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<document ID-DOI="http://doi.org/10.5281/zenodo.3728406" ID-GBIF-Dataset="5b53b2fd-98cb-4450-96cc-b8e26c1e093f" ID-GBIF-Taxon="162716777" ID-Zenodo-Dep="3728406" approvalRequired="24" approvalRequired_for_document="1" approvalRequired_for_matCits="2" approvalRequired_for_textStreams="21" checkinTime="1584986693040" checkinUser="valdenar" docAuthor="Solis, M. Alma, Rodríguez, Eugenie Phillips-, Hallwachs, Winnie, Dapkey, Tanya &amp; Janzen, Daniel H." docDate="2020" docId="03E087B3DF268A3DFD11A402FB07FB5C" docLanguage="en" docName="PROC.ENTOMOL.SOC.WASH.122.1.147.pdf.imf" docOrigin="Proceedings of the Entomological Society of Washington 122 (1)" docStyle="DocumentStyle{}" docStyleName="procentomolsocwash.2010.journal_article" docTitle="Asturodes Amsel 1956" docType="treatment" docVersion="6" lastPageId="6" lastPageNumber="153" masterDocId="FFD9FFCBDF228A3BFFF6A034FFBDFFE5" masterDocTitle="Asturodes Amsel (Lepidoptera: Crambidae: Spilomelinae): Three New Species From The Western Hemisphere And Food Plant Records From Area De Conservación Guanacaste, Costa Rica" masterLastPageNumber="171" masterPageNumber="147" pageId="4" pageNumber="151" updateTime="1643439281101" updateUser="ExternalLinkService">
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<mods:title>Asturodes Amsel (Lepidoptera: Crambidae: Spilomelinae): Three New Species From The Western Hemisphere And Food Plant Records From Area De Conservación Guanacaste, Costa Rica</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Solis, M. Alma</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rodríguez, Eugenie Phillips-</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hallwachs, Winnie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dapkey, Tanya</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Janzen, Daniel H.</mods:namePart>
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<mods:title>Proceedings of the Entomological Society of Washington</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-02-29</mods:number>
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<mods:number>122</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.4289/0013-8797.122.1.147</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3728406" ID-GBIF-Taxon="162716777" ID-Zenodo-Dep="3728406" LSID="urn:lsid:plazi:treatment:03E087B3DF268A3DFD11A402FB07FB5C" httpUri="http://treatment.plazi.org/id/03E087B3DF268A3DFD11A402FB07FB5C" lastPageId="6" lastPageNumber="153" pageId="4" pageNumber="151">
<subSubSection box="[743,1170,1077,1102]" pageId="4" pageNumber="151" type="nomenclature">
<paragraph blockId="4.[743,1170,1077,1102]" box="[743,1170,1077,1102]" pageId="4" pageNumber="151">
<heading box="[743,1170,1077,1102]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="151" reason="4">
KEY TO
<taxonomicName authorityName="Amsel" authorityYear="1956" box="[866,1039,1077,1101]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis box="[866,1039,1077,1101]" italics="true" pageId="4" pageNumber="151">ASTURODES</emphasis>
</taxonomicName>
SPECIES
</heading>
</paragraph>
</subSubSection>
<keyStep pageId="4" pageNumber="151">
<subSubSection pageId="4" pageNumber="151" type="key">
<paragraph blockId="4.[702,1212,1126,1549]" pageId="4" pageNumber="151">
<keyLead pageId="4" pageNumber="151">
<key pageId="4" pageNumber="151">
1. Forewing with dark brown markings in radial area between reniform spot and postmedial line (
<figureCitation box="[938,1044,1192,1217]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 17. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="4" pageNumber="151">Figs.1, 3</figureCitation>
,
<figureCitation box="[1064,1129,1192,1217]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 811. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="4" pageNumber="151">8, 10</figureCitation>
); male genitalia with fibula sclerotized, finger-like process curving toward costal margin (
<figureCitation box="[745,895,1292,1317]" captionStart="Figs" captionStartId="7.[140,186,1638,1658]" captionTargetBox="[112,1154,198,1605]" captionTargetId="figure@7.[111,1159,196,1606]" captionTargetPageId="7" captionText="Figs. 1219. Asturodes male genitalia and phallus. 1213, A. junkoshimurae, USNM slide 115489. 1415, A. fimbriauralis, USNM slide 115491. 1617, A. bioalfae, USNM slide 115493. 18-19, A. encisoensis, USNM slide 115502, 115504." figureDoi="http://doi.org/10.5281/zenodo.3724754" httpUri="https://zenodo.org/record/3724754/files/figure.png" pageId="4" pageNumber="151">Figs. 12, 16</figureCitation>
); female genitalia without signa (
<figureCitation box="[811,947,1325,1350]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 2023. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="4" pageNumber="151">Figs. 20, 22</figureCitation>
) ............................... 2
</key>
</keyLead>
</paragraph>
</subSubSection>
<paragraph blockId="4.[702,1212,1126,1549]" pageId="4" pageNumber="151">
<keyLead pageId="4" pageNumber="151">
Forewing without dark brown markings in radial area between discal spot and postmedial line (
<figureCitation box="[794,903,1425,1450]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 17. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="4" pageNumber="151">Figs. 2, 4</figureCitation>
,
<figureCitation box="[919,978,1425,1450]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 811. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="4" pageNumber="151">9, 11</figureCitation>
); male genitalia with fibula short, not curving toward costal margin (
<figureCitation box="[743,884,1491,1516]" captionStart="Figs" captionStartId="7.[140,186,1638,1658]" captionTargetBox="[112,1154,198,1605]" captionTargetId="figure@7.[111,1159,196,1606]" captionTargetPageId="7" captionText="Figs. 1219. Asturodes male genitalia and phallus. 1213, A. junkoshimurae, USNM slide 115489. 1415, A. fimbriauralis, USNM slide 115491. 1617, A. bioalfae, USNM slide 115493. 18-19, A. encisoensis, USNM slide 115502, 115504." figureDoi="http://doi.org/10.5281/zenodo.3724754" httpUri="https://zenodo.org/record/3724754/files/figure.png" pageId="4" pageNumber="151">Figs. 14, 18</figureCitation>
); female genitalia with signa (
<figureCitation box="[743,870,1524,1549]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 2023. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="4" pageNumber="151">Figs. 21, 23</figureCitation>
) ................................................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep lastPageId="6" lastPageNumber="153" pageId="4" pageNumber="151">
<keyLead lastPageId="6" lastPageNumber="153" pageId="4" pageNumber="151">
<paragraph blockId="4.[702,1212,1558,1749]" pageId="4" pageNumber="151">
2. A light brown smudge in radial area between reniform spot and postmedial line; second lengthwise line of zigzag between Rs
<subScript attach="left" box="[1154,1164,1635,1651]" fontSize="7" pageId="4" pageNumber="151">4</subScript>
and M
<subScript attach="left" box="[761,771,1668,1684]" fontSize="7" pageId="4" pageNumber="151">1</subScript>
1/4 as long as first lengthwise line (
<figureCitation box="[745,826,1690,1715]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 17. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="4" pageNumber="151">Figs. 1</figureCitation>
,
<figureCitation box="[843,858,1690,1715]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 811. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="4" pageNumber="151">8</figureCitation>
); female genitalia with ductus bursae anterior third not of equal width
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3724750" ID-Zenodo-Dep="3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="5" pageNumber="152" startId="5.[140,186,1589,1609]" targetBox="[124,1143,202,1554]" targetPageId="5">
<paragraph blockId="5.[109,1161,1589,1693]" pageId="5" pageNumber="152">
Figs. 17. Adults,
<taxonomicName authorityName="Amsel" authorityYear="1956" box="[344,441,1590,1610]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis box="[344,441,1590,1610]" italics="true" pageId="5" pageNumber="152">Asturodes</emphasis>
</taxonomicName>
. 1,
<taxonomicName authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[479,653,1590,1610]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="junkoshimurae">
<emphasis box="[479,653,1590,1610]" italics="true" pageId="5" pageNumber="152">A. junkoshimurae</emphasis>
</taxonomicName>
, USNMENT01462908, forewing length=10.0 mm. 2,
<taxonomicName authorityName="Guenee" authorityYear="1854" box="[132,287,1618,1638]" class="Insecta" family="Crambidae" genus="Astura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="fimbriauralis">
<emphasis box="[132,287,1618,1638]" italics="true" pageId="5" pageNumber="152">A. fimbriauralis</emphasis>
</taxonomicName>
, USNMENT01462896, forewing length=11.0 mm. 3,
<taxonomicName authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[818,925,1618,1638]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="bioalfae">
<emphasis box="[818,925,1618,1638]" italics="true" pageId="5" pageNumber="152">A. bioalfae</emphasis>
</taxonomicName>
, USNMENT01462909, forewing length 0.9 mm. 4,
<emphasis box="[388,538,1645,1665]" italics="true" pageId="5" pageNumber="152">
<taxonomicName authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[388,533,1645,1665]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="encisoensis">A. encisoensis</taxonomicName>
,
</emphasis>
USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of
<taxonomicName authorityName="Guenee" authorityYear="1854" box="[194,355,1673,1693]" class="Insecta" family="Crambidae" genus="Astura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="fimbriauralis">
<emphasis box="[194,355,1673,1693]" italics="true" pageId="5" pageNumber="152">A. fimbriauralis</emphasis>
</taxonomicName>
[BMNH]. 6,
<taxonomicName authorityName="Guenee" authorityYear="1854" box="[495,655,1673,1693]" class="Insecta" family="Crambidae" genus="Astura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="fimbriauralis">
<emphasis box="[495,655,1673,1693]" italics="true" pageId="5" pageNumber="152">A. fimbriauralis</emphasis>
</taxonomicName>
, head,
<emphasis box="[731,751,1673,1693]" italics="true" pageId="5" pageNumber="152">lp</emphasis>
=labial palpi. 7,
<taxonomicName authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[920,1065,1673,1693]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="encisoensis">
<emphasis box="[920,1065,1673,1693]" italics="true" pageId="5" pageNumber="152">A. encisoensis</emphasis>
</taxonomicName>
, head.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3724752" ID-Zenodo-Dep="3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="6" pageNumber="153" startId="6.[191,237,903,923]" targetBox="[163,1209,197,870]" targetPageId="6">
<paragraph blockId="6.[159,1212,903,951]" pageId="6" pageNumber="153">
Figs. 811.
<taxonomicName authorityName="Amsel" authorityYear="1956" box="[330,430,903,923]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis box="[330,430,903,923]" italics="true" pageId="6" pageNumber="153">Asturodes</emphasis>
</taxonomicName>
stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8,
<taxonomicName authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[289,468,931,951]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="junkoshimurae">
<emphasis box="[289,468,931,951]" italics="true" pageId="6" pageNumber="153">A. junkoshimurae</emphasis>
</taxonomicName>
. 9,
<taxonomicName authorityName="Guenee" authorityYear="1854" box="[508,668,931,951]" class="Insecta" family="Crambidae" genus="Astura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="fimbriauralis">
<emphasis box="[508,668,931,951]" italics="true" pageId="6" pageNumber="153">A. fimbriauralis</emphasis>
</taxonomicName>
. 10,
<taxonomicName authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[722,834,931,951]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="bioalfae">
<emphasis box="[722,834,931,951]" italics="true" pageId="6" pageNumber="153">A. bioalfae</emphasis>
</taxonomicName>
. 11,
<taxonomicName authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[887,1032,931,951]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="encisoensis">
<emphasis box="[887,1032,931,951]" italics="true" pageId="6" pageNumber="153">A. encisoensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="6.[163,671,1007,1339]" pageId="6" pageNumber="153">
throughout (
<figureCitation box="[333,415,1007,1032]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 2023. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="6" pageNumber="153">Fig. 20</figureCitation>
) ................................ ...............
<taxonomicName authority="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey, 2020" authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[320,517,1040,1064]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="junkoshimurae" status="new species">
<emphasis box="[320,517,1040,1064]" italics="true" pageId="6" pageNumber="153">A. junkoshimurae</emphasis>
</taxonomicName>
,
<emphasis bold="true" box="[532,670,1039,1064]" pageId="6" pageNumber="153">
<taxonomicNameLabel box="[532,670,1039,1064]" pageId="6" pageNumber="153" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</keyLead>
<paragraph blockId="6.[163,671,1007,1339]" pageId="6" pageNumber="153">
<keyLead pageId="6" pageNumber="153">
A dark brown line in radial area between reniform spot and postmedial line; second lengthwise line of zigzag between Rs
<subScript attach="left" box="[659,669,1152,1168]" fontSize="7" pageId="6" pageNumber="153">4</subScript>
and M
<subScript attach="left" box="[267,277,1186,1202]" fontSize="7" pageId="6" pageNumber="153">1</subScript>
1/2 as long as the first lengthwise line (
<figureCitation box="[258,342,1210,1235]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 17. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="6" pageNumber="153">Figs. 3</figureCitation>
,
<figureCitation box="[361,392,1210,1235]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 811. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="6" pageNumber="153">10</figureCitation>
); female genitalia with ductus bursae anterior third of equal width throughout (
<figureCitation box="[330,410,1279,1304]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 2023. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="6" pageNumber="153">Fig. 22</figureCitation>
) .................................. ............................
