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<document id="137AA4B565E75247EDFF2E7CF7C6444B" ID-DOI="10.5281/zenodo.177713" ID-GBIF-Dataset="a00002c5-bbdf-4c1c-9e51-0a8aa5e78c72" ID-ISSN="1175-5326" ID-Zenodo-Dep="177713" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459870769079" checkinUser="plazi" docAuthor="Zhang, Kuiyan, Yang, Ding &amp; Ren, Dong" docDate="2007" docId="03E087B72003FFECDCDC2B923626FA0A" docLanguage="en" docName="zt01530p032.pdf" docOrigin="Zootaxa 1530" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Protapiocera convergens Zhang, Yang &amp; Ren, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="30" masterDocId="FFD9FFCF2001FFEFDC4B2B053776FF8A" masterDocTitle="Notes on the extinct family Protapioceridae, with description of a new species from China (Insecta: Diptera: Asiloidea)" masterLastPageNumber="32" masterPageNumber="27" pageNumber="29" updateTime="1698226581307" updateUser="plazi">
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<mods:title id="0BC5FA869A4B2D4772B30054BC2800BE">Notes on the extinct family Protapioceridae, with description of a new species from China (Insecta: Diptera: Asiloidea)</mods:title>
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<mods:namePart id="249186B8862C2ECDE7FE09318535B08A">Zhang, Kuiyan</mods:namePart>
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<mods:namePart id="46692DFDB721184ADE803D09B7F2AC52">Yang, Ding</mods:namePart>
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<mods:namePart id="4624A25E19B9C6857829D5014D0CEEB5">Ren, Dong</mods:namePart>
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<mods:part id="0E3F7E4CFB4E0F007C34AE67BF3D3163">
<mods:date id="986301FDC346015404D005E705A8FDD5">2007</mods:date>
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<treatment id="03E087B72003FFECDCDC2B923626FA0A" ID-DOI="http://doi.org/10.5281/zenodo.6244653" ID-GBIF-Taxon="119652080" ID-Zenodo-Dep="6244653" LSID="urn:lsid:plazi:treatment:03E087B72003FFECDCDC2B923626FA0A" httpUri="http://treatment.plazi.org/id/03E087B72003FFECDCDC2B923626FA0A" lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29">
<subSubSection id="C353652A2003FFEDDCDC2B923661FF5E" pageId="2" pageNumber="29" type="nomenclature">
<paragraph id="8BF636A12003FFEDDCDC2B923551FF3B" blockId="2.[151,551,151,212]" box="[151,551,151,177]" pageId="2" pageNumber="29">
<heading id="D0BE81CD2003FFEDDCDC2B923551FF3B" bold="true" box="[151,551,151,177]" fontSize="11" level="1" pageId="2" pageNumber="29" reason="1">
<emphasis id="B93DEAB32003FFEDDCDC2B923551FF3B" bold="true" box="[151,551,151,177]" pageId="2" pageNumber="29">
<taxonomicName id="4C494D222003FFEDDCDC2B9236B5FF3B" ID-CoL="8DHG8" box="[151,451,151,177]" class="Insecta" family="Protapioceridae" genus="Protapiocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="convergens" status="sp. nov.">
<emphasis id="B93DEAB32003FFEDDCDC2B9236B5FF3B" bold="true" box="[151,451,151,177]" italics="true" pageId="2" pageNumber="29">Protapiocera convergens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57C82003FFEDDD812B923551FF3B" box="[458,551,151,177]" pageId="2" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636A12003FFEDDCDC2BBF3661FF5E" blockId="2.[151,551,151,212]" box="[151,279,186,212]" pageId="2" pageNumber="29">
(
<figureCitation id="13722A242003FFEDDCD42BBF3679FF5E" box="[159,271,186,212]" captionStart-0="FIGURE 2" captionStart-1="FIGURES 3 4" captionStartId-0="2.[151,255,1805,1829]" captionStartId-1="3.[151,269,903,927]" captionTargetBox-0="[223,1363,1126,1781]" captionTargetBox-1="[194,1392,190,872]" captionTargetId-0="figure@2.[223,1363,1126,1781]" captionTargetId-1="figure@3.[156,1431,108,907]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 2. Protapiocera convergens sp. nov. Body, photograph." captionText-1="FIGURES 3 4. Protapiocera convergens sp. nov. 3. Body, camera lucida drawing. 4. Antennae." httpUri-0="https://zenodo.org/record/177715/files/figure.png" httpUri-1="https://zenodo.org/record/177716/files/figure.png" pageId="2" pageNumber="29">Figs. 24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C353652A2003FFEDDCDC2A0F3384FEAE" box="[151,1266,266,296]" pageId="2" pageNumber="29" type="diagnosis">
<paragraph id="8BF636A12003FFEDDCDC2A0F3384FEAE" blockId="2.[151,1437,266,1070]" box="[151,1266,266,296]" pageId="2" pageNumber="29">
<emphasis id="B93DEAB32003FFEDDCDC2A0F3661FEAE" bold="true" box="[151,279,266,292]" pageId="2" pageNumber="29">Diagnosis.</emphasis>
Ve i ns R 2+3 nearly converged with R4; base of cell m2 acute; apex of vein A2 obscure.
</paragraph>
</subSubSection>
<subSubSection id="C353652A2003FFEDDC8C2A3035FEFC04" pageId="2" pageNumber="29" type="description">
<paragraph id="8BF636A12003FFEDDC8C2A3036AFFE65" blockId="2.[151,1437,266,1070]" pageId="2" pageNumber="29">
<emphasis id="B93DEAB32003FFEDDC8C2A303629FEC5" bold="true" box="[199,351,309,335]" pageId="2" pageNumber="29">Description.</emphasis>
Body length 19.5 mm; wing length
<quantity id="4CB19B442003FFEDDF552A303401FEC5" box="[798,887,309,335]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="29" unit="mm" value="12.0">12 mm</quantity>
, width 3.5 mm. Body large, stout, elongated. Head large and round, as long as high. Eyes thinly pilose, with clear facets. Antenna short, missing basally; first flagellomere somewhat thick basally, its apical portion [= arista] long and thin. Thorax robust with hairs, about 1.3 times as high as head. Most parts of legs missed.
<collectingCountry id="F35E76312003FFEDDF122AA834E2FE4D" box="[857,916,429,455]" name="India" pageId="2" pageNumber="29">Hind</collectingCountry>
femur swollen, with dense short hairs; tibia narrow and slightly curved.
</paragraph>
<paragraph id="8BF636A12003FFEDDC8C2AF83354FCB4" blockId="2.[151,1437,266,1070]" pageId="2" pageNumber="29">Wing long, nearly as long as abdomen; venation on basal and middle part distinct, but absent or obscure on apical part of wing in natural state (not caused by preservation). Vein C ending nearly at wing tip. Vein Sc long, reaching vein C, far beyond middle of wing. Vein R1 ending nearly at apical 1/6 of wing. Vein Rs very short, originating from vein R1 at middle of wing; veins R2+3 and R4 short, curved forward and ending on vein R1 apically; veins R2+3 converged with R4; apex of vein R5 obsolete. Crossvein r-m situated at basal 2/5 of cell d. Vein M with 3 branches, all indistinct apically, thus all posterior cells open. Vein CuA1 arising from cell bm. Ve in C u P p re s en t. Ve in C uA 2 short and strongly curved backward. Apex of vein A2 obscure. Base of cell m2 obviously acute. Cell cup closed, nearly as wide as cell bm, apically truncate.</paragraph>
<paragraph id="8BF636A12003FFEDDC8C284935FEFC04" blockId="2.[151,1437,266,1070]" pageId="2" pageNumber="29">Abdomen long and subcylindrical, with 8 visible segments. Segment I shortest; segments II and III thickest, almost 3.3 times as wide as segment I.</paragraph>
</subSubSection>
<subSubSection id="C353652A2003FFEDDC8C289E3402FBA4" pageId="2" pageNumber="29" type="materials_examined">
<paragraph id="8BF636A12003FFEDDC8C289E32E0FC54" blockId="2.[151,1437,266,1070]" pageId="2" pageNumber="29">
<emphasis id="B93DEAB32003FFEDDC8C289E3636FC3F" bold="true" box="[199,320,923,949]" pageId="2" pageNumber="29">
<typeStatus id="54F288032003FFEDDC8C289E364DFC3F" box="[199,315,923,949]" pageId="2" pageNumber="29" type="holotype">Holotype</typeStatus>
.
</emphasis>
Body with wing in lateral view, part and counterpart. No. CNU-DB-LB2006001, 2006002, housed in Key Lab of Insect Evolution &amp; Environmental Changes, Capital Normal University, Beijing,
<collectingCountry id="F35E76312003FFEDD90328C132E4FC54" box="[1352,1426,964,990]" name="China" pageId="2" pageNumber="29">China</collectingCountry>
.
</paragraph>
<paragraph id="8BF636A12003FFEDDC8C28EE3402FBA4" blockId="2.[151,1437,266,1070]" pageId="2" pageNumber="29">
<emphasis id="B93DEAB32003FFEDDC8C28EE36A5FB8F" bold="true" box="[199,467,1003,1029]" pageId="2" pageNumber="29">Locality and horizon.</emphasis>
Collected from near Chaomidian Village, Beipiao City, Liaoning Province,
<collectingCountry id="F35E76312003FFEDD90528E932EEFB8C" box="[1358,1432,1004,1030]" name="China" pageId="2" pageNumber="29">China</collectingCountry>
. Upper Jurassic or Early Cretaceous Yixian Formation outcrop.
</paragraph>
</subSubSection>
<caption id="DF3666292003FFEDDCDC2C083424F8AF" box="[151,850,1805,1829]" httpUri="https://zenodo.org/record/177715/files/figure.png" pageId="2" pageNumber="29" targetBox="[223,1363,1126,1781]" targetPageId="2">
<paragraph id="8BF636A12003FFEDDCDC2C083424F8AF" blockId="2.[151,850,1805,1829]" box="[151,850,1805,1829]" pageId="2" pageNumber="29">
<emphasis id="B93DEAB32003FFEDDCDC2C08366FF8AF" bold="true" box="[151,281,1805,1829]" pageId="2" pageNumber="29">FIGURE 2.</emphasis>
<taxonomicName id="4C494D222003FFEDDD6B2C0B355AF8AF" box="[288,556,1806,1829]" class="Insecta" family="Protapioceridae" genus="Protapiocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="convergens" status="sp. nov.">
<emphasis id="B93DEAB32003FFEDDD6B2C0B355AF8AF" box="[288,556,1806,1829]" italics="true" pageId="2" pageNumber="29">Protapiocera convergens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57C82003FFEDDE782C0B35F3F8AF" box="[563,645,1806,1829]" pageId="2" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
Body, photograph.
</paragraph>
</caption>
<caption id="DF3666292002FFECDCDC288233EBFC14" box="[151,1181,903,927]" httpUri="https://zenodo.org/record/177716/files/figure.png" pageId="3" pageNumber="30" targetBox="[194,1392,190,872]" targetPageId="3">
<paragraph id="8BF636A12002FFECDCDC288233EBFC14" blockId="3.[151,1181,903,927]" box="[151,1181,903,927]" pageId="3" pageNumber="30">
<emphasis id="B93DEAB32002FFECDCDC28823635FC15" bold="true" box="[151,323,903,927]" pageId="3" pageNumber="30">FIGURES 34.</emphasis>
<taxonomicName id="4C494D222002FFECDD0228823523FC14" box="[329,597,903,926]" class="Insecta" family="Protapioceridae" genus="Protapiocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="convergens" status="sp. nov.">
<emphasis id="B93DEAB32002FFECDD0228823523FC14" box="[329,597,903,926]" italics="true" pageId="3" pageNumber="30">Protapiocera convergens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57C82002FFECDE17288235D9FC14" box="[604,687,903,926]" pageId="3" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
3. Body, camera lucida drawing. 4. Antennae.
</paragraph>
</caption>
<subSubSection id="C353652A2002FFECDC8C28EC3338FB8E" box="[199,1102,1001,1032]" pageId="3" pageNumber="30" type="etymology">
<paragraph id="8BF636A12002FFECDC8C28EC3338FB8E" blockId="3.[151,1436,1001,1408]" box="[199,1102,1001,1032]" pageId="3" pageNumber="30">
<emphasis id="B93DEAB32002FFECDC8C28EC3622FB89" bold="true" box="[199,340,1001,1027]" pageId="3" pageNumber="30">Etymology.</emphasis>
The species name refers to the convergence of veins R2+3 and R4.
