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<document id="E2648E3DDA31F96EC2B2F18CA770D226" ID-CLB-Dataset="28431" ID-DOI="10.11646/zootaxa.4564.1.2" ID-GBIF-Dataset="d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588621" ID-ZooBank="27F70B5F-24DD-4C9E-BBA7-2B87476BD49C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552162070752" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03832713FFB48D52A3DAFAF55019951A" docLanguage="en" docName="zootaxa.4564.1.2.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus pecki Gunter &amp; Weir 2019, new species" docType="treatment" docUuid="2A9D39E4-2808-40FC-B863-E4C849ED5A42" docUuidSource="ZooBank" docVersion="9" lastPageNumber="73" masterDocId="FFBA5F6BFFAB8D72A34DFFB65569910A" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group" masterLastPageNumber="80" masterPageNumber="41" pageNumber="72" updateTime="1698700752634" updateUser="ExternalLinkService">
<mods:mods id="5AA0B8DAC49F55102D454C8DFA74D6A8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E49C353BA48AB25B2C10714DF595B31D">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
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<mods:namePart id="CA44ADF3C7B8BCFC05D5A719BA19DF24">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="ABE6DDB3184E2F5152D3E059D31F4344">Weir, Thomas A.</mods:namePart>
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<mods:title id="E394878CB9B763AF49B89AC0227ED71D">Zootaxa</mods:title>
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<mods:date id="90022E2141F177A06A30C391FF5C8ED5">2019</mods:date>
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<mods:number id="9A2C84F89D509489EF079E9A9069D2EA">2019-03-05</mods:number>
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<mods:number id="A3138F2A9308EA6DC3B9EF906B3EA709">4564</mods:number>
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<mods:identifier id="49C416F54FE2A48B62458A91FE801B70" type="DOI">10.11646/zootaxa.4564.1.2</mods:identifier>
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<treatment id="03832713FFB48D52A3DAFAF55019951A" ID-DOI="http://doi.org/10.5281/zenodo.5922826" ID-GBIF-Taxon="154366353" ID-Zenodo-Dep="5922826" LSID="urn:lsid:zoobank.org:act:2A9D39E4-2808-40FC-B863-E4C849ED5A42" httpUri="http://treatment.plazi.org/id/03832713FFB48D52A3DAFAF55019951A" lastPageId="32" lastPageNumber="73" pageId="31" pageNumber="72">
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<heading id="D0DD2169FFB48D6DA3DAFAF557C09454" bold="true" box="[151,681,1347,1374]" fontSize="11" level="1" pageId="31" pageNumber="72" reason="1">
<emphasis id="B95E4A17FFB48D6DA3DAFAF557C09454" bold="true" box="[151,681,1347,1374]" pageId="31" pageNumber="72">
<taxonomicName id="4C2AED86FFB48D6DA3DAFAF557659454" ID-CoL="3T8VC" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,524,1347,1374]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="72" phylum="Arthropoda" rank="species" species="pecki" status="sp. nov.">
<emphasis id="B95E4A17FFB48D6DA3DAFAF5542C9457" bold="true" box="[151,325,1347,1373]" italics="true" pageId="31" pageNumber="72">Lepanus pecki</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFB48D6DA154FAF257C09454" box="[537,681,1348,1374]" pageId="31" pageNumber="72" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="8B959605FFB48D6DA3DAFAD057459475" blockId="31.[151,681,1347,1407]" box="[151,556,1382,1407]" pageId="31" pageNumber="72">
(
<figureCitation id="13118A80FFB48D6DA3D2FAD055859475" box="[159,236,1382,1407]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="31" pageNumber="72">Figs. 4</figureCitation>
D–E, 4I–J, 6D, 9A, 11E, 14)
</paragraph>
</subSubSection>
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<emphasis id="B95E4A17FFB48D6DA3DAFA1854C094CD" bold="true" box="[151,425,1454,1479]" pageId="31" pageNumber="72">
Type series.
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<typeStatus id="549128A7FFB48D6DA27EFA1854CD94CD" box="[307,420,1454,1479]" pageId="31" pageNumber="72" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D2C5D8CFFB48D6DA2F8FA1954A394CC" box="[437,458,1455,1478]" pageId="31" pageNumber="72" type="male">♂</specimenCount>
“AUST: NT; Darwin CSIRO, McMilliansRd
<date id="FF94B0C5FFB48D6DA0A0FA1851E794CD" box="[1005,1166,1454,1479]" pageId="31" pageNumber="72" value="1993-12-01" valueMax="1993-12-25" valueMin="1993-12-01">
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</date>
FIT S&amp;J Peck 93-87” [
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,
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] /“Canadian Museum of NATURE
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” / “ANIC Database 25-065551” / “
<taxonomicName id="4C2AED86FFB48D6DA3E2FA4154799705" authorityName=", Matthews" authorityYear="1974" box="[175,272,1527,1551]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
NT6 det.
<bibRefCitation id="EFBBEBF4FFB48D6DA2CCFA4057449705" author="Gunter, N. L. &amp; Monteith, G. B. &amp; Cameron, S. L. &amp; Weir, T." box="[385,557,1526,1551]" pageId="31" pageNumber="72" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. &amp; Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">T.A.Weir 2018</bibRefCitation>
” / “ANIC Image” in ANIC.
<materialsCitation id="3B429C58FFB48D6DA026FA40507B97ED" ID-GBIF-Occurrence="2012745371" collectingDate="1992-01-11" collectingDateMax="1994-01-10" collectingDateMin="1992-01-11" collectionCode="CSIRO" collectorName="S. B. &amp; J. Peck &amp; McMillans Road &amp; D. Bickel" county="Darwin" latitude="-12.633333" location="Byrne" longLatPrecision="1293" longitude="131.26666" municipality="McMillans Road" pageId="31" pageNumber="72" specimenCode="CMNEN-00030265-00030267, 2, CMNC 5, CMNEN-00030286-00030287, QM 1, CMNEN-00030271-79, ANIC 9, CMNEN-00030252-00030258, CMNC 1, CMNEN- 00030259-00030260, QM 2, CMNEN-00030262-00030263, ANIC 1, ANIC 25-060801, AM 1" specimenCount="13" specimenCount-female="9" specimenCount-male="4" stateProvince="Northern Territory" typeStatus="paratype">
<emphasis id="B95E4A17FFB48D6DA026FA4051889705" bold="true" box="[875,1249,1526,1551]" pageId="31" pageNumber="72">
<typeStatus id="549128A7FFB48D6DA026FA40568E9705" box="[875,999,1526,1551]" pageId="31" pageNumber="72" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="49EE58E7FFB48D6DA0BEFA4051B59705" box="[1011,1244,1526,1551]" country="Australia" name="Northern Territory" pageId="31" pageNumber="72">Northern Territory</collectingRegion>
:
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<collectingCounty id="62F4EE89FFB48D6DA7A5FA4050559705" box="[1256,1340,1526,1551]" pageId="31" pageNumber="72">Darwin</collectingCounty>
<collectionCode id="ED3B0EC0FFB48D6DA60FFA4150FE9705" box="[1346,1431,1527,1551]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="31" pageNumber="72" type="Museum">CSIRO</collectionCode>
,
<collectingMunicipality id="6BF10C7FFFB48D6DA3DAF9AC54309739" box="[151,345,1562,1587]" pageId="31" pageNumber="72">McMillans Road</collectingMunicipality>
, [
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,
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<collectingDate id="EFD0492DFFB48D6DA1EBF9AC562C9739" box="[678,837,1562,1587]" pageId="31" pageNumber="72" value="1993-12-01" valueMax="1993-12-25" valueMin="1993-12-01">1–25.xii.1993</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFB48D6DA01FF9AD56929739" box="[850,1019,1562,1587]" pageId="31" pageNumber="72">S.B. &amp; J. Peck</collectorName>
(
<specimenCode id="DB8C3E7EFFB48D6DA741F9AC50009739" box="[1036,1385,1562,1587]" collectionCode="CMNEN-" pageId="31" pageNumber="72">CMNEN-00030265-00030267</specimenCode>
,
<specimenCode id="DB8C3E7EFFB48D6DA638F9AD50EA9739" box="[1397,1411,1563,1587]" collectionCode="CMNEN-" pageId="31" pageNumber="72">2</specimenCode>
×
<specimenCount id="9D2C5D8CFFB48D6DA3DAF98955C5975C" box="[151,172,1599,1622]" pageId="31" pageNumber="72" type="female">♀</specimenCount>
, 1 ×
<specimenCount id="9D2C5D8CFFB48D6DA3A8F9895595975C" box="[229,252,1599,1622]" pageId="31" pageNumber="72" type="male">♂</specimenCount>
, 0 0 0 30269
<specimenCount id="9D2C5D8CFFB48D6DA233F98954FA975C" box="[382,403,1599,1622]" pageId="31" pageNumber="72" type="female">♀</specimenCount>
, 00030281–00030285 in
<specimenCode id="DB8C3E7EFFB48D6DA1FEF9895673975D" box="[691,794,1598,1623]" collectionCode="CMNC" pageId="31" pageNumber="72">CMNC 5</specimenCode>
×
<specimenCount id="9D2C5D8CFFB48D6DA07AF9895625975C" box="[823,844,1599,1622]" pageId="31" pageNumber="72" type="female">♀</specimenCount>
,
<specimenCode id="DB8C3E7EFFB48D6DA017F98851DE975D" box="[858,1207,1598,1623]" collectionCode="CMNEN-" pageId="31" pageNumber="72">CMNEN-00030286–00030287</specimenCode>
in
<specimenCode id="DB8C3E7EFFB48D6DA797F9895075975D" box="[1242,1308,1599,1623]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="31" pageNumber="72" type="Museum">QM 1</specimenCode>
×
<specimenCount id="9D2C5D8CFFB48D6DA674F9895027975C" box="[1337,1358,1599,1622]" pageId="31" pageNumber="72" type="male">♂</specimenCount>
1 ×
<specimenCount id="9D2C5D8CFFB48D6DA6CDF98950FC975C" box="[1408,1429,1599,1622]" pageId="31" pageNumber="72" type="female">♀</specimenCount>
,
<specimenCode id="DB8C3E7EFFB48D6DA3DAF9D554F39771" box="[151,410,1635,1659]" collectionCode="CMNEN-" pageId="31" pageNumber="72">
CMNEN-00030271-
<quantity id="4CD23BE0FFB48D6DA232F9D554F39771" box="[383,410,1635,1659]" metricMagnitude="0" metricUnit="m" metricValue="2.0065999999999997" pageId="31" pageNumber="72" unit="in" value="79.0">79</quantity>
</specimenCode>
in
<specimenCode id="DB8C3E7EFFB48D6DA298F9D557539771" box="[469,570,1635,1659]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="31" pageNumber="72">ANIC 9</specimenCode>
×
<specimenCount id="9D2C5D8CFFB48D6DA123F9D557EB9771" box="[622,642,1635,1659]" pageId="31" pageNumber="72" type="female">♀</specimenCount>
); Darwin
<collectionCode id="ED3B0EC0FFB48D6DA043F9D5560A9771" box="[782,867,1635,1659]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="31" pageNumber="72" type="Museum">CSIRO</collectionCode>
,
<collectorName id="26DFF3D3FFB48D6DA034F9D4512C9771" box="[889,1093,1634,1659]" pageId="31" pageNumber="72">McMillans Road</collectorName>
, [
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,
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],
<date id="FF94B0C5FFB48D6DA3DAF93054709795" box="[151,281,1670,1695]" pageId="31" pageNumber="72" value="1993-12-25">
<collectingDate id="EFD0492DFFB48D6DA3DAF93054709795" box="[151,281,1670,1695]" pageId="31" pageNumber="72" value="1993-12-25">25.xii.1993</collectingDate>
</date>
–
<date id="FF94B0C5FFB48D6DA265F93054F89795" box="[296,401,1670,1695]" pageId="31" pageNumber="72" value="1994-01-10">
<collectingDate id="EFD0492DFFB48D6DA265F93054F89795" box="[296,401,1670,1695]" pageId="31" pageNumber="72" value="1994-01-10">10.i.1994</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFB48D6DA2E9F93157359795" box="[420,604,1670,1695]" pageId="31" pageNumber="72">S.B. &amp; J. Peck</collectorName>
(
<specimenCode id="DB8C3E7EFFB48D6DA13EF93056B89795" box="[627,977,1670,1695]" collectionCode="CMNEN-" pageId="31" pageNumber="72">CMNEN-00030252–00030258</specimenCode>
in
<specimenCode id="DB8C3E7EFFB48D6DA74EF931511B9795" box="[1027,1138,1671,1695]" collectionCode="CMNC" pageId="31" pageNumber="72">CMNC 1</specimenCode>
×
<specimenCount id="9D2C5D8CFFB48D6DA7D2F93151DF9794" box="[1183,1206,1671,1694]" pageId="31" pageNumber="72" type="male">♂</specimenCount>
, 6 ×
<specimenCount id="9D2C5D8CFFB48D6DA64EF93150719794" box="[1283,1304,1671,1694]" pageId="31" pageNumber="72" type="female">♀</specimenCount>
,
<specimenCode id="DB8C3E7EFFB48D6DA663F93154EC97C9" collectionCode="CMNEN-" pageId="31" pageNumber="72">CMNEN- 00030259–00030260</specimenCode>
in
<specimenCode id="DB8C3E7EFFB48D6DA2E2F91D549D97C9" box="[431,500,1707,1731]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="31" pageNumber="72" type="Museum">QM 2</specimenCode>
×
<specimenCount id="9D2C5D8CFFB48D6DA155F91D574497C8" box="[536,557,1707,1730]" pageId="31" pageNumber="72" type="female">♀</specimenCount>
,
<specimenCode id="DB8C3E7EFFB48D6DA173F91C56F397C9" box="[574,922,1706,1731]" collectionCode="CMNEN-" pageId="31" pageNumber="72">CMNEN-00030262–00030263</specimenCode>
in
<specimenCode id="DB8C3E7EFFB48D6DA089F91D517797C9" box="[964,1054,1707,1731]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="31" pageNumber="72">ANIC 1</specimenCode>
♂,
<specimenCount id="9D2C5D8CFFB48D6DA70BF91D511B97C9" box="[1094,1138,1707,1731]" pageId="31" pageNumber="72" type="female">1 ♀</specimenCount>
);
<locationDeviation id="9A278F27FFB48D6DA7C2F91C50F197C8" box="[1167,1432,1706,1731]" location="Humpty Doo" pageId="31" pageNumber="72">
<quantity id="4CD23BE0FFB48D6DA7C2F91C51A297C9" box="[1167,1227,1706,1731]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="72" unit="km" value="5.0">5 km</quantity>
SE Humpty Doo
</locationDeviation>
,
<location id="8EF5C0DEFFB48D6DA3DAF97955B797ED" LSID="urn:lsid:plazi:treatment:03832713FFB48D52A3DAFAF55019951A:8EF5C0DEFFB48D6DA3DAF97955B797ED" box="[151,222,1743,1767]" county="Darwin" latitude="-12.633333" longLatPrecision="1293" longitude="131.26666" municipality="McMillans Road" name="Byrne" pageId="31" pageNumber="72" stateProvince="Northern Territory">Byrne</location>
, [
<geoCoordinate id="EE1EF0C2FFB48D6DA3BCF979542597ED" box="[241,332,1743,1767]" direction="south" orientation="latitude" pageId="31" pageNumber="72" precision="925" value="-12.633333">12°38'S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFB48D6DA215F97854AD97ED" box="[344,452,1742,1767]" direction="east" orientation="longitude" pageId="31" pageNumber="72" precision="925" value="131.26666">131°16'E</geoCoordinate>
],
<date id="FF94B0C5FFB48D6DA29AF978570597ED" box="[471,620,1742,1767]" pageId="31" pageNumber="72" value="1992-01-11" valueMax="1992-01-13" valueMin="1992-01-11">
<collectingDate id="EFD0492DFFB48D6DA29AF978570597ED" box="[471,620,1742,1767]" pageId="31" pageNumber="72" value="1992-01-11" valueMax="1992-01-13" valueMin="1992-01-11">11–13.i.1992</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFB48D6DA135F979578A97ED" box="[632,739,1742,1767]" pageId="31" pageNumber="72">D. Bickel</collectorName>
(K266973,
<specimenCode id="DB8C3E7EFFB48D6DA027F979514697ED" box="[874,1071,1742,1767]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="31" pageNumber="72">
ANIC 25-
<date id="FF94B0C5FFB48D6DA095F978514697ED" box="[984,1071,1742,1767]" pageId="31" pageNumber="72" value="01-06-08">060801</date>
</specimenCode>
in
<specimenCode id="DB8C3E7EFFB48D6DA71FF97951FD97ED" box="[1106,1172,1743,1767]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="31" pageNumber="72" type="Museum">AM 1</specimenCode>
×
<specimenCount id="9D2C5D8CFFB48D6DA7FFF97951A097EC" box="[1202,1225,1743,1766]" pageId="31" pageNumber="72" type="male">♂</specimenCount>
,
<date id="FF94B0C5FFB48D6DA798F979519C97ED" box="[1237,1269,1743,1767]" pageId="31" pageNumber="72">
<collectingDate id="EFD0492DFFB48D6DA798F979519C97ED" box="[1237,1269,1743,1767]" pageId="31" pageNumber="72">1x</collectingDate>
</date>
<specimenCount id="9D2C5D8CFFB48D6DA7B9F979506F97ED" box="[1268,1286,1743,1767]" pageId="31" pageNumber="72" type="female">♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFB48D52A38AF94457BE936A" lastPageId="32" lastPageNumber="73" pageId="31" pageNumber="72" type="description">
<paragraph id="8B959605FFB48D6DA38AF94457D49625" blockId="31.[151,1437,1454,2019]" pageId="31" pageNumber="72">
<emphasis id="B95E4A17FFB48D6DA38AF94454309601" bold="true" box="[199,345,1778,1803]" pageId="31" pageNumber="72">Description.</emphasis>
Either uniformly dark brown or bicoloured with head and pronotum dark brown and rest of the body light brown. Antennal clubs grey to yellow.
</paragraph>
<paragraph id="8B959605FFB48D6DA38AF88C519B9658" blockId="31.[151,1437,1454,2019]" box="[199,1266,1850,1875]" pageId="31" pageNumber="72">
Total length: 2.0–
<quantity id="4CD23BE0FFB48D6DA2C2F88D548F9658" box="[399,486,1851,1875]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="31" pageNumber="72" unit="mm" value="2.1">2.1 mm</quantity>
.
<typeStatus id="549128A7FFB48D6DA2BDF88C57319659" box="[496,600,1850,1875]" pageId="31" pageNumber="72" type="holotype">Holotype</typeStatus>
measurements: Total length 2.0 mm, elytral width
<quantity id="4CD23BE0FFB48D6DA7D8F88D51879658" box="[1173,1262,1851,1875]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="31" pageNumber="72" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
<paragraph id="8B959605FFB48D6DA38AF8E857B896E9" blockId="31.[151,1437,1454,2019]" pageId="31" pageNumber="72">Male: Head: U-shaped between clypeal teeth, which are short, blunt, and upturned. Margin slightly angulate at clypeogenal suture. Genal angle weakly defined. Basal carina present and interrupted medially. Dorsal part of eyes narrow in size, separated by interocular space approximately 15 times eye width (30:2); eye canthus nearly dividing eye. Head 1.15 × wider than long (37:32).</paragraph>
<paragraph id="8B959605FF8B8D52A38AFF21573491DF" blockId="32.[151,1437,151,1040]" pageId="32" pageNumber="73">Prothorax: Hypomeral surface finely reticulate. Hypomeral striae about half the length of hypomeron. Pronotum 1.7 × wider than long (60:35).</paragraph>
<paragraph id="8B959605FF8B8D52A38AFF6951BF91F2" blockId="32.[151,1437,151,1040]" box="[199,1238,223,248]" pageId="32" pageNumber="73">Elytra: Striae 1–3 narrow and double; striae 4–7 single. Elytra 1.3 × wider than long (78:60).</paragraph>
<paragraph id="8B959605FF8B8D52A38AFEB356E9906F" blockId="32.[151,1437,151,1040]" pageId="32" pageNumber="73">Legs: Three teeth on the outer edge. Front edge of protibiae deeply angulate; apical digit somewhat elongate and sharp. Inner edge of metatibiae expanded approximately midlength, creating a distinct flange. Metatarsi with 2nd tarsomere subequal to the 5th. Tarsal claws small and toothed.</paragraph>
<paragraph id="8B959605FF8B8D52A38AFED9516D90C5" blockId="32.[151,1437,151,1040]" pageId="32" pageNumber="73">Abdomen: Pygidium with almost ovoid depression with angulate upper edge and containing a round tubercle in upper third. Depression occupies at least half size of disc and just extending under the angulate upper edge of pygidium. Abdominal surface smooth and non-reticulate. Segment 6 punctate.</paragraph>
<paragraph id="8B959605FF8B8D52A38AFE6A51BE9337" blockId="32.[151,1437,151,1040]" pageId="32" pageNumber="73">Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Lateral lobe of metaventrite smooth and punctate with setae. Meso-metaventral suture very slightly arched. Mesoventrite with slight notch anteriorly. Mesoventrite and mesepimeron smooth. Metanepisternum reticulate.</paragraph>
<paragraph id="8B959605FF8B8D52A38AFDF157BE936A" blockId="32.[151,1437,151,1040]" box="[199,727,583,608]" pageId="32" pageNumber="73">
Female: As per the
<taxonomicName id="4C2AED86FF8B8D52A2EFFDFF5759936A" authorityName="in Matthews" authorityYear="1974" box="[418,560,584,608]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FF8B8D52A2EFFDFF54D8936A" box="[418,433,585,608]" italics="true" pageId="32" pageNumber="73">L</emphasis>
.
<emphasis id="B95E4A17FF8B8D52A28DFDFF5759936A" box="[448,560,585,608]" italics="true" pageId="32" pageNumber="73">pygmaeus</emphasis>
</taxonomicName>
species group.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8B8D52A38AFDDA55B093C7" pageId="32" pageNumber="73" type="etymology">
<paragraph id="8B959605FF8B8D52A38AFDDA55B093C7" blockId="32.[151,1437,151,1040]" pageId="32" pageNumber="73">
<emphasis id="B95E4A17FF8B8D52A38AFDDA5427938F" bold="true" box="[199,334,620,645]" pageId="32" pageNumber="73">Etymology.</emphasis>
Named in honour of Stewart Peck, Canadian Museum of Nature, Ottawa,
<collectingCountry id="F33DD695FF8B8D52A7D1FDDA519A938F" box="[1180,1267,620,645]" name="Canada" pageId="32" pageNumber="73">Canada</collectingCountry>
, who collected the majority of known specimens and many other
<taxonomicName id="4C2AED86FF8B8D52A191FD27565293A2" authorityName=", Matthews" authorityYear="1974" box="[732,827,657,680]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF8B8D52A191FD27565293A2" box="[732,827,657,680]" italics="true" pageId="32" pageNumber="73">Lepanus</emphasis>
</taxonomicName>
from the Northern Territory that are included in this study.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8B8D52A38AFD61574F9257" pageId="32" pageNumber="73" type="distribution">
<paragraph id="8B959605FF8B8D52A38AFD61574F9257" blockId="32.[151,1437,151,1040]" pageId="32" pageNumber="73">
<emphasis id="B95E4A17FF8B8D52A38AFD61540993FA" bold="true" box="[199,352,727,752]" pageId="32" pageNumber="73">Distribution.</emphasis>
Known only from mixed eucalypt woodland and in monsoon woodlands of Darwin and Humpty Doo in the
<collectingRegion id="49EE58E7FF8B8D52A26DFD4A549A921F" box="[288,499,764,789]" country="Australia" name="Northern Territory" pageId="32" pageNumber="73">Northern Territory</collectingRegion>
IBRA region DAC (
<figureCitation id="13118A80FF8B8D52A1A0FD4A562C921E" box="[749,837,764,789]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="32" pageNumber="73">Fig. 14</figureCitation>
). Very limited sampling of dung beetles have been conducted in DAC and the neighbouring VIB, DAB, PCK, ARP, and ARC IBRA regions and
<taxonomicName id="4C2AED86FF8B8D52A7E1FC97506D9232" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1196,1284,799,824]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B95E4A17FF8B8D52A7E1FC97506D9232" box="[1196,1284,799,824]" italics="true" pageId="32" pageNumber="73">L. pecki</emphasis>
</taxonomicName>
may be more widespread than currently reported.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8B8D52A38AFCD15019951A" pageId="32" pageNumber="73" type="discussion">
<paragraph id="8B959605FF8B8D52A38AFCD15019951A" blockId="32.[151,1437,151,1040]" pageId="32" pageNumber="73">
<emphasis id="B95E4A17FF8B8D52A38AFCD15426928A" bold="true" box="[199,335,871,896]" pageId="32" pageNumber="73">Comments.</emphasis>
<taxonomicName id="4C2AED86FF8B8D52A21BFCDF549E928A" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[342,503,871,896]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B95E4A17FF8B8D52A21BFCDF549E928A" box="[342,503,871,896]" italics="true" pageId="32" pageNumber="73">Lepanus pecki</emphasis>
</taxonomicName>
was unknown at the time when the informal nomenclature for undescribed species of
<taxonomicName id="4C2AED86FF8B8D52A3FAFC3B547F92AE" authorityName=", Matthews" authorityYear="1974" box="[183,278,909,932]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF8B8D52A3FAFC3B547F92AE" box="[183,278,909,932]" italics="true" pageId="32" pageNumber="73">Lepanus</emphasis>
</taxonomicName>
was reported (
<bibRefCitation id="EFBBEBF4FF8B8D52A28EFC3A57FD92AE" author="Yeates, D. &amp; Gunter, N. L. &amp; Weir, T. &amp; Cameron, S. L." box="[451,660,908,933]" pageId="32" pageNumber="73" pagination="92 - 99" refId="ref30201" refString="Yeates, D. K, Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: accelerating research and discovery of Australia's biodiversity, Taxonomy Research and Identification Network Final Report- March, 2011, pp. 92 - 99. Available from: https: // www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 12 December 2018)" type="journal article" year="2011">
Yeates
<emphasis id="B95E4A17FF8B8D52A155FC3B573A92AF" box="[536,595,908,933]" italics="true" pageId="32" pageNumber="73">et al.</emphasis>
2011
</bibRefCitation>
) but was designated the informal name “
<taxonomicName id="4C2AED86FF8B8D52A723FC3A51A692AE" authorityName=", Matthews" authorityYear="1974" box="[1134,1231,908,932]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
NT6” within this revision. This species has been collected in flight-intercept traps and pan traps. It is differentiated from all other
<taxonomicName id="4C2AED86FF8B8D52A3DAFC63559F92E6" authorityName=", Matthews" authorityYear="1974" box="[151,246,981,1004]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF8B8D52A3DAFC63559F92E6" box="[151,246,981,1004]" italics="true" pageId="32" pageNumber="73">Lepanus</emphasis>
</taxonomicName>
species (except
<emphasis id="B95E4A17FF8B8D52A2E3FC63572992E7" box="[430,576,980,1005]" italics="true" pageId="32" pageNumber="73">
<taxonomicName id="4C2AED86FF8B8D52A2E3FC63575192E7" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[430,568,980,1005]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="guthrieae">L. guthrieae</taxonomicName>
)
</emphasis>
by males having a distinctive flange on the metatibiae. Furthermore it is one of only two species in the
<taxonomicName id="4C2AED86FF8B8D52A2F0FC4F5723951A" authorityName="in Matthews" authorityYear="1974" box="[445,586,1016,1040]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FF8B8D52A2F0FC4F54A5951A" box="[445,460,1017,1040]" italics="true" pageId="32" pageNumber="73">L</emphasis>
.
<emphasis id="B95E4A17FF8B8D52A297FC4F5723951A" box="[474,586,1017,1040]" italics="true" pageId="32" pageNumber="73">pygmaeus</emphasis>
</taxonomicName>
species group that have three protibial teeth, the other being
<emphasis id="B95E4A17FF8B8D52A7BBFC4F5019951A" box="[1270,1392,1017,1040]" italics="true" pageId="32" pageNumber="73">
<taxonomicName id="4C2AED86FF8B8D52A7BBFC4F5005951A" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1270,1388,1017,1040]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="73" phylum="Arthropoda" rank="species" species="sauroni">L. sauroni</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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