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<document id="B8521BE36EEE70865B2AFBDCDDA92E20" ID-CLB-Dataset="8099" ID-DOI="10.5252/z2010n2a5" ID-GBIF-Dataset="f76e17a3-3bc3-43a8-ae33-6655aba3fa57" ID-ISSN="1638-9387" ID-Zenodo-Dep="4521008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612809505872" checkinUser="felipe" docAuthor="Peña Cantero, Álvaro L. &amp; Vervoort, Willem" docDate="2010" docId="03E5636AFF85FFBEFCFF54D475C6F92E" docLanguage="en" docName="Zoosystema.32.2.267-332.pdf" docOrigin="Zoosystema 32 (2)" docSource="http://www.bioone.org/doi/abs/10.5252/z2010n2a5" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Acryptolaria intermedia Peña Cantero &amp; Vervoort 2010, n. sp." docType="treatment" docVersion="8" lastPageNumber="293" masterDocId="FFDC1B12FF9CFFA4FFF35744764DFF8A" masterDocTitle="Species of Acryptolaria Norman, 1875 (Cnidaria, Hydrozoa, Lafoeidae) collected in the Western Pacific by various French expeditions, with the description of nineteen new species" masterLastPageNumber="332" masterPageNumber="267" pageNumber="292" updateTime="1698928392101" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="0391E45082ABC6DDAECDD10C125375CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="62923E8A34E03E96481229F4C3103F7E">
<mods:title id="B4FB2B74996DA2E54E525EC303DE1C9A">Species of Acryptolaria Norman, 1875 (Cnidaria, Hydrozoa, Lafoeidae) collected in the Western Pacific by various French expeditions, with the description of nineteen new species</mods:title>
</mods:titleInfo>
<mods:name id="3120ADE927FC663D8CF77FDC16D76986" type="personal">
<mods:role id="AC0C07094FCBBFC606F354DB52852FFE">
<mods:roleTerm id="AE3AB4F7502AB01435C7901D5796B701">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="039880C3504FFCD8C1544DA96030F70C">Peña Cantero, Álvaro L.</mods:namePart>
<mods:affiliation id="FC9AFE7DAA6CD33C38CD2EA3C05E8D86">Instituto Cavanilles de Biodiversidad y Biología Evolutiva, Universidad de Valencia / Fundación General, Universidad de Valencia, Apdo. Correos 22085, E- 46071 Valencia (Spain) alvaro. l. pena @ uv. es</mods:affiliation>
<mods:nameIdentifier id="C55F80C33F0129143888A4AD644D76A3" type="email">pena@uv.es</mods:nameIdentifier>
</mods:name>
<mods:name id="0F45A7376EE3252673904AC83659C919" type="personal">
<mods:role id="C85D5DB00B7BCF224B167BB3DC1EEB70">
<mods:roleTerm id="C39D581C2628081DF93EAFCA27073ECE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="03D1D56428CE549FD7EE8EE0EA1D0776">Vervoort, Willem</mods:namePart>
<mods:affiliation id="D298B9C10AFC5192F7780CA7A4056F72">National Museum of Natural History, P. O. Box 9517, NL- 2300 RA Leiden (The Netherlands) vervoort @ naturalis. nnm. nl</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EBF4DF50C838C5B10C5F58949170DC12">text</mods:typeOfResource>
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<mods:title id="A02D1C304B85E1BC7A784768BBD4675E">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="5C4750F4A55B5F1EFB3CFF1B2D518D36">
<mods:date id="3EB36EE7B7416C9948907816CABBE6F4">2010</mods:date>
<mods:detail id="6C1EA7843894283F26B4FDADEFC406DD" type="pubDate">
<mods:number id="8C9DFD059A46893CD8A7D99C84C55783">2010-06-30</mods:number>
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<mods:number id="999A064680D8414734E5CC11CA5512CE">32</mods:number>
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<mods:number id="B7A9066B0E5FC26ECE5E55942BDBB693">2</mods:number>
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<mods:start id="1AC7749C2163C7220B14B9DF20A6234C">267</mods:start>
<mods:end id="ADF1EAE7DC9DB2E40A45AD2F4A336883">332</mods:end>
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<mods:url id="37223DD275BF34F97F55B94CA5E163C7">http://www.bioone.org/doi/abs/10.5252/z2010n2a5</mods:url>
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<mods:classification id="576B96287CCD300FDCFAB00B80CE70E9">journal article</mods:classification>
<mods:identifier id="D3C8339A9679D0B69E4C21FE4E1401D1" type="CLB-Dataset">8099</mods:identifier>
<mods:identifier id="12A7D15D784C0999CD1BAF2CE6E05B8C" type="DOI">10.5252/z2010n2a5</mods:identifier>
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<treatment id="03E5636AFF85FFBEFCFF54D475C6F92E" ID-DOI="http://doi.org/10.5281/zenodo.4551878" ID-GBIF-Taxon="177429094" ID-Zenodo-Dep="4551878" LSID="urn:lsid:plazi:treatment:03E5636AFF85FFBEFCFF54D475C6F92E" httpUri="http://treatment.plazi.org/id/03E5636AFF85FFBEFCFF54D475C6F92E" lastPageId="26" lastPageNumber="293" pageId="25" pageNumber="292">
<subSubSection id="C35681F7FF85FFBDFCFF54D4721EFC26" box="[780,1107,912,940]" pageId="25" pageNumber="292" type="nomenclature">
