154A893E4" box="[356,449,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="22" pageNumber="63" unit="mm" value="2.1">2.1 mm</quantity>
|
||
.
|
||
<typeStatus id="549128A7FFBD8D64A281FD60575093E5" box="[460,569,726,751]" pageId="22" pageNumber="63" type="holotype">Holotype</typeStatus>
|
||
measurements: Total length
|
||
<quantity id="4CD23BE0FFBD8D64A0C6FD61568193E4" box="[907,1000,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="22" pageNumber="63" unit="mm" value="2.1">2.1 mm</quantity>
|
||
, elytral width
|
||
<quantity id="4CD23BE0FFBD8D64A7DDFD61518793E4" box="[1168,1262,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="63" unit="mm" value="1.4">1.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B959605FFBD8D64A38AFD4C56579275" blockId="22.[151,1437,474,1759]" pageId="22" pageNumber="63">Male: Head: U-shaped between clypeal teeth, which are short and upturned. Margin slightly angulate at clypeogenal suture. Genal angle weakly defined. Basal carina present but interrupted medially. Dorsal part of eyes narrow in size, separated by interocular space approximately 15 times eye width (30: 2); eye canthus nearly dividing eye. Head 1.05 × wider than long (37:35).</paragraph>
|
||
<paragraph id="8B959605FFBD8D64A38AFC3C56EF92CD" blockId="22.[151,1437,474,1759]" pageId="22" pageNumber="63">Prothorax: Hypomeral surface finely reticulate with light punctation. Hypomeral striae at least half the length of hypomeron. Pronotum 1.55 × wider than long (58:37).</paragraph>
|
||
<paragraph id="8B959605FFBD8D64A38AFC6450FB9505" blockId="22.[151,1437,474,1759]" pageId="22" pageNumber="63">Elytra: Striae 1–5 narrow and double, with very fine punctate edges. All striae on disc not equal width, striae 6–7 are weaker and narrower; elytral flange and groove narrow. Elytra 1.15 × wider than long (70:60).</paragraph>
|
||
<paragraph id="8B959605FFBD8D64A38AFBAD54199571" blockId="22.[151,1437,474,1759]" pageId="22" pageNumber="63">Legs: Front edge of protibiae deeply angulate; apical digit somewhat elongate and sharp; 2 teeth on outer edge. Inner edge of metatibiae almost straight. Metatarsi with 2nd tarsomere subequal to the 5th. Tarsal claws small and toothed.</paragraph>
|
||
<paragraph id="8B959605FFBD8D64A38AFB3057DE9400" blockId="22.[151,1437,474,1759]" pageId="22" pageNumber="63">Abdomen: Pygidium with tear-drop shape depression containing an inverted V-shape tubercle, positioned in upper third of the depression. Depression occupies at least half size of disc and just extending under the virtually straight upper edge of pygidium; secondary basal line not evident. Abdominal surface smooth with reticulations right across. Segment 6 punctate.</paragraph>
|
||
<paragraph id="8B959605FFBD8D64A38AFAA054889491" blockId="22.[151,1437,474,1759]" pageId="22" pageNumber="63">Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Lateral lobe of metaventrite smooth and punctate with setae. Meso-metaventral suture very slightly arched. Mesoventrite with slight notch anteriorly. Mesoventrite smooth, virtually impunctate. Mesepimeron smooth. Metanepisternum reticulate.</paragraph>
|
||
<paragraph id="8B959605FFBD8D64A38AFA10579994B5" blockId="22.[151,1437,474,1759]" box="[199,752,1446,1471]" pageId="22" pageNumber="63">
|
||
Female: As per the
|
||
<taxonomicName id="4C2AED86FFBD8D64A2E1FA11572894B4" authorityName=": Matthews" authorityYear="1974" box="[428,577,1447,1470]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="pygmaeus">
|
||
<emphasis id="B95E4A17FFBD8D64A2E1FA1154D294B4" box="[428,443,1447,1470]" italics="true" pageId="22" pageNumber="63">L</emphasis>
|
||
.
|
||
<emphasis id="B95E4A17FFBD8D64A281FA11572894B4" box="[460,577,1447,1470]" italics="true" pageId="22" pageNumber="63">pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330C58EFFBD8D64A38AFA7C54989721" pageId="22" pageNumber="63" type="etymology">
|
||
<paragraph id="8B959605FFBD8D64A38AFA7C54989721" blockId="22.[151,1437,474,1759]" pageId="22" pageNumber="63">
|
||
<emphasis id="B95E4A17FFBD8D64A38AFA7C543A94E9" bold="true" box="[199,339,1482,1507]" pageId="22" pageNumber="63">Etymology.</emphasis>
|
||
Named for the type locality Gubara, Kakadu National Park. The locality label of the
|
||
<typeStatus id="549128A7FFBD8D64A67BFA7C50F594E9" box="[1334,1436,1482,1507]" pageId="22" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
lists “Gubarra Spring”, however the correct spelling for the locality is Gubara (
|
||
<bibRefCitation id="EFBBEBF4FFBD8D64A777FA58505C970D" author="Parks Australia" box="[1082,1333,1518,1543]" pageId="22" pageNumber="63" refId="ref29873" refString="Parks Australia. (2017) Kakadu National Park Visitor Guide. Available from: https: // parksaustralia. gov. au / kakadu / pub / visitorguide. pdf (accessed 15 June 2018)" type="url" year="2017">
|
||
Parks
|
||
<collectingCountry id="F33DD695FFBD8D64A7C9FA585199970D" box="[1156,1264,1518,1543]" name="Australia" pageId="22" pageNumber="63">Australia</collectingCountry>
|
||
2017
|
||
</bibRefCitation>
|
||
; Kakadu National Park Visitor Guide).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330C58EFFBD8D64A38AF980575097D5" pageId="22" pageNumber="63" type="distribution">
|
||
<paragraph id="8B959605FFBD8D64A38AF980575097D5" blockId="22.[151,1437,474,1759]" pageId="22" pageNumber="63">
|
||
<emphasis id="B95E4A17FFBD8D64A38AF980540E9745" bold="true" box="[199,359,1590,1615]" pageId="22" pageNumber="63">Distribution.</emphasis>
|
||
Known from only four specimens collected from Kakadu National Park and Nitmiluk National Park from the
|
||
<collectingRegion id="49EE58E7FFBD8D64A2E6F9EC57EF9779" box="[427,646,1626,1651]" country="Australia" name="Northern Territory" pageId="22" pageNumber="63">Northern Territory</collectingRegion>
|
||
, in the ARP bioregion (
|
||
<figureCitation id="13118A80FFBD8D64A0D6F9EC569A9779" box="[923,1011,1626,1651]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="22" pageNumber="63">Fig. 14</figureCitation>
|
||
). All specimens have been collected in rainforest or open forest. Given the limited collecting in the ARP bioregion and surrounding PCK, DAC, ARC, CEA, and GFU bioregions, and presence in both open and closed forest,
|
||
<taxonomicName id="4C2AED86FFBD8D64A723F912518F97B1" authorityName="Gunter & Weir" authorityYear="2019" box="[1134,1254,1698,1723]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="63" phylum="Arthropoda" rank="species" species="gubara">
|
||
<emphasis id="B95E4A17FFBD8D64A723F912518F97B1" box="[1134,1254,1698,1723]" italics="true" pageId="22" pageNumber="63">L. gubara</emphasis>
|
||
</taxonomicName>
|
||
is likely more widespread than currently reported.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF55C68DFFBC8D65A3DAF8C3567F96ED" httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="23" pageNumber="64" startId="23.[151,250,1909,1931]" targetBox="[167,1333,208,1895]" targetPageId="23">
|
||
<paragraph id="8B959605FFBC8D65A3DAF8C3567F96ED" blockId="23.[151,1437,1909,2023]" pageId="23" pageNumber="64">
|
||
<emphasis id="B95E4A17FFBC8D65A3DAF8C354759680" bold="true" box="[151,284,1909,1931]" pageId="23" pageNumber="64">FIGURE 10</emphasis>
|
||
. Left, dorsal, and right view of aedeagi of species within the
|
||
<taxonomicName id="4C2AED86FFBC8D65A0FBF8C051139681" authorityName=": Matthews" authorityYear="1974" box="[950,1146,1910,1931]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="pygmaeus">
|
||
<emphasis id="B95E4A17FFBC8D65A0FBF8C051139681" box="[950,1146,1910,1931]" italics="true" pageId="23" pageNumber="64">Lepanus pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B95E4A17FFBC8D65A669F8C3505F9680" bold="true" box="[1316,1334,1909,1930]" pageId="23" pageNumber="64">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFBC8D65A60BF8C0544296A0" authorityName="Gunter & Weir" authorityYear="2019" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="andersonorum">
|
||
<emphasis id="B95E4A17FFBC8D65A60BF8C0544296A0" italics="true" pageId="23" pageNumber="64">Lepanus andersonorum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFBC8D65A276F822542596A3" bold="true" box="[315,332,1940,1961]" pageId="23" pageNumber="64">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFBC8D65A216F823577596A0" authorityName="Gunter & Weir" authorityYear="2019" box="[347,540,1940,1962]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="crenidens">
|
||
<emphasis id="B95E4A17FFBC8D65A216F823577596A0" box="[347,540,1940,1962]" italics="true" pageId="23" pageNumber="64">Lepanus crenidens</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFBC8D65A161F822575796A0" bold="true" box="[556,574,1940,1962]" pageId="23" pageNumber="64">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFBC8D65A103F823579196A0" authorityName="Gunter & Weir" authorityYear="2019" box="[590,760,1940,1962]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="feehani">
|
||
<emphasis id="B95E4A17FFBC8D65A103F823579196A0" box="[590,760,1940,1962]" italics="true" pageId="23" pageNumber="64">Lepanus feehani</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFBC8D65A045F822567396A3" bold="true" box="[776,794,1940,1961]" pageId="23" pageNumber="64">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFBC8D65A067F82356BB96A0" authorityName="Gunter & Weir" authorityYear="2019" box="[810,978,1940,1962]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="gubara">
|
||
<emphasis id="B95E4A17FFBC8D65A067F82356BB96A0" box="[810,978,1940,1962]" italics="true" pageId="23" pageNumber="64">Lepanus gubara</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFBC8D65A0AFF822569A96A3" bold="true" box="[994,1011,1940,1961]" pageId="23" pageNumber="64">E</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFBC8D65A74EF82351AA96A0" authorityName="Gunter & Weir" authorityYear="2019" box="[1027,1219,1940,1962]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="64" phylum="Arthropoda" rank="species" species="guthrieae">
|
||
<emphasis id="B95E4A17FFBC8D65A74EF82351AA96A0" box="[1027,1219,1940,1962]" italics="true" pageId="23" pageNumber="64">Lepanus guthrieae</emphasis>
|
||
</taxonomicName>
|
||
. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF55C68DFFB38D6AA3DAF8C757C596E9" httpUri="https://zenodo.org/record/2588643/files/figure.png" pageId="24" pageNumber="65" startId="24.[151,250,1905,1927]" targetBox="[195,1359,208,1854]" targetPageId="24">
|
||
<paragraph id="8B959605FFB38D6AA3DAF8C757C596E9" blockId="24.[151,1436,1905,2019]" pageId="24" pageNumber="65">
|
||
<emphasis id="B95E4A17FFB38D6AA3DAF8C75471968C" bold="true" box="[151,280,1905,1927]" pageId="24" pageNumber="65">FIGURE 11</emphasis>
|
||
. Left, dorsal, and right view of aedeagi of species within the
|
||
<taxonomicName id="4C2AED86FFB38D6AA0CBF8C4512E968D" authorityName=": Matthews" authorityYear="1974" box="[902,1095,1906,1927]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="pygmaeus">
|
||
<emphasis id="B95E4A17FFB38D6AA0CBF8C4512E968D" box="[902,1095,1906,1927]" italics="true" pageId="24" pageNumber="65">Lepanus pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B95E4A17FFB38D6AA7A8F8C7519E968C" bold="true" box="[1253,1271,1905,1926]" pageId="24" pageNumber="65">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB38D6AA649F8C450FC968D" authorityName="Gunter & Weir" authorityYear="2019" box="[1284,1429,1905,1927]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="lentil">
|
||
<emphasis id="B95E4A17FFB38D6AA649F8C450FC968D" box="[1284,1429,1905,1927]" italics="true" pageId="24" pageNumber="65">Lepanus lentil</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFB38D6AA3DAF82655C196AF" bold="true" box="[151,168,1936,1957]" pageId="24" pageNumber="65">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB38D6AA3F8F827540A96AC" authorityName="Gunter & Weir" authorityYear="2019" box="[181,355,1936,1958]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="lingziae">
|
||
<emphasis id="B95E4A17FFB38D6AA3F8F827540A96AC" box="[181,355,1936,1958]" italics="true" pageId="24" pageNumber="65">Lepanus lingziae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFB38D6AA23CF82654EA96AC" bold="true" box="[369,387,1936,1958]" pageId="24" pageNumber="65">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB38D6AA2DCF827573A96AC" authorityName="Gunter & Weir" authorityYear="2019" box="[401,595,1936,1958]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="mckenziei">
|
||
