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<document id="7C246C42C4144CA0E9D2EF400558F818" ID-DOI="10.11646/zootaxa.5453.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="11233890" ID-ZooBank="DC7283A1-7BB1-4FD4-83FA-7F8A8779AD7E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1716282806434" checkinUser="plazi" docAuthor="Salinas-Velasco, Héctor Vicente, Valdez-Mondragón, Alejandro &amp; Villegas, Julián Bueno-" docDate="2024" docId="03E087A9BE41FFF9FF6FFE72FC91A1B1" docLanguage="en" docName="zootaxa.5453.2.4.pdf" docOrigin="Zootaxa 5453 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5453.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mexentypesa chiapas , Raven 1987" docType="treatment" docVersion="4" lastPageNumber="243" masterDocId="FFD9FFD1BE49FFF3FFF8FFA9FF80A37C" masterDocTitle="A new species of the spider genus Mexentypesa Raven (Araneae: Nemesiidae) from Hidalgo, Mexico and taxonomic comments on M. chiapas Raven" masterLastPageNumber="244" masterPageNumber="233" pageNumber="241" updateTime="1716396315016" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="25CEE48DF4A59268C9B1E8E839F5592A">A new species of the spider genus Mexentypesa Raven (Araneae: Nemesiidae) from Hidalgo, Mexico and taxonomic comments on M. chiapas Raven</mods:title>
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<mods:namePart id="6028D0080287D3B9B38FB22D159DC7A4">Salinas-Velasco, Héctor Vicente</mods:namePart>
<mods:affiliation id="6B9F60A5B4E73C5C6088B35D05DC36C9">Maestría en Ciencias en Biodiversidad y Conservación, Instituto de Ciencias Básicas e Ingeniería, Universidad Autónoma del Estado de Hidalgo. Carretera Pachuca-Tulancingo km 4.5, Mineral de la Reforma, C. P. 42184, Hidalgo, Mexico</mods:affiliation>
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<mods:namePart id="5D498DB2385F9A9BC7CC2B3AA8A5C3CE">Valdez-Mondragón, Alejandro</mods:namePart>
<mods:affiliation id="B92289C439A979912F3B3D4D255EC11F">Colección de Aracnología (CARCIB), Programa Académico de Planeación Ambiental y Conservación (PLAYCO), Centro de Investigaciones Biológicas del Noroeste CIBNOR, S. C. Km 1 Carretera a San Juan de La Costa “ El Comitán ”, La Paz, Baja California Sur, C. P. 23205, Mexico</mods:affiliation>
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<mods:namePart id="11C99045C331ED309C34B5461D2EA2A6">Villegas, Julián Bueno-</mods:namePart>
<mods:affiliation id="FF3465DBCB8EF571570DDC2A0B60ABA0">Laboratorio de Sistemática Animal, Centro de Investigaciones Biológicas, Universidad Autónoma del Estado de Hidalgo. Carretera Pachuca-Tulancingo km 4.5, Mineral de la Reforma, C. P. 42184, Hidalgo, Mexico</mods:affiliation>
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<treatment id="03E087A9BE41FFF9FF6FFE72FC91A1B1" LSID="urn:lsid:plazi:treatment:03E087A9BE41FFF9FF6FFE72FC91A1B1" httpUri="http://treatment.plazi.org/id/03E087A9BE41FFF9FF6FFE72FC91A1B1" lastPageId="10" lastPageNumber="243" pageId="8" pageNumber="241">
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<taxonomicName id="4C494D3CBE41FFFBFF6FFE72FDBAA288" ID-CoL="42KPL" authority="Raven, 1987" authorityName=", Raven" authorityYear="1987" box="[151,570,474,501]" class="Arachnida" family="Nemesiidae" genus="Mexentypesa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="241" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B93DEAADBE41FFFBFF6FFE72FDBAA288" bold="true" box="[151,570,474,501]" pageId="8" pageNumber="241">
<emphasis id="B93DEAADBE41FFFBFF6FFE72FE1AA289" bold="true" box="[151,410,474,501]" italics="true" pageId="8" pageNumber="241">Mexentypesa chiapas</emphasis>
<bibRefCitation id="EFD84B4EBE41FFFBFE59FE72FDBAA288" author="Raven, R. J." box="[417,570,474,501]" pageId="8" pageNumber="241" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven, 1987</bibRefCitation>
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<subSubSection id="C3536534BE41FFFBFF6FFE56FE9BA165" box="[151,283,511,537]" pageId="8" pageNumber="241" type="description">
<paragraph id="8BF636BFBE41FFFBFF6FFE56FE9BA165" blockId="8.[151,570,474,537]" box="[151,283,511,537]" pageId="8" pageNumber="241">
<figureCitation id="13722A3ABE41FFFBFF6FFE56FE9BA165" box="[151,283,511,537]" captionStart-0="FIGURES 28–31" captionStart-1="FIGURES 32–36" captionStart-2="FIGURES 37–44" captionStartId-0="7.[151,264,1936,1960]" captionStartId-1="8.[151,264,1792,1816]" captionStartId-2="9.[151,264,1288,1312]" captionTargetBox-0="[321,1267,741,1913]" captionTargetBox-1="[321,1267,658,1767]" captionTargetBox-2="[323,1264,181,1263]" captionTargetId-0="figure-288@7.[321,1267,741,1913]" captionTargetId-1="figure-168@8.[321,1267,658,1767]" captionTargetId-2="figure-18@9.[323,1264,181,1264]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURES 28–31. Mexentypesa chiapas Raven, 1987. Male: 28–29, Habitus, dorsal and ventral views, respectively. 30, Carapace, dorsal view. 31, Prosoma, ventral view showing coxae, sternum, labium, and endites (arrow indicates cuspules). Scale bars 0.5 mm (28, 29), 0.2 mm (30, 31). Modified of Valdez-Mondragón and Cortez-Roldán (2016)." captionText-1="FIGURES 32–36. Mexentypesa chiapas Raven, 1987. Male: 32–33, Left palp, prolateral and retrolateral views, respectively. 34, Left palp, ventral view. 35, Left palp, prolateral-apical view (arrow indicates apical spines on cymbium). 36, Detail of the bulb and embolus, ventral view. Scale bars 0.5 mm (35), 1 mm (32–34, 36). Modified of Valdez-Mondragón and Cortez-Roldán (2016)." captionText-2="FIGURES 37–44. Mexentypesa chiapas Raven, 1987. Male: 37, Tibia I, retrolateral view. 38, Tarsus I, retrolateral view. 39, Metatarsus and tarsus I, dorsal view (arrow indicates the small finger-shaped projection on retrolateral-ventral part of metatarsus). 40, Metatarsus I, retrolateral view (ditto). 41, Detail of the small finger-shaped projection on retrolateral-ventral part of metatarsus I. 42, Spination pattern on legs III and IV. 43, Endites, ventral view; detail of the cuspules. Female: 44, Seminal receptacles, dorsal view. Scale bars 0.2 mm (41), 0.5 mm (38, 40, 43), 1 mm (37, 39), 2 mm (42). Modified of Valdez- Mondragón and Cortez-Roldán (2016) (Figs 37–43), Modified of Raven (1987) (Fig 44)." figureDoi-0="http://doi.org/10.5281/zenodo.11233902" figureDoi-1="http://doi.org/10.5281/zenodo.11233904" figureDoi-2="http://doi.org/10.5281/zenodo.11233906" httpUri-0="https://zenodo.org/record/11233902/files/figure.png" httpUri-1="https://zenodo.org/record/11233904/files/figure.png" httpUri-2="https://zenodo.org/record/11233906/files/figure.png" pageId="8" pageNumber="241">Figs. 28–44</figureCitation>
</paragraph>
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<treatmentCitationGroup id="AB591191BE41FFFBFF6FFDEDFC35A127" box="[151,949,579,603]" pageId="8" pageNumber="241">
<treatmentCitation id="0AE810AEBE41FFFBFF6FFDEDFDACA127" author="Raven, R. J." box="[151,556,579,603]" page="358" pageId="8" pageNumber="241" year="1987">
<taxonomicName id="4C494D3CBE41FFFBFF6FFDEDFDACA127" ID-CoL="42KPL" authority="Raven, 1987: 358" authorityName=", Raven" authorityPageNumber="358" authorityYear="1987" box="[151,556,579,603]" class="Arachnida" family="Nemesiidae" genus="Mexentypesa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="241" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B93DEAADBE41FFFBFF6FFDEDFEF1A127" box="[151,369,579,603]" italics="true" pageId="8" pageNumber="241">Mexentypesa chiapas</emphasis>
<bibRefCitation id="EFD84B4EBE41FFFBFE8FFDEDFDACA127" author="Raven, R. J." box="[375,556,579,603]" pageId="8" pageNumber="241" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven, 1987: 358</bibRefCitation>
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, f. 1–9 (Description male and female).
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<treatmentCitationGroup id="AB591191BE41FFFAFF6FFDCBFC74A518" lastPageId="9" lastPageNumber="242" pageId="8" pageNumber="241">
<emphasis id="B93DEAADBE41FFFBFF6FFDCBFF25A106" box="[151,165,610,634]" italics="true" pageId="8" pageNumber="241">=</emphasis>
<treatmentCitation id="0AE810AEBE41FFFBFF5DFDCBFCDAA106" LSID="urn:lsid:zoobank.org:act:E78270D0-B3A8-4BC7-B2E3-FF617CBFA6A3" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." box="[165,858,610,634]" httpUri="http://treatment.plazi.org/id/86FE803C2B7476362BE9ACB0D9152EEF" page="84" pageId="8" pageNumber="241" year="2016">
<taxonomicName id="4C494D3CBE41FFFBFF5DFDCBFCDAA106" ID-CoL="682D7" authority="Valdez-Mondragon &amp; Cortez-Roldan, 2016: 84" authorityName="Valdez-Mondragon &amp; Cortez-Roldan" authorityPageNumber="84" authorityYear="2016" box="[165,858,610,634]" class="Arachnida" family="Euctenizidae" genus="Aptostichus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="241" phylum="Arthropoda" rank="species" species="sabinae">
<emphasis id="B93DEAADBE41FFFBFF5DFDCBFEF4A106" box="[165,372,610,634]" italics="true" pageId="8" pageNumber="241">Aptostichus sabinae</emphasis>
<materialsCitation id="3B213CE2BE41FFFBFE82FDCBFCDAA106" box="[378,858,610,634]" collectingDate="1950-06-25" collectorName="C. and N. Goodnight &amp; L. J. Stannard Cols." country="Mexico" elevation="900" location="Description" pageId="8" pageNumber="241" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="holotype">
<bibRefCitation id="EFD84B4EBE41FFFBFE82FDCBFCDAA106" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." box="[378,858,610,634]" pageId="8" pageNumber="241" pagination="81" refId="ref5872" refString="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R. (2016) On the trapdoor spiders of Mexico: description of the first new species of the spider genus Aptostichus from Mexico and the description of the female of Eucteniza zapatista (Araneae, Mygalomorphae, Euctenizidae). ZooKeys, 641, 81. https: // doi. org / 10.3897 / zookeys. 641.10521" type="journal article" year="2016">Valdez-Mondragón &amp; Cortez-Roldán, 2016: 84</bibRefCitation>
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, f. 1–20 (
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male and female), syn. n.
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<collectingCountry id="F35E762FBE40FFFAFEB3F98EFE3EA53D" box="[331,446,1575,1601]" name="Mexico" pageId="9" pageNumber="242">MEXICO</collectingCountry>
:
<collectingRegion id="498DF85DBE40FFFAFE30F98FFDA7A53C" box="[456,551,1574,1600]" country="Mexico" name="Chiapas" pageId="9" pageNumber="242">Chiapas</collectingRegion>
:
<specimenCount id="9D4FFD36BE40FFFAFDC8F98EFDD3A53D" box="[560,595,1575,1601]" count="1" pageId="9" pageNumber="242" type="male">1♂</specimenCount>
male
<typeStatus id="54F2881DBE40FFFAFD6FF98FFD7CA53C" box="[663,764,1574,1600]" pageId="9" pageNumber="242" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D4FFD36BE40FFFAFCFEF98EFCA9A53D" box="[774,809,1575,1601]" count="1" pageId="9" pageNumber="242" type="female">1♀</specimenCount>
female
<typeStatus id="54F2881DBE40FFFAFC7BF98EFC63A53D" box="[899,995,1575,1601]" pageId="9" pageNumber="242" type="paratype">paratype</typeStatus>
(AMNH), from Ocosingo (altitude
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<elevation id="0064D18CBE40FFFAFA8AF98EFF30A519" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="242" unit="m" value="900.0">900 m</elevation>
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;
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<collectingDate id="EFB3E997BE40FFFAFF44F9E2FEDBA519" box="[188,347,1610,1637]" pageId="9" pageNumber="242" value="1950-06-25">June 25, 1950</collectingDate>
</date>
,
<collectorName id="26BC5369BE40FFFAFE9EF9E2FDD3A518" box="[358,595,1610,1637]" pageId="9" pageNumber="242">C. and N. Goodnight</collectorName>
,
<collectorName id="26BC5369BE40FFFAFDA6F9E2FCBCA518" box="[606,828,1610,1637]" pageId="9" pageNumber="242">L. J. Stannard Cols.</collectorName>
(not examined).
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FIGURES 32–36.
<taxonomicName id="4C494D3CBE41FFFBFEA3F8A9FD47A464" authority="Raven, 1987" authorityName=", Raven" authorityYear="1987" box="[347,711,1791,1816]" class="Arachnida" family="Nemesiidae" genus="Mexentypesa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="241" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B93DEAADBE41FFFBFEA3F8A9FDBAA46B" bold="true" box="[347,570,1791,1815]" italics="true" pageId="8" pageNumber="241">Mexentypesa chiapas</emphasis>
<bibRefCitation id="EFD84B4EBE41FFFBFDB9F8A9FD47A464" author="Raven, R. J." box="[577,711,1792,1816]" pageId="8" pageNumber="241" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven, 1987</bibRefCitation>
</taxonomicName>
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. Male: 32–33, Left palp, prolateral and retrolateral views, respectively. 34, Left palp, ventral view. 35, Left palp, prolateral-apical view (arrow indicates apical spines on cymbium). 36, Detail of the bulb and embolus, ventral view. Scale bars 0.5 mm (35), 1 mm (32–34, 36). Modified of
<bibRefCitation id="EFD84B4EBE41FFFBFBEBF8E1FF5DA4F8" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." pageId="8" pageNumber="241" pagination="81" refId="ref5872" refString="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R. (2016) On the trapdoor spiders of Mexico: description of the first new species of the spider genus Aptostichus from Mexico and the description of the female of Eucteniza zapatista (Araneae, Mygalomorphae, Euctenizidae). ZooKeys, 641, 81. https: // doi. org / 10.3897 / zookeys. 641.10521" type="journal article" year="2016">Valdez-Mondragón and Cortez-Roldán (2016)</bibRefCitation>
.