<paragraph id="8BF3D27CFF85FFBDFCFF54D4721EFC26" blockId="25.[780,1107,912,940]" box="[780,1107,912,940]" pageId="25" pageNumber="292">
<heading id="D0BB6510FF85FFBDFCFF54D4721EFC26" box="[780,1107,912,940]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="292" reason="2">
<taxonomicName id="4C4CA9FFFF85FFBDFCFF54D4725CFC21" ID-CoL="9RX4" authority="Peña Cantero &amp; Vervoort, 2010" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[780,1041,912,939]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="292" phylum="Cnidaria" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B9380E6EFF85FFBDFCFF54D4725CFC21" bold="true" box="[780,1041,912,939]" italics="true" pageId="25" pageNumber="292">Acryptolaria intermedia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20BB315FF85FFBDFBEB54D5721EFC26" box="[1048,1107,913,940]" pageId="25" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35681F7FF85FFBDFCE854F4720EFC41" box="[795,1091,944,971]" pageId="25" pageNumber="292" type="description">
<paragraph id="8BF3D27CFF85FFBDFCE854F4720EFC41" blockId="25.[795,1091,944,971]" box="[795,1091,944,971]" pageId="25" pageNumber="292">
<heading id="D0BB6510FF85FFBDFCE854F4720EFC41" box="[795,1091,944,971]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="292" reason="2">
(
<figureCitation id="1377CEF9FF85FFBDFCD154F47539FC41" box="[802,884,944,971]" captionStart="FIG" captionStartId="26.[677,688,1404,1421]" captionTargetBox="[679,1208,669,1364]" captionTargetId="figure-17@26.[677,1210,666,1366]" captionTargetPageId="26" captionText="FIG. 12.— Acryptolaria intermedia n.sp., from CHALCAL 2 stn DW 76: A, B, branch fragments showing hydrothecal arrangement and hydrothecae; C, D, hydrothecae. Scale bar: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.4521042" httpUri="https://zenodo.org/record/4521042/files/figure.png" pageId="25" pageNumber="292">Figs 12</figureCitation>
;
<figureCitation id="1377CEF9FF85FFBDFC7354F475D3FC41" box="[896,926,944,971]" captionStart="FIG" captionStartId="59.[113,124,1540,1557]" captionTargetBox="[116,1194,153,1518]" captionTargetId="figure-17@59.[118,1191,203,1564]" captionTargetPageId="59" captionText="FIG. 30. — Outline drawings of the large type nematocysts of Acryptolaria species discussed in the present report. Scale bar: 30 μm." figureDoi="http://doi.org/10.5281/zenodo.4521094" httpUri="https://zenodo.org/record/4521094/files/figure.png" pageId="25" pageNumber="292">30</figureCitation>
;
<figureCitation id="1377CEF9FF85FFBDFC5954F4759CFC41" box="[938,977,944,971]" captionStart="FIG" captionStartId="60.[113,124,1567,1584]" captionTargetBox="[231,1092,215,1538]" captionTargetId="figure-27@60.[620,725,541,909]" captionTargetPageId="60" captionText="FIG. 31. — Colony shape and branching in Acryptolaria Norman, 1875: A, A. angulata (Bale, 1914), from MUSORSTOM 6 stn DW 487;B, A. bathyalis n. sp., holotype, MUSORSTOM 6 stn CP 419; C, A. bulbosa (Stechow, 1932), from MUSORSTOM 6 stn CP 419; D, A.disordinata n.sp.,holotype,MUSORSTOM 4 stn DW 197;E, A. encarnae n. sp., holotype,MUSORSTOM 6 stn CP 464;F, A.flabelloides n. sp., from SMIB 4 stn DW 36; G, A. gemini n. sp., holotype, GEMINI stn DW 60; H, A. infinita n. sp., holotype, MUSORSTOM 4 stn DW 197; I, A. intermedia n. sp., holotype, CHALCAL 2 stn DW 76; J, A. inversa n. sp., holotype, MUSORSTOM 6 stn DW 476; K, A. laertesi n. sp., holotype, MUSORSTOM 6 stn DW 425. Scale bars: A, C, F, G, I, 10 mm; B, D, E, H, J, K, 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4521098" httpUri="https://zenodo.org/record/4521098/files/figure.png" pageId="25" pageNumber="292">31I</figureCitation>
;
<tableCitation id="C6CEE7C7FF85FFBDFC2F54F47277FC41" box="[988,1082,944,971]" captionStart="TABLE" captionStartId="26.[303,314,219,236]" captionText="TABLE 13. — Measurements of Acryptolaria intermedia n. sp. from the holotype (in μm)." pageId="25" pageNumber="292">Table 13</tableCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35681F7FF85FFBDFD5654AE7205FBE1" pageId="25" pageNumber="292" type="materials_examined">
<paragraph id="8BF3D27CFF85FFBDFD5654AE7205FBE1" blockId="25.[675,1210,1001,1131]" pageId="25" pageNumber="292">
<materialsCitation id="3B24D821FF85FFBDFD5654AE7239FBDB" ID-GBIF-Occurrence="3038494336" collectingDate="1986-10-30" collectionCode="TYPE, MATERIAL" elevation="470" latitude="-23.675" longLatPrecision="12" longitude="167.75333" pageId="25" pageNumber="292" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED5D4AB9FF85FFBDFD5654AE7574FB8A" box="[677,825,1002,1026]" pageId="25" pageNumber="292">TYPE MATERIAL</collectionCode>