<emphasis id="B95E4A17FFB38D6AA2DCF827573A96AC" box="[401,595,1936,1958]" italics="true" pageId="24" pageNumber="65">Lepanus mckenziei</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFB38D6AA12CF826571A96AF" bold="true" box="[609,627,1936,1957]" pageId="24" pageNumber="65">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB38D6AA1CCF827563396AC" authorityName="Gunter & Weir" authorityYear="2019" box="[641,858,1936,1958]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="menendezae">
|
||
<emphasis id="B95E4A17FFB38D6AA1CCF827563396AC" box="[641,858,1936,1958]" italics="true" pageId="24" pageNumber="65">Lepanus menendezae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFB38D6AA025F826561096AF" bold="true" box="[872,889,1936,1957]" pageId="24" pageNumber="65">E</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB38D6AA0CAF827517096AC" authorityName="Gunter & Weir" authorityYear="2019" box="[903,1049,1936,1958]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="pecki">
|
||
<emphasis id="B95E4A17FFB38D6AA0CAF827517096AC" box="[903,1049,1936,1958]" italics="true" pageId="24" pageNumber="65">Lepanus pecki</emphasis>
|
||
</taxonomicName>
|
||
. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF55C68DFFB28D6BA3DAF8C7578D96E9" httpUri="https://zenodo.org/record/2588645/files/figure.png" pageId="25" pageNumber="66" startId="25.[151,250,1905,1927]" targetBox="[231,1341,208,1744]" targetPageId="25">
|
||
<paragraph id="8B959605FFB28D6BA3DAF8C7578D96E9" blockId="25.[151,1436,1905,2019]" pageId="25" pageNumber="66">
|
||
<emphasis id="B95E4A17FFB28D6BA3DAF8C75475968C" bold="true" box="[151,284,1905,1927]" pageId="25" pageNumber="66">FIGURE 12</emphasis>
|
||
. Left, dorsal, and right view of aedeagi of species within the
|
||
<taxonomicName id="4C2AED86FFB28D6BA0FBF8C45113968D" authorityName=": Matthews" authorityYear="1974" box="[950,1146,1906,1927]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="66" phylum="Arthropoda" rank="species" species="pygmaeus">
|
||
<emphasis id="B95E4A17FFB28D6BA0FBF8C45113968D" box="[950,1146,1906,1927]" italics="true" pageId="25" pageNumber="66">Lepanus pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B95E4A17FFB28D6BA669F8C7505F968C" bold="true" box="[1316,1334,1905,1926]" pageId="25" pageNumber="66">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB28D6BA60BF8C4559196AC" authorityName="Gunter & Weir" authorityYear="2019" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="66" phylum="Arthropoda" rank="species" species="podocarp">
|
||
<emphasis id="B95E4A17FFB28D6BA60BF8C4559196AC" italics="true" pageId="25" pageNumber="66">Lepanus podocarp</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFB28D6BA248F826547F96AF" bold="true" box="[261,278,1936,1957]" pageId="25" pageNumber="66">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB28D6BA26FF827548D96AC" authorityName="Gunter & Weir" authorityYear="2019" box="[290,484,1936,1958]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="66" phylum="Arthropoda" rank="species" species="pungalina">
|
||
<emphasis id="B95E4A17FFB28D6BA26FF827548D96AC" box="[290,484,1936,1958]" italics="true" pageId="25" pageNumber="66">Lepanus pungalina</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFB28D6BA2BCF826576A96AC" bold="true" box="[497,515,1936,1958]" pageId="25" pageNumber="66">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB28D6BA142F82757B996AC" authorityName=": Matthews" authorityYear="1974" box="[527,720,1937,1958]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="66" phylum="Arthropoda" rank="species" species="pygmaeus">
|
||
<emphasis id="B95E4A17FFB28D6BA142F82757B996AC" box="[527,720,1937,1958]" italics="true" pageId="25" pageNumber="66">Lepanus pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFB28D6BA190F826578696AF" bold="true" box="[733,751,1936,1957]" pageId="25" pageNumber="66">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C2AED86FFB28D6BA1B1F82756CA96AC" authorityName="Gunter & Weir" authorityYear="2019" box="[764,931,1937,1958]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="66" phylum="Arthropoda" rank="species" species="sauroni">
|
||
<emphasis id="B95E4A17FFB28D6BA1B1F82756CA96AC" box="[764,931,1937,1958]" italics="true" pageId="25" pageNumber="66">Lepanus sauroni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B95E4A17FFB28D6BA0FCF82656AB96AF" bold="true" box="[945,962,1936,1957]" pageId="25" pageNumber="66">E</emphasis>
|
||
:
|
||
<taxonomicName id="4C2AED86FFB28D6BA083F82751E796AC" authorityName="Gunter & Weir" authorityYear="2019" box="[974,1166,1937,1958]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="66" phylum="Arthropoda" rank="species" species="tozerensis">
|
||
<emphasis id="B95E4A17FFB28D6BA083F82751E796AC" box="[974,1166,1937,1958]" italics="true" pageId="25" pageNumber="66">Lepanus tozerensis</emphasis>
|
||
</taxonomicName>
|
||
. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C330C58EFFB18D68A38AFF2150FC906E" pageId="26" pageNumber="67" type="discussion">
|
||
<paragraph id="8B959605FFB18D68A38AFF2150FC906E" blockId="26.[151,1437,151,357]" pageId="26" pageNumber="67">
|
||
<emphasis id="B95E4A17FFB18D68A38AFF21542691BA" bold="true" box="[199,335,151,176]" pageId="26" pageNumber="67">Comments.</emphasis>
|
||
<taxonomicName id="4C2AED86FFB18D68A216FF2F577191BA" authorityName="Gunter & Weir" authorityYear="2019" box="[347,536,151,176]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="species" species="gubara">
|
||
<emphasis id="B95E4A17FFB18D68A216FF2F577191BA" box="[347,536,151,176]" italics="true" pageId="26" pageNumber="67">Lepanus gubara</emphasis>
|
||
</taxonomicName>
|
||
was unknown at the time when the informal nomenclature for undescribed species of
|
||
<taxonomicName id="4C2AED86FFB18D68A240FF0B540591DE" authorityName=", Matthews" authorityYear="1974" box="[269,364,189,212]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E4A17FFB18D68A240FF0B540591DE" box="[269,364,189,212]" italics="true" pageId="26" pageNumber="67">Lepanus</emphasis>
|
||
</taxonomicName>
|
||
was reported (
|
||
<bibRefCitation id="EFBBEBF4FFB18D68A15CFF0A57B591DE" author="Yeates, D. & Gunter, N. L. & Weir, T. & Cameron, S. L." box="[529,732,188,213]" pageId="26" pageNumber="67" pagination="92 - 99" refId="ref30201" refString="Yeates, D. K, Gunter, N. L., Weir, T. & Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: accelerating research and discovery of Australia's biodiversity, Taxonomy Research and Identification Network Final Report- March, 2011, pp. 92 - 99. Available from: https: // www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 12 December 2018)" type="journal article" year="2011">
|
||
Yeates
|
||
<emphasis id="B95E4A17FFB18D68A129FF0B57F591DF" box="[612,668,188,213]" italics="true" pageId="26" pageNumber="67">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
) but was designated the informal name “
|
||
<taxonomicName id="4C2AED86FFB18D68A7EAFF0B506F91DE" authorityName=", Matthews" authorityYear="1974" box="[1191,1286,189,212]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95E4A17FFB18D68A7EAFF0B506F91DE" box="[1191,1286,189,212]" italics="true" pageId="26" pageNumber="67">Lepanus</emphasis>
|
||
</taxonomicName>
|
||
NT8” within this revision. The only known specimens were collected in flight-intercept traps and by pyrethrum. This species is superficially similar to
|
||
<taxonomicName id="4C2AED86FFB18D68A2E2FEB3572D9017" authorityName="Gunter & Weir" authorityYear="2019" box="[431,580,260,285]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="species" species="pungalina">
|
||
<emphasis id="B95E4A17FFB18D68A2E2FEB3572D9017" box="[431,580,260,285]" italics="true" pageId="26" pageNumber="67">L. pungalina</emphasis>
|
||
</taxonomicName>
|
||
but both males and females
|
||
<taxonomicName id="4C2AED86FFB18D68A0E0FEB3514B9017" authorityName="Gunter & Weir" authorityYear="2019" box="[941,1058,260,285]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="species" species="gubara">
|
||
<emphasis id="B95E4A17FFB18D68A0E0FEB3514B9017" box="[941,1058,260,285]" italics="true" pageId="26" pageNumber="67">L. gubara</emphasis>
|
||
</taxonomicName>
|
||
can diagnosed by the virtually impunctate mesoventrite, secondary basal line of pygidium not evident, absence of reticulation on the head between clypeogenal suture and apical edge, and punctation of the sixth abdominal ventrite (heavier in
|
||
<taxonomicName id="4C2AED86FFB18D68A65AFEFB50EC906F" authorityName="Gunter & Weir" authorityYear="2019" box="[1303,1413,332,357]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="species" species="gubara">
|
||
<emphasis id="B95E4A17FFB18D68A65AFEFB50EC906F" box="[1303,1413,332,357]" italics="true" pageId="26" pageNumber="67">L. gubara</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |