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<document id="B33FFC5C9162AF54CC6B3743A7B5FFE5" ID-ISSN="1536-9307" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638241281660" checkinUser="felipe" docAuthor="Ayrey, Richard F." docDate="2013" docId="03E087F74B61FF914C9CF93EFEA1F557" docLanguage="en" docName="Euscorpius.176.1-13.pdf" docOrigin="Euscorpius 176" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Vaejovis trinityae Ayrey 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="FFD9FF8F4B62FF984F39FF98FFA7F012" masterDocTitle="A new species of Vaejovis from the Mogollon Rim of northern Arizona (Scorpiones: Vaejovidae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="2" updateTime="1699438346649" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="E106696FDD004B7CA06445F1EA2B8DC5">A new species of Vaejovis from the Mogollon Rim of northern Arizona (Scorpiones: Vaejovidae)</mods:title>
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<mods:namePart id="A1053CF8C7844291D25F9B2F3FE2E7CC">Ayrey, Richard F.</mods:namePart>
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<mods:title id="E82C943555777EAFB7FDDF70B7D4EF9D">Euscorpius</mods:title>
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<mods:date id="B8A5DA716BD62EA52D48A519929755FB">2013</mods:date>
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<mods:number id="DCCE03D064511541937DCE241CDCD9D1">176</mods:number>
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<treatment id="03E087F74B61FF914C9CF93EFEA1F557" LSID="urn:lsid:zoobank.org:act:670320DB-F16C-4209-98AF-2773956248E4" httpUri="http://treatment.plazi.org/id/03E087F74B61FF914C9CF93EFEA1F557" lastPageId="9" lastPageNumber="7" pageId="3" pageNumber="2">
<subSubSection id="C353656A4B61FF9B4C9CF93EFA9EF6D2" box="[933,1337,1702,1728]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8BF636E14B61FF9B4C9CF93EFA9EF6D2" blockId="3.[839,1432,1702,1820]" box="[933,1337,1702,1728]" pageId="3" pageNumber="2">
<heading id="D0BE818D4B61FF9B4C9CF93EFA9EF6D2" box="[933,1337,1702,1728]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="2" reason="2">
<emphasis id="B93DEAF34B61FF9B4C9CF93EFA9EF6D2" bold="true" box="[933,1337,1702,1728]" pageId="3" pageNumber="2">
<taxonomicName id="4C494D624B61FF9B4C9CF93EFB6AF6D2" ID-CoL="7F8QD" authority="Ayrey" authorityName="Ayrey" authorityYear="2013" box="[933,1229,1702,1728]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="trinityae" status="sp. nov.">
<emphasis id="B93DEAF34B61FF9B4C9CF93EFBDFF6D2" bold="true" box="[933,1144,1702,1728]" italics="true" pageId="3" pageNumber="2">Vaejovis trinityae</emphasis>
Ayrey
</taxonomicName>
,
<taxonomicNameLabel id="A20E57884B61FF9B4BE3F93EFA9EF6D2" box="[1242,1337,1702,1728]" pageId="3" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C353656A4B61FF9B4CD5F95EFA99F70E" pageId="3" pageNumber="2" type="description">
<paragraph id="8BF636E14B61FF9B4CD5F95EFA3FF6EC" blockId="3.[839,1432,1702,1820]" pageId="3" pageNumber="2">
(
<figureCitation id="13722A644B61FF9B4CCDF95EFBCFF6CC" box="[1012,1128,1734,1758]" captionStart-0="Figures 12" captionStart-1="Figure 3" captionStart-2="Figures 49" captionStart-3="Figure 10" captionStart-4="Figure 11" captionStart-5="Figures 1213" captionStartId-0="4.[192,278,1853,1877]" captionStartId-1="5.[192,267,1518,1542]" captionStartId-2="6.[192,278,1472,1496]" captionStartId-3="7.[192,267,1798,1822]" captionStartId-4="8.[192,267,902,926]" captionStartId-5="10.[192,278,1846,1870]" captionTargetBox-0="[191,1440,168,1825]" captionTargetBox-1="[212,1424,188,1477]" captionTargetBox-2="[214,1427,204,1418]" captionTargetBox-3="[202,1389,199,1752]" captionTargetBox-4="[191,1441,167,870]" captionTargetBox-5="[191,1441,191,1814]" captionTargetId-0="figure-25@4.[192,1439,169,1824]" captionTargetId-1="figure-161@5.[192,1439,180,1488]" captionTargetId-2="figure-195@6.[192,1441,192,1441]" captionTargetId-3="figure-48@7.[192,1437,192,1768]" captionTargetId-4="figure-504@8.[192,1440,168,869]" captionTargetId-5="figure-29@10.[192,1440,192,1813]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="10" captionText-0="Figures 12: Vaejovis trinityae sp. nov. female and male paratypes in natural habitat." captionText-1="Figure 3: Vaejovis trinityae sp. nov. holotype female dorsal and ventral views." captionText-2="Figures 49: Vaejovis trinityae sp. nov. paratype male. 4. Carapace. 5. Dentition of fixed and movable fingers of chela. 6. Chelicera, dorsal view. 7. Telson, ventral and lateral views. 8. Metasomal segment V, lateral and ventral views. 