</paragraph>
</subSubSection>
<subSubSection id="C353652A2002FFECDC8C2F113626FA0A" pageId="3" pageNumber="30" type="discussion">
<paragraph id="8BF636A12002FFECDC8C2F113626FA0A" blockId="3.[151,1436,1001,1408]" pageId="3" pageNumber="30">
<emphasis id="B93DEAB32002FFECDC8C2F113636FBA4" bold="true" box="[199,320,1044,1070]" pageId="3" pageNumber="30">Remarks.</emphasis>
The new species is similar to the two known species:
<taxonomicName id="4C494D222002FFECDF8C2F1033B0FBA4" box="[967,1222,1045,1070]" class="Insecta" family="Protapioceridae" genus="Protapiocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="megista">
<emphasis id="B93DEAB32002FFECDF8C2F1033B0FBA4" box="[967,1222,1045,1070]" italics="true" pageId="3" pageNumber="30">Protapiocera megista</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D222002FFECD94A2F10379BFBDC" class="Insecta" family="Protapioceridae" genus="Protapiocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="ischyra">
<emphasis id="B93DEAB32002FFECD94A2F10379BFBDC" italics="true" pageId="3" pageNumber="30">Protapiocera ischyra</emphasis>
</taxonomicName>
, but can be easily separated from
<emphasis id="B93DEAB32002FFECDECA2F383580FBDC" box="[641,758,1085,1110]" italics="true" pageId="3" pageNumber="30">P. m e g i s t a</emphasis>
by having the smaller body (body length 19.5 mm), vein R2+3 converged with vein R4, and the base of cell m2 obviously acute. In
<emphasis id="B93DEAB32002FFECDFB72F603304FBF4" box="[1020,1138,1125,1150]" italics="true" pageId="3" pageNumber="30">P. m e g i s t a</emphasis>
, the body is large (body length
<quantity id="4CB19B442002FFECDCA12F953632FB20" box="[234,324,1168,1194]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="30" unit="mm" value="31.0">31 mm</quantity>
), veins R2+3 and R4 are distinctly separated apically, and cell m2 is subquadrate basally (Ren 1998). Also,
<taxonomicName id="4C494D222002FFECDD7B2FBE36A7FB5E" box="[304,465,1211,1236]" class="Insecta" family="Protapioceridae" genus="Protapiocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="convergens" status="sp. nov.">
<emphasis id="B93DEAB32002FFECDD7B2FBE36A7FB5E" box="[304,465,1211,1236]" italics="true" pageId="3" pageNumber="30">P. convergens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57C82002FFECDD912FBF3543FB5E" box="[474,565,1210,1236]" pageId="3" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from
<emphasis id="B93DEAB32002FFECDF0D2FBE34C0FB5E" box="[838,950,1211,1236]" italics="true" pageId="3" pageNumber="30">P. i s c h y r a</emphasis>
by having vein R2+3 converged with vein R4, crossvein r-m located at basal 2/5 of cell d, cell br and cell bm subequal in width, vein A2 very short, not exceeding 1/2 length of cell bm. In
<emphasis id="B93DEAB32002FFECDE7F2E1435D4FAA0" box="[564,674,1297,1322]" italics="true" pageId="3" pageNumber="30">P. i s c h y r a</emphasis>
, veins R2+3 and R4 are distinctly separated apically, crossvein r-m is located at basal 1/4 of cell d, cell br is wider than cell bm, and vein A2 is long, exceeding 2/3 length of cell bm (Ren 1998).
</paragraph>
</subSubSection>
</treatment>
</document>