190 lines
21 KiB
XML
190 lines
21 KiB
XML
<document id="E911F60B3800E23B64F321D0CCB70B77" ID-CLB-Dataset="55724" ID-DOI="10.5281/zenodo.4507307" ID-GBIF-Dataset="72a1f456-f29c-471e-baf8-6c026cc900a1" ID-ISSN="0253-116X" ID-RefBankID="1643FFB2ED42FFC2FFE4FFE28E27FFAF" ID-Zenodo-Dep="4507307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612537541968" checkinUser="felipe" docAuthor="Assing, V." docDate="2013" docId="03E087C369717A26DDA829F70702FA7B" docLanguage="en" docName="LinzerbiolBeitr.45.1.0171-0190.pdf" docOrigin="Linzer biologische Beiträge 45 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Rugilus (Rugilus) curvatus Assing 2013, nov.sp." docType="treatment" docVersion="10" lastPageNumber="176" masterDocId="FFD9FFBB69747A23DD0B2B5B0548FFA4" masterDocTitle="A revision of Palaearctic and Oriental Rugilus. III. Five new species from the Palaearctic region and additional records (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="190" masterPageNumber="171" pageNumber="176" updateTime="1698867118016" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9775C0C85102F8F95DE30F3E67988286">A revision of Palaearctic and Oriental Rugilus. III. Five new species from the Palaearctic region and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="89D247E1F5C1B5D9BD4BDE7EAFB7B911">Assing, V.</mods:namePart>
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<mods:title id="AB37C6C47C66F58287BC83D7500FA697">Linzer biologische Beiträge</mods:title>
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<mods:part id="D3DE56ECA94D8CCF9CA4DC4CC5978222">
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<mods:date id="A183AF27D15B452EF973F1F23F989529">2013</mods:date>
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<mods:number id="9F9B266A9B3D5B11C596272E27D51805">2013-07-31</mods:number>
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<treatment id="03E087C369717A26DDA829F70702FA7B" ID-DOI="http://doi.org/10.5281/zenodo.6521058" ID-GBIF-Taxon="195191795" ID-Zenodo-Dep="6521058" LSID="urn:lsid:plazi:treatment:03E087C369717A26DDA829F70702FA7B" httpUri="http://treatment.plazi.org/id/03E087C369717A26DDA829F70702FA7B" lastPageNumber="176" pageId="5" pageNumber="176">
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<subSubSection id="C353655E69717A26DDA829F70758FD67" box="[163,528,684,707]" pageId="5" pageNumber="176" type="nomenclature">
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<paragraph id="8BF636D569717A26DDA829F70758FD67" blockId="5.[163,647,684,707]" box="[163,528,684,707]" pageId="5" pageNumber="176">
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<heading id="D0BE81B969717A26DDA829F70758FD67" box="[163,528,684,707]" fontSize="8" level="2" pageId="5" pageNumber="176" reason="2">
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<emphasis id="B93DEAC769717A26DDA829F70758FD67" bold="true" box="[163,528,684,707]" pageId="5" pageNumber="176">
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<taxonomicName id="4C494D5669717A26DDA829F704F6FD66" ID-CoL="4TNVY" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[163,446,684,707]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="curvatus" status="sp. nov." subGenus="Rugilus">
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<emphasis id="B93DEAC769717A26DDA829F705BCFD66" bold="true" box="[163,244,684,706]" italics="true" pageId="5" pageNumber="176">Rugilus</emphasis>
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(
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<emphasis id="B93DEAC769717A26DC0829F7041DFD66" bold="true" box="[259,341,684,706]" italics="true" pageId="5" pageNumber="176">Rugilus</emphasis>
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)
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<emphasis id="B93DEAC769717A26DC6F29F604F6FD66" bold="true" box="[356,446,685,706]" italics="true" pageId="5" pageNumber="176">curvatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E57BC69717A26DCCF29F60758FD67" box="[452,528,685,707]" pageId="5" pageNumber="176" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69717A26DF1D29F607CFFD67" box="[534,647,685,707]" pageId="5" pageNumber="176" type="description">
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<paragraph id="8BF636D569717A26DF1D29F607CFFD67" blockId="5.[163,647,684,707]" box="[534,647,685,707]" pageId="5" pageNumber="176">
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<heading id="D0BE81B969717A26DF1D29F607CFFD67" box="[534,647,685,707]" fontSize="8" level="2" pageId="5" pageNumber="176" reason="2">
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(
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<figureCitation id="13722A5069717A26DF1629F60737FD67" box="[541,639,685,707]" captionStart="Figs 8-12" captionStartId="6.[163,203,1060,1080]" captionTargetBox="[163,1046,197,1029]" captionTargetId="figure-210@6.[163,1098,197,1033]" captionTargetPageId="6" captionText="Figs 8-12: Rugilus curvatus nov.sp.: (8) forebody; (9) male sternite VII; (10) male sternite VIII; (11) aedeagus in lateral view; (12) aedeagus in ventral view. Scale bars: 8: 1.0 mm; 9-12: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4507311" httpUri="https://zenodo.org/record/4507311/files/figure.png" pageId="5" pageNumber="176">Figs 8-12</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69717A26DDA8298807B0FC8F" pageId="5" pageNumber="176" type="materials_examined">
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<paragraph id="8BF636D569717A26DDA8298807B0FC8F" blockId="5.[163,1071,722,811]" pageId="5" pageNumber="176">
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Type material:
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<materialsCitation id="3B213C8869717A26DC922988048DFCB1" ID-GBIF-Occurrence="3762949354" collectionCode="SNSD" collectorName="Schmidt & Jager" country="Nepal-Himal" determinerName="V. Assing" elevation="2400" location="Annapurna" pageId="5" pageNumber="176" specimenCount="1" specimenCount-male="1" stateProvince="N-Pokhara" typeStatus="holotype">
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<typeStatus id="54F2887769717A26DC92298804A5FD43" box="[409,493,723,743]" pageId="5" pageNumber="176" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D4FFD5C69717A26DCFE29890740FD43" box="[501,520,722,743]" pageId="5" pageNumber="176" type="male">♂</specimenCount>
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: "
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<collectingCountry id="F35E764569717A26DF16298807DEFD43" box="[541,662,723,743]" pageId="5" pageNumber="176">Nepal-Himal</collectingCountry>
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1996,
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<location id="8E96600E69717A26DFD329880675FD43" LSID="urn:lsid:plazi:treatment:03E087C369717A26DDA829F70702FA7B:8E96600E69717A26DFD329880675FD43" box="[728,829,723,743]" country="Nepal-Himal" name="Annapurna" pageId="5" pageNumber="176" stateProvince="N-Pokhara">Annapurna</location>
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mts., lg.
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<collectorName id="26BC530369717A26DE92298806AFFD43" box="[921,999,723,743]" pageId="5" pageNumber="176">Schmidt</collectorName>
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,
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<collectorName id="26BC530369717A26DEF92988016AFD43" box="[1010,1058,723,743]" pageId="5" pageNumber="176">Jäger</collectorName>
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/
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<collectingRegion id="498DF83769717A26DDA829B10440FD5A" box="[163,264,746,766]" pageId="5" pageNumber="176">N-Pokhara</collectingRegion>
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, oberh.
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<location id="8E96600E69717A26DC5829B104C6FD5A" LSID="urn:lsid:plazi:treatment:03E087C369717A26DDA829F70702FA7B:8E96600E69717A26DC5829B104C6FD5A" box="[339,398,746,766]" country="Nepal-Himal" name="Chipli" pageId="5" pageNumber="176" stateProvince="N-Pokhara">Chipli</location>
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,
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<quantity id="4CB19B3069717A26DC9329B104A9FD5A" box="[408,481,746,766]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="176" unit="m" value="2400.0">
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<elevation id="0064D1E669717A26DC9329B104A9FD5A" box="[408,481,746,766]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="176" unit="m" value="2400.0">2400 m</elevation>
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</quantity>
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, 22.4., Nyaulikharka BG / Holotypus ♂
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<originalDetermination id="A8578E2969717A26DE5029B30162FD5A" box="[859,1066,744,766]" pageId="5" pageNumber="176">
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<taxonomicName id="4C494D5669717A26DE5029B306BEFD5A" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[859,1014,744,766]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="curvatus" status="sp. nov.">
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<emphasis id="B93DEAC769717A26DE5029B306BEFD5A" box="[859,1014,744,766]" italics="true" pageId="5" pageNumber="176">Rugilus curvatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E57BC69717A26DEF629B10162FD5A" box="[1021,1066,746,766]" pageId="5" pageNumber="176" rank="species">sp.n.</taxonomicNameLabel>
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</originalDetermination>
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, det.
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<determinerName id="7E2ADA1169717A26DDC5285A0466FCB1" box="[206,302,769,789]" pageId="5" pageNumber="176">V. Assing</determinerName>
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2012" (
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<collectionCode id="ED58AE1069717A26DC74285A04F6FCB1" box="[383,446,769,789]" pageId="5" pageNumber="176">SNSD</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B213C8869717A26DCD9285A07B0FC8F" ID-GBIF-Occurrence="3762949361" pageId="5" pageNumber="176" typeStatus="paratype">
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<typeStatus id="54F2887769717A26DCD9285A076AFCB1" box="[466,546,769,789]" pageId="5" pageNumber="176" type="paratype">Paratype</typeStatus>
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♂: "Nepal-Himal 1996, Annapurna mts., lg. Schmidt, Jäger /
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<quantity id="4CB19B3069717A26DDEE284C045BFC8F" box="[229,275,791,811]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="176" unit="km" value="5.0">5 km</quantity>
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n. Siklis, Kyojo Kharka, 2600 mNN, 29.IV." (
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<collectionCode id="ED58AE1069717A26DFB6284C07A6FC8F" box="[701,750,791,811]" pageId="5" pageNumber="176">cAss</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69717A26DDA8286007EBFCCB" pageId="5" pageNumber="176" type="etymology">
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<paragraph id="8BF636D569717A26DDA8286007EBFCCB" blockId="5.[163,1070,827,879]" pageId="5" pageNumber="176">Etymology: The specific epithet (Latin, adjective: curved) refers to the shape of the ventral process of the aedeagus in lateral view.</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69717A26DDA828DB0621FBE4" pageId="5" pageNumber="176" type="description">
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<paragraph id="8BF636D569717A26DDA828DB064BFC11" blockId="5.[163,1071,896,949]" pageId="5" pageNumber="176">
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Description: Body length
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<quantity id="4CB19B3069717A26DF2E28DB07E1FC32" box="[549,681,896,918]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.3" metricValueMin="5.2" pageId="5" pageNumber="176" unit="mm" value="5.25" valueMax="5.3" valueMin="5.2">5.2-5.3 mm</quantity>
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; length of forebody 2.9-3.0 mm. Externally similar to
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<taxonomicName id="4C494D5669717A26DC7228FB049CFC11" authorityName="Assing" authorityYear="2013" box="[377,468,928,949]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="rectus">
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<emphasis id="B93DEAC769717A26DC7228FB049CFC11" box="[377,468,928,949]" italics="true" pageId="5" pageNumber="176">R. rectus</emphasis>
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</taxonomicName>
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, but distinguished as follows:
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</paragraph>
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<paragraph id="8BF636D569717A26DDA8289D014CFC78" blockId="5.[163,1028,966,988]" box="[163,1028,966,988]" pageId="5" pageNumber="176">
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Head (
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<figureCitation id="13722A5069717A26DDED289D046BFC78" box="[230,291,966,988]" captionStart="Figs 8-12" captionStartId="6.[163,203,1060,1080]" captionTargetBox="[163,1046,197,1029]" captionTargetId="figure-210@6.[163,1098,197,1033]" captionTargetPageId="6" captionText="Figs 8-12: Rugilus curvatus nov.sp.: (8) forebody; (9) male sternite VII; (10) male sternite VIII; (11) aedeagus in lateral view; (12) aedeagus in ventral view. Scale bars: 8: 1.0 mm; 9-12: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4507311" httpUri="https://zenodo.org/record/4507311/files/figure.png" pageId="5" pageNumber="176">Fig. 8</figureCitation>
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) 1.06-1.08 times as broad as long; punctation less extensively confluent.