. —
<emphasis id="B9380E6EFF85FFBDFC9B54AD724DFB8B" bold="true" box="[872,1024,1001,1025]" pageId="25" pageNumber="292">Norfolk Ridge.</emphasis>
CHALCAL 2, stn DW 76,
<geoCoordinate id="EE78B4BBFF85FFBDFD0F53407526FB96" box="[764,875,1028,1052]" degrees="23" direction="south" minutes="40.50" orientation="latitude" pageId="25" pageNumber="292" precision="9" value="-23.675">23°40.50’S</geoCoordinate>
,
<geoCoordinate id="EE78B4BBFF85FFBDFC86534075BEFB96" box="[885,1011,1028,1052]" degrees="167" direction="east" minutes="45.20" orientation="longitude" pageId="25" pageNumber="292" precision="9" value="167.75333">167°45.20’E</geoCoordinate>
,
<quantity id="4CB47F99FF85FFBDFC0E5340720DFB96" box="[1021,1088,1028,1052]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="25" pageNumber="292" unit="m" value="470.0">
<elevation id="0061354FFF85FFBDFC0E5340720DFB96" box="[1021,1088,1028,1052]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="25" pageNumber="292" unit="m" value="470.0">470 m</elevation>
</quantity>
,
<date id="FFF2F4BCFF85FFBDFBB9534072FBFB96" box="[1098,1206,1028,1052]" pageId="25" pageNumber="292" value="1986-10-30">
<collectingDate id="EFB60D54FF85FFBDFBB9534072FBFB96" box="[1098,1206,1028,1052]" pageId="25" pageNumber="292" value="1986-10-30">30.X.1986</collectingDate>
</date>
, several stems, up to
<quantity id="4CB47F99FF85FFBDFCAB535A75D1FBBC" box="[856,924,1054,1078]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="292" unit="mm" value="24.0">24 mm</quantity>
high, from communal stolonal fibers: 1 stem
<typeStatus id="54F76CDEFF85FFBDFCD7537D753AFBDB" box="[804,887,1081,1105]" pageId="25" pageNumber="292" type="holotype">holotype</typeStatus>
(MNHN-Hy.2009-0163)
</materialsCitation>
; several stems
<typeStatus id="54F76CDEFF85FFBDFD2C5317757FFBE1" box="[735,818,1107,1131]" pageId="25" pageNumber="292" type="paratype">paratype</typeStatus>
(RMNH-Coel. no. 31510).
</paragraph>
</subSubSection>
<subSubSection id="C35681F7FF85FFBDFD5653C772C0FB5A" pageId="25" pageNumber="292" type="etymology">
<paragraph id="8BF3D27CFF85FFBDFD5653C772C0FB5A" blockId="25.[675,1210,1155,1232]" pageId="25" pageNumber="292">
ETYMOLOGY. — The specific name “
<taxonomicName id="4C4CA9FFFF85FFBDFC0853C77212FB11" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[1019,1119,1155,1179]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="292" phylum="Cnidaria" rank="species" species="intermedia">
<emphasis id="B9380E6EFF85FFBDFC0853C77212FB11" box="[1019,1119,1155,1179]" italics="true" pageId="25" pageNumber="292">intermedia</emphasis>
</taxonomicName>
” refers to the presence of intermediate features between
<taxonomicName id="4C4CA9FFFF85FFBDFBA353DA72F4FB3F" baseAuthorityName="Bale" baseAuthorityYear="1914" box="[1104,1209,1181,1205]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="292" phylum="Cnidaria" rank="species" species="angulata">
<emphasis id="B9380E6EFF85FFBDFBA353DA72F4FB3F" box="[1104,1209,1181,1205]" italics="true" pageId="25" pageNumber="292">A. angulata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4CA9FFFF85FFBDFD3C53FC757FFB5A" baseAuthorityName="Stechow" baseAuthorityYear="1932" box="[719,818,1208,1232]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="292" phylum="Cnidaria" rank="species" species="bulbosa">
<emphasis id="B9380E6EFF85FFBDFD3C53FC757FFB5A" box="[719,818,1208,1232]" italics="true" pageId="25" pageNumber="292">A. bulbosa</emphasis>
</taxonomicName>
; it is an adjective in femine gender.
</paragraph>
</subSubSection>
<subSubSection id="C35681F7FF85FFBDFD5653AC755FFABE" pageId="25" pageNumber="292" type="biology_ecology">
<paragraph id="8BF3D27CFF85FFBDFD5653AC755FFABE" blockId="25.[677,1209,1255,1332]" pageId="25" pageNumber="292">
ECOLOGY AND DISTRIBUTION. —
<taxonomicName id="4C4CA9FFFF85FFBDFC1353A372F4FB75" authority="Peña Cantero &amp; Vervoort, 2010" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[992,1209,1255,1279]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="25" pageNumber="292" phylum="Cnidaria" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B9380E6EFF85FFBDFC1353A372F4FB75" box="[992,1209,1255,1279]" italics="true" pageId="25" pageNumber="292">Acryptolaria intermedia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20BB315FF85FFBDFD5652467497FA90" box="[677,730,1282,1306]" pageId="25" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
was collected at a depth of
<quantity id="4CB47F99FF85FFBDFC155246726BFA90" box="[998,1062,1282,1306]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="25" pageNumber="292" unit="m" value="470.0">470 m</quantity>
in the Norfolk Ridge area.
</paragraph>
</subSubSection>
<subSubSection id="C35681F7FF85FFBEFD56522877C5FB23" lastPageId="26" lastPageNumber="293" pageId="25" pageNumber="292" type="description">
<paragraph id="8BF3D27CFF85FFBDFD5652287578FA0F" blockId="25.[675,1212,1388,1700]" box="[677,821,1388,1415]" pageId="25" pageNumber="292">DESCRIPTION</paragraph>
<paragraph id="8BF3D27CFF85FFBDFD5652C87525F9AF" blockId="25.[675,1212,1388,1700]" pageId="25" pageNumber="292">
Stems up to
<quantity id="4CB47F99FF85FFBDFCC652C875CAFA2D" box="[821,903,1420,1447]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="292" unit="mm" value="24.0">24 mm</quantity>
high (
<figureCitation id="1377CEF9FF85FFBDFC2052C8727CFA2D" box="[979,1073,1420,1447]" captionStart="FIG" captionStartId="60.[113,124,1567,1584]" captionTargetBox="[231,1092,215,1538]" captionTargetId="figure-27@60.[620,725,541,909]" captionTargetPageId="60" captionText="FIG. 31. — Colony shape and branching in Acryptolaria Norman, 1875: A, A. angulata (Bale, 1914), from MUSORSTOM 6 stn DW 487;B, A. bathyalis n. sp., holotype, MUSORSTOM 6 stn CP 419; C, A. bulbosa (Stechow, 1932), from MUSORSTOM 6 stn CP 419; D, A.disordinata n.sp.,holotype,MUSORSTOM 4 stn DW 197;E, A. encarnae n. sp., holotype,MUSORSTOM 6 stn CP 464;F, A.flabelloides n. sp., from SMIB 4 stn DW 36; G, A. gemini n. sp., holotype, GEMINI stn DW 60; H, A. infinita n. sp., holotype, MUSORSTOM 4 stn DW 197; I, A. intermedia n. sp., holotype, CHALCAL 2 stn DW 76; J, A. inversa n. sp., holotype, MUSORSTOM 6 stn DW 476; K, A. laertesi n. sp., holotype, MUSORSTOM 6 stn DW 425. Scale bars: A, C, F, G, I, 10 mm; B, D, E, H, J, K, 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4521098" httpUri="https://zenodo.org/record/4521098/files/figure.png" pageId="25" pageNumber="292">Fig. 31I</figureCitation>
). Branching irregular, but usually in one plane, and scarce, though branches up to third order are present. Branches straight or slightly sinuous (
<figureCitation id="1377CEF9FF85FFBDFBC152AF72E3F98C" box="[1074,1198,1515,1542]" captionStart="FIG" captionStartId="26.[677,688,1404,1421]" captionTargetBox="[679,1208,669,1364]" captionTargetId="figure-17@26.[677,1210,666,1366]" captionTargetPageId="26" captionText="FIG. 12.— Acryptolaria intermedia n.sp., from CHALCAL 2 stn DW 76: A, B, branch fragments showing hydrothecal arrangement and hydrothecae; C, D, hydrothecae. Scale bar: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.4521042" httpUri="https://zenodo.org/record/4521042/files/figure.png" pageId="25" pageNumber="292">Fig. 12A, B</figureCitation>
), with anastomoses.
</paragraph>
<paragraph id="8BF3D27CFF85FFBEFD48516E77AEFBC0" blockId="25.[675,1212,1388,1700]" lastBlockId="26.[111,648,659,1193]" lastPageId="26" lastPageNumber="293" pageId="25" pageNumber="292">
Hydrothecae alternately arranged in one plane (
<figureCitation id="1377CEF9FF85FFBDFD5D510E7562F9EF" box="[686,815,1610,1637]" captionStart="FIG" captionStartId="26.[677,688,1404,1421]" captionTargetBox="[679,1208,669,1364]" captionTargetId="figure-17@26.[677,1210,666,1366]" captionTargetPageId="26" captionText="FIG. 12.— Acryptolaria intermedia n.sp., from CHALCAL 2 stn DW 76: A, B, branch fragments showing hydrothecal arrangement and hydrothecae; C, D, hydrothecae. Scale bar: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.4521042" httpUri="https://zenodo.org/record/4521042/files/figure.png" pageId="25" pageNumber="292">Fig. 12A, B</figureCitation>
), tubular, roughly cylindrical, with a distinct decrease in diameter at their base (
<figureCitation id="1377CEF9FF85FFBDFBAC512D72E0F90E" box="[1119,1197,1641,1668]" captionStart="FIG" captionStartId="26.[677,688,1404,1421]" captionTargetBox="[679,1208,669,1364]" captionTargetId="figure-17@26.[677,1210,666,1366]" captionTargetPageId="26" captionText="FIG. 12.— Acryptolaria intermedia n.sp., from CHALCAL 2 stn DW 76: A, B, branch fragments showing hydrothecal arrangement and hydrothecae; C, D, hydrothecae. Scale bar: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.4521042" httpUri="https://zenodo.org/record/4521042/files/figure.png" pageId="25" pageNumber="292">Fig. 12</figureCitation>
). Hydrotheca curved twice: strongly outwards at distal part of adnate portion, becoming more or less perpendicular to adnate part, and markedly upwards consecutively, usually shortly after the adnate wall becomes free. Abcauline wall with a distinct inflexion point at about half its length, forming a shallow embayment at that point, accompanied by an internal perisarc cusp (
<figureCitation id="1377CEF9FF86FFBEFE8854157784FCE6" box="[379,457,849,876]" captionStart="FIG" captionStartId="26.[677,688,1404,1421]" captionTargetBox="[679,1208,669,1364]" captionTargetId="figure-17@26.[677,1210,666,1366]" captionTargetPageId="26" captionText="FIG. 12.— Acryptolaria intermedia n.sp., from CHALCAL 2 stn DW 76: A, B, branch fragments showing hydrothecal arrangement and hydrothecae; C, D, hydrothecae. Scale bar: 250 μm." figureDoi="http://doi.org/10.5281/zenodo.4521042" httpUri="https://zenodo.org/record/4521042/files/figure.png" pageId="26" pageNumber="293">Fig. 12</figureCitation>
). Basal part of abcauline wall straight, parallel to branch; distal part slightly convex or straight too. Adcauline wall adnate for over half of its length (adnate/free ratio
<emphasis id="B9380E6EFF86FFBEFDCE54F57403FC40" box="[573,590,945,970]" italics="true" pageId="26" pageNumber="293">c.</emphasis>
1.7); adnate part convex; free part concave proximally becoming straight distally. Hydrothecal aperture circular, oblique and slightly directed upwards; rim even, sometimes with renovations.