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</caption>
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<paragraph id="8BF636BFBE40FFFAFF6FFAA1FB9DA6A8" blockId="9.[151,1437,1288,1492]" pageId="9" pageNumber="242">
<emphasis id="B93DEAADBE40FFFAFF6FFAA1FD5BA65C" bold="true" box="[151,731,1288,1312]" pageId="9" pageNumber="242">
FIGURES 37–44.
<taxonomicName id="4C494D3CBE40FFFAFE99FAA0FD56A65C" authority="Raven, 1987" authorityName=", Raven" authorityYear="1987" box="[353,726,1288,1312]" class="Arachnida" family="Nemesiidae" genus="Mexentypesa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="242" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B93DEAADBE40FFFAFE99FAA0FDC3A65C" bold="true" box="[353,579,1288,1312]" italics="true" pageId="9" pageNumber="242">Mexentypesa chiapas</emphasis>
<bibRefCitation id="EFD84B4EBE40FFFAFDB5FAA1FD56A65C" author="Raven, R. J." box="[589,726,1288,1312]" pageId="9" pageNumber="242" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven, 1987</bibRefCitation>
</taxonomicName>
.
</emphasis>
Male: 37, Tibia I, retrolateral view. 38, Tarsus I, retrolateral view. 39, Metatarsus and tarsus I, dorsal view (arrow indicates the small finger-shaped projection on retrolateral-ventral part of metatarsus). 40, Metatarsus I, retrolateral view (ditto). 41, Detail of the small finger-shaped projection on retrolateral-ventral part of metatarsus I. 42, Spination pattern on legs III and IV. 43, Endites, ventral view; detail of the cuspules. Female: 44, Seminal receptacles, dorsal view. Scale bars 0.2 mm (41), 0.5 mm (38, 40, 43), 1 mm (37, 39), 2 mm (42). Modified of ValdezMondragón and Cortez-Roldán (2016) (Figs 37–43), Modified of
<bibRefCitation id="EFD84B4EBE40FFFAFCCBFA14FC3DA6A8" author="Raven, R. J." box="[819,957,1468,1492]" pageId="9" pageNumber="242" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven (1987)</bibRefCitation>
(Fig 44).
</paragraph>
</caption>
<subSubSection id="C3536534BE40FFFAFF3FF9C7FE47A5AC" pageId="9" pageNumber="242" type="materials_examined">
<paragraph id="8BF636BFBE40FFFAFF3FF9C7FE47A5AC" blockId="9.[151,1437,1574,2033]" pageId="9" pageNumber="242">
<emphasis id="B93DEAADBE40FFFAFF3FF9C7FE76A5F4" bold="true" box="[199,502,1646,1672]" pageId="9" pageNumber="242">Other material examined</emphasis>
.
<materialsCitation id="3B213CE2BE40FFFAFDFDF9C6FE43A5AC" collectingDate="2014-04-12" collectorName="J. Mendoza &amp; J. Cruz &amp; S. Davlantes &amp; M. Minkton Cols." country="Mexico" elevation="1919" location="Municipio Huautla de Jimenez" municipality="Cueva Li Nita" pageId="9" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
<collectingCountry id="F35E762FBE40FFFAFDFDF9C6FDF8A5F5" box="[517,632,1647,1673]" name="Mexico" pageId="9" pageNumber="242">MEXICO</collectingCountry>
:
<collectingRegion id="498DF85DBE40FFFAFD7CF9C6FD5CA5F5" box="[644,732,1647,1673]" country="Mexico" name="Oaxaca" pageId="9" pageNumber="242">Oaxaca</collectingRegion>
:
<specimenCount id="9D4FFD36BE40FFFAFD10F9C6FC8AA5F5" box="[744,778,1647,1673]" count="1" pageId="9" pageNumber="242" type="male">1♂</specimenCount>
male (CNAN-T1121) from
<collectingMunicipality id="6B92ACC5BE40FFFAFBB3F9C6FB6BA5F4" box="[1099,1259,1646,1673]" pageId="9" pageNumber="242">Cueva Li Nita</collectingMunicipality>
(lat 18.14767°, lon -96.79844°,
<quantity id="4CB19B5ABE40FFFAFEAAF93AFE2CA5D1" box="[338,428,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.919" pageId="9" pageNumber="242" unit="m" value="1919.0">
<elevation id="0064D18CBE40FFFAFEAAF93AFE2CA5D1" box="[338,428,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.919" pageId="9" pageNumber="242" unit="m" value="1919.0">1919 m</elevation>
</quantity>
),
<location id="8E966064BE40FFFAFE3AF93BFCA4A5D0" LSID="urn:lsid:plazi:treatment:03E087A9BE41FFF9FF6FFE72FC91A1B1:8E966064BE40FFFAFE3AF93BFCA4A5D0" box="[450,804,1682,1708]" country="Mexico" municipality="Cueva Li Nita" name="Municipio Huautla de Jimenez" pageId="9" pageNumber="242" stateProvince="Oaxaca">Municipio Huautla de Jiménez</location>
,
<date id="FFF7107FBE40FFFAFCD7F93BFC5BA5D1" box="[815,987,1682,1709]" pageId="9" pageNumber="242" value="2014-04-12">
<collectingDate id="EFB3E997BE40FFFAFCD7F93BFC5BA5D1" box="[815,987,1682,1709]" pageId="9" pageNumber="242" value="2014-04-12">April 12, 2014</collectingDate>
</date>
,
<collectorName id="26BC5369BE40FFFAFC10F93AFBEEA5D0" box="[1000,1134,1682,1709]" pageId="9" pageNumber="242">J. Mendoza</collectorName>
,
<collectorName id="26BC5369BE40FFFAFB83F93AFB4FA5D1" box="[1147,1231,1683,1709]" pageId="9" pageNumber="242">J. Cruz</collectorName>
,
<collectorName id="26BC5369BE40FFFAFB25F93AFAEFA5D0" box="[1245,1391,1682,1709]" pageId="9" pageNumber="242">S. Davlantes</collectorName>
,
<collectorName id="26BC5369BE40FFFAFA84F93AFEB9A5AC" pageId="9" pageNumber="242">M. Minkton Cols.</collectorName>
(examined)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536534BE40FFFAFF3FF973FC1DA588" box="[199,925,1754,1781]" pageId="9" pageNumber="242" type="diagnosis">
<paragraph id="8BF636BFBE40FFFAFF3FF973FC1DA588" blockId="9.[151,1437,1574,2033]" box="[199,925,1754,1781]" pageId="9" pageNumber="242">
<emphasis id="B93DEAADBE40FFFAFF3FF973FEC2A588" bold="true" box="[199,322,1754,1780]" pageId="9" pageNumber="242">Diagnosis.</emphasis>
See comparative diagnosis of
<taxonomicName id="4C494D3CBE40FFFAFD60F972FCCCA588" authorityName="Salinas-Velasco &amp; Valdez-Mondragón &amp; Villegas" authorityYear="2024" box="[664,844,1754,1780]" class="Arachnida" family="Nemesiidae" genus="Mexentypesa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="242" phylum="Arthropoda" rank="species" species="hidalguensis">
<emphasis id="B93DEAADBE40FFFAFD60F972FCCCA588" box="[664,844,1754,1780]" italics="true" pageId="9" pageNumber="242">M. hidalguensis</emphasis>
</taxonomicName>
above.
</paragraph>
</subSubSection>
<subSubSection id="C3536534BE40FFFAFF3FF957FB08A440" pageId="9" pageNumber="242" type="description">
<paragraph id="8BF636BFBE40FFFAFF3FF957FB08A440" blockId="9.[151,1437,1574,2033]" pageId="9" pageNumber="242">
<emphasis id="B93DEAADBE40FFFAFF3FF957FED9A464" bold="true" box="[199,345,1790,1816]" pageId="9" pageNumber="242">Description.</emphasis>
See
<bibRefCitation id="EFD84B4EBE40FFFAFE65F957FC33A464" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." box="[413,947,1790,1816]" pageId="9" pageNumber="242" pagination="81" refId="ref5872" refString="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R. (2016) On the trapdoor spiders of Mexico: description of the first new species of the spider genus Aptostichus from Mexico and the description of the female of Eucteniza zapatista (Araneae, Mygalomorphae, Euctenizidae). ZooKeys, 641, 81. https: // doi. org / 10.3897 / zookeys. 641.10521" type="journal article" year="2016">Valdez-Mondragón and Cortez-Roldán (2016</bibRefCitation>
: figs 1–20) for a complete and detailed description of the male, and
<bibRefCitation id="EFD84B4EBE40FFFAFE2EF88AFDEAA441" author="Raven, R. J." box="[470,618,1827,1853]" pageId="9" pageNumber="242" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven (1987</bibRefCitation>
: fig. 8) for a complete description of the female.
</paragraph>
</subSubSection>
<subSubSection id="C3536534BE40FFF9FF3FF8EEFB19A1D5" lastPageId="10" lastPageNumber="243" pageId="9" pageNumber="242" type="discussion">
<paragraph id="8BF636BFBE40FFF9FF3FF8EEFE7AA165" blockId="9.[151,1437,1574,2033]" lastBlockId="10.[151,1437,151,717]" lastPageId="10" lastPageNumber="243" pageId="9" pageNumber="242">
<emphasis id="B93DEAADBE40FFFAFF3FF8EEFEBBA41D" bold="true" box="[199,315,1863,1889]" pageId="9" pageNumber="242">Remarks.</emphasis>
Based on the original description and illustrations by
<bibRefCitation id="EFD84B4EBE40FFFAFC61F8EFFA18A41C" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." box="[921,1432,1862,1888]" pageId="9" pageNumber="242" pagination="81" refId="ref5872" refString="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R. (2016) On the trapdoor spiders of Mexico: description of the first new species of the spider genus Aptostichus from Mexico and the description of the female of Eucteniza zapatista (Araneae, Mygalomorphae, Euctenizidae). ZooKeys, 641, 81. https: // doi. org / 10.3897 / zookeys. 641.10521" type="journal article" year="2016">Valdez-Mondragón and Cortez-Roldán (2016</bibRefCitation>
: figs 1–20), the male
<typeStatus id="54F2881DBE40FFFAFE87F8C3FE61A4F8" box="[383,481,1898,1924]" pageId="9" pageNumber="242" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C494D3CBE40FFFAFDFEF8C3FB55A4F8" authority="Valdez-Mondragon &amp; Cortez-Roldan, 2016" authorityName="Valdez-Mondragon &amp; Cortez-Roldan" authorityYear="2016" box="[518,1237,1898,1925]" class="Arachnida" family="Euctenizidae" genus="Aptostichus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="242" phylum="Arthropoda" rank="species" species="sabinae">
<emphasis id="B93DEAADBE40FFFAFDFEF8C3FD67A4F8" box="[518,743,1898,1924]" italics="true" pageId="9" pageNumber="242">Aptostichus sabinae</emphasis>
<bibRefCitation id="EFD84B4EBE40FFFAFD16F8C3FB55A4F8" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." box="[750,1237,1898,1925]" pageId="9" pageNumber="242" pagination="81" refId="ref5872" refString="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R. (2016) On the trapdoor spiders of Mexico: description of the first new species of the spider genus Aptostichus from Mexico and the description of the female of Eucteniza zapatista (Araneae, Mygalomorphae, Euctenizidae). ZooKeys, 641, 81. https: // doi. org / 10.3897 / zookeys. 641.10521" type="journal article" year="2016">Valdez-Mondragón &amp; Cortez-Roldán, 2016</bibRefCitation>
</taxonomicName>
is identical to the male
<typeStatus id="54F2881DBE40FFFAFF2CF827FEB6A4D4" box="[212,310,1934,1960]" pageId="9" pageNumber="242" type="holotype">holotype</typeStatus>
and description by
<bibRefCitation id="EFD84B4EBE40FFFAFDEAF826FD2BA4D5" author="Raven, R. J." box="[530,683,1935,1961]" pageId="9" pageNumber="242" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven (1987)</bibRefCitation>
of
<taxonomicName id="4C494D3CBE40FFFAFD28F826FBD7A4D5" authority="Raven, 1987" authorityName=", Raven" authorityYear="1987" box="[720,1111,1934,1961]" class="Arachnida" family="Nemesiidae" genus="Mexentypesa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="242" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B93DEAADBE40FFFAFD28F826FC40A4D4" box="[720,960,1934,1960]" italics="true" pageId="9" pageNumber="242">Mexentypesa chiapas</emphasis>
<bibRefCitation id="EFD84B4EBE40FFFAFC3FF826FBD7A4D5" author="Raven, R. J." box="[967,1111,1935,1961]" pageId="9" pageNumber="242" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven, 1987</bibRefCitation>
</taxonomicName>
. For this reason, we consider
<taxonomicName id="4C494D3CBE40FFFAFF6FF81AFE8EA4B0" authorityName="Valdez-Mondragon &amp; Cortez-Roldan" authorityYear="2016" box="[151,270,1970,1997]" class="Arachnida" family="Euctenizidae" genus="Aptostichus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="242" phylum="Arthropoda" rank="species" species="sabinae">
<emphasis id="B93DEAADBE40FFFAFF6FF81AFE8EA4B0" box="[151,270,1970,1997]" italics="true" pageId="9" pageNumber="242">A. sabinae</emphasis>
</taxonomicName>
as a junior synonym of
<emphasis id="B93DEAADBE40FFFAFDDCF81AFD29A4B0" box="[548,681,1970,1996]" italics="true" pageId="9" pageNumber="242">
<taxonomicName id="4C494D3CBE40FFFAFDDCF81AFD26A4B0" authorityName=", Raven" authorityYear="1987" box="[548,678,1970,1996]" class="Arachnida" family="Nemesiidae" genus="Mexentypesa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="242" phylum="Arthropoda" rank="species" species="chiapas">M. chiapas</taxonomicName>
.
</emphasis>
Although
<bibRefCitation id="EFD84B4EBE40FFFAFCDBF81AFC3DA4B1" author="Raven, R. J." box="[803,957,1971,1997]" pageId="9" pageNumber="242" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven (1987)</bibRefCitation>
reports that the male
<typeStatus id="54F2881DBE40FFFAFB4EF81BFA98A4B0" box="[1206,1304,1970,1996]" pageId="9" pageNumber="242" type="holotype">holotype</typeStatus>
used in the description of the species had both embolus broken, the rest of the shape of the male palp, as well as the other body features, are the same (Robert Raven, pers. com.): a unique embolus, very long, slender, and sigmoidal, the palp tibiae with ventral and prolateral massive spines (
<figureCitation id="13722A3ABE43FFF9FCF8FF12FC02A3A9" box="[768,898,187,213]" captionStart="FIGURES 32–36" captionStartId="8.[151,264,1792,1816]" captionTargetBox="[321,1267,658,1767]" captionTargetId="figure-168@8.[321,1267,658,1767]" captionTargetPageId="8" captionText="FIGURES 32–36. Mexentypesa chiapas Raven, 1987. Male: 32–33, Left palp, prolateral and retrolateral views, respectively. 34, Left palp, ventral view. 35, Left palp, prolateral-apical view (arrow indicates apical spines on cymbium). 36, Detail of the bulb and embolus, ventral view. Scale bars 0.5 mm (35), 1 mm (32–34, 36). Modified of Valdez-Mondragón and Cortez-Roldán (2016)." figureDoi="http://doi.org/10.5281/zenodo.11233904" httpUri="https://zenodo.org/record/11233904/files/figure.png" pageId="10" pageNumber="243">Figs 32–36</figureCitation>
) (broken in
<bibRefCitation id="EFD84B4EBE43FFF9FBF6FF12FB19A3A9" author="Raven, R. J." box="[1038,1177,187,213]" pageId="10" pageNumber="243" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven 1987</bibRefCitation>
: fig. 4); the metatarsus I with a retrolateral-ventral small finger-shaped projection (
<figureCitation id="13722A3ABE43FFF9FCB8FF76FC45A385" box="[832,965,223,249]" captionStart="FIGURES 37–44" captionStartId="9.[151,264,1288,1312]" captionTargetBox="[323,1264,181,1263]" captionTargetId="figure-18@9.[323,1264,181,1264]" captionTargetPageId="9" captionText="FIGURES 37–44. Mexentypesa chiapas Raven, 1987. Male: 37, Tibia I, retrolateral view. 38, Tarsus I, retrolateral view. 39, Metatarsus and tarsus I, dorsal view (arrow indicates the small finger-shaped projection on retrolateral-ventral part of metatarsus). 40, Metatarsus I, retrolateral view (ditto). 41, Detail of the small finger-shaped projection on retrolateral-ventral part of metatarsus I. 42, Spination pattern on legs III and IV. 43, Endites, ventral view; detail of the cuspules. Female: 44, Seminal receptacles, dorsal view. Scale bars 0.2 mm (41), 0.5 mm (38, 40, 43), 1 mm (37, 39), 2 mm (42). Modified of Valdez- Mondragón and Cortez-Roldán (2016) (Figs 37–43), Modified of Raven (1987) (Fig 44)." figureDoi="http://doi.org/10.5281/zenodo.11233906" httpUri="https://zenodo.org/record/11233906/files/figure.png" pageId="10" pageNumber="243">Figs 39–41</figureCitation>
), also described by
<bibRefCitation id="EFD84B4EBE43FFF9FB53FF76FAC1A385" author="Raven, R. J." box="[1195,1345,223,249]" pageId="10" pageNumber="243" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven (1987</bibRefCitation>
: fig. 2); the tibiae and metatarsi III and IV have many ventral spines (
<figureCitation id="13722A3ABE43FFF9FCB1FEAAFC1DA261" box="[841,925,259,285]" captionStart="FIGURES 37–44" captionStartId="9.[151,264,1288,1312]" captionTargetBox="[323,1264,181,1263]" captionTargetId="figure-18@9.[323,1264,181,1264]" captionTargetPageId="9" captionText="FIGURES 37–44. Mexentypesa chiapas Raven, 1987. Male: 37, Tibia I, retrolateral view. 38, Tarsus I, retrolateral view. 39, Metatarsus and tarsus I, dorsal view (arrow indicates the small finger-shaped projection on retrolateral-ventral part of metatarsus). 40, Metatarsus I, retrolateral view (ditto). 41, Detail of the small finger-shaped projection on retrolateral-ventral part of metatarsus I. 42, Spination pattern on legs III and IV. 43, Endites, ventral view; detail of the cuspules. Female: 44, Seminal receptacles, dorsal view. Scale bars 0.2 mm (41), 0.5 mm (38, 40, 43), 1 mm (37, 39), 2 mm (42). Modified of Valdez- Mondragón and Cortez-Roldán (2016) (Figs 37–43), Modified of Raven (1987) (Fig 44)." figureDoi="http://doi.org/10.5281/zenodo.11233906" httpUri="https://zenodo.org/record/11233906/files/figure.png" pageId="10" pageNumber="243">Fig. 42</figureCitation>
), as described by
<bibRefCitation id="EFD84B4EBE43FFF9FB9EFEAAFA83A261" author="Raven, R. J." box="[1126,1283,259,285]" pageId="10" pageNumber="243" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven (1987)</bibRefCitation>
; similar body coloration as described by
<bibRefCitation id="EFD84B4EBE43FFF9FE33FE8EFDE8A23D" author="Raven, R. J." box="[459,616,295,321]" pageId="10" pageNumber="243" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven (1987)</bibRefCitation>
, with reddish brown coloration, even the abdomen dorsally brown with white mottling, ventrally with brown areas on pallid yellow as also reported by
<bibRefCitation id="EFD84B4EBE43FFF9FBCCFEE2FEBDA2F5" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." pageId="10" pageNumber="243" pagination="81" refId="ref5872" refString="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R. (2016) On the trapdoor spiders of Mexico: description of the first new species of the spider genus Aptostichus from Mexico and the description of the female of Eucteniza zapatista (Araneae, Mygalomorphae, Euctenizidae). ZooKeys, 641, 81. https: // doi. org / 10.3897 / zookeys. 641.10521" type="journal article" year="2016">Valdez-Mondragón and Cortez-Roldán (2016)</bibRefCitation>
(
<figureCitation id="13722A3ABE43FFF9FEB7FEC6FE56A2F5" box="[335,470,367,393]" captionStart="FIGURES 28–31" captionStartId="7.[151,264,1936,1960]" captionTargetBox="[321,1267,741,1913]" captionTargetId="figure-288@7.[321,1267,741,1913]" captionTargetPageId="7" captionText="FIGURES 28–31. Mexentypesa chiapas Raven, 1987. Male: 28–29, Habitus, dorsal and ventral views, respectively. 30, Carapace, dorsal view. 31, Prosoma, ventral view showing coxae, sternum, labium, and endites (arrow indicates cuspules). Scale bars 0.5 mm (28, 29), 0.2 mm (30, 31). Modified of Valdez-Mondragón and Cortez-Roldán (2016)." figureDoi="http://doi.org/10.5281/zenodo.11233902" httpUri="https://zenodo.org/record/11233902/files/figure.png" pageId="10" pageNumber="243">Figs 28–31</figureCitation>
). Although around
<quantity id="4CB19B5ABE43FFF9FD4FFEC7FC8FA2F5" box="[695,783,366,393]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="243" unit="km" value="500.0">500 km</quantity>
separate the type locality (Ocosingo,
<collectingRegion id="498DF85DBE43FFF9FB33FEC6FAA7A2F5" box="[1227,1319,367,393]" country="Mexico" name="Chiapas" pageId="10" pageNumber="243">Chiapas</collectingRegion>
) from the locality (Cueva Li Nita, Huautla de Jiménez,
<collectingRegion id="498DF85DBE43FFF9FD58FE3AFD76A2D1" box="[672,758,403,429]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="243">Oaxaca</collectingRegion>
) reported by
<bibRefCitation id="EFD84B4EBE43FFF9FC69FE3AFA1CA2D1" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." box="[913,1436,403,429]" pageId="10" pageNumber="243" pagination="81" refId="ref5872" refString="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R. (2016) On the trapdoor spiders of Mexico: description of the first new species of the spider genus Aptostichus from Mexico and the description of the female of Eucteniza zapatista (Araneae, Mygalomorphae, Euctenizidae). ZooKeys, 641, 81. https: // doi. org / 10.3897 / zookeys. 641.10521" type="journal article" year="2016">Valdez-Mondragón and Cortez-Roldán (2016)</bibRefCitation>
(
<figureCitation id="13722A3ABE43FFF9FF67FE1EFF72A2AD" box="[159,242,438,465]" captionStart="FIGURE 45" captionStartId="10.[151,250,1382,1406]" captionTargetBox="[318,1270,741,1359]" captionTargetId="figure-264@10.[318,1270,741,1359]" captionTargetPageId="10" captionText="FIGURE 45. Known records of Mexentypesa hidalguensis sp. nov. (red circle) and M. chiapas (red diamonds)." figureDoi="http://doi.org/10.5281/zenodo.11233910" httpUri="https://zenodo.org/record/11233910/files/figure.png" pageId="10" pageNumber="243">Fig. 45</figureCitation>
), as well as different habitats, we cannot know the geological histories of these areas. Additional molecular evidence might help to corroborate the morphology between specimens from both localities and understand these distantly allopatric populations.
</paragraph>
<paragraph id="8BF636BFBE43FFF9FF3FFD8AFB19A1D5" blockId="10.[151,1437,151,717]" pageId="10" pageNumber