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</paragraph>
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<paragraph id="8BF636D569717A26DDA828B60621FBE4" blockId="5.[163,1070,1004,1088]" pageId="5" pageNumber="176">
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♂: sternite VII (
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<figureCitation id="13722A5069717A26DC4C28B704CCFBA6" box="[327,388,1004,1026]" captionStart="Figs 8-12" captionStartId="6.[163,203,1060,1080]" captionTargetBox="[163,1046,197,1029]" captionTargetId="figure-210@6.[163,1098,197,1033]" captionTargetPageId="6" captionText="Figs 8-12: Rugilus curvatus nov.sp.: (8) forebody; (9) male sternite VII; (10) male sternite VIII; (11) aedeagus in lateral view; (12) aedeagus in ventral view. Scale bars: 8: 1.0 mm; 9-12: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4507311" httpUri="https://zenodo.org/record/4507311/files/figure.png" pageId="5" pageNumber="176">Fig. 9</figureCitation>
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) not distinctly modified; sternite VIII (
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<figureCitation id="13722A5069717A26DE1928B70613FBA6" box="[786,859,1004,1026]" captionStart="Figs 8-12" captionStartId="6.[163,203,1060,1080]" captionTargetBox="[163,1046,197,1029]" captionTargetId="figure-210@6.[163,1098,197,1033]" captionTargetPageId="6" captionText="Figs 8-12: Rugilus curvatus nov.sp.: (8) forebody; (9) male sternite VII; (10) male sternite VIII; (11) aedeagus in lateral view; (12) aedeagus in ventral view. Scale bars: 8: 1.0 mm; 9-12: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4507311" httpUri="https://zenodo.org/record/4507311/files/figure.png" pageId="5" pageNumber="176">Fig. 10</figureCitation>
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) with not very deep, broadly V-shaped posterior excision; aedeagus (
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<figureCitation id="13722A5069717A26DFA62F50066BFB85" box="[685,803,1035,1057]" captionStart="Figs 8-12" captionStartId="6.[163,203,1060,1080]" captionTargetBox="[163,1046,197,1029]" captionTargetId="figure-210@6.[163,1098,197,1033]" captionTargetPageId="6" captionText="Figs 8-12: Rugilus curvatus nov.sp.: (8) forebody; (9) male sternite VII; (10) male sternite VIII; (11) aedeagus in lateral view; (12) aedeagus in ventral view. Scale bars: 8: 1.0 mm; 9-12: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4507311" httpUri="https://zenodo.org/record/4507311/files/figure.png" pageId="5" pageNumber="176">Figs 11-12</figureCitation>
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)
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<quantity id="4CB19B3069717A26DE332F5006D1FB85" box="[824,921,1035,1057]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="176" unit="mm" value="0.65">0.65 mm</quantity>
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long; ventral process strongly curved in lateral view, apically acute in ventral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69717A26DDA82F0A05B1FADF" pageId="5" pageNumber="176" type="discussion">
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<paragraph id="8BF636D569717A26DDA82F0A05B1FADF" blockId="5.[163,1071,1105,1403]" pageId="5" pageNumber="176">
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Comparative notes: As can be inferred from the similar external and male sexual characters,
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<taxonomicName id="4C494D5669717A26DC6A2F2B049EFB21" authorityName="Assing" authorityYear="2013" box="[353,470,1136,1157]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="curvatus">
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<emphasis id="B93DEAC769717A26DC6A2F2B049EFB21" box="[353,470,1136,1157]" italics="true" pageId="5" pageNumber="176">R. curvatus</emphasis>
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</taxonomicName>
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is closely allied to
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<taxonomicName id="4C494D5669717A26DFAE2F2B06AAFB21" authority="(Dhaulagiri)" baseAuthorityName="Dhaulagiri" baseAuthorityYear="1975" box="[677,994,1135,1157]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="quadridentatus">
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<emphasis id="B93DEAC769717A26DFAE2F2B0614FB20" box="[677,860,1135,1157]" italics="true" pageId="5" pageNumber="176">R. quadridentatus</emphasis>
|
|
(
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<collectingRegion id="498DF83769717A26DE672F340693FB21" box="[876,987,1135,1157]" country="Nepal" name="Dhawalagiri" pageId="5" pageNumber="176">Dhaulagiri</collectingRegion>
|
|
)
|
|
</taxonomicName>
|
|
and
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|
<taxonomicName id="4C494D5669717A26D9132F2B0438FB00" authority="(Annapurna)" baseAuthorityName="Annapurna" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="rectus">
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<emphasis id="B93DEAC769717A26D9132F2B05A9FB00" italics="true" pageId="5" pageNumber="176">R. rectus</emphasis>
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|
(Annapurna)
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|
</taxonomicName>
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|
. From the former, it is distinguished by the absence of a median posterior excision of the male sternite VII, the different shape of the posterior excision of the male sternite VIII, and by the differently shaped ventral process of the aedeagus. From the latter, it differs by the much shallower posterior excision of the male sternite VIII and by the much shorter and distinctly curved ventral process of the aedeagus. For additional characters separating it from
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<taxonomicName id="4C494D5669717A26DF6E2E730781FA99" box="[613,713,1320,1341]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="rectus">
|
|
<emphasis id="B93DEAC769717A26DF6E2E730781FA99" box="[613,713,1320,1341]" italics="true" pageId="5" pageNumber="176">R. rectus</emphasis>
|
|
</taxonomicName>
|
|
see the description above. For illustrations of the male sexual characters of
|
|
<taxonomicName id="4C494D5669717A26DF742E1C067FFAFF" baseAuthorityName="COIFFAIT" baseAuthorityYear="1975" box="[639,823,1350,1372]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="quadridentatus">
|
|
<emphasis id="B93DEAC769717A26DF742E1C067FFAFF" box="[639,823,1350,1372]" italics="true" pageId="5" pageNumber="176">R. quadridentatus</emphasis>
|
|
</taxonomicName>
|
|
see
|
|
<figureCitation id="13722A5069717A26DE602E1D06E2FAF8" box="[875,938,1350,1372]" captionStart="Figs 1-7" captionStartId="4.[163,203,1098,1118]" captionTargetBox="[171,1067,199,1075]" captionTargetId="figure-251@4.[162,1069,197,1082]" captionTargetPageId="4" captionText="Figs 1-7: Rugilus rectus nov.sp. (1-6) and R. quadridentatus (COIFFAIT) (7): (1) habitus; (2) forebody; (3, 7) male sternite VII; (4) male sternite VIII; (5) aedeagus in lateral view; (6) ventral process of aedeagus in ventral view. Scale bars: 1-2: 1.0 mm; 3-7: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4507309" httpUri="https://zenodo.org/record/4507309/files/figure.png" pageId="5" pageNumber="176">Fig. 7</figureCitation>
|
|
and
|
|
<bibRefCitation id="EFD84B2469717A26DEE92E1D05BCFADF" author="ASSING V" pageId="5" pageNumber="176" pagination="115 - 190" refId="ref11068" refString="ASSING V. (2012 a): The Rugilus species of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 5: 115 - 190." type="journal article" year="2012">ASSING (2012a)</bibRefCitation>
|
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.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C353655E69717A26DDA82ED70702FA7B" pageId="5" pageNumber="176" type="distribution">
|
|
<paragraph id="8BF636D569717A26DDA82ED70702FA7B" blockId="5.[163,1070,1420,1503]" pageId="5" pageNumber="176">
|
|
Distribution and natural history The
|
|
<typeStatus id="54F2887769717A26DE422ED70625FA05" box="[841,877,1420,1441]" pageId="5" pageNumber="176">type</typeStatus>
|
|
specimens were found in two localities in the southern Annapurna range, some
|
|
<quantity id="4CB19B3069717A26DE242EF106D9FA64" box="[815,913,1450,1472]" metricMagnitude="4" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="5" pageNumber="176" unit="km" value="17.5" valueMax="20.0" valueMin="15.0">15-20 km</quantity>
|
|
to the north of Pokhara, at altitudes of 2400 and
|
|
<quantity id="4CB19B3069717A26DCFC2E92070EFA7B" box="[503,582,1481,1503]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="176" unit="m" value="2600.0">2600 m</quantity>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |