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<document id="0A17A72509C614B45657D726CBC2995B" ID-CLB-Dataset="8517" ID-DOI="10.11646/zootaxa.4915.3.1" ID-GBIF-Dataset="76c6e3ae-e102-4522-816f-b5058a089bb0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4456421" ID-ZooBank="4DD891E2-10E7-453B-A935-875976A32611" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611302567007" checkinUser="plazi" docAuthor="Köhler, Gunther, Khaing, Khin Pa Pa, Than, Ni Lar, Baranski, Damian, Schell, Tilman, Greve, Carola, Janke, Axel &amp; Pauls, Steffen U." docDate="2021" docId="03E087CF023EFFE8FF319713799FF999" docLanguage="en" docName="zootaxa.4915.3.1.pdf" docOrigin="Zootaxa 4915 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Myanophis thanlyinensis Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls 2021, gen. nov." docType="treatment" docUuid="68106D4B-425B-4094-A6D3-81C4C42FEFB7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="311" masterDocId="FFD9FFB70239FFE2FFA69531797FFFCA" masterDocTitle="A new genus and species of mud snake from Myanmar (Reptilia, Squamata Homalopsidae)" masterLastPageNumber="325" masterPageNumber="301" pageNumber="308" updateTime="1698925108602" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FD95BB6A8C9A73741C66A174DA4C3705">A new genus and species of mud snake from Myanmar (Reptilia, Squamata Homalopsidae)</mods:title>
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<mods:namePart id="E1E7947F6FEB3BD0C415655A15E96BD5">Köhler, Gunther</mods:namePart>
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<mods:namePart id="FA22CCC214873EEC4010A90FFDEE04E5">Khaing, Khin Pa Pa</mods:namePart>
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<mods:affiliation id="9EEAC414773B698747623C60BC799F78">Senckenberg Forschungsinstitut und Naturmuseum, Senckenberganlage 25, 60325 Frankfurt a. M., Germany. &amp; papakhaing @ web. de; https: // orcid. org / 0000 - 0002 - 5926 - 4794</mods:affiliation>
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<mods:namePart id="F1F82B594281384F587E0248A4B58EFB">Than, Ni Lar</mods:namePart>
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<mods:affiliation id="8F00965F1A773DD1FE133C25B160630C">East Yangon University, Thanlyin, Yangon, Myanmar. &amp; nilarthan 65 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1098 - 7406</mods:affiliation>
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<mods:namePart id="B566DB92F53041C3A10B723AFA796E35">Baranski, Damian</mods:namePart>
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<mods:affiliation id="034AA0FD650FD3F35B33C19ADB0FD2A4">LOEWE Centre for Translational Biodiversity Genomics (LOEWE-TBG), Senckenberganlage 25, 60325 Frankfurt a. M., Germany. Senckenberg Biodiversity and Climate Research Centre, Georg-Voigt-Strasse 14 - 16, 60325 Frankfurt a. M., Germany. &amp; damian. baranski @ senckenberg. de; https: // orcid. org / 0000 - 0001 - 9671 - 0565</mods:affiliation>
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<mods:namePart id="696C79DF2B83CA03533DD5EA35B4A4C8">Schell, Tilman</mods:namePart>
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<mods:namePart id="F4775E09B78E23EA037458FDB4543948">Greve, Carola</mods:namePart>
<mods:affiliation id="4BC24771204EA55AA68D1C9331B9E441">LOEWE Centre for Translational Biodiversity Genomics (LOEWE-TBG), Senckenberganlage 25, 60325 Frankfurt a. M., Germany. Senckenberg Biodiversity and Climate Research Centre, Georg-Voigt-Strasse 14 - 16, 60325 Frankfurt a. M., Germany.</mods:affiliation>
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<mods:namePart id="F385E5C940545E9BBA526806E2ECDA2A">Janke, Axel</mods:namePart>
<mods:affiliation id="E10A4BDFE086BBA36D8EA2557E01480A">LOEWE Centre for Translational Biodiversity Genomics (LOEWE-TBG), Senckenberganlage 25, 60325 Frankfurt a. M., Germany. Senckenberg Biodiversity and Climate Research Centre, Georg-Voigt-Strasse 14 - 16, 60325 Frankfurt a. M., Germany.</mods:affiliation>
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<mods:namePart id="83514AFDB7F0D88806DFCFA42E871695">Pauls, Steffen U.</mods:namePart>
<mods:affiliation id="4E6717FEE57EC06092461FA6C2FD1657">Senckenberg Forschungsinstitut und Naturmuseum, Senckenberganlage 25, 60325 Frankfurt a. M., Germany. &amp; LOEWE Centre for Translational Biodiversity Genomics (LOEWE-TBG), Senckenberganlage 25, 60325 Frankfurt a. M., Germany. Senckenberg Biodiversity and Climate Research Centre, Georg-Voigt-Strasse 14 - 16, 60325 Frankfurt a. M., Germany.</mods:affiliation>
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<treatment id="03E087CF023EFFE8FF319713799FF999" ID-DOI="http://doi.org/10.5281/zenodo.4495448" ID-GBIF-Taxon="177206219" ID-Zenodo-Dep="4495448" LSID="urn:lsid:zoobank.org:act:68106D4B-425B-4094-A6D3-81C4C42FEFB7" httpUri="http://treatment.plazi.org/id/03E087CF023EFFE8FF319713799FF999" lastPageId="10" lastPageNumber="311" pageId="7" pageNumber="308">
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<taxonomicName id="4C494D5A023EFFE5FF31971378C3FDF6" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[151,444,546,572]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB023EFFE5FF31971378C3FDF6" box="[151,444,546,572]" italics="true" pageId="7" pageNumber="308">Myanophis thanlyinensis</emphasis>
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, sp. nov.
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<emphasis id="B93DEACB023EFFE5FF31975B7A72FD4E" bold="true" box="[151,781,618,644]" pageId="7" pageNumber="308">Gunther Köhler, Khin Pa Pa Khaing, Ni Lar Than</emphasis>
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ZOOBANK
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urn:lsid:zoobank.org: act:
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<figureCitation id="13722A5C023EFFE5FF3197E0787CFD26" box="[151,259,721,748]" captionStart-0="FIG" captionStart-1="FIG" captionStartId-0="7.[151,198,1663,1687]" captionStartId-1="9.[151,196,1686,1710]" captionTargetBox-0="[189,1398,915,1639]" captionTargetBox-1="[249,1338,181,1662]" captionTargetId-0="figure-204@7.[189,1398,915,1639]" captionTargetId-1="figure-19@9.[249,1338,181,1662]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIG. 5. Holotype (SMF 100707) of Myanophis thanlyinensis gen. nov., sp. nov. in life. Photograph by G.K." captionText-1="FIG. 6. Holotype (SMF 100707) of Myanophis thanlyinensis gen. nov., sp. nov.: (A) dorsal view; (B) ventral view; (C) left lateral head; (D) right lateral head; (E) dorsal head; (F) ventral head; (G) lateral body; (H) cloacal region with everted hemipenes. Scale bars equal 10.0 mm in A and B, and 5.0 mm in C to H. Photographs by G.K." figureDoi-0="http://doi.org/10.5281/zenodo.4456431" figureDoi-1="http://doi.org/10.5281/zenodo.4456433" httpUri-0="https://zenodo.org/record/4456431/files/figure.png" httpUri-1="https://zenodo.org/record/4456433/files/figure.png" pageId="7" pageNumber="308">Figs. 56</figureCitation>
;
<tableCitation id="C6CB0362023EFFE5FEA897E0781EFD21" box="[270,353,721,747]" captionStart="TABLE 2" captionStartId="8.[151,239,1088,1112]" captionTargetBox="[163,1360,1166,1952]" captionTargetPageId="8" captionText="TABLE 2. Variation in selected characters of external morphology in type series of Myanophis thanlyinensis gen. nov., sp. nov. For abbreviations see main text." httpUri="http://table.plazi.org/id/DF3666510231FFEAFF3191717B36FBB6" pageId="7" pageNumber="308" tableUuid="DF3666510231FFEAFF3191717B36FBB6">Table 2</tableCitation>
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<typeStatus id="54F2887B023EFFE5FF319628787BFCF9" box="[151,260,793,819]" pageId="7" pageNumber="308" type="holotype">Holotype</typeStatus>
</emphasis>
. SMF 100707, an adult male from wetlands near campus of East
<collectingRegion id="498DF83B023EFFE5FC7296287D55FCF9" box="[980,1066,793,819]" country="Myanmar" name="Yangon" pageId="7" pageNumber="308">Yangon</collectingRegion>
University (
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,
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;
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a.s.l.),
<collectingCountry id="F35E7649023EFFE5FEBF960C78FEFC9D" box="[281,385,829,855]" name="Myanmar" pageId="7" pageNumber="308">Thanlyin</collectingCountry>
,
<collectingRegion id="498DF83B023EFFE5FE2B960C7899FC9D" box="[397,486,829,855]" country="Myanmar" name="Yangon" pageId="7" pageNumber="308">Yangon</collectingRegion>
,
<collectingCountry id="F35E7649023EFFE5FE55960C7B1DFC9D" box="[499,610,829,855]" name="Myanmar" pageId="7" pageNumber="308">Myanmar</collectingCountry>
, collected on 5 of
<date id="FFF71019023EFFE5FC9E960C7A90FC9D" box="[824,1007,829,855]" pageId="7" pageNumber="308" value="2019-11">November 2019</date>
by Ni Lar Than and Gunther Köhler. Original field tag GK-7730.
</paragraph>
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<paragraph id="8BF636D9023EFFE5FF31934E7D9DF95D" blockId="7.[151,1250,1663,1687]" box="[151,1250,1663,1687]" pageId="7" pageNumber="308">
<emphasis id="B93DEACB023EFFE5FF31934E799BF95D" bold="true" box="[151,228,1663,1687]" pageId="7" pageNumber="308">FIG. 5.</emphasis>
Holotype (SMF 100707) of
<taxonomicName id="4C494D5A023EFFE5FDA1934E7A7CF95D" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[519,771,1663,1687]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB023EFFE5FDA1934E7A7CF95D" box="[519,771,1663,1687]" italics="true" pageId="7" pageNumber="308">Myanophis thanlyinensis</emphasis>
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<taxonomicNameLabel id="A20E57B0023EFFE5FCAF934E7A15F95D" box="[777,874,1663,1687]" pageId="7" pageNumber="308" rank="species">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
in life. Photograph by G.K.
</paragraph>
</caption>
<paragraph id="8BF636D9023EFFE5FF6193F47CF3F8A5" blockId="7.[151,1437,1733,2012]" pageId="7" pageNumber="308">
<emphasis id="B93DEACB023EFFE5FF6193F47840F915" bold="true" box="[199,319,1733,1759]" pageId="7" pageNumber="308">
<typeStatus id="54F2887B023EFFE5FF6193F47840F915" box="[199,319,1733,1759]" pageId="7" pageNumber="308" type="paratype">Paratypes</typeStatus>
</emphasis>
. EYU PaPa-0023 and SMF 100709 (original field tag PaPa-0019), two adult females from wetlands near campus of East
<collectingRegion id="498DF83B023EFFE5FED993DB78AAF8CE" box="[383,469,1770,1796]" country="Myanmar" name="Yangon" pageId="7" pageNumber="308">Yangon</collectingRegion>
University (
<geoCoordinate id="EE7D501E023EFFE5FDC693D87BA4F8C9" box="[608,731,1769,1795]" degrees="16.740297" direction="north" orientation="latitude" pageId="7" pageNumber="308" precision="1" value="16.740297">16.740297</geoCoordinate>
,
<geoCoordinate id="EE7D501E023EFFE5FD4093D87A1EF8CE" box="[742,865,1769,1796]" degrees="96.285184" direction="east" orientation="longitude" pageId="7" pageNumber="308" precision="1" value="96.28519">96.285184</geoCoordinate>
;
<quantity id="4CB19B3C023EFFE5FCCA93D87AE8F8CE" box="[876,919,1769,1796]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="308" unit="m" value="6.0">6 m</quantity>
a.s.l.),
<collectingCountry id="F35E7649023EFFE5FC4E93D87D2FF8C9" box="[1000,1104,1769,1795]" name="Myanmar" pageId="7" pageNumber="308">Thanlyin</collectingCountry>
,
<collectingRegion id="498DF83B023EFFE5FBFC93DB7DCCF8CE" box="[1114,1203,1770,1796]" country="Myanmar" name="Yangon" pageId="7" pageNumber="308">Yangon</collectingRegion>
,
<collectingCountry id="F35E7649023EFFE5FB1893DB7C52F8CE" box="[1214,1325,1770,1796]" name="Myanmar" pageId="7" pageNumber="308">Myanmar</collectingCountry>
, collected on 14 of
<date id="FFF71019023EFFE5FF5D923C78CDF8ED" box="[251,434,1805,1831]" pageId="7" pageNumber="308" value="2018-09">September 2018</date>
by a local fishermen and received by Khin Pa Pa Khaing. SMF 100708 (original field tag GK-7731), an adult male from wetlands near campus of East
<collectingRegion id="498DF83B023EFFE5FCE992037ADAF886" box="[847,933,1842,1868]" country="Myanmar" name="Yangon" pageId="7" pageNumber="308">Yangon</collectingRegion>
University (
<geoCoordinate id="EE7D501E023EFFE5FB9492007DE0F886" box="[1074,1183,1841,1868]" degrees="16.74275" direction="north" orientation="latitude" pageId="7" pageNumber="308" precision="1" value="16.74275">16.74275</geoCoordinate>
,
<geoCoordinate id="EE7D501E023EFFE5FB0A92007C66F886" box="[1196,1305,1841,1868]" degrees="96.31250" direction="east" orientation="longitude" pageId="7" pageNumber="308" precision="1" value="96.3125">96.31250</geoCoordinate>
;
<quantity id="4CB19B3C023EFFE5FA8292007C2FF886" box="[1316,1360,1841,1868]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="308" unit="m" value="6.0">6 m</quantity>
a.s.l.),
<collectingCountry id="F35E7649023EFFE5FF3192647980F8A5" box="[151,255,1877,1903]" name="Myanmar" pageId="7" pageNumber="308">Thanlyin</collectingCountry>
,
<collectingRegion id="498DF83B023EFFE5FEAF9267781DF8BA" box="[265,354,1878,1904]" country="Myanmar" name="Yangon" pageId="7" pageNumber="308">Yangon</collectingRegion>
,
<collectingCountry id="F35E7649023EFFE5FECB926778A3F8BA" box="[365,476,1878,1904]" name="Myanmar" pageId="7" pageNumber="308">Myanmar</collectingCountry>
, collected on 15 of
<date id="FFF71019023EFFE5FD1192647A2FF8A5" box="[695,848,1877,1903]" pageId="7" pageNumber="308" value="2019-10">October 2019</date>
by a local fishermen and received by Ni Lar Than.
</paragraph>
</subSubSection>
<subSubSection id="C3536552023EFFEAFF6192487BC4FD4F" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308" type="diagnosis">
<paragraph id="8BF636D9023EFFEAFF6192487BC4FD4F" blockId="7.[151,1437,1733,2012]" lastBlockId="8.[151,1437,150,1041]" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308">
<emphasis id="B93DEACB023EFFE5FF6192487845F859" bold="true" box="[199,314,1913,1939]" pageId="7" pageNumber="308">Diagnosis</emphasis>
. A species of the genus
<taxonomicName id="4C494D5A023EFFE5FDFC92487BA9F859" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[602,726,1913,1939]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="genus">
<emphasis id="B93DEACB023EFFE5FDFC92487BA9F859" box="[602,726,1913,1939]" italics="true" pageId="7" pageNumber="308">Myanophis</emphasis>
</taxonomicName>
that differs from all other homalopsid species by the following combination of characters: (1) dorsal scales smooth, row formula 212119 or 212117; (2) tail short, ratio TaL/SVL
<quantity id="4CB19B3C023EFFE5FEAE92F078CDF811" box="[264,434,1985,2012]" metricMagnitude="-3" metricUnit="m" metricValue="4.9403" metricValueMax="5.1815999999999995" metricValueMin="4.699" pageId="7" pageNumber="308" unit="in" value="0.1945" valueMax="0.204" valueMin="0.185">
<geoCoordinate id="EE7D501E023EFFE5FEAE92F07839F816" box="[264,326,1985,2012]" degrees="0.185" direction="north" orientation="latitude" pageId="7" pageNumber="308" precision="55" value="0.185">0.185</geoCoordinate>
<geoCoordinate id="EE7D501E023EFFE5FEE092F078ECF816" box="[326,403,1985,2012]" degrees="0.204" direction="west" orientation="longitude" pageId="7" pageNumber="308" precision="55" value="-0.204">0.204</geoCoordinate>
in
</quantity>
males,
<quantity id="4CB19B3C023EFFE5FDAC92F07BCBF811" box="[522,692,1985,2011]" metricMagnitude="-3" metricUnit="m" metricValue="4.1529" metricValueMax="4.2418" metricValueMin="4.064" pageId="7" pageNumber="308" unit="in" value="0.1635" valueMax="0.167" valueMin="0.16">
<geoCoordinate id="EE7D501E023EFFE5FDAC92F07B36F811" box="[522,585,1985,2011]" degrees="0.160" direction="north" orientation="latitude" pageId="7" pageNumber="308" precision="55" value="0.16">0.160</geoCoordinate>
<geoCoordinate id="EE7D501E023EFFE5FDEF92F07BE9F811" box="[585,662,1985,2011]" degrees="0.167" direction="west" orientation="longitude" pageId="7" pageNumber="308" precision="55" value="-0.167">0.167</geoCoordinate>
in
</quantity>
females; (3) nasal scales separated; (4) 125126 ventral scales in males,
<quantity id="4CB19B3C0231FFEAFF4F95A67814FF7B" box="[233,363,151,177]" metricMagnitude="0" metricUnit="m" metricValue="3.0734" metricValueMax="3.0987999999999998" metricValueMin="3.048" pageId="8" pageNumber="309" unit="in" value="121.0" valueMax="122.0" valueMin="120.0">120122 in</quantity>
females; (5) 3839 subcaudal scales in males,
<quantity id="4CB19B3C0231FFEAFC3695A67A8AFF7B" box="[912,1013,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" metricValueMax="8.636" metricValueMin="8.128" pageId="8" pageNumber="309" unit="in" value="33.0" valueMax="34.0" valueMin="32.0">3234 in</quantity>
females; and (6) hemipenis bilobed. Phylogenetically, (based on analyses of 16S and
<emphasis id="B93DEACB0231FFEAFD1C958A7A64FF1F" box="[698,795,187,213]" italics="true" pageId="8" pageNumber="309">MT-CYB</emphasis>
sequences),
<taxonomicName id="4C494D5A0231FFEAFC0A958A7C49FF1F" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[940,1334,187,213]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB0231FFEAFC0A958A7DBBFF1F" box="[940,1220,187,213]" italics="true" pageId="8" pageNumber="309">Myanophis thanlyinensis</emphasis>
<taxonomicNameLabel id="A20E57B00231FFEAFB6D958A7C49FF1F" box="[1227,1334,187,213]" pageId="8" pageNumber="309" rank="species">
<emphasis id="B93DEACB0231FFEAFB6D958A7C49FF1F" bold="true" box="[1227,1334,187,213]" pageId="8" pageNumber="309">gen. nov.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B93DEACB0231FFEAFA90958A7CE3FF1F" bold="true" box="[1334,1436,187,213]" pageId="8" pageNumber="309">, sp. nov.</emphasis>
differs from its closest relatives,
<taxonomicName id="4C494D5A0231FFEAFDAB95EE7BFDFF33" box="[525,642,223,249]" class="Reptilia" family="Homalopsidae" genus="Myrrophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="B93DEACB0231FFEAFDAB95EE7BFDFF33" box="[525,642,223,249]" italics="true" pageId="8" pageNumber="309">Myrrophis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D5A0231FFEAFD1A95EE7A5EFF33" authorityName="Murphy &amp; Voris" authorityYear="2014" box="[700,801,223,249]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="B93DEACB0231FFEAFD1A95EE7A5EFF33" box="[700,801,223,249]" italics="true" pageId="8" pageNumber="309">Gyiophis</emphasis>
</taxonomicName>
, by having a bilobed hemipenis (vs. unilobed). Further,
<taxonomicName id="4C494D5A0231FFEAFF3194327B62FED7" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[151,541,259,285]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB0231FFEAFF31943278D1FED7" box="[151,430,259,285]" italics="true" pageId="8" pageNumber="309">Myanophis thanlyinensis</emphasis>
<taxonomicNameLabel id="A20E57B00231FFEAFE1594327B62FED7" box="[435,541,259,285]" pageId="8" pageNumber="309" rank="species">
<emphasis id="B93DEACB0231FFEAFE1594327B62FED7" bold="true" box="[435,541,259,285]" pageId="8" pageNumber="309">gen. nov.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B93DEACB0231FFEAFDBB94327BFEFED7" bold="true" box="[541,641,259,285]" pageId="8" pageNumber="309">, sp. nov.</emphasis>
differs from the species of
<taxonomicName id="4C494D5A0231FFEAFC0894327D5CFED7" box="[942,1059,259,285]" class="Reptilia" family="Homalopsidae" genus="Myrrophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="B93DEACB0231FFEAFC0894327D5CFED7" box="[942,1059,259,285]" italics="true" pageId="8" pageNumber="309">Myrrophis</emphasis>
</taxonomicName>
by having 125126 ventral scales in males,
<quantity id="4CB19B3C0231FFEAFEA0941678F8FE8B" box="[262,391,295,321]" metricMagnitude="0" metricUnit="m" metricValue="3.0734" metricValueMax="3.0987999999999998" metricValueMin="3.048" pageId="8" pageNumber="309" unit="in" value="121.0" valueMax="122.0" valueMin="120.0">120122 in</quantity>
females (vs. 137162, 137164, respectively), and 3839 subcaudal scales in males,
<quantity id="4CB19B3C0231FFEAFAF0941679D2FEAF" metricMagnitude="-1" metricUnit="m" metricValue="8.382" metricValueMax="8.636" metricValueMin="8.128" pageId="8" pageNumber="309" unit="in" value="33.0" valueMax="34.0" valueMin="32.0">3234 in</quantity>
females (vs. 3955, 3752, respectively).
<taxonomicName id="4C494D5A0231FFEAFD32947A7D5DFEAF" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[660,1058,331,357]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB0231FFEAFD32947A7AD2FEAF" box="[660,941,331,357]" italics="true" pageId="8" pageNumber="309">Myanophis thanlyinensis</emphasis>
<taxonomicNameLabel id="A20E57B00231FFEAFC10947A7D5DFEAF" box="[950,1058,331,357]" pageId="8" pageNumber="309" rank="species">
<emphasis id="B93DEACB0231FFEAFC10947A7D5DFEAF" bold="true" box="[950,1058,331,357]" pageId="8" pageNumber="309">gen. nov.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B93DEACB0231FFEAFB84947A7DF3FEAF" bold="true" box="[1058,1164,331,357]" pageId="8" pageNumber="309">, sp. nov.</emphasis>
differs further from the species of
<taxonomicName id="4C494D5A0231FFEAFEAA945E780EFE43" authorityName="Murphy &amp; Voris" authorityYear="2014" box="[268,369,367,393]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="B93DEACB0231FFEAFEAA945E780EFE43" box="[268,369,367,393]" italics="true" pageId="8" pageNumber="309">Gyiophis</emphasis>
</taxonomicName>
by lacking dark blotches along flank (vs. having blotches), and by having 21 dorsal scales rows at midbody (vs. 25 scales). The following other homalopsid species share with
<taxonomicName id="4C494D5A0231FFEAFBAA94A27DBAFE67" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[1036,1221,403,429]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov., sp. nov.">
<emphasis id="B93DEACB0231FFEAFBAA94A27DBAFE67" box="[1036,1221,403,429]" italics="true" pageId="8" pageNumber="309">M. thanlyinensis</emphasis>
</taxonomicName>
<emphasis id="B93DEACB0231FFEAFB6A94A27CE3FE67" bold="true" box="[1228,1436,403,429]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A20E57B00231FFEAFB6A94A27CE3FE67" box="[1228,1436,403,429]" pageId="8" pageNumber="309" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
the combination of having smooth dorsal scales in 21 rows at midbody, reducing to 17 or 19 rows anterior to vent and fewer than 130 ventral scales:
<emphasis id="B93DEACB0231FFEAFD8594EA7D5AFE3E" box="[547,1061,475,501]" italics="true" pageId="8" pageNumber="309">
<taxonomicName id="4C494D5A0231FFEAFD8594EA7B88FE3F" authorityName="Murphy &amp; Voris" authorityYear="2005" box="[547,759,475,501]" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="chanardi">Enhydris chanardi</taxonomicName>
,
<taxonomicName id="4C494D5A0231FFEAFCA594EA7ADAFE3E" baseAuthorityName="Morice" baseAuthorityYear="1875" box="[771,933,475,500]" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="innominata">E. innominata</taxonomicName>
,
<taxonomicName id="4C494D5A0231FFEAFC1494EA7D5EFE3E" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[946,1057,475,500]" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="jagorii">E. jagorii</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C494D5A0231FFEAFBF894EA7D83FE3F" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[1118,1276,475,501]" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="B93DEACB0231FFEAFBF894EA7D83FE3F" box="[1118,1276,475,501]" italics="true" pageId="8" pageNumber="309">E. longicauda</emphasis>
</taxonomicName>
. However, all species have more than 50 subcaudal scales in males and more than
<quantity id="4CB19B3C0231FFEAFC3194CE7AAEFDD3" box="[919,977,511,537]" metricMagnitude="0" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="309" unit="in" value="40.0">40 in</quantity>
females (vs. 3839, 3234, respectively, in
<taxonomicName id="4C494D5A0231FFEAFF1297127811FDF7" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[180,366,547,573]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov., sp. nov.">
<emphasis id="B93DEACB0231FFEAFF1297127811FDF7" box="[180,366,547,573]" italics="true" pageId="8" pageNumber="309">M. thanlyinensis</emphasis>
</taxonomicName>
<emphasis id="B93DEACB0231FFEAFED397127B38FDF7" bold="true" box="[373,583,547,573]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A20E57B00231FFEAFED397127B38FDF7" box="[373,583,547,573]" pageId="8" pageNumber="309" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
), and a TaL/SVL of&gt;
<quantity id="4CB19B3C0231FFEAFC9997127AF1FDF7" box="[831,910,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="5.587999999999999" pageId="8" pageNumber="309" unit="in" value="0.22">0.22 in</quantity>
males, and&gt;
<quantity id="4CB19B3C0231FFEAFB8297127D0CFDF7" box="[1060,1139,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="4.318" pageId="8" pageNumber="309" unit="in" value="0.17">0.17 in</quantity>
females (vs. 0.1850.204, 0.1600.167, respectively).Also,
<taxonomicName id="4C494D5A0231FFEAFDA897767BB9FDAA" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[526,710,583,609]" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="jagorii">
<emphasis id="B93DEACB0231FFEAFDA897767BB9FDAA" box="[526,710,583,609]" italics="true" pageId="8" pageNumber="309">Enhydris jagorii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D5A0231FFEAFD5C97767AE8FDAB" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[762,919,583,609]" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="B93DEACB0231FFEAFD5C97767AE8FDAB" box="[762,919,583,609]" italics="true" pageId="8" pageNumber="309">E. longicauda</emphasis>
</taxonomicName>
have 2327 dorsal scales rows on neck (vs.
<quantity id="4CB19B3C0231FFEAFA26977679D2FD4F" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="8" pageNumber="309" unit="in" value="21.0">21 in</quantity>
<taxonomicName id="4C494D5A0231FFEAFF12975A7B3BFD4F" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[180,580,619,645]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB0231FFEAFF12975A78ACFD4F" box="[180,467,619,645]" italics="true" pageId="8" pageNumber="309">Myanophis Thanlyinensis</emphasis>
<taxonomicNameLabel id="A20E57B00231FFEAFE7C975A7B3BFD4F" box="[474,580,619,645]" pageId="8" pageNumber="309" rank="species">
<emphasis id="B93DEACB0231FFEAFE7C975A7B3BFD4F" bold="true" box="[474,580,619,645]" pageId="8" pageNumber="309">gen. nov.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B93DEACB0231FFEAFDE2975A7BD4FD4F" bold="true" box="[580,683,619,645]" pageId="8" pageNumber="309">, sp. nov.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C35365520231FFE8FF6197BF7B5EFF32" lastPageId="10" lastPageNumber="311" pageId="8" pageNumber="309" type="description">
<paragraph id="8BF636D90231FFEAFF6197BF7AA1FBDB" blockId="8.[151,1437,150,1041]" pageId="8" pageNumber="309">
<emphasis id="B93DEACB0231FFEAFF6197BF7B73FD63" bold="true" box="[199,524,654,681]" pageId="8" pageNumber="309">
Description of the
<typeStatus id="54F2887B0231FFEAFE0397BE7B73FD63" box="[421,524,655,681]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="13722A5C0231FFEAFDBD97BE7B16FD63" box="[539,617,654,681]" captionStart="FIG" captionStartId="7.[151,198,1663,1687]" captionTargetBox="[189,1398,915,1639]" captionTargetId="figure-204@7.[189,1398,915,1639]" captionTargetPageId="7" captionText="FIG. 5. Holotype (SMF 100707) of Myanophis thanlyinensis gen. nov., sp. nov. in life. Photograph by G.K." figureDoi="http://doi.org/10.5281/zenodo.4456431" httpUri="https://zenodo.org/record/4456431/files/figure.png" pageId="8" pageNumber="309">Figs. 5</figureCitation>
&amp;
<figureCitation id="13722A5C0231FFEAFD2897BE7BE1FD60" box="[654,670,655,682]" captionStart="FIG" captionStartId="9.[151,196,1686,1710]" captionTargetBox="[249,1338,181,1662]" captionTargetId="figure-19@9.[249,1338,181,1662]" captionTargetPageId="9" captionText="FIG. 6. Holotype (SMF 100707) of Myanophis thanlyinensis gen. nov., sp. nov.: (A) dorsal view; (B) ventral view; (C) left lateral head; (D) right lateral head; (E) dorsal head; (F) ventral head; (G) lateral body; (H) cloacal region with everted hemipenes. Scale bars equal 10.0 mm in A and B, and 5.0 mm in C to H. Photographs by G.K." figureDoi="http://doi.org/10.5281/zenodo.4456433" httpUri="https://zenodo.org/record/4456433/files/figure.png" pageId="8" pageNumber="309">6</figureCitation>
). Adult male, indicated by fully developed everted hemipenes; head depressed, distinct from neck; snout short and rounded, rostrum tapers downwards; rostral scale pentagonal, nearly as broad as tall, visible from above; eye moderate in size, pupils round; nasals divided with nasal groove contacting first supralabial; internasal squarish, not in contact with loreal; loreals 1/1, quadrangular, contacting first three supralabials; preocular 1/1; supraocular 1/1; postoculars 2/2; prefrontals 2, in broad contact with each other, frontal, internasal, loreals, preoculars, supraoculars, and nasals; frontal roughly pentagonal, 1.4 times longer than supraocular; parietals elongate; temporal scale formula 1 + 2 + 3; supralabials 8/8, 4th supralabial entering orbit; infralabials 10/10, first five in contact with anterior chin shield; anterior pair of chin shields larger than second pair, the former in contact with each other, the latter separated medially from each other by two smaller scales; 21 ASR, 21 MSR, 19 PSR; dorsal scales smooth without scale pits; 126 ventrals; cloacal plate divided; subcaudal scales 39. Body short, somewhat stocky; tail short (TaL/SVL 0.2038); SVL
<quantity id="4CB19B3C0231FFEAFD4C96C67A35FBDB" box="[746,842,1015,1041]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="309" unit="mm" value="260.0">260 mm</quantity>
; TaL
<quantity id="4CB19B3C0231FFEAFC2196C77AA6FBDB" box="[903,985,1014,1041]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="8" pageNumber="309" unit="mm" value="53.0">53 mm</quantity>
.
</paragraph>
<caption id="DF3666510231FFEAFF3191717B36FBB6" ID-Table-UUID="DF3666510231FFEAFF3191717B36FBB6" httpUri="http://table.plazi.org/id/DF3666510231FFEAFF3191717B36FBB6" pageId="8" pageNumber="309" startId="8.[151,239,1088,1112]" subCaptionStartIDs="8.[287,431,1124,1148]" subCaptionStarts="abbr" targetBox="[163,1360,1166,1952]" targetIsTable="true" targetPageId="8">
<paragraph id="8BF636D90231FFEAFF3191717B36FBB6" blockId="8.[151,1436,1088,1113]" lastBlockId="8.[151,585,1124,1149]" pageId="8" pageNumber="309">
<emphasis id="B93DEACB0231FFEAFF3191717874FB92" bold="true" box="[151,267,1088,1112]" pageId="8" pageNumber="309">TABLE 2.</emphasis>
Variation in selected characters of external morphology in type series of
<taxonomicName id="4C494D5A0231FFEAFBB991717C55FB92" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[1055,1322,1088,1112]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB0231FFEAFBB991717C55FB92" box="[1055,1322,1088,1112]" italics="true" pageId="8" pageNumber="309">Myanophis thanlyinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57B00231FFEAFA9791707CE8FB93" box="[1329,1431,1089,1113]" pageId="8" pageNumber="309">gen. nov.</taxonomicNameLabel>
,
<emphasis id="B93DEACB0231FFEAFF3191547992FBB7" bold="true" box="[151,237,1125,1149]" pageId="8" pageNumber="309">sp. nov.</emphasis>
For abbreviations see main text.
</paragraph>
</caption>
<paragraph id="8BF636D90231FFEAFE2891BF7DA4F86A" pageId="8" pageNumber="309">
<table id="F949C4790231001DFF0591BF7C2FF86A" box="[163,1360,1166,1952]" gridcols="5" gridrows="19" pageId="8" pageNumber="309">
<tr id="3579349B0231001DFF0591BF7C2FFB00" box="[163,1360,1166,1226]" gridrow="0" pageId="8" pageNumber="309" rowspan-0="1">
<th id="76A85DE70231001DFE2891BF7B6AFB00" box="[398,533,1166,1226]" gridcol="1" gridrow="0" pageId="8" pageNumber="309">
<emphasis id="B93DEACB0231FFEAFE2891BF788EFB00" bold="true" pageId="8" pageNumber="309">SMF 100707 Holotype</emphasis>
</th>
<th id="76A85DE70231001DFD3591BF7A65FB00" box="[659,794,1166,1226]" gridcol="2" gridrow="0" pageId="8" pageNumber="309">
<emphasis id="B93DEACB0231FFEAFD3591BF7B8AFB00" bold="true" pageId="8" pageNumber="309">SMF 100708 Paratype</emphasis>
</th>
<th id="76A85DE70231001DFC3191BF7D61FB00" box="[919,1054,1166,1226]" gridcol="3" gridrow="0" pageId="8" pageNumber="309">
<emphasis id="B93DEACB0231FFEAFC3191BF7A86FB00" bold="true" pageId="8" pageNumber="309">SMF 100709 Paratype</emphasis>
</th>
<th id="76A85DE70231001DFB0791BF7C2FFB00" box="[1185,1360,1166,1226]" gridcol="4" gridrow="0" pageId="8" pageNumber="309">
<emphasis id="B93DEACB0231FFEAFB0791BF7C7CFB00" bold="true" pageId="8" pageNumber="309">EYU PaPa-0023 Paratype</emphasis>
</th>
</tr>
<tr id="3579349B0231001DFF0591ED7C2FFB3E" box="[163,1360,1244,1268]" gridrow="1" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF0591ED787DFB3E" box="[163,258,1244,1268]" gridcol="0" gridrow="1" pageId="8" pageNumber="309">Sex</th>
<td id="76A85DE70231001DFE2891ED7B6AFB3E" box="[398,533,1244,1268]" gridcol="1" gridrow="1" pageId="8" pageNumber="309">male</td>
<td id="76A85DE70231001DFD3591ED7A65FB3E" box="[659,794,1244,1268]" gridcol="2" gridrow="1" pageId="8" pageNumber="309">male</td>
<td id="76A85DE70231001DFC3191ED7D61FB3E" box="[919,1054,1244,1268]" gridcol="3" gridrow="1" pageId="8" pageNumber="309">female</td>
<td id="76A85DE70231001DFB0791ED7C2FFB3E" box="[1185,1360,1244,1268]" gridcol="4" gridrow="1" pageId="8" pageNumber="309">female</td>
</tr>
<tr id="3579349B0231001DFF0590357C2FFAD6" box="[163,1360,1284,1308]" gridrow="2" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF059035787DFAD6" box="[163,258,1284,1308]" gridcol="0" gridrow="2" pageId="8" pageNumber="309">SVL</th>
<td id="76A85DE70231001DFE2890357B6AFAD6" box="[398,533,1284,1308]" gridcol="1" gridrow="2" pageId="8" pageNumber="309">260</td>
<td id="76A85DE70231001DFD3590357A65FAD6" box="[659,794,1284,1308]" gridcol="2" gridrow="2" pageId="8" pageNumber="309">335</td>
<td id="76A85DE70231001DFC3190357D61FAD6" box="[919,1054,1284,1308]" gridcol="3" gridrow="2" pageId="8" pageNumber="309">362</td>
<td id="76A85DE70231001DFB0790357C2FFAD6" box="[1185,1360,1284,1308]" gridcol="4" gridrow="2" pageId="8" pageNumber="309">300</td>
</tr>
<tr id="3579349B0231001DFF05901D7C2FFA8E" box="[163,1360,1324,1348]" gridrow="3" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF05901D787DFA8E" box="[163,258,1324,1348]" gridcol="0" gridrow="3" pageId="8" pageNumber="309">TaL</th>
<td id="76A85DE70231001DFE28901D7B6AFA8E" box="[398,533,1324,1348]" gridcol="1" gridrow="3" pageId="8" pageNumber="309">53</td>
<td id="76A85DE70231001DFD35901D7A65FA8E" box="[659,794,1324,1348]" gridcol="2" gridrow="3" pageId="8" pageNumber="309">62</td>
<td id="76A85DE70231001DFC31901D7D61FA8E" box="[919,1054,1324,1348]" gridcol="3" gridrow="3" pageId="8" pageNumber="309">58</td>
<td id="76A85DE70231001DFB07901D7C2FFA8E" box="[1185,1360,1324,1348]" gridcol="4" gridrow="3" pageId="8" pageNumber="309">50</td>
</tr>
<tr id="3579349B0231001DFF0590657C2FFAA7" box="[163,1360,1364,1389]" gridrow="4" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF059065787DFAA7" box="[163,258,1364,1389]" gridcol="0" gridrow="4" pageId="8" pageNumber="309">TaL/SVL</th>
<td id="76A85DE70231001DFE2890657B6AFAA7" box="[398,533,1364,1389]" gridcol="1" gridrow="4" pageId="8" pageNumber="309">0.204</td>
<td id="76A85DE70231001DFD3590657A65FAA7" box="[659,794,1364,1389]" gridcol="2" gridrow="4" pageId="8" pageNumber="309">0.185</td>
<td id="76A85DE70231001DFC3190657D61FAA7" box="[919,1054,1364,1389]" gridcol="3" gridrow="4" pageId="8" pageNumber="309">0.160</td>
<td id="76A85DE70231001DFB0790657C2FFAA7" box="[1185,1360,1364,1389]" gridcol="4" gridrow="4" pageId="8" pageNumber="309">0.167</td>
</tr>
<tr id="3579349B0231001DFF05904C7C2FFA5F" box="[163,1360,1405,1429]" gridrow="5" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF05904C787DFA5F" box="[163,258,1405,1429]" gridcol="0" gridrow="5" pageId="8" pageNumber="309">SV</th>
<td id="76A85DE70231001DFE28904C7B6AFA5F" box="[398,533,1405,1429]" gridcol="1" gridrow="5" pageId="8" pageNumber="309">126</td>
<td id="76A85DE70231001DFD35904C7A65FA5F" box="[659,794,1405,1429]" gridcol="2" gridrow="5" pageId="8" pageNumber="309">125</td>
<td id="76A85DE70231001DFC31904C7D61FA5F" box="[919,1054,1405,1429]" gridcol="3" gridrow="5" pageId="8" pageNumber="309">122</td>
<td id="76A85DE70231001DFB07904C7C2FFA5F" box="[1185,1360,1405,1429]" gridcol="4" gridrow="5" pageId="8" pageNumber="309">120</td>
</tr>
<tr id="3579349B0231001DFF0590947C2FFA77" box="[163,1360,1445,1469]" gridrow="6" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF059094787DFA77" box="[163,258,1445,1469]" gridcol="0" gridrow="6" pageId="8" pageNumber="309">ScS</th>
<td id="76A85DE70231001DFE2890947B6AFA77" box="[398,533,1445,1469]" gridcol="1" gridrow="6" pageId="8" pageNumber="309">39</td>
<td id="76A85DE70231001DFD3590947A65FA77" box="[659,794,1445,1469]" gridcol="2" gridrow="6" pageId="8" pageNumber="309">38</td>
<td id="76A85DE70231001DFC3190947D61FA77" box="[919,1054,1445,1469]" gridcol="3" gridrow="6" pageId="8" pageNumber="309">34</td>
<td id="76A85DE70231001DFB0790947C2FFA77" box="[1185,1360,1445,1469]" gridcol="4" gridrow="6" pageId="8" pageNumber="309">32</td>
</tr>
<tr id="3579349B0231001DFF0590FC7C2FFA2F" box="[163,1360,1485,1509]" gridrow="7" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF0590FC787DFA2F" box="[163,258,1485,1509]" gridcol="0" gridrow="7" pageId="8" pageNumber="309">PrOc</th>
<td id="76A85DE70231001DFE2890FC7B6AFA2F" box="[398,533,1485,1509]" gridcol="1" gridrow="7" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFD3590FC7A65FA2F" box="[659,794,1485,1509]" gridcol="2" gridrow="7" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFC3190FC7D61FA2F" box="[919,1054,1485,1509]" gridcol="3" gridrow="7" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFB0790FC7C2FFA2F" box="[1185,1360,1485,1509]" gridcol="4" gridrow="7" pageId="8" pageNumber="309">1</td>
</tr>
<tr id="3579349B0231001DFF0590C77C2FF9C4" box="[163,1360,1526,1550]" gridrow="8" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF0590C7787DF9C4" box="[163,258,1526,1550]" gridcol="0" gridrow="8" pageId="8" pageNumber="309">LoS</th>
<td id="76A85DE70231001DFE2890C77B6AF9C4" box="[398,533,1526,1550]" gridcol="1" gridrow="8" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFD3590C77A65F9C4" box="[659,794,1526,1550]" gridcol="2" gridrow="8" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFC3190C77D61F9C4" box="[919,1054,1526,1550]" gridcol="3" gridrow="8" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFB0790C77C2FF9C4" box="[1185,1360,1526,1550]" gridcol="4" gridrow="8" pageId="8" pageNumber="309">1</td>
</tr>
<tr id="3579349B0231001DFF05932F7C2FF9FC" box="[163,1360,1566,1590]" gridrow="9" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF05932F787DF9FC" box="[163,258,1566,1590]" gridcol="0" gridrow="9" pageId="8" pageNumber="309">PoOc</th>
<td id="76A85DE70231001DFE28932F7B6AF9FC" box="[398,533,1566,1590]" gridcol="1" gridrow="9" pageId="8" pageNumber="309">2</td>
<td id="76A85DE70231001DFD35932F7A65F9FC" box="[659,794,1566,1590]" gridcol="2" gridrow="9" pageId="8" pageNumber="309">2</td>
<td id="76A85DE70231001DFC31932F7D61F9FC" box="[919,1054,1566,1590]" gridcol="3" gridrow="9" pageId="8" pageNumber="309">2</td>
<td id="76A85DE70231001DFB07932F7C2FF9FC" box="[1185,1360,1566,1590]" gridcol="4" gridrow="9" pageId="8" pageNumber="309">2</td>
</tr>
<tr id="3579349B0231001DFF0593777C2FF994" box="[163,1360,1606,1630]" gridrow="10" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF059377787DF994" box="[163,258,1606,1630]" gridcol="0" gridrow="10" pageId="8" pageNumber="309">TempA</th>
<td id="76A85DE70231001DFE2893777B6AF994" box="[398,533,1606,1630]" gridcol="1" gridrow="10" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFD3593777A65F994" box="[659,794,1606,1630]" gridcol="2" gridrow="10" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFC3193777D61F994" box="[919,1054,1606,1630]" gridcol="3" gridrow="10" pageId="8" pageNumber="309">1</td>
<td id="76A85DE70231001DFB0793777C2FF994" box="[1185,1360,1606,1630]" gridcol="4" gridrow="10" pageId="8" pageNumber="309">1</td>
</tr>
<tr id="3579349B0231001DFF05935F7C2FF94C" box="[163,1360,1646,1670]" gridrow="11" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF05935F787DF94C" box="[163,258,1646,1670]" gridcol="0" gridrow="11" pageId="8" pageNumber="309">TempP</th>
<td id="76A85DE70231001DFE28935F7B6AF94C" box="[398,533,1646,1670]" gridcol="1" gridrow="11" pageId="8" pageNumber="309">2</td>
<td id="76A85DE70231001DFD35935F7A65F94C" box="[659,794,1646,1670]" gridcol="2" gridrow="11" pageId="8" pageNumber="309">2</td>
<td id="76A85DE70231001DFC31935F7D61F94C" box="[919,1054,1646,1670]" gridcol="3" gridrow="11" pageId="8" pageNumber="309">2</td>
<td id="76A85DE70231001DFB07935F7C2FF94C" box="[1185,1360,1646,1670]" gridcol="4" gridrow="11" pageId="8" pageNumber="309">2</td>
</tr>
<tr id="3579349B0231001DFF0593A67C2FF965" box="[163,1360,1687,1711]" gridrow="12" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF0593A6787DF965" box="[163,258,1687,1711]" gridcol="0" gridrow="12" pageId="8" pageNumber="309">SPL</th>
<td id="76A85DE70231001DFE2893A67B6AF965" box="[398,533,1687,1711]" gridcol="1" gridrow="12" pageId="8" pageNumber="309">8</td>
<td id="76A85DE70231001DFD3593A67A65F965" box="[659,794,1687,1711]" gridcol="2" gridrow="12" pageId="8" pageNumber="309">8</td>
<td id="76A85DE70231001DFC3193A67D61F965" box="[919,1054,1687,1711]" gridcol="3" gridrow="12" pageId="8" pageNumber="309">8</td>
<td id="76A85DE70231001DFB0793A67C2FF965" box="[1185,1360,1687,1711]" gridcol="4" gridrow="12" pageId="8" pageNumber="309">8</td>
</tr>
<tr id="3579349B0231001DFF05938E7C2FF91D" box="[163,1360,1727,1751]" gridrow="13" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF05938E787DF91D" box="[163,258,1727,1751]" gridcol="0" gridrow="13" pageId="8" pageNumber="309">SPL-Oc</th>
<td id="76A85DE70231001DFE28938E7B6AF91D" box="[398,533,1727,1751]" gridcol="1" gridrow="13" pageId="8" pageNumber="309">4</td>
<td id="76A85DE70231001DFD35938E7A65F91D" box="[659,794,1727,1751]" gridcol="2" gridrow="13" pageId="8" pageNumber="309">4</td>
<td id="76A85DE70231001DFC31938E7D61F91D" box="[919,1054,1727,1751]" gridcol="3" gridrow="13" pageId="8" pageNumber="309">4</td>
<td id="76A85DE70231001DFB07938E7C2FF91D" box="[1185,1360,1727,1751]" gridcol="4" gridrow="13" pageId="8" pageNumber="309">4</td>
</tr>
<tr id="3579349B0231001DFF0593D67C2FF935" box="[163,1360,1767,1791]" gridrow="14" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF0593D6787DF935" box="[163,258,1767,1791]" gridcol="0" gridrow="14" pageId="8" pageNumber="309">IFL</th>
<td id="76A85DE70231001DFE2893D67B6AF935" box="[398,533,1767,1791]" gridcol="1" gridrow="14" pageId="8" pageNumber="309">10</td>
<td id="76A85DE70231001DFD3593D67A65F935" box="[659,794,1767,1791]" gridcol="2" gridrow="14" pageId="8" pageNumber="309">10/9</td>
<td id="76A85DE70231001DFC3193D67D61F935" box="[919,1054,1767,1791]" gridcol="3" gridrow="14" pageId="8" pageNumber="309">10</td>
<td id="76A85DE70231001DFB0793D67C2FF935" box="[1185,1360,1767,1791]" gridcol="4" gridrow="14" pageId="8" pageNumber="309">10</td>
</tr>
<tr id="3579349B0231001DFF05923E7C2FF8ED" box="[163,1360,1807,1831]" gridrow="15" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF05923E787DF8ED" box="[163,258,1807,1831]" gridcol="0" gridrow="15" pageId="8" pageNumber="309">ASR</th>
<td id="76A85DE70231001DFE28923E7B6AF8ED" box="[398,533,1807,1831]" gridcol="1" gridrow="15" pageId="8" pageNumber="309">21</td>
<td id="76A85DE70231001DFD35923E7A65F8ED" box="[659,794,1807,1831]" gridcol="2" gridrow="15" pageId="8" pageNumber="309">21</td>
<td id="76A85DE70231001DFC31923E7D61F8ED" box="[919,1054,1807,1831]" gridcol="3" gridrow="15" pageId="8" pageNumber="309">21</td>
<td id="76A85DE70231001DFB07923E7C2FF8ED" box="[1185,1360,1807,1831]" gridcol="4" gridrow="15" pageId="8" pageNumber="309">21</td>
</tr>
<tr id="3579349B0231001DFF0592097C2FF89A" box="[163,1360,1848,1872]" gridrow="16" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF059209787DF89A" box="[163,258,1848,1872]" gridcol="0" gridrow="16" pageId="8" pageNumber="309">MSR</th>
<td id="76A85DE70231001DFE2892097B6AF89A" box="[398,533,1848,1872]" gridcol="1" gridrow="16" pageId="8" pageNumber="309">21</td>
<td id="76A85DE70231001DFD3592097A65F89A" box="[659,794,1848,1872]" gridcol="2" gridrow="16" pageId="8" pageNumber="309">21</td>
<td id="76A85DE70231001DFC3192097D61F89A" box="[919,1054,1848,1872]" gridcol="3" gridrow="16" pageId="8" pageNumber="309">21</td>
<td id="76A85DE70231001DFB0792097C2FF89A" box="[1185,1360,1848,1872]" gridcol="4" gridrow="16" pageId="8" pageNumber="309">21</td>
</tr>
<tr id="3579349B0231001DFF0592517C2FF8B2" box="[163,1360,1888,1912]" gridrow="17" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF059251787DF8B2" box="[163,258,1888,1912]" gridcol="0" gridrow="17" pageId="8" pageNumber="309">PSR</th>
<td id="76A85DE70231001DFE2892517B6AF8B2" box="[398,533,1888,1912]" gridcol="1" gridrow="17" pageId="8" pageNumber="309">19</td>
<td id="76A85DE70231001DFD3592517A65F8B2" box="[659,794,1888,1912]" gridcol="2" gridrow="17" pageId="8" pageNumber="309">19</td>
<td id="76A85DE70231001DFC3192517D61F8B2" box="[919,1054,1888,1912]" gridcol="3" gridrow="17" pageId="8" pageNumber="309">17</td>
<td id="76A85DE70231001DFB0792517C2FF8B2" box="[1185,1360,1888,1912]" gridcol="4" gridrow="17" pageId="8" pageNumber="309">17</td>
</tr>
<tr id="3579349B0231001DFF0592B97C2FF86A" box="[163,1360,1928,1952]" gridrow="18" pageId="8" pageNumber="309">
<th id="76A85DE70231001DFF0592B9787DF86A" box="[163,258,1928,1952]" gridcol="0" gridrow="18" pageId="8" pageNumber="309">ClS</th>
<td id="76A85DE70231001DFE2892B97B6AF86A" box="[398,533,1928,1952]" gridcol="1" gridrow="18" pageId="8" pageNumber="309">entire</td>
<td id="76A85DE70231001DFD3592B97A65F86A" box="[659,794,1928,1952]" gridcol="2" gridrow="18" pageId="8" pageNumber="309">entire</td>
<td id="76A85DE70231001DFC3192B97D61F86A" box="[919,1054,1928,1952]" gridcol="3" gridrow="18" pageId="8" pageNumber="309">entire</td>
<td id="76A85DE70231001DFB0792B97C2FF86A" box="[1185,1360,1928,1952]" gridcol="4" gridrow="18" pageId="8" pageNumber="309">entire</td>
</tr>
</table>
</paragraph>
<caption id="DF3666510230FFEBFF3193A77AA5F93C" ID-DOI="http://doi.org/10.5281/zenodo.4456433" ID-Zenodo-Dep="4456433" httpUri="https://zenodo.org/record/4456433/files/figure.png" pageId="9" pageNumber="310" startId="9.[151,196,1686,1710]" targetBox="[249,1338,181,1662]" targetPageId="9">
<paragraph id="8BF636D90230FFEBFF3193A77AA5F93C" blockId="9.[151,1437,1686,1783]" pageId="9" pageNumber="310">
<emphasis id="B93DEACB0230FFEBFF3193A7799CF964" bold="true" box="[151,227,1686,1710]" pageId="9" pageNumber="310">FIG. 6.</emphasis>
Holotype (SMF 100707) of
<taxonomicName id="4C494D5A0230FFEBFDA293A77A7FF964" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[516,768,1686,1710]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB0230FFEBFDA293A77A7FF964" box="[516,768,1686,1710]" italics="true" pageId="9" pageNumber="310">Myanophis thanlyinensis</emphasis>
</taxonomicName>
<emphasis id="B93DEACB0230FFEBFCA093A77ABEF964" bold="true" box="[774,961,1686,1710]" pageId="9" pageNumber="310">
<taxonomicNameLabel id="A20E57B00230FFEBFCA093A77A19F964" box="[774,870,1686,1710]" pageId="9" pageNumber="310" rank="species">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
: (A) dorsal view; (B) ventral view; (C) left lateral head; (D) right lateral head; (E) dorsal head; (F) ventral head; (G) lateral body; (H) cloacal region with everted hemipenes. Scale bars equal 10.0 mm in A and B, and 5.0 mm in C to H. Photographs by G.K.
</paragraph>
</caption>
<paragraph id="8BF636D90230FFEBFF6192157AFEF861" blockId="9.[151,1436,1828,2034]" pageId="9" pageNumber="310">
Coloration in life (
<figureCitation id="13722A5C0230FFEBFE3D9215789FF8F5" box="[411,480,1828,1855]" captionStart="FIG" captionStartId="7.[151,198,1663,1687]" captionTargetBox="[189,1398,915,1639]" captionTargetId="figure-204@7.[189,1398,915,1639]" captionTargetPageId="7" captionText="FIG. 5. Holotype (SMF 100707) of Myanophis thanlyinensis gen. nov., sp. nov. in life. Photograph by G.K." figureDoi="http://doi.org/10.5281/zenodo.4456431" httpUri="https://zenodo.org/record/4456431/files/figure.png" pageId="9" pageNumber="310">Fig. 5</figureCitation>
; colors in parentheses according to
<bibRefCitation id="EFD84B280230FFEBFCD192157D74F8F5" author="Kohler, G." box="[887,1035,1828,1855]" pageId="9" pageNumber="310" refId="ref10893" refString="Kohler, G. (2012) Color Catalogue for Field Biologists. Herpeton, Offenbach, 49 pp." type="book" year="2012">Köhler 2012</bibRefCitation>
): Dorsal ground color Olive Brown (color 278), scale rows 4 and 5 slightly darker (Sepia 286) and scale rows 1 to 3 Pale Cinnamon (55) with Hair Brown (277) blotch on anterior portion of each scale; ventral scales Pale Cinnamon (55) with Hair Brown (277) suffusion of anterior portion of each scale; iris Olive Brown (278).
</paragraph>
<paragraph id="8BF636D90230FFEBFF619285785FF838" blockId="9.[151,1436,1828,2034]" pageId="9" pageNumber="310">Coloration after about three months in preservation in 70% ethanol agrees well with the description of coloration in life.</paragraph>
<paragraph id="8BF636D90233FFE8FF6195A77B5EFF32" blockId="10.[151,1437,150,609]" pageId="10" pageNumber="311">
<emphasis id="B93DEACB0233FFE8FF6195A77847FF7A" bold="true" box="[199,312,150,176]" pageId="10" pageNumber="311">Variation</emphasis>
.
<typeStatus id="54F2887B0233FFE8FEE095A678CCFF7B" box="[326,435,151,177]" pageId="10" pageNumber="311" type="paratype">Paratypes</typeStatus>
agree well with the
<typeStatus id="54F2887B0233FFE8FD3D95A77B82FF7A" box="[667,765,150,176]" pageId="10" pageNumber="311" type="holotype">holotype</typeStatus>
in general appearance, morphometrics and scalation (
<tableCitation id="C6CB03620233FFE8FAFB95A779D7FF1E" captionStart="TABLE 2" captionStartId="8.[151,239,1088,1112]" captionTargetBox="[163,1360,1166,1952]" captionTargetPageId="8" captionText="TABLE 2. Variation in selected characters of external morphology in type series of Myanophis thanlyinensis gen. nov., sp. nov. For abbreviations see main text." httpUri="http://table.plazi.org/id/DF3666510231FFEAFF3191717B36FBB6" pageId="10" pageNumber="311" tableUuid="DF3666510231FFEAFF3191717B36FBB6">Table 2</tableCitation>
). Strong sexual dimorphism is evident in the number of ventral and subcaudal scales that show non-overlapping values between males and females.
</paragraph>
</subSubSection>
<subSubSection id="C35365520233FFE8FF619433787BFEAE" pageId="10" pageNumber="311" type="etymology">
<paragraph id="8BF636D90233FFE8FF619433787BFEAE" blockId="10.[151,1437,150,609]" pageId="10" pageNumber="311">
<emphasis id="B93DEACB0233FFE8FF6194337837FED6" bold="true" box="[199,328,258,284]" pageId="10" pageNumber="311">Etymology</emphasis>
. The species epithet refers to the city of
<collectingCountry id="F35E76490233FFE8FC8494337AF5FED6" box="[802,906,258,284]" name="Myanmar" pageId="10" pageNumber="311">Thanlyin</collectingCountry>
, a major port city of
<collectingCountry id="F35E76490233FFE8FB2694327D90FED7" box="[1152,1263,259,285]" name="Myanmar" pageId="10" pageNumber="311">Myanmar</collectingCountry>
, located across the
<collectingRegion id="498DF83B0233FFE8FF6494167982FE8B" box="[194,253,295,321]" country="Myanmar" name="Bago" pageId="10" pageNumber="311">Bago</collectingRegion>
River southeast of the city of
<collectingRegion id="498DF83B0233FFE8FDF094167BD0FE8B" box="[598,687,295,321]" country="Myanmar" name="Yangon" pageId="10" pageNumber="311">Yangon</collectingRegion>
, where the
<typeStatus id="54F2887B0233FFE8FC9494167A1DFE8B" box="[818,866,295,321]" pageId="10" pageNumber="311">type</typeStatus>
series of
<taxonomicName id="4C494D5A0233FFE8FC7494167DF3FE8A" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[978,1164,294,320]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov., sp. nov.">
<emphasis id="B93DEACB0233FFE8FC7494167DF3FE8A" box="[978,1164,294,320]" italics="true" pageId="10" pageNumber="311">M. thanlyinensis</emphasis>
</taxonomicName>
<emphasis id="B93DEACB0233FFE8FB3294177C16FE8A" bold="true" box="[1172,1385,294,320]" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A20E57B00233FFE8FB3294177C16FE8A" box="[1172,1385,294,320]" pageId="10" pageNumber="311" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="C35365520233FFE8FF61945F7BE5FDAA" pageId="10" pageNumber="311" type="biology_ecology">
<paragraph id="8BF636D90233FFE8FF61945F7BE5FDAA" blockId="10.[151,1437,150,609]" pageId="10" pageNumber="311">
<emphasis id="B93DEACB0233FFE8FF61945F78BDFE42" bold="true" box="[199,450,366,392]" pageId="10" pageNumber="311">Natural history notes</emphasis>
. The
<typeStatus id="54F2887B0233FFE8FDA7945E7B4EFE43" box="[513,561,367,393]" pageId="10" pageNumber="311">type</typeStatus>
locality is open grassland (
<figureCitation id="13722A5C0233FFE8FCF8945F7ADEFE43" box="[862,929,366,393]" captionStart="FIG" captionStartId="10.[151,198,1378,1402]" captionTargetBox="[151,1436,727,1355]" captionTargetId="figure-218@10.[151,1436,632,1355]" captionTargetPageId="10" captionText="FIG. 7. Habitat of Myanophis thanlyinensis gen. nov., sp. nov. near the type locality (near campus of East Yangon University, Thanlyin, Yangon, Myanmar). Photograph by G.K, taken on 23 June 2019." figureDoi="http://doi.org/10.5281/zenodo.4456435" httpUri="https://zenodo.org/record/4456435/files/figure.png" pageId="10" pageNumber="311">Fig. 7</figureCitation>
), mostly flooded during the rainy season with only a few scattered wooden houses, adjacent to the campus of East
<collectingRegion id="498DF83B0233FFE8FC2194A27AA2FE67" box="[903,989,403,429]" country="Myanmar" name="Yangon" pageId="10" pageNumber="311">Yangon</collectingRegion>
University (EYU),
<collectingCountry id="F35E76490233FFE8FB1E94A37C5FFE66" box="[1208,1312,402,428]" name="Myanmar" pageId="10" pageNumber="311">Thanlyin</collectingCountry>
,
<collectingCountry id="F35E76490233FFE8FA8C94A27CE6FE67" box="[1322,1433,403,429]" name="Myanmar" pageId="10" pageNumber="311">Myanmar</collectingCountry>
. The snakes were caught at night with the help of local fishermen in shallow water (&lt;
<quantity id="4CB19B3C0233FFE8FB9394877D05FE1B" box="[1077,1146,438,465]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="311" unit="cm" value="50.0">50 cm</quantity>
depth) with an abundance of grassy vegetation. Syntopic with
<taxonomicName id="4C494D5A0233FFE8FD9694EA7B94FE3E" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[560,747,474,500]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov., sp. nov.">
<emphasis id="B93DEACB0233FFE8FD9694EA7B94FE3E" box="[560,747,474,500]" italics="true" pageId="10" pageNumber="311">M. thanlyinensis</emphasis>
</taxonomicName>
<emphasis id="B93DEACB0233FFE8FD5594EB7AB8FE3E" bold="true" box="[755,967,474,500]" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A20E57B00233FFE8FD5594EB7AB8FE3E" box="[755,967,474,500]" pageId="10" pageNumber="311" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
we collected the homalopsid species
<taxonomicName id="4C494D5A0233FFE8FAD294EA783AFDD2" authority="enhydris" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="enhydris">
<emphasis id="B93DEACB0233FFE8FAD294EA783AFDD2" italics="true" pageId="10" pageNumber="311">Enhydris enhydris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D5A0233FFE8FEDA94CF7B56FDD2" baseAuthorityName="Murphy" baseAuthorityYear="2007" box="[380,553,510,536]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="vorisi">
<emphasis id="B93DEACB0233FFE8FEDA94CF7B56FDD2" box="[380,553,510,536]" italics="true" pageId="10" pageNumber="311">Gyiophis vorisi</emphasis>
</taxonomicName>
(
<figureCitation id="13722A5C0233FFE8FD9E94CF7B04FDD2" box="[568,635,510,536]" captionStart="FIG" captionStartId="12.[151,198,1754,1778]" captionTargetBox="[227,1361,181,1730]" captionTargetId="figure-21@12.[227,1361,181,1730]" captionTargetPageId="12" captionText="FIG. 8. Homalopsid species in life, collected at the type locality (near campus of East Yangon University, Thanlyin, Yangon, Myanmar) of Myanophis thanlyinensis gen. nov., sp. nov.: (A) Gyiophis vorisi (SMF 100700); (B) Enhydris enhydris (GK- 7158). Photographs by G.K." figureDoi="http://doi.org/10.5281/zenodo.4456437" httpUri="https://zenodo.org/record/4456437/files/figure.png" pageId="10" pageNumber="311">Fig. 8</figureCitation>
) as well the semiaquatic colubrids
<taxonomicName id="4C494D5A0233FFE8FBAF94CF7D8CFDD2" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1033,1267,510,536]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="stolatum">
<emphasis id="B93DEACB0233FFE8FBAF94CF7D8CFDD2" box="[1033,1267,510,536]" italics="true" pageId="10" pageNumber="311">Amphiesma stolatum</emphasis>
</taxonomicName>
,
<emphasis id="B93DEACB0233FFE8FAA494CF7988FDF6" italics="true" pageId="10" pageNumber="311">
<taxonomicName id="4C494D5A0233FFE8FAA494CF798DFDF6" class="Reptilia" family="Colubridae" genus="Xenochrophis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="bellulus">Xenochrophis bellulus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C494D5A0233FFE8FE8A971278D8FDF6" baseAuthorityName="Schneider" baseAuthorityYear="1799" box="[300,423,547,572]" class="Reptilia" family="Colubridae" genus="Xenochrophis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="piscator">
<emphasis id="B93DEACB0233FFE8FE8A971278D8FDF6" box="[300,423,547,572]" italics="true" pageId="10" pageNumber="311">X. piscator</emphasis>
</taxonomicName>
. Of these syntopic species,
<taxonomicName id="4C494D5A0233FFE8FD7D97127A25FDF6" box="[731,858,546,572]" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="enhydris">
<emphasis id="B93DEACB0233FFE8FD7D97127A25FDF6" box="[731,858,546,572]" italics="true" pageId="10" pageNumber="311">E. enhydris</emphasis>
</taxonomicName>
was the by far most abundant snake at this site with&gt;20 individuals encountered per night-search.
</paragraph>
</subSubSection>
<caption id="DF3666510233FFE8FF3190537AF1FA55" ID-DOI="http://doi.org/10.5281/zenodo.4456435" ID-Zenodo-Dep="4456435" httpUri="https://zenodo.org/record/4456435/files/figure.png" pageId="10" pageNumber="311" startId="10.[151,198,1378,1402]" targetBox="[151,1436,727,1355]" targetPageId="10">
<paragraph id="8BF636D90233FFE8FF3190537AF1FA55" blockId="10.[151,1437,1378,1439]" pageId="10" pageNumber="311">
<emphasis id="B93DEACB0233FFE8FF319053799BFAB1" bold="true" box="[151,228,1378,1403]" pageId="10" pageNumber="311">FIG. 7.</emphasis>
Habitat of
<taxonomicName id="4C494D5A0233FFE8FEFE90537BC3FAB1" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[344,700,1378,1403]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB0233FFE8FEFE90537B2AFAB0" box="[344,597,1378,1402]" italics="true" pageId="10" pageNumber="311">Myanophis thanlyinensis</emphasis>
<taxonomicNameLabel id="A20E57B00233FFE8FDFD90527BC3FAB1" box="[603,700,1379,1403]" pageId="10" pageNumber="311" rank="species">
<emphasis id="B93DEACB0233FFE8FDFD90527BC3FAB1" bold="true" box="[603,700,1379,1403]" pageId="10" pageNumber="311">gen. nov.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B93DEACB0233FFE8FD1A90527A66FAB1" bold="true" box="[700,793,1379,1403]" pageId="10" pageNumber="311">, sp. nov.</emphasis>
near the type locality (near campus of East Yangon University, Thanlyin, Yangon, Myanmar). Photograph by G.K, taken on 23 June 2019.
</paragraph>
</caption>
<subSubSection id="C35365520233FFE8FF6190FD799FF999" pageId="10" pageNumber="311" type="distribution">
<paragraph id="8BF636D90233FFE8FF6190FD799FF999" blockId="10.[151,1437,1484,1619]" pageId="10" pageNumber="311">
<emphasis id="B93DEACB0233FFE8FF6190FD7BADFA2C" bold="true" box="[199,722,1484,1511]" pageId="10" pageNumber="311">Geographic Distribution and Conservation</emphasis>
. As currently known,
<taxonomicName id="4C494D5A0233FFE8FC7E90FC7D8CFA2D" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[984,1267,1485,1511]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="309" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov.,">
<emphasis id="B93DEACB0233FFE8FC7E90FC7D8CFA2D" box="[984,1267,1485,1511]" italics="true" pageId="10" pageNumber="311">Myanophis thanlyinensis</emphasis>
</taxonomicName>
<emphasis id="B93DEACB0233FFE8FB5B90FC79B7F9C1" bold="true" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A20E57B00233FFE8FB5B90FC7C14FA2D" box="[1277,1387,1485,1511]" pageId="10" pageNumber="311" rank="species">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
is only known from the vicinity of its
<typeStatus id="54F2887B0233FFE8FD2090C07BC9F9C1" box="[646,694,1521,1547]" pageId="10" pageNumber="311">type</typeStatus>
locality. Given the little we know about this species, we classify
<taxonomicName id="4C494D5A0233FFE8FF319327782EF9E5" authority="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls, 2021" authorityName="Köhler &amp; Khaing &amp; Than &amp; Baranski &amp; Schell &amp; Greve &amp; Janke &amp; Pauls" authorityYear="2021" box="[151,337,1557,1583]" class="Reptilia" family="Homalopsidae" genus="Myanophis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="thanlyinensis" status="gen. nov., sp. nov.">
<emphasis id="B93DEACB0233FFE8FF319327782EF9E5" box="[151,337,1557,1583]" italics="true" pageId="10" pageNumber="311">M. thanlyinensis</emphasis>
</taxonomicName>
<emphasis id="B93DEACB0233FFE8FEFE93247B55F9E5" bold="true" box="[344,554,1557,1583]" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A20E57B00233FFE8FEFE93247B55F9E5" box="[344,554,1557,1583]" pageId="10" pageNumber="311" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
</emphasis>
as Data Deficient based on the IUCN Red List Categories and Criteria (
<bibRefCitation id="EFD84B280233FFE8FAF0932479ABF999" author="IUCN" pageId="10" pageNumber="311" refId="ref10541" refString="IUCN (2012) IUCN Red List Categories and Criteria, Version 3.1. 2 nd Edition. Available from: https: // portals. ucn. org / library / node / 10315 (accessed 17 November 2018)" type="url" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>