</paragraph>
<caption id="DF3382F4FF86FFBEFEDC579F75B3FF66" box="[303,1022,219,236]" pageId="26" pageNumber="293">
<paragraph id="8BF3D27CFF86FFBEFEDC579F75B3FF66" blockId="26.[303,1022,219,236]" box="[303,1022,219,236]" pageId="26" pageNumber="293">
TABLE 13. — Measurements of
<taxonomicName id="4C4CA9FFFF86FFBEFDDC579F74A3FF66" authority="Peña Cantero &amp; Vervoort, 2010" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[559,750,219,236]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B9380E6EFF86FFBEFDDC579F74A3FF66" box="[559,750,219,236]" italics="true" pageId="26" pageNumber="293">Acryptolaria intermedia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20BB315FF86FFBEFD07579F7569FF66" box="[756,804,219,236]" pageId="26" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
from the holotype (in μm).
</paragraph>
</caption>
<paragraph id="8BF3D27CFF86FFBEFD69566374A8FDC5" pageId="26" pageNumber="293">
<table id="F94C20DCFF86005BFF8256637222FDC5" box="[113,1135,295,591]" colsContinueIn="53.[113,1147,295,567]" gridcols="3" gridrows="12" pageId="26" pageNumber="293">
<tr id="357CD03EFF86005BFF8256637222FEB0" box="[113,1135,295,314]" gridrow="0" pageId="26" pageNumber="293" rowspan-0="1">
<th id="76ADB942FF86005BFD85566374B2FEB0" box="[630,767,295,314]" gridcol="1" gridrow="0" pageId="26" pageNumber="293">
<emphasis id="B9380E6EFF86FFBEFD6956637491FEB0" bold="true" box="[666,732,295,314]" pageId="26" pageNumber="293">Range</emphasis>
</th>
<th id="76ADB942FF86005BFC5D56637222FEB0" box="[942,1135,295,314]" gridcol="2" gridrow="0" pageId="26" pageNumber="293">
<emphasis id="B9380E6EFF86FFBEFC5C56637223FEB0" bold="true" box="[943,1134,295,314]" pageId="26" pageNumber="293">Mean ± SD (n = 10)</emphasis>
</th>
</tr>
<tr id="357CD03EFF86005BFF8256017222FED3" box="[113,1135,325,345]" gridrow="1" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF8256017222FED3" box="[113,1135,325,345]" colspan="3" colspanRight="2" gridcol="0" gridrow="1" pageId="26" pageNumber="293">Hydrothecae</th>
</tr>
<tr id="357CD03EFF86005BFF8256267222FEFC" box="[113,1135,354,374]" gridrow="2" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF82562677F4FEFC" box="[113,441,354,374]" gridcol="0" gridrow="2" pageId="26" pageNumber="293">Length of abcauline wall</th>
<td id="76ADB942FF86005BFD85562674B2FEFC" box="[630,767,354,374]" gridcol="1" gridrow="2" pageId="26" pageNumber="293">490-600</td>
<td id="76ADB942FF86005BFC5D56267222FEFC" box="[942,1135,354,374]" gridcol="2" gridrow="2" pageId="26" pageNumber="293">547.5 ± 33.4</td>
</tr>
<tr id="357CD03EFF86005BFF82563E7222FE04" box="[113,1135,378,398]" gridrow="3" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF82563E77F4FE04" box="[113,441,378,398]" gridcol="0" gridrow="3" pageId="26" pageNumber="293">Length of adcauline wall</th>
<td id="76ADB942FF86005BFD85563E74B2FE04" box="[630,767,378,398]" gridcol="1" gridrow="3" pageId="26" pageNumber="293">550-690</td>
<td id="76ADB942FF86005BFC5D563E7222FE04" box="[942,1135,378,398]" gridcol="2" gridrow="3" pageId="26" pageNumber="293">621.5 ± 39.2</td>
</tr>
<tr id="357CD03EFF86005BFF8256D67222FE2C" box="[113,1135,402,422]" gridrow="4" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF8256D677F4FE2C" box="[113,441,402,422]" gridcol="0" gridrow="4" pageId="26" pageNumber="293">Length of free adcauline wall</th>
<td id="76ADB942FF86005BFD8556D674B2FE2C" box="[630,767,402,422]" gridcol="1" gridrow="4" pageId="26" pageNumber="293">190-270</td>
<td id="76ADB942FF86005BFC5D56D67222FE2C" box="[942,1135,402,422]" gridcol="2" gridrow="4" pageId="26" pageNumber="293">235.5 ± 24.1</td>
</tr>
<tr id="357CD03EFF86005BFF8256EF7222FE35" box="[113,1135,427,447]" gridrow="5" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF8256EF77F4FE35" box="[113,441,427,447]" gridcol="0" gridrow="5" pageId="26" pageNumber="293">Length of adnate adcauline wall</th>
<td id="76ADB942FF86005BFD8556EF74B2FE35" box="[630,767,427,447]" gridcol="1" gridrow="5" pageId="26" pageNumber="293">360-420</td>
<td id="76ADB942FF86005BFC5D56EF7222FE35" box="[942,1135,427,447]" gridcol="2" gridrow="5" pageId="26" pageNumber="293">386.0 ± 18.1</td>
</tr>
<tr id="357CD03EFF86005BFF8256877222FE5D" box="[113,1135,451,471]" gridrow="6" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF82568777F4FE5D" box="[113,441,451,471]" gridcol="0" gridrow="6" pageId="26" pageNumber="293">Ratio adnate/free adcauline wall</th>
<td id="76ADB942FF86005BFD85568774B2FE5D" box="[630,767,451,471]" gridcol="1" gridrow="6" pageId="26" pageNumber="293">1.4-1.9</td>
<td id="76ADB942FF86005BFC5D56877222FE5D" box="[942,1135,451,471]" gridcol="2" gridrow="6" pageId="26" pageNumber="293">1.7 ± 0.1</td>
</tr>
<tr id="357CD03EFF86005BFF82569F7222FE65" box="[113,1135,475,495]" gridrow="7" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF82569F77F4FE65" box="[113,441,475,495]" gridcol="0" gridrow="7" pageId="26" pageNumber="293">Diameter at aperture</th>
<td id="76ADB942FF86005BFD85569F74B2FE65" box="[630,767,475,495]" gridcol="1" gridrow="7" pageId="26" pageNumber="293">95-110</td>
<td id="76ADB942FF86005BFC5D569F7222FE65" box="[942,1135,475,495]" gridcol="2" gridrow="7" pageId="26" pageNumber="293">102.5 ± 4.0</td>
</tr>
<tr id="357CD03EFF86005BFF8256B77222FD8D" box="[113,1135,499,519]" gridrow="8" pageId="26" pageNumber="293" rowspan-1="1" rowspan-2="1">
<th id="76ADB942FF86005BFF8256B777F4FD8D" box="[113,441,499,519]" gridcol="0" gridrow="8" pageId="26" pageNumber="293">Nematocysts</th>
</tr>
<tr id="357CD03EFF86005BFF82554F7222FD95" box="[113,1135,523,543]" gridrow="9" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF82554F77F4FD95" box="[113,441,523,543]" gridcol="0" gridrow="9" pageId="26" pageNumber="293">Larger group</th>
<td id="76ADB942FF86005BFD85554F74B2FD95" box="[630,767,523,543]" gridcol="1" gridrow="9" pageId="26" pageNumber="293">8.5-10 × 6-6.5</td>
<td id="76ADB942FF86005BFC5D554F7222FD95" box="[942,1135,523,543]" gridcol="2" gridrow="9" pageId="26" pageNumber="293">9.1 ± 0.4 × 6.1 ± 0.2</td>
</tr>
<tr id="357CD03EFF86005BFF8255677222FDBD" box="[113,1135,547,567]" gridrow="10" pageId="26" pageNumber="293">
<th id="76ADB942FF86005BFF82556777F4FDBD" box="[113,441,547,567]" gridcol="0" gridrow="10" pageId="26" pageNumber="293">Ratio</th>
<td id="76ADB942FF86005BFD85556774B2FDBD" box="[630,767,547,567]" gridcol="1" gridrow="10" pageId="26" pageNumber="293">1.4-1.5</td>
<td id="76ADB942FF86005BFC5D55677222FDBD" box="[942,1135,547,567]" gridcol="2" gridrow="10" pageId="26" pageNumber="293">1.48 ± 0.04</td>
</tr>
<tr id="357CD03EFF86005BFF82557F7222FDC5" box="[113,1135,571,591]" gridrow="11" pageId="26" pageNumber="293" rowspan-2="1">
<th id="76ADB942FF86005BFF82557F77F4FDC5" box="[113,441,571,591]" gridcol="0" gridrow="11" pageId="26" pageNumber="293">Smaller group</th>
<td id="76ADB942FF86005BFD85557F74B2FDC5" box="[630,767,571,591]" gridcol="1" gridrow="11" pageId="26" pageNumber="293">5.5 × 2.5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF3D27CFF86FFBEFF7B530B76B6FB00" blockId="26.[111,648,659,1193]" pageId="26" pageNumber="293">
Large nematocysts relatively very small and spherical (
<figureCitation id="1377CEF9FF86FFBEFF6D532B76A2FB00" box="[158,239,1135,1162]" captionStart="FIG" captionStartId="59.[113,124,1540,1557]" captionTargetBox="[116,1194,153,1518]" captionTargetId="figure-17@59.[118,1191,203,1564]" captionTargetPageId="59" captionText="FIG. 30. — Outline drawings of the large type nematocysts of Acryptolaria species discussed in the present report. Scale bar: 30 μm." figureDoi="http://doi.org/10.5281/zenodo.4521094" httpUri="https://zenodo.org/record/4521094/files/figure.png" pageId="26" pageNumber="293">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="8BF3D27CFF86FFBEFF7B53CA77C5FB23" blockId="26.[111,648,659,1193]" box="[136,392,1166,1193]" pageId="26" pageNumber="293">Coppinia not observed.</paragraph>
</subSubSection>
<subSubSection id="C35681F7FF86FFBEFF82538A75C6F92E" pageId="26" pageNumber="293" type="discussion">
<paragraph id="8BF3D27CFF86FFBEFF82538A7698FB6D" blockId="26.[111,647,1230,1700]" box="[113,213,1230,1257]" pageId="26" pageNumber="293">REMARKS</paragraph>
<paragraph id="8BF3D27CFF86FFBEFF8253A9741FF9EF" blockId="26.[111,647,1230,1700]" pageId="26" pageNumber="293">
At first glance,
<taxonomicName id="4C4CA9FFFF86FFBEFEF753A977A5FA8D" authority="Peña Cantero &amp; Vervoort, 2010" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[260,488,1261,1287]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B9380E6EFF86FFBEFEF753A977A5FA8D" box="[260,488,1261,1287]" italics="true" pageId="26" pageNumber="293">Acryptolaria intermedia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20BB315FF86FFBEFE1F53A9746FFA82" box="[492,546,1261,1288]" pageId="26" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="4C4CA9FFFF86FFBEFF9C52497696FAAD" baseAuthorityName="Stechow" baseAuthorityYear="1932" box="[111,219,1293,1319]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="bulbosa">
<emphasis id="B9380E6EFF86FFBEFF9C52497696FAAD" box="[111,219,1293,1319]" italics="true" pageId="26" pageNumber="293">A. bulbosa</emphasis>
</taxonomicName>
in the shape of the hydrothecae. Nevertheless they are clearly different species as the latter has bigger hydrothecae, much bigger nematocysts (20.7 ×
<quantity id="4CB47F99FF86FFBEFF3252287740FA0D" box="[193,269,1388,1415]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="26" pageNumber="293" unit="mm" value="6.7">6.7 µm</quantity>
) and the internal perisarc cusp at the abcauline hydrothecal embayment is absent. In the size of the nematocysts,
<taxonomicName id="4C4CA9FFFF86FFBEFE9A52E877B9FA4F" authority="Peña Cantero &amp; Vervoort, 2010" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[361,500,1451,1478]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B9380E6EFF86FFBEFE9A52E877B9FA4F" box="[361,500,1451,1478]" italics="true" pageId="26" pageNumber="293">A. intermedia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20BB315FF86FFBEFE0A52EF747CFA4C" box="[505,561,1451,1478]" pageId="26" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
is closer to
<taxonomicName id="4C4CA9FFFF86FFBEFF7C528F7744FA6F" baseAuthorityName="Bale" baseAuthorityYear="1914" box="[143,265,1483,1509]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="angulata">
<emphasis id="B9380E6EFF86FFBEFF7C528F7744FA6F" box="[143,265,1483,1509]" italics="true" pageId="26" pageNumber="293">A. angulata</emphasis>
</taxonomicName>
(15.9 ×
<quantity id="4CB47F99FF86FFBEFE94528F77F7FA6C" box="[359,442,1483,1510]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="26" pageNumber="293" unit="mm" value="8.6">8.6 µm</quantity>
), but they differ in the shape and size of the hydrotheca;
<taxonomicName id="4C4CA9FFFF86FFBEFE0A52AF74CAF98F" authority="Peña Cantero &amp; Vervoort, 2010" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[505,647,1515,1541]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B9380E6EFF86FFBEFE0A52AF74CAF98F" box="[505,647,1515,1541]" italics="true" pageId="26" pageNumber="293">A. intermedia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20BB315FF86FFBEFF82514E76E6F9AF" box="[113,171,1546,1573]" pageId="26" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
lacks the characteristic sharp invagination of the adnate adcauline wall of
<taxonomicName id="4C4CA9FFFF86FFBEFE5D516E7467F9CE" baseAuthorityName="Bale" baseAuthorityYear="1914" box="[430,554,1578,1604]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="angulata">
<emphasis id="B9380E6EFF86FFBEFE5D516E7467F9CE" box="[430,554,1578,1604]" italics="true" pageId="26" pageNumber="293">A. angulata</emphasis>
</taxonomicName>
and has distinctly smaller hydrothecae (cf.
<tableCitation id="C6CEE7C7FF86FFBEFE17510E7408F9EF" box="[484,581,1610,1637]" captionStart="TABLE" captionStartId="58.[113,124,220,237]" captionText="TABLE 31. — Comparative morphometric data (in μm) of the type material of the species of Acryptolaria Norman, 1875. Note: 1, from Jarvis’s (1922) figures." pageId="26" pageNumber="293">Table 31</tableCitation>
).
</paragraph>
<caption id="DF3382F4FF86FFBEFD565238721AFA31" ID-DOI="http://doi.org/10.5281/zenodo.4521042" ID-Zenodo-Dep="4521042" httpUri="https://zenodo.org/record/4521042/files/figure.png" pageId="26" pageNumber="293" startId="26.[677,688,1404,1421]" targetBox="[679,1208,669,1364]" targetPageId="26">
<paragraph id="8BF3D27CFF86FFBEFD565238721AFA31" blockId="26.[675,1209,1404,1467]" pageId="26" pageNumber="293">
FIG. 12.—
<taxonomicName id="4C4CA9FFFF86FFBEFD09523875F8FA07" authority="Peña Cantero &amp; Vervoort, 2010" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[762,949,1404,1421]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B9380E6EFF86FFBEFD09523875F8FA07" box="[762,949,1404,1421]" italics="true" pageId="26" pageNumber="293">Acryptolaria intermedia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20BB315FF86FFBEFC4A523875A7FA07" box="[953,1002,1404,1421]" pageId="26" pageNumber="293" rank="species">n.sp.</taxonomicNameLabel>
, from CHALCAL 2 stn DW 76:
<emphasis id="B9380E6EFF86FFBEFD3152D07482FA2F" bold="true" box="[706,719,1428,1445]" pageId="26" pageNumber="293">A</emphasis>
,
<emphasis id="B9380E6EFF86FFBEFD2452D074A9FA2F" bold="true" box="[727,740,1428,1445]" pageId="26" pageNumber="293">B</emphasis>
, branch fragments showing hydrothecal arrangement and hydrothecae;
<emphasis id="B9380E6EFF86FFBEFCE452EE7568FA31" bold="true" box="[791,805,1450,1467]" pageId="26" pageNumber="293">C</emphasis>
,
<emphasis id="B9380E6EFF86FFBEFCDC52EE7570FA31" bold="true" box="[815,829,1450,1467]" pageId="26" pageNumber="293">D</emphasis>
, hydrothecae. Scale bar: 250 μm.
</paragraph>
</caption>
<paragraph id="8BF3D27CFF86FFBEFF7B512D75C6F92E" blockId="26.[111,647,1230,1700]" lastBlockId="26.[675,1210,1546,1700]" pageId="26" pageNumber="293">
Above, when dealing with
<taxonomicName id="4C4CA9FFFF86FFBEFE43512E7452F909" baseAuthorityName="Stechow" baseAuthorityYear="1932" box="[432,543,1641,1668]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="bulbosa">
<emphasis id="B9380E6EFF86FFBEFE43512E7452F909" box="[432,543,1641,1668]" italics="true" pageId="26" pageNumber="293">A. bulbosa</emphasis>
</taxonomicName>
, we have indicated that material described either as
<emphasis id="B9380E6EFF86FFBEFDB251CD7546F9AE" box="[577,779,1546,1699]" italics="true" pageId="26" pageNumber="293">A. rectangularis</emphasis>
(cf. Gravier-Bonnet 1979;
<bibRefCitation id="EFDDAF8DFF86FFBEFBD5514E72F8F9AF" author="MILLARD N. A. H." box="[1062,1205,1546,1573]" pageId="26" pageNumber="293" pagination="168 - 194" refId="ref37636" refString="MILLARD N. A. H. 1967. - Hydroids from the south-west Indian Ocean. Annals of the South African Museum 50 (9): 168 - 194." type="journal article" year="1967">Millard 1967</bibRefCitation>
,
<bibRefCitation id="EFDDAF8DFF86FFBEFD50516E7491F9CF" author="MILLARD N. A. H." box="[675,732,1578,1605]" pageId="26" pageNumber="293" pagination="251 - 288" refId="ref37668" refString="MILLARD N. A. H. 1968. - South African hydroids from Dr Th. Mortensen's Java-South Africa expedition, 1929 - 1930. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 131: 251 - 288." type="journal article" year="1968">1968</bibRefCitation>
) or as
<taxonomicName id="4C4CA9FFFF86FFBEFCD6516E75D2F9CE" baseAuthorityName="Bale" baseAuthorityYear="1914" box="[805,927,1578,1604]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="angulata">
<emphasis id="B9380E6EFF86FFBEFCD6516E75D2F9CE" box="[805,927,1578,1604]" italics="true" pageId="26" pageNumber="293">A. angulata</emphasis>
</taxonomicName>
(cf. Hirohito 1995) could belong to
<taxonomicName id="4C4CA9FFFF86FFBEFCE2510E7530F9EE" baseAuthorityName="Stechow" baseAuthorityYear="1932" box="[785,893,1610,1636]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="bulbosa">
<emphasis id="B9380E6EFF86FFBEFCE2510E7530F9EE" box="[785,893,1610,1636]" italics="true" pageId="26" pageNumber="293">A. bulbosa</emphasis>
</taxonomicName>
or to
<taxonomicName id="4C4CA9FFFF86FFBEFC4D510E7200F9EE" authority="Peña Cantero &amp; Vervoort, 2010" authorityName="Peña Cantero &amp; Vervoort" authorityYear="2010" box="[958,1101,1610,1636]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="26" pageNumber="293" phylum="Cnidaria" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B9380E6EFF86FFBEFC4D510E7200F9EE" box="[958,1101,1610,1636]" italics="true" pageId="26" pageNumber="293">A. intermedia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20BB315FF86FFBEFBA7510E72C3F9EF" box="[1108,1166,1610,1637]" pageId="26" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
but that it is necessary to study the cnidome to properly identify this material.
</paragraph>
</subSubSection>
</treatment>
</document>
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