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<document id="1AA061112819A994A63447831796B90E" ID-DOI="10.5281/zenodo.5167847" ID-ISSN="1942-1354" ID-Zenodo-Dep="5167847" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628271438169" checkinUser="felipe" docAuthor="Gillett, Conrad P. D. T." docDate="2009" docId="03E087DB4D59487AD89DFEF8CF05FA45" docLanguage="en" docName="InsectaMundi.0099.1-9.pdf" docOrigin="Insecta Mundi 2009 (99)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Coelosis biloba" docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="FFD9FFA34D5D487ED85DFFBFCB55FFBB" masterDocTitle="New records of dynastine scarab beetles in the tribes Oryctini, Agaocephalini and Dynastini from Cayo district, Belize (Coleoptera: Scarabaeidae: Dynastinae)" masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1699038676687" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="B5AB130C4573EB2CA808DB2B76510466">New records of dynastine scarab beetles in the tribes Oryctini, Agaocephalini and Dynastini from Cayo district, Belize (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
</mods:titleInfo>
<mods:name id="F02EBF51E7EE506B9A6E0CBEED6110B4" type="personal">
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</mods:role>
<mods:namePart id="72616FC86DDFF7A89B59A686E25909EC">Gillett, Conrad P. D. T.</mods:namePart>
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<mods:typeOfResource id="729E6983A47D1EBFEC6BAFEE971AB445">text</mods:typeOfResource>
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<mods:title id="F5325EA1F59F374C01DFEE3B1CB3D0A9">Insecta Mundi</mods:title>
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<mods:part id="BC978CD6F5EDA34B51B7065E07557E43">
<mods:date id="C141B46FF6DCF306C47E56AB393A2D80">2009</mods:date>
<mods:detail id="988B26C3B7588E3C64476E31FBFC434B" type="pubDate">
<mods:number id="9D9AB08796CD258679F28A5424BCDF7E">2009-10-25</mods:number>
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<mods:detail id="06F80FDF7946021CA687C3552D4694FC" type="volume">
<mods:number id="EACF0CA3E871D5FF009A81A6EF39F0E7">2009</mods:number>
</mods:detail>
<mods:detail id="355CB098491E3505A7D393E7787EB0A1" type="issue">
<mods:number id="BC4B539F96299FB49C0997B65D7B0A97">99</mods:number>
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<mods:classification id="4936D17DD138423FC767588523544F9D">journal article</mods:classification>
<mods:identifier id="86307C80653A043D6F51F69B9F293A81" type="DOI">10.5281/zenodo.5167847</mods:identifier>
<mods:identifier id="5B33757101473CB9BA69DB6BAACCE8D0" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="EEF68A367131B73A0C12CFC1BE5C85B8" type="Zenodo-Dep">5167847</mods:identifier>
</mods:mods>
<treatment id="03E087DB4D59487AD89DFEF8CF05FA45" LSID="urn:lsid:plazi:treatment:03E087DB4D59487AD89DFEF8CF05FA45" httpUri="http://treatment.plazi.org/id/03E087DB4D59487AD89DFEF8CF05FA45" lastPageNumber="3" pageId="4" pageNumber="3">
<subSubSection id="C35365464D59487AD89DFEF8C847FE25" pageId="4" pageNumber="3" type="nomenclature">
<paragraph id="8BF636CD4D59487AD89DFEF8C93DFEE5" blockId="4.[192,616,326,350]" box="[192,616,326,350]" pageId="4" pageNumber="3">
<heading id="D0BE81A14D59487AD89DFEF8C93DFEE5" bold="true" box="[192,616,326,350]" fontSize="10" level="2" pageId="4" pageNumber="3" reason="3">
<taxonomicName id="4C494D4E4D59487AD89DFEF8C93DFEE5" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[192,616,326,350]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="biloba">
<emphasis id="B93DEADF4D59487AD89DFEF8C93DFEE5" bold="true" box="[192,616,326,350]" pageId="4" pageNumber="3">
<emphasis id="B93DEADF4D59487AD89DFEF8CADDFEE5" bold="true" box="[192,392,327,350]" italics="true" pageId="4" pageNumber="3">Coelosis biloba</emphasis>
(Linnaeus, 1767)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFE39C847FE25" blockId="4.[192,786,390,446]" box="[192,786,390,414]" pageId="4" pageNumber="3">
<emphasis id="B93DEADF4D59487AD89DFE39CAB3FE25" bold="true" box="[192,486,390,414]" pageId="4" pageNumber="3">Specimens examined.</emphasis>
3 (
<specimenCount id="9D4FFD444D59487ADA5AFE39C936FE25" box="[519,611,390,414]" pageId="4" pageNumber="3" type="male">2 males</specimenCount>
and
<specimenCount id="9D4FFD444D59487ADAC1FE39C853FE25" box="[668,774,390,414]" pageId="4" pageNumber="3" type="female">1 female</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C35365464D59487AD89DFE18CAAEFD85" pageId="4" pageNumber="3" type="description">
<paragraph id="8BF636CD4D59487AD89DFE18CAF9FE05" blockId="4.[192,786,390,446]" box="[192,428,422,446]" pageId="4" pageNumber="3">BMNH and MGCB.</paragraph>
<paragraph id="8BF636CD4D59487AD89DFE59C95FFE45" blockId="4.[192,522,486,510]" box="[192,522,486,510]" pageId="4" pageNumber="3">
<emphasis id="B93DEADF4D59487AD89DFE59CA05FE45" bold="true" box="[192,336,486,510]" pageId="4" pageNumber="3">Localities.</emphasis>
Las Cuevas (3)
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFD99CAAEFD85" blockId="4.[192,507,550,574]" box="[192,507,550,574]" pageId="4" pageNumber="3">
<emphasis id="B93DEADF4D59487AD89DFD99CAC7FD85" bold="true" box="[192,402,550,574]" pageId="4" pageNumber="3">Temporal data.</emphasis>
June (3)
</paragraph>
</subSubSection>
<subSubSection id="C35365464D59487AD89DFDD9C93BFD45" pageId="4" pageNumber="3" type="distribution">
<paragraph id="8BF636CD4D59487AD89DFDD9C847FDC5" blockId="4.[192,786,614,766]" box="[192,786,614,638]" pageId="4" pageNumber="3">
<emphasis id="B93DEADF4D59487AD89DFDD9CA25FDC5" bold="true" box="[192,368,614,638]" pageId="4" pageNumber="3">Distribution.</emphasis>
Central
<collectingRegion id="498DF82F4D59487AD98AFDD9C979FDC5" box="[471,556,614,638]" country="Mexico" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingRegion>
to southern Argen-
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFD39C844FD25" blockId="4.[192,786,614,766]" box="[192,785,646,670]" pageId="4" pageNumber="3">
tina.
<bibRefCitation id="EFD84B3C4D59487AD8A3FD38CAC1FD25" author="Bates, H. W." box="[254,404,646,670]" pageId="4" pageNumber="3" pagination="296 - 342" refId="ref4388" refString="Bates, H. W. 1888. Pectinicornia and Lamellicornia, Family Dynastinae. Biologia Centrali-Americana. Insecta, Coleoptera 2 (part 2): 296 - 342." type="journal article" year="1888">Bates (1888)</bibRefCitation>
recorded the species from
<collectingRegion id="498DF82F4D59487ADA90FD39C858FD25" box="[717,781,646,670]" country="Belize" name="Cayo" pageId="4" pageNumber="3">Cayo</collectingRegion>
,
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFD19C847FD05" blockId="4.[192,786,614,766]" box="[192,786,678,702]" pageId="4" pageNumber="3">
and
<bibRefCitation id="EFD84B3C4D59487AD8ABFD19CAB0FD05" author="Blackwelder, R. E." box="[246,485,678,702]" pageId="4" pageNumber="3" pagination="189 - 341" refId="ref4419" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous Insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the U. S. National Museum 185: 189 - 341." type="journal article" year="1944">Blackwelder (1944)</bibRefCitation>
recorded it from
<collectingCountry id="F35E765D4D59487ADAE7FD19C847FD05" box="[698,786,678,702]" name="Belize" pageId="4" pageNumber="3">British</collectingCountry>
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFD79C847FD65" blockId="4.[192,786,614,766]" box="[192,786,710,734]" pageId="4" pageNumber="3">
Honduras. As discussed above,
<bibRefCitation id="EFD84B3C4D59487ADA6BFD79C9BFFD65" author="Endrodi, S." box="[566,746,710,734]" pageId="4" pageNumber="3" pagination="9 - 174" refId="ref4611" refString="Endrodi, S. 1976. Monographie der Dynastinae 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthinae). Folia Entomologica Hungarica (series nova) 29: 9 - 174." type="journal article" year="1976">Endrödi (1976)</bibRefCitation>
re-
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFD59C93BFD45" blockId="4.[192,786,614,766]" box="[192,622,742,766]" pageId="4" pageNumber="3">
corded it from
<collectingRegion id="498DF82F4D59487AD933FD59C93EFD45" box="[366,619,742,766]" country="Belize" name="Stann Creek" pageId="4" pageNumber="3">Stann Creek district</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C35365464D59487AD89DFC99CF05FA45" pageId="4" pageNumber="3" type="discussion">
<paragraph id="8BF636CD4D59487AD89DFC99C847FC84" blockId="4.[192,1440,806,1534]" box="[192,786,806,831]" pageId="4" pageNumber="3">
<emphasis id="B93DEADF4D59487AD89DFC99CA10FC85" bold="true" box="[192,325,806,830]" pageId="4" pageNumber="3">Remarks.</emphasis>
This species is known to be myrmeco-
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFCF9C847FCE5" blockId="4.[192,1440,806,1534]" box="[192,786,838,862]" pageId="4" pageNumber="3">philous, having been associated with leaf-cutter</paragraph>
<paragraph id="8BF636CD4D59487AD89DFCD9C847FCC5" blockId="4.[192,1440,806,1534]" box="[192,786,870,894]" pageId="4" pageNumber="3">
attine ants (
<bibRefCitation id="EFD84B3C4D59487AD911FCD9C95EFCC5" author="Lachaume, G." box="[332,523,870,894]" pageId="4" pageNumber="3" pagination="83 - 89" refId="ref4678" refString="Lachaume, G. 1992. Dynastinae americains. Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14. Sciences Nat, Venette; France. p. 1 - 56, 83 - 89, pl. 1 - 11." type="journal article" year="1992">Lachaume 1992</bibRefCitation>
; Ratcliffe 2006), which
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFC38C847FC25" blockId="4.[192,1440,806,1534]" box="[192,786,902,926]" pageId="4" pageNumber="3">are abundant at Las Cuevas. Two specimens were</paragraph>
<paragraph id="8BF636CD4D59487AD89DFC19C844FC05" blockId="4.[192,1440,806,1534]" box="[192,785,934,958]" pageId="4" pageNumber="3">collected dead in 2006. Although no biological data</paragraph>
<paragraph id="8BF636CD4D59487AD89DFC79C847FC65" blockId="4.[192,1440,806,1534]" box="[192,786,966,990]" pageId="4" pageNumber="3">is available for these specimens, it is notable that</paragraph>
<paragraph id="8BF636CD4D59487AD89DFC59C844FC45" blockId="4.[192,1440,806,1534]" box="[192,785,998,1022]" pageId="4" pageNumber="3">in more than two weeks of light trapping at Las</paragraph>
<paragraph id="8BF636CD4D59487AD89DFBB9C847FBA5" blockId="4.[192,1440,806,1534]" box="[192,786,1030,1054]" pageId="4" pageNumber="3">Cuevas in 2006, not a single specimen of this spe-</paragraph>
<paragraph id="8BF636CD4D59487AD89DFB99C844FB85" blockId="4.[192,1440,806,1534]" box="[192,785,1062,1086]" pageId="4" pageNumber="3">cies was captured, possibly indicating that the main</paragraph>
<paragraph id="8BF636CD4D59487AD89DFBF9C847FBE5" blockId="4.[192,1440,806,1534]" box="[192,786,1094,1118]" pageId="4" pageNumber="3">period of adult activity for this species had passed</paragraph>
<paragraph id="8BF636CD4D59487AD89DFBD9C847FBC5" blockId="4.[192,1440,806,1534]" box="[192,786,1126,1150]" pageId="4" pageNumber="3">by the time the two dead specimens above were</paragraph>
<paragraph id="8BF636CD4D59487AD89DFB39C844FB25" blockId="4.[192,1440,806,1534]" box="[192,785,1158,1183]" pageId="4" pageNumber="3">collected. This species can be amongst the com-</paragraph>
<paragraph id="8BF636CD4D59487AD89DFB18C844FB05" blockId="4.[192,1440,806,1534]" box="[192,785,1190,1214]" pageId="4" pageNumber="3">
monest