9. Stigma IV, left side." captionText-3="Figure 10: Vaejovis trinityae, sp. nov., paratype male. Trichobothrial pattern." captionText-4="Figure 11: Right hemispermatophore and mating plug of Vaejovis trinityae, sp. nov., paratype male (photographed submerged in alcohol). Left. Dorsal view. Upper Left. Closeup of median area, dorsal and externodorsal views. Right. Ventral view. Upper Right. Closeup of median area, internoventral and ventral views. Bottom. Mating plug, dorsal and ventral views, showing smooth barb (in dorsal view). Note in all ventral views of the hemispermatophore the embedded matinig plug is visible, the barb positioned towards the internal edge. Due to the translucency of the lamina, the ventral trough edge is visible from the dorsal view and the bifurcated lamellar hook, which emanates from the dorsal trough edge, is visible from the ventral view." captionText-5="Figures 1213: Vaejovis trinityae sp. nov. 12 (top). Habitat. 13 (bottom). Paratype female with newborn." pageId="3" pageNumber="2">Figs. 113</figureCitation>
;
<tableCitation id="C6CB035A4B61FF9B4B4DF95EFB4CF6CC" box="[1140,1259,1734,1758]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="9.[192,257,1234,1258]" captionStartId-1="11.[192,257,1303,1327]" captionText-0="Table 1: Morphometrics (mm) of Vaejovis trinityae, sp. nov." captionText-1="Table 2: Morphometrics (mm) of female V. trinityae versus other select “vorhiesi” group species. Bold underlined comparisons are those for which there is no overlap with V. trinityae sp. nov. V. lapidicola data from the original description and redescription (Stahnke, 1940 &amp; Graham, 2006). Pectinal tooth count data stated as min-max (mean) [N]. Number in parentheses following species name is number of samples used in the morphometric comparisons. CA_L = carapace length, FF_L = fixed finger length." pageId="3" pageNumber="2">Tables 12</tableCitation>
)
<emphasis id="B93DEAF34B61FF9B4C7EF97EFA3FF6EC" bold="true" box="[839,1432,1766,1790]" pageId="3" pageNumber="2">http://zoobank.org/urn:lsid:zoobank.org:act:670320</emphasis>
</paragraph>
<paragraph id="8BF636E14B61FF9B4C99F89CFA99F70E" blockId="3.[839,1432,1702,1820]" box="[928,1342,1796,1820]" pageId="3" pageNumber="2">
<emphasis id="B93DEAF34B61FF9B4C99F89CFA99F70E" bold="true" box="[928,1342,1796,1820]" pageId="3" pageNumber="2">DB-F16C-4209-98AF-2773956248E4</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C353656A4B61FF9D4C06F8A6FE52F6C8" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8BF636E14B61FF9D4C06F8A6FE52F6C8" blockId="3.[831,1440,1854,1908]" lastBlockId="5.[192,802,1577,1754]" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
<emphasis id="B93DEAF34B61FF9B4C06F8A6FC14F744" bold="true" box="[831,947,1854,1878]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
Small (
<quantity id="4CB19B044B61FF9B4B25F8A6FB3CF744" box="[1052,1179,1854,1878]" metricMagnitude="-2" metricUnit="m" metricValue="2.355" pageId="3" pageNumber="2" unit="mm" value="23.55">23.55 mm</quantity>
) scorpions. Color is brown, lighter on the legs, with underlying mottling on carapace and mesosoma. (see
<figureCitation id="13722A644B67FF9D4D00F9B1FD07F653" box="[569,672,1577,1601]" captionStart="Figures 12" captionStartId="4.[192,278,1853,1877]" captionTargetBox="[191,1440,168,1825]" captionTargetId="figure-25@4.[192,1439,169,1824]" captionTargetPageId="4" captionText="Figures 12: Vaejovis trinityae sp. nov. female and male paratypes in natural habitat." pageId="5" pageNumber="4">Figure 1</figureCitation>
). Pedipalp movable finger usually with 7
<emphasis id="B93DEAF34B67FF9D4D1DF9D1FDE7F672" box="[548,576,1609,1632]" italics="true" pageId="5" pageNumber="4">ID</emphasis>
denticles and fixed finger usually with 6. Carapace of female is longer than the fifth metasomal segment. Pectinal tooth count for females 11.72 [n = 32], males 13.00 [n = 10]. Small, obsolete subaculear tubercle.
</paragraph>
</subSubSection>
<caption id="DF3666694B66FF9C4FF9F8A5FBAAF746" box="[192,1037,1853,1877]" pageId="4" pageNumber="3" startId="4.[192,278,1853,1877]" targetBox="[191,1440,168,1825]" targetPageId="4">
<paragraph id="8BF636E14B66FF9C4FF9F8A5FBAAF746" blockId="4.[192,1037,1853,1877]" box="[192,1037,1853,1877]" pageId="4" pageNumber="3">
<emphasis id="B93DEAF34B66FF9C4FF9F8A5FEE9F747" bold="true" box="[192,334,1853,1877]" pageId="4" pageNumber="3">Figures 12:</emphasis>
<taxonomicName id="4C494D624B66FF9C4E63F8A7FDA4F746" authority="Ayrey, 2013" authorityName="Ayrey" authorityYear="2013" box="[346,515,1855,1876]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="trinityae" status="sp. nov.">
<emphasis id="B93DEAF34B66FF9C4E63F8A7FDA4F746" box="[346,515,1855,1876]" italics="true" pageId="4" pageNumber="3">Vaejovis trinityae</emphasis>
</taxonomicName>
<emphasis id="B93DEAF34B66FF9C4D33F8A6FDF0F741" bold="true" box="[522,599,1854,1875]" pageId="4" pageNumber="3">
<taxonomicNameLabel id="A20E57884B66FF9C4D33F8A6FDF0F741" box="[522,599,1854,1875]" pageId="4" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female and male paratypes in natural habitat.
</paragraph>
</caption>
<caption id="DF3666694B67FF9D4FF9FA76FC63F617" box="[192,964,1518,1542]" pageId="5" pageNumber="4" startId="5.[192,267,1518,1542]" targetBox="[212,1424,188,1477]" targetPageId="5">
<paragraph id="8BF636E14B67FF9D4FF9FA76FC63F617" blockId="5.[192,964,1518,1542]" box="[192,964,1518,1542]" pageId="5" pageNumber="4">
<emphasis id="B93DEAF34B67FF9D4FF9FA76FE8FF614" bold="true" box="[192,296,1518,1542]" pageId="5" pageNumber="4">Figure 3:</emphasis>
<taxonomicName id="4C494D624B67FF9D4E16FA68FE7FF617" authority="Ayrey, 2013" authorityName="Ayrey" authorityYear="2013" box="[303,472,1520,1541]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="trinityae" status="sp. nov.">
<emphasis id="B93DEAF34B67FF9D4E16FA68FE7FF617" box="[303,472,1520,1541]" italics="true" pageId="5" pageNumber="4">Vaejovis trinityae</emphasis>
</taxonomicName>
<emphasis id="B93DEAF34B67FF9D4EE6FA77FD8BF616" bold="true" box="[479,556,1519,1540]" pageId="5" pageNumber="4">
<taxonomicNameLabel id="A20E57884B67FF9D4EE6FA77FD8BF616" box="[479,556,1519,1540]" pageId="5" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype female dorsal and ventral views.
</paragraph>
</caption>
<subSubSection id="C353656A4B67FF9D4FF9F898FBC2F672" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BF636E14B67FF9D4FF9F898FBC2F672" blockId="5.[192,802,1791,1908]" lastBlockId="5.[831,1439,1577,1632]" pageId="5" pageNumber="4">
<emphasis id="B93DEAF34B67FF9D4FF9F898FEDEF70A" bold="true" box="[192,377,1792,1816]" pageId="5" pageNumber="4">Type material.</emphasis>
<materialsCitation id="3B213CBC4B67FF9D4EA9F967FE26F766" collectingDate="2011-09-11" collectionCode="R, USNM" collectorName="F. Ayrey" country="United States of America" county="Coconino County" latitude="34.5201" location="Mogollon Rim" longLatPrecision="6" longitude="-111.5074" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="Arizona" typeStatus="holotype">
<typeStatus id="54F288434B67FF9D4EA9F967FE54F705" box="[400,499,1791,1815]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4FFD684B67FF9D4D32F898FDB8F705" box="[523,543,1792,1815]" pageId="5" pageNumber="4" type="female"></specimenCount>
, (
<geoCoordinate id="EE7D50264B67FF9D4D7CF967FD19F705" box="[581,702,1791,1815]" degrees="34.5201" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="5" value="34.5201">N34.5201°</geoCoordinate>
,
<geoCoordinate id="EE7D50264B67FF9D4DE3F967FEA3F724" degrees="111. 5074" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="5" value="-111.5074">W111. 5074°</geoCoordinate>
),
<collectingCountry id="F35E76714B67FF9D4E19F886FEFEF724" box="[288,345,1822,1846]" name="United States of America" pageId="5" pageNumber="4">USA</collectingCountry>
:
<collectingRegion id="498DF8034B67FF9D4E54F886FE60F724" box="[365,455,1822,1846]" country="United States of America" name="Arizona" pageId="5" pageNumber="4">Arizona</collectingRegion>
:
<collectingCounty id="62974E6D4B67FF9D4EE3F886FD03F724" box="[474,676,1822,1846]" pageId="5" pageNumber="4">Coconino County</collectingCounty>
,
<location id="8E96603A4B67FF9D4D81F886FF57F747" LSID="urn:lsid:plazi:treatment:03E087F74B61FF914C9CF93EFEA1F557:8E96603A4B67FF9D4D81F886FF57F747" country="United States of America" county="Coconino County" latitude="34.5201" longLatPrecision="6" longitude="-111.5074" name="Mogollon Rim" pageId="5" pageNumber="4" stateProvince="Arizona">Mogollon Rim</location>
,
<date id="FFF710214B67FF9D4E3BF8A5FE45F747" box="[258,482,1853,1877]" pageId="5" pageNumber="4" value="2011-09-11">
<collectingDate id="EFB3E9C94B67FF9D4E3BF8A5FE45F747" box="[258,482,1853,1877]" pageId="5" pageNumber="4" value="2011-09-11">11 September 2011</collectingDate>
</date>
, leg.
<collectionCode id="ED58AE244B67FF9D4D10F8A6FD99F747" box="[553,574,1854,1877]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="4">R</collectionCode>
.
<collectorName id="26BC53374B67FF9D4D7BF8A6FD0DF747" box="[578,682,1853,1877]" pageId="5" pageNumber="4">F. Ayrey</collectorName>
, specimen RA269 (
<collectionCode id="ED58AE244B67FF9D4E1FF8C4FEDEF766" box="[294,377,1884,1908]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="4" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B213CBC4B67FF9D4EACF8C4FBBAF653" box="[405,1053,1577,1908]" collectingDate="2011-09-16" collectionCode="R" collectorName="F. Ayrey" country="United States of America" county="Coconino County" latitude="34.5201" location="Mogollon Rim" longLatPrecision="6" longitude="-111.5074" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Arizona" typeStatus="paratype">
<typeStatus id="54F288434B67FF9D4EACF8C4FDA7F766" box="[405,512,1884,1908]" pageId="5" pageNumber="4" type="paratype">Paratypes</typeStatus>
: same locality,
<date id="FFF710214B67FF9D4DFDF8C4FC6DF653" box="[708,970,1577,1908]" pageId="5" pageNumber="4" value="2011-09-16">
<collectingDate id="EFB3E9C94B67FF9D4DFDF8C4FC6DF653" box="[708,970,1577,1908]" pageId="5" pageNumber="4" value="2011-09-16">16 September 2011</collectingDate>
</date>
, leg.
<collectionCode id="ED58AE244B67FF9D4B31F9B2FBBAF653" box="[1032,1053,1578,1601]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="4">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B213CBC4B67FF9D4B18F9B2FBC5F672" collectionCode="RFA" collectorName="F. Ayrey" country="United States of America" pageId="5" pageNumber="4" specimenCode="RA580, RA579" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Arizona" typeStatus="paratype">
<collectorName id="26BC53374B67FF9D4B18F9B2FB22F653" box="[1057,1157,1577,1601]" pageId="5" pageNumber="4">F. Ayrey</collectorName>
,
<specimenCount id="9D4FFD684B67FF9D4BABF9B1FB1AF653" box="[1170,1213,1577,1601]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
(
<specimenCode id="DBEF9E9A4B67FF9D4BF7F9B1FAB8F653" box="[1230,1311,1577,1601]" collectionCode="RA" pageId="5" pageNumber="4">RA580</specimenCode>
, RA581),
<specimenCount id="9D4FFD684B67FF9D4AABF9B1FCF4F672" pageId="5" pageNumber="4" type="female">2 ♀</specimenCount>
(
<specimenCode id="DBEF9E9A4B67FF9D4C58F9D0FC15F672" box="[865,946,1608,1632]" collectionCode="RA" pageId="5" pageNumber="4">RA579</specimenCode>
, RA583) (
<collectionCode id="ED58AE244B67FF9D4B19F9D0FBFEF672" box="[1056,1113,1608,1632]" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" pageId="5" pageNumber="4" type="Herbarium">RFA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C353656A4B67FF9D4C06F91DFAFFF6AE" pageId="5" pageNumber="4" type="etymology">
<paragraph id="8BF636E14B67FF9D4C06F91DFAFFF6AE" blockId="5.[831,1439,1669,1724]" pageId="5" pageNumber="4">
<emphasis id="B93DEAF34B67FF9D4C06F91DFC67F68F" bold="true" box="[831,960,1669,1693]" pageId="5" pageNumber="4">Etymology.</emphasis>
This species is named in honor of Trinity Frances Ayrey, my granddaughter and inspiration.
</paragraph>
</subSubSection>
<subSubSection id="C353656A4B67FF9D4C06F979FAB6F705" pageId="5" pageNumber="4" type="distribution">
<paragraph id="8BF636E14B67FF9D4C06F979FAB6F705" blockId="5.[831,1440,1761,1815]" pageId="5" pageNumber="4">
<emphasis id="B93DEAF34B67FF9D4C06F979FC76F6EB" bold="true" box="[831,977,1761,1785]" pageId="5" pageNumber="4">Distribution.</emphasis>
Known only from the
<typeStatus id="54F288434B67FF9D4BF4F97AFB5CF6E8" box="[1229,1275,1762,1786]" pageId="5" pageNumber="4">type</typeStatus>
locality:
<collectingCountry id="F35E76714B67FF9D4A5AF979FA3BF6EB" box="[1379,1436,1761,1785]" name="United States of America" pageId="5" pageNumber="4">USA</collectingCountry>
:
<collectingRegion id="498DF8034B67FF9D4C06F967FC3EF705" box="[831,921,1791,1815]" country="United States of America" name="Arizona" pageId="5" pageNumber="4">Arizona</collectingRegion>
: Coconino County, Mogollon Rim.
</paragraph>
</subSubSection>
<subSubSection id="C353656A4B67FF914C06F8A5FEA1F557" lastPageId="9" lastPageNumber="8" pageId="5" pageNumber="4" type="description">
<paragraph id="8BF636E14B67FF9D4C06F8A5FA3DF766" blockId="5.[831,1440,1853,1908]" pageId="5" pageNumber="4">
<emphasis id="B93DEAF34B67FF9D4C06F8A5FC6EF747" bold="true" box="[831,969,1853,1877]" pageId="5" pageNumber="4">Description.</emphasis>
Based on
<typeStatus id="54F288434B67FF9D4B69F8A5FB0AF747" box="[1104,1197,1853,1877]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
female, unless otherwise noted. See
<figureCitation id="13722A644B67FF9D4CCBF8C4FB97F766" box="[1010,1072,1884,1908]" captionStart="Figure 3" captionStartId="5.[192,267,1518,1542]" captionTargetBox="[212,1424,188,1477]" captionTargetId="figure-161@5.[192,1439,180,1488]" captionTargetPageId="5" captionText="Figure 3: Vaejovis trinityae sp. nov. holotype female dorsal and ventral views." pageId="5" pageNumber="4">Fig. 3</figureCitation>
for dorsal and ventral views and
</paragraph>
<caption id="DF3666694B64FF9E4FF9FA58FF49F61D" pageId="6" pageNumber="5" startId="6.[192,278,1472,1496]" targetBox="[214,1427,204,1418]" targetPageId="6">
<paragraph id="8BF636E14B64FF9E4FF9FA58FF49F61D" blockId="6.[192,1440,1472,1551]" pageId="6" pageNumber="5">
<emphasis id="B93DEAF34B64FF9E4FF9FA58FEF7F5CA" bold="true" box="[192,336,1472,1496]" pageId="6" pageNumber="5">Figures 49:</emphasis>
<taxonomicName id="4C494D624B64FF9E4E63FA5AFDA1F5C5" authority="Ayrey, 2013" authorityName="Ayrey" authorityYear="2013" box="[346,518,1474,1495]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="trinityae" status="sp. nov.">
<emphasis id="B93DEAF34B64FF9E4E63FA5AFDA1F5C5" box="[346,518,1474,1495]" italics="true" pageId="6" pageNumber="5">Vaejovis trinityae</emphasis>
</taxonomicName>
<emphasis id="B93DEAF34B64FF9E4D36FA5AFDC7F5C5" bold="true" box="[527,608,1474,1495]" pageId="6" pageNumber="5">
<taxonomicNameLabel id="A20E57884B64FF9E4D36FA5AFDC7F5C5" box="[527,608,1474,1495]" pageId="6" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype male.
<emphasis id="B93DEAF34B64FF9E4C3AFA5AFCB2F5C5" bold="true" box="[771,789,1474,1495]" pageId="6" pageNumber="5">4.</emphasis>
Carapace.
<emphasis id="B93DEAF34B64FF9E4CBFFA5AFC3FF5C5" bold="true" box="[902,920,1474,1495]" pageId="6" pageNumber="5">5.</emphasis>
Dentition of fixed and movable fingers of chela.
<emphasis id="B93DEAF34B64FF9E4AB7FA5AFA07F5C5" bold="true" box="[1422,1440,1474,1495]" pageId="6" pageNumber="5">6.</emphasis>
Chelicera, dorsal view.
<emphasis id="B93DEAF34B64FF9E4E9DFA46FE10F5E1" bold="true" box="[420,439,1502,1523]" pageId="6" pageNumber="5">7.</emphasis>
Telson, ventral and lateral views.
<emphasis id="B93DEAF34B64FF9E4C3BFA46FCB2F5E1" bold="true" box="[770,789,1502,1523]" pageId="6" pageNumber="5">8.</emphasis>
Metasomal segment V, lateral and ventral views.
<emphasis id="B93DEAF34B64FF9E4BC1FA46FAADF5E1" bold="true" box="[1272,1290,1502,1523]" pageId="6" pageNumber="5">9.</emphasis>
Stigma IV, left side.
</paragraph>
</caption>
<paragraph id="8BF636E14B64FF9E4FF9F9DBFF58F668" blockId="6.[192,802,1603,1658]" pageId="6" pageNumber="5">
<tableCitation id="C6CB035A4B64FF9E4FF9F9DBFEB1F649" box="[192,278,1603,1627]" captionStart="Table 1" captionStartId="9.[192,257,1234,1258]" captionText="Table 1: Morphometrics (mm) of Vaejovis trinityae, sp. nov." pageId="6" pageNumber="5">Table 1</tableCitation>
for measurements of
<typeStatus id="54F288434B64FF9E4D2CF9DBFDD5F649" box="[533,626,1603,1627]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D4FFD684B64FF9E4D88F9DBFF5CF668" pageId="6" pageNumber="5" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
.
</paragraph>
<paragraph id="8BF636E14B64FF9E4FF9F938FD17F6C4" blockId="6.[192,801,1695,1750]" pageId="6" pageNumber="5">
<emphasis id="B93DEAF34B64FF9E4FF9F938FEA6F6AA" bold="true" box="[192,257,1696,1720]" pageId="6" pageNumber="5">Color</emphasis>
. Color is brown, lighter on the legs. Faint underlying mottling on the carapace and mesosoma.
</paragraph>
<paragraph id="8BF636E14B64FF9E4FF9F963FBC0F6A5" blockId="6.[192,802,1787,1903]" lastBlockId="6.[831,1440,1603,1720]" pageId="6" pageNumber="5">
<emphasis id="B93DEAF34B64FF9E4FF9F963FE31F701" bold="true" box="[192,406,1787,1811]" pageId="6" pageNumber="5">
Carapace (
<figureCitation id="13722A644B64FF9E4E78F963FE2FF701" box="[321,392,1787,1811]" captionStart="Figures 49" captionStartId="6.[192,278,1472,1496]" captionTargetBox="[214,1427,204,1418]" captionTargetId="figure-195@6.[192,1441,192,1441]" captionTargetPageId="6" captionText="Figures 49: Vaejovis trinityae sp. nov. paratype male. 4. Carapace. 5. Dentition of fixed and movable fingers of chela. 6. Chelicera, dorsal view. 7. Telson, ventral and lateral views. 8. Metasomal segment V, lateral and ventral views. 9. Stigma IV, left side." pageId="6" pageNumber="5">Fig. 4</figureCitation>
).
</emphasis>
Anterior margin of carapace moderately emarginated, posterior margin slightly emarginated. Carapace finely granular. Three lateral eyes on each side. Median furrow moderate and traverses entire length of carapace. Ratio of median eyes location from anterior edge/carapace length 0.69; carapace length/ width at median eyes 1.43. Carapace of female is longer than metasomal segment V.
</paragraph>
<paragraph id="8BF636E14B64FF9F4C06F945FB2CF76A" blockId="6.[831,1440,1756,1903]" lastBlockId="7.[831,1439,1857,1912]" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5">
<emphasis id="B93DEAF34B64FF9E4C06F945FBB8F6E7" bold="true" box="[831,1055,1757,1781]" pageId="6" pageNumber="5">
Mesosoma (
<figureCitation id="13722A644B64FF9E4CF0F945FBB7F6E7" box="[969,1040,1757,1781]" captionStart="Figures 49" captionStartId="6.[192,278,1472,1496]" captionTargetBox="[214,1427,204,1418]" captionTargetId="figure-195@6.[192,1441,192,1441]" captionTargetPageId="6" captionText="Figures 49: Vaejovis trinityae sp. nov. paratype male. 4. Carapace. 5. Dentition of fixed and movable fingers of chela. 6. Chelicera, dorsal view. 7. Telson, ventral and lateral views. 8. Metasomal segment V, lateral and ventral views. 9. Stigma IV, left side." pageId="6" pageNumber="5">Fig. 9</figureCitation>
).
</emphasis>
Tergites finely granular with vestigial median carina on Tergites I-VI. Tergite VII with weak median carina on anterior third and strong dorsal lateral and lateral supramedian granular carinae. Sternites III-VI finely granular and without carinae. Sternite VII with granular ventral lateral carinae on middle third. Presternites smooth. Spiracles ovoid with median side rotated 35º from posterior sternite margin. Sternites with variable number of microsetae.
</paragraph>
<caption id="DF3666694B65FF9F4FF9F89EFC1BF70F" box="[192,956,1798,1822]" pageId="7" pageNumber="6" startId="7.[192,267,1798,1822]" targetBox="[202,1389,199,1752]" targetPageId="7">
<paragraph id="8BF636E14B65FF9F4FF9F89EFC1BF70F" blockId="7.[192,956,1798,1822]" box="[192,956,1798,1822]" pageId="7" pageNumber="6">
<emphasis id="B93DEAF34B65FF9F4FF9F89EFE91F70C" bold="true" box="[192,310,1798,1822]" pageId="7" pageNumber="6">Figure 10:</emphasis>
<taxonomicName id="4C494D624B65FF9F4E05F890FE41F70F" authority="Ayrey, 2013" authorityName="Ayrey" authorityYear="2013" box="[316,486,1800,1821]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="trinityae" status="sp. nov.">
<emphasis id="B93DEAF34B65FF9F4E05F890FE41F70F" box="[316,486,1800,1821]" italics="true" pageId="7" pageNumber="6">Vaejovis trinityae</emphasis>
</taxonomicName>
,
<emphasis id="B93DEAF34B65FF9F4ECBF89FFD98F70E" bold="true" box="[498,575,1799,1820]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A20E57884B65FF9F4ECBF89FFD98F70E" box="[498,575,1799,1820]" pageId="7" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male. Trichobothrial pattern.
</paragraph>
</caption>
<caption id="DF3666694B6AFF904FF9FC1EFA87F40B" pageId="8" pageNumber="7" startId="8.[192,267,902,926]" targetBox="[191,1441,167,870]" targetPageId="8">
<paragraph id="8BF636E14B6AFF904FF9FC1EFA87F40B" blockId="8.[192,1440,902,1049]" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904FF9FC1EFE90F38C" bold="true" box="[192,311,902,926]" pageId="8" pageNumber="7">Figure 11:</emphasis>
Right hemispermatophore and mating plug of
<taxonomicName id="4C494D624B6AFF904C39FC10FC0CF38F" authority="Ayrey, 2013" authorityName="Ayrey" authorityYear="2013" box="[768,939,904,925]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="trinityae" status="sp. nov.">
<emphasis id="B93DEAF34B6AFF904C39FC10FC0CF38F" box="[768,939,904,925]" italics="true" pageId="8" pageNumber="7">Vaejovis trinityae</emphasis>
</taxonomicName>
,
<emphasis id="B93DEAF34B6AFF904C8EFC1FFBA2F38E" bold="true" box="[951,1029,903,924]" pageId="8" pageNumber="7">
<taxonomicNameLabel id="A20E57884B6AFF904C8EFC1FFBA2F38E" box="[951,1029,903,924]" pageId="8" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male (photographed submerged in alcohol).
<emphasis id="B93DEAF34B6AFF904E0DFC38FEC3F3A7" bold="true" box="[308,356,928,949]" pageId="8" pageNumber="7">Left.</emphasis>
Dorsal view.
<emphasis id="B93DEAF34B6AFF904ED2FC38FDC4F3A7" bold="true" box="[491,611,928,949]" pageId="8" pageNumber="7">Upper Left.</emphasis>
Closeup of median area, dorsal and externodorsal views.
<emphasis id="B93DEAF34B6AFF904BA8FC38FB77F3A7" bold="true" box="[1169,1232,928,949]" pageId="8" pageNumber="7">Right.</emphasis>
Ventral view.
<emphasis id="B93DEAF34B6AFF904A66FC38FF58F3DC" bold="true" pageId="8" pageNumber="7">Upper Right.</emphasis>
Closeup of median area, internoventral and ventral views.
<emphasis id="B93DEAF34B6AFF904C59FC21FC16F3DC" bold="true" box="[864,945,953,974]" pageId="8" pageNumber="7">Bottom.</emphasis>
Mating plug, dorsal and ventral views, showing smooth barb (in dorsal view). Note in all ventral views of the hemispermatophore the embedded matinig plug is visible, the barb positioned towards the internal edge. Due to the translucency of the lamina, the ventral trough edge is visible from the dorsal view and the bifurcated lamellar hook, which emanates from the dorsal trough edge, is visible from the ventral view.
</paragraph>
</caption>
<paragraph id="8BF636E14B6AFF904FF9FBD0FE20F46C" blockId="8.[192,801,1096,1150]" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904FF9FBD0FE8DF472" bold="true" box="[192,298,1096,1120]" pageId="8" pageNumber="7">Sternum.</emphasis>
Sternum is
<typeStatus id="54F288434B6AFF904E8BFBD0FE47F472" box="[434,480,1096,1120]" pageId="8" pageNumber="7">type</typeStatus>
2 (Soleglad &amp; Fet, 2003) and is wider than long.
</paragraph>
<paragraph id="8BF636E14B6AFF904FF9FB3CFE82F4C9" blockId="8.[192,801,1188,1243]" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904FF9FB3CFE0BF4AE" bold="true" box="[192,428,1188,1212]" pageId="8" pageNumber="7">Genital Operculum.</emphasis>
Sclerites separated on posterior one-fifth.
</paragraph>
<paragraph id="8BF636E14B6AFF904FF9FA98FE04F580" blockId="8.[192,802,1280,1427]" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904FF9FA98FE83F50A" bold="true" box="[192,292,1280,1304]" pageId="8" pageNumber="7">Pectines.</emphasis>
Pectinal tooth count for the
<typeStatus id="54F288434B6AFF904D49FA98FD6AF50A" box="[624,717,1280,1304]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
female 10/10 and 14/15 for the
<typeStatus id="54F288434B6AFF904EFDFA87FD87F525" box="[452,544,1311,1335]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
male. All pectinal teeth have exterodistal angling with large sensorial area. Middle lamellae 6/6. Fulcra are present. Each fulcrum with one central seta.
</paragraph>
<paragraph id="8BF636E14B6AFF904FF9FA2AFAFDF4C9" blockId="8.[192,801,1458,1912]" lastBlockId="8.[831,1440,1096,1243]" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904FF9FA2AFE3AF5D8" bold="true" box="[192,413,1458,1482]" pageId="8" pageNumber="7">
Metasoma (
<figureCitation id="13722A644B6AFF904E71FA2AFE28F5D8" box="[328,399,1458,1482]" captionStart="Figures 49" captionStartId="6.[192,278,1472,1496]" captionTargetBox="[214,1427,204,1418]" captionTargetId="figure-195@6.[192,1441,192,1441]" captionTargetPageId="6" captionText="Figures 49: Vaejovis trinityae sp. nov. paratype male. 4. Carapace. 5. Dentition of fixed and movable fingers of chela. 6. Chelicera, dorsal view. 7. Telson, ventral and lateral views. 8. Metasomal segment V, lateral and ventral views. 9. Stigma IV, left side." pageId="8" pageNumber="7">Fig. 8</figureCitation>
).
</emphasis>
Carapace of female is longer than the fifth metasomal segment. Ratio of segment I length/ width 0.74;.of segment II length/width 0.97; of segment III length/width 0.99; of segment IV length/ width 1.34; of segment V length/width 2.05. Segments I-IV: dorsolateral carinae moderate, granular. Lateral supramedian carinae I-IV strong and granular with enlarged spinoid distal denticle on I &amp; II. Lateral inframedian carinae moderately granular on segment I, posterior 3/5 of II, 2/5 of III, and obsolete on IV. Ventrolateral carinae I weak and granular; on II-III moderate, granular; on IV strong, granular. Ventral submedian carinae weak to moderate on segment I, moderate on II, moderate, granular on III and moderate to strong, granular on IV. Dorsal and lateral intercarinal spaces very finely granular. Segment I-IV ventral submedian setae 3/3. Segment V: Dorsolateral carinae weak, distally crenulate, basally granular Lateromedian carinae moderate, granular. Ventrolateral and ventromedian carinae moderate. Intercarinal spaces finely granular. Segment V ventrolateral setae 4/4.
</paragraph>
<paragraph id="8BF636E14B6AFF904C06FA9EFB0AF568" blockId="8.[831,1440,1286,1402]" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904C06FA9EFC57F50C" bold="true" box="[831,1008,1286,1310]" pageId="8" pageNumber="7">
Telson (
<figureCitation id="13722A644B6AFF904CA5FA9EFC45F50C" box="[924,994,1286,1310]" captionStart="Figures 49" captionStartId="6.[192,278,1472,1496]" captionTargetBox="[214,1427,204,1418]" captionTargetId="figure-195@6.[192,1441,192,1441]" captionTargetPageId="6" captionText="Figures 49: Vaejovis trinityae sp. nov. paratype male. 4. Carapace. 5. Dentition of fixed and movable fingers of chela. 6. Chelicera, dorsal view. 7. Telson, ventral and lateral views. 8. Metasomal segment V, lateral and ventral views. 9. Stigma IV, left side." pageId="8" pageNumber="7">Fig. 7</figureCitation>
).
</emphasis>
Smooth with 4 pairs of large setae on the ventral surface, 3 large setae along both lateral edges of the vesicle and numerous smaller setae. Small, obsolete subaculear tubercle present.
</paragraph>
<paragraph id="8BF636E14B6AFF904C06FA3EFAFDF5E9" blockId="8.[831,1440,1445,1531]" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904C06FA3EFBBDF5AC" bold="true" box="[831,1050,1446,1470]" pageId="8" pageNumber="7">
Chelicerae (
<figureCitation id="13722A644B6AFF904CF1FA3EFBABF5AC" box="[968,1036,1446,1470]" captionStart="Figures 49" captionStartId="6.[192,278,1472,1496]" captionTargetBox="[214,1427,204,1418]" captionTargetId="figure-195@6.[192,1441,192,1441]" captionTargetPageId="6" captionText="Figures 49: Vaejovis trinityae sp. nov. paratype male. 4. Carapace. 5. Dentition of fixed and movable fingers of chela. 6. Chelicera, dorsal view. 7. Telson, ventral and lateral views. 8. Metasomal segment V, lateral and ventral views. 9. Stigma IV, left side." pageId="8" pageNumber="7">Fig. 6</figureCitation>
).
</emphasis>
Dorsal edge of movable cheliceral finger with two subdistal (sd) denticles. Ventral edge is smooth, with well developed serrula on distal half.
</paragraph>
<paragraph id="8BF636E14B6AFF904C06F9BFFCD2F6C5" blockId="8.[831,1440,1574,1751]" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904C06F9BFFBEAF62D" bold="true" box="[831,1101,1575,1599]" pageId="8" pageNumber="7">
Pedipalps (
<figureCitation id="13722A644B6AFF904CF8F9BFFBB6F62D" box="[961,1041,1575,1599]" captionStart="Figures 49" captionStartId="6.[192,278,1472,1496]" captionTargetBox="[214,1427,204,1418]" captionTargetId="figure-195@6.[192,1441,192,1441]" captionTargetPageId="6" captionText="Figures 49: Vaejovis trinityae sp. nov. paratype male. 4. Carapace. 5. Dentition of fixed and movable fingers of chela. 6. Chelicera, dorsal view. 7. Telson, ventral and lateral views. 8. Metasomal segment V, lateral and ventral views. 9. Stigma IV, left side." pageId="8" pageNumber="7">Figs. 5</figureCitation>
,
<figureCitation id="13722A644B6AFF904B1AF9BFFBE7F62D" box="[1059,1088,1575,1599]" captionStart="Figure 10" captionStartId="7.[192,267,1798,1822]" captionTargetBox="[202,1389,199,1752]" captionTargetId="figure-48@7.[192,1437,192,1768]" captionTargetPageId="7" captionText="Figure 10: Vaejovis trinityae, sp. nov., paratype male. Trichobothrial pattern." pageId="8" pageNumber="7">10</figureCitation>
).
</emphasis>
Trichobothrial pattern
<typeStatus id="54F288434B6AFF904A6DF9BFFA25F62D" box="[1364,1410,1575,1599]" pageId="8" pageNumber="7">type</typeStatus>
C (
<bibRefCitation id="EFD84B104B6AFF904C7FF9DDFC43F64F" author="VACHON" box="[838,996,1605,1629]" pageId="8" pageNumber="7" pagination="857 - 958" refId="ref6405" refString="VACHON, M. 1974. Etude des caracteres utilizes pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum National d'Historie Naturelle, Paris, (Ser. 3), 140 (Zool. 104): 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
) (see
<figureCitation id="13722A644B6AFF904B09F9DDFB23F64F" box="[1072,1156,1605,1629]" captionStart="Figure 10" captionStartId="7.[192,267,1798,1822]" captionTargetBox="[202,1389,199,1752]" captionTargetId="figure-48@7.[192,1437,192,1768]" captionTargetPageId="7" captionText="Figure 10: Vaejovis trinityae, sp. nov., paratype male. Trichobothrial pattern." pageId="8" pageNumber="7">Fig. 10</figureCitation>
). Trichobothria
<emphasis id="B93DEAF34B6AFF904A04F9DDFAF5F64F" box="[1341,1362,1605,1629]" italics="true" pageId="8" pageNumber="7">ib</emphasis>
and
<emphasis id="B93DEAF34B6AFF904AA8F9DEFA07F64F" box="[1425,1440,1606,1629]" italics="true" pageId="8" pageNumber="7">it</emphasis>
located at base of fixed finger. Pedipalp ratios: chela length/width 3.88; femur length/width 3.39; patella length/width 3.14; fixed finger length/carapace length 0.72.
</paragraph>
<paragraph id="8BF636E14B6AFF914C06F89BFEA1F557" blockId="8.[831,1440,1795,1911]" lastBlockId="9.[192,801,1294,1349]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="B93DEAF34B6AFF904C06F89BFC57F709" bold="true" box="[831,1008,1795,1819]" pageId="8" pageNumber="7">
Chela (
<figureCitation id="13722A644B6AFF904CAEF89BFC45F709" box="[919,994,1795,1819]" captionStart="Figures 49" captionStartId="6.[192,278,1472,1496]" captionTargetBox="[214,1427,204,1418]" captionTargetId="figure-195@6.[192,1441,192,1441]" captionTargetPageId="6" captionText="Figures 49: Vaejovis trinityae sp. nov. paratype male. 4. Carapace. 5. Dentition of fixed and movable fingers of chela. 6. Chelicera, dorsal view. 7. Telson, ventral and lateral views. 8. Metasomal segment V, lateral and ventral views. 9. Stigma IV, left side." pageId="8" pageNumber="7">Fig. 5</figureCitation>
).
</emphasis>
Carinae moderate. Fixed finger with median (
<emphasis id="B93DEAF34B6AFF904C91F8BBFC75F728" box="[936,978,1827,1850]" italics="true" pageId="8" pageNumber="7">MD</emphasis>
) denticles aligned and divided into 6 subrows by 5 outer (
<emphasis id="B93DEAF34B6AFF904B0AF8D9FBFEF74A" box="[1075,1113,1857,1880]" italics="true" pageId="8" pageNumber="7">OD</emphasis>
) denticles and usually 6
<emphasis id="B93DEAF34B6AFF904ABDF8D9FA07F74A" box="[1412,1440,1857,1880]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
denticles. Movable finger with 6 subrows, 5
<emphasis id="B93DEAF34B6AFF904A16F8F8FAF2F765" box="[1327,1365,1888,1911]" italics="true" pageId="8" pageNumber="7">OD</emphasis>
denticles and usually 7
<emphasis id="B93DEAF34B6BFF914E98FA97FE1AF534" box="[417,445,1295,1318]" italics="true" pageId="9" pageNumber="8">ID</emphasis>
denticles (Soleglad &amp; Sissom, 2001).
</paragraph>
</subSubSection>
</treatment>
</document>