<taxonomicName authority="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey, 2020" authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[399,520,1314,1338]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="bioalfae" status="new species">
<emphasis box="[399,520,1314,1338]" italics="true" pageId="6" pageNumber="153">A. bioalfae</emphasis>
</taxonomicName>
,
<emphasis bold="true" box="[535,669,1314,1339]" pageId="6" pageNumber="153">
<taxonomicNameLabel box="[535,669,1314,1339]" pageId="6" pageNumber="153" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<paragraph blockId="6.[160,670,1347,1749]" pageId="6" pageNumber="153">
<keyLead pageId="6" pageNumber="153">
<keyStep pageId="6" pageNumber="153">
3. Forewing with a “C” pattern on both sides of M
<subScript attach="left" box="[218,228,1391,1407]" fontSize="7" pageId="6" pageNumber="153">2</subScript>
near outer margin (
<figureCitation box="[441,506,1380,1405]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 811. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="6" pageNumber="153">Fig. 9</figureCitation>
, see red circle) with a dark brown line to reniform spot; reniform spot with three dark brown circles (
<figureCitation box="[202,282,1484,1509]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 17. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="6" pageNumber="153">Figs. 2</figureCitation>
,
<figureCitation box="[298,313,1484,1509]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 811. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="6" pageNumber="153">9</figureCitation>
); male genitalia with uncus tip round (
<figureCitation box="[273,354,1519,1544]" captionStart="Figs" captionStartId="7.[140,186,1638,1658]" captionTargetBox="[112,1154,198,1605]" captionTargetId="figure@7.[111,1159,196,1606]" captionTargetPageId="7" captionText="Figs. 1219. Asturodes male genitalia and phallus. 1213, A. junkoshimurae, USNM slide 115489. 1415, A. fimbriauralis, USNM slide 115491. 1617, A. bioalfae, USNM slide 115493. 18-19, A. encisoensis, USNM slide 115502, 115504." figureDoi="http://doi.org/10.5281/zenodo.3724754" httpUri="https://zenodo.org/record/3724754/files/figure.png" pageId="6" pageNumber="153">Fig. 14</figureCitation>
); female genitalia with signa on corpus bursae a small circle (
<figureCitation box="[574,656,1553,1578]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 2023. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="6" pageNumber="153">Fig. 21</figureCitation>
); food plant only
<taxonomicName box="[365,473,1588,1612]" class="Magnoliopsida" family="Rhamnaceae" genus="Colubrina" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="153" phylum="Tracheophyta" rank="genus">
<emphasis box="[365,473,1588,1612]" italics="true" pageId="6" pageNumber="153">Colubrina</emphasis>
</taxonomicName>
............................. .................................
<taxonomicName authority="Guenee" authorityName="Guenee" authorityYear="1854" box="[415,669,1623,1647]" class="Insecta" family="Crambidae" genus="Astura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="fimbriauralis">
<emphasis box="[415,582,1623,1647]" italics="true" pageId="6" pageNumber="153">A. fimbriauralis</emphasis>
Guenée
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="6.[160,670,1347,1749]" pageId="6" pageNumber="153">
<keyLead pageId="6" pageNumber="153">
Forewing with a dark brown line on M
<subScript attach="left" box="[607,617,1667,1683]" fontSize="7" pageId="6" pageNumber="153">2</subScript>
near margin, only a spot on M
<subScript attach="left" box="[497,507,1700,1716]" fontSize="7" pageId="6" pageNumber="153">1</subScript>
with no dark brown bar to reniform spot, reniform spot a
</keyLead>
</paragraph>
<paragraph blockId="6.[736,1212,1007,1209]" pageId="6" pageNumber="153">
dark brown “upside down U” over a “rightside up U” (
<figureCitation box="[869,946,1043,1068]" captionStart="Figs" captionStartId="5.[140,186,1589,1609]" captionTargetBox="[124,1143,202,1554]" captionTargetId="figure@5.[121,1148,196,1557]" captionTargetPageId="5" captionText="Figs. 17. Adults, Asturodes. 1, A. junkoshimurae, USNMENT01462908, forewing length=10.0 mm. 2, A. fimbriauralis, USNMENT01462896, forewing length=11.0 mm. 3, A. bioalfae, USNMENT01462909, forewing length 0.9 mm. 4, A. encisoensis, USNMENT01116302, forewing length=0.8 mm. 5, Holotype male of A. fimbriauralis [BMNH]. 6, A. fimbriauralis, head, lp=labial palpi. 7, A. encisoensis, head." figureDoi="http://doi.org/10.5281/zenodo.3724750" httpUri="https://zenodo.org/record/3724750/files/figure.png" pageId="6" pageNumber="153">Figs. 4</figureCitation>
,
<figureCitation box="[959,987,1043,1068]" captionStart="Figs" captionStartId="6.[191,237,903,923]" captionTargetBox="[163,1209,197,870]" captionTargetId="figure@6.[162,1210,196,871]" captionTargetPageId="6" captionText="Figs. 811. Asturodes stylized wing patterns, red circles indicate distinguishing patterns (see text and key). 8, A. junkoshimurae. 9, A. fimbriauralis. 10, A. bioalfae. 11, A. encisoensis." figureDoi="http://doi.org/10.5281/zenodo.3724752" httpUri="https://zenodo.org/record/3724752/files/figure.png" pageId="6" pageNumber="153">11</figureCitation>
); male genitalia with uncus tip pointed (
<figureCitation box="[939,1019,1079,1104]" captionStart="Figs" captionStartId="7.[140,186,1638,1658]" captionTargetBox="[112,1154,198,1605]" captionTargetId="figure@7.[111,1159,196,1606]" captionTargetPageId="7" captionText="Figs. 1219. Asturodes male genitalia and phallus. 1213, A. junkoshimurae, USNM slide 115489. 1415, A. fimbriauralis, USNM slide 115491. 1617, A. bioalfae, USNM slide 115493. 18-19, A. encisoensis, USNM slide 115502, 115504." figureDoi="http://doi.org/10.5281/zenodo.3724754" httpUri="https://zenodo.org/record/3724754/files/figure.png" pageId="6" pageNumber="153">Fig. 18</figureCitation>
); female genitalia with signa on corpus bursae in two dorsal longitudinal lines (
<figureCitation box="[930,1006,1149,1174]" captionStart="Figs" captionStartId="8.[191,237,1290,1310]" captionTargetBox="[163,1209,197,1257]" captionTargetId="figure@8.[162,1210,196,1258]" captionTargetPageId="8" captionText="Figs. 2023. Asturodes female genitalia. 20, A. junkoshimurae, USNM slide 115488. 21, A. fimbriauralis, USNM slide 115490. 22, A. bioalfae, USNM slide 115492. 23, A. encisoensis, USNM slide 115503." figureDoi="http://doi.org/10.5281/zenodo.3724756" httpUri="https://zenodo.org/record/3724756/files/figure.png" pageId="6" pageNumber="153">Fig. 23</figureCitation>
) .............................. ............................
<taxonomicName authority="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey, 2020" authorityName="Guanacaste &amp; Alma Solis &amp; Rodríguez &amp; Hallwachs &amp; Dapkey" authorityYear="2020" box="[918,1064,1185,1209]" class="Insecta" family="Crambidae" genus="Asturodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="153" phylum="Arthropoda" rank="species" species="encisoensis" status="new species">
<emphasis box="[918,1064,1185,1209]" italics="true" pageId="6" pageNumber="153">A. encisoensis</emphasis>
</taxonomicName>
,
<emphasis bold="true" box="[1078,1210,1184,1209]" pageId="6" pageNumber="153">
<taxonomicNameLabel box="[1078,1206,1184,1209]" pageId="6" pageNumber="153" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
</paragraph>
</treatment>
</document>