<taxonomicName id="4C494D4E4D59487AD943FB19CAD6FB05" authorityName="LeConte &amp; Horn" authorityYear="1883" box="[286,387,1190,1214]" class="Insecta" family="Dynastidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Oryctini">Oryctini</taxonomicName>
attracted to artificial light in cer-
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFB79C9E7FB65" blockId="4.[192,1440,806,1534]" box="[192,690,1222,1246]" pageId="4" pageNumber="3">tain areas (Yannig Ponchel pers. comm.).</paragraph>
<paragraph id="8BF636CD4D59487AD8ADFB59C847FB45" blockId="4.[192,1440,806,1534]" box="[240,786,1254,1278]" pageId="4" pageNumber="3">
<bibRefCitation id="EFD84B3C4D59487AD8ADFB59CAFFFB45" author="Endrodi, S." box="[240,426,1254,1278]" pageId="4" pageNumber="3" pagination="9 - 174" refId="ref4611" refString="Endrodi, S. 1976. Monographie der Dynastinae 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthinae). Folia Entomologica Hungarica (series nova) 29: 9 - 174." type="journal article" year="1976">Endrödi (1976)</bibRefCitation>
mentioned a locality for this
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFAB9C847FAA5" blockId="4.[192,1440,806,1534]" box="[192,786,1286,1310]" pageId="4" pageNumber="3">
species in
<collectingCountry id="F35E765D4D59487AD966FAB9CAEDFAA5" box="[315,440,1286,1310]" name="Honduras" pageId="4" pageNumber="3">Honduras</collectingCountry>
, which actually represents a
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFA99CFB3FA24" blockId="4.[192,1440,806,1534]" pageId="4" pageNumber="3">
record from
<collectingCountry id="F35E765D4D59487AD93DFA99CAE4FA85" box="[352,433,1318,1342]" name="Belize" pageId="4" pageNumber="3">Belize</collectingCountry>
. The locality in question is
<emphasis id="B93DEADF4D59487ADB6EFA8CC8F2FAF3" bold="true" box="[819,935,1331,1353]" pageId="4" pageNumber="3">Figure 3.</emphasis>
Map indicating the three sites (numbered) Middlesex, Stann Creck (sic) district.
<collectingRegion id="498DF82F4D59487ADA26FAF9C844FAE5" box="[635,785,1350,1374]" country="Belize" name="Stann Creek" pageId="4" pageNumber="3">Stann Creek</collectingRegion>
where collecting was undertaken in
<collectingRegion id="498DF82F4D59487ADC9CFAEFCE05FADD" box="[1217,1360,1360,1382]" country="Belize" name="Cayo" pageId="4" pageNumber="3">Cayo district</collectingRegion>
during is a district in eastern
<collectingCountry id="F35E765D4D59487AD998FAD9C95AFAC5" box="[453,527,1382,1406]" name="Belize" pageId="4" pageNumber="3">Belize</collectingCountry>
(the country formerly
<date id="FFF7100D4D59487ADB6EFAD2CF44FA39" box="[819,1041,1388,1410]" pageId="4" pageNumber="3" value="2006-05" valueMax="2006-06-31" valueMin="2006-05-00">May and June 2006</date>
. 1) Las Cuevas Research Station. 2) known as
<collectingCountry id="F35E765D4D59487AD964FA39C941FA25" box="[313,532,1414,1438]" name="Belize" pageId="4" pageNumber="3">British Honduras</collectingCountry>
), which undoubtedly Pooks Hill Lodge. 3) San Ignacio town.
</paragraph>
<paragraph id="8BF636CD4D59487AD89DFA19C844FA05" blockId="4.[192,1440,806,1534]" box="[192,785,1446,1470]" pageId="4" pageNumber="3">explains the confusion. Other records from Hon-</paragraph>
<paragraph id="8BF636CD4D59487AD89DFA79CF05FA45" blockId="4.[192,1440,806,1534]" pageId="4" pageNumber="3">
duras reported in Endrödis monographs could also possibly actually relate to
<collectingCountry id="F35E765D4D59487ADC34FA79CE6BFA65" box="[1129,1342,1478,1502]" name="Belize" pageId="4" pageNumber="3">British Honduras</collectingCountry>
(
<collectingCountry id="F35E765D4D59487ADD11FA79CECCFA65" box="[1356,1433,1478,1502]" name="Belize" pageId="4" pageNumber="3">Belize</collectingCountry>
) and care should be taken when interpreting them (e.g., Ratcliffe 2003: 225).
</paragraph>
</subSubSection>
</treatment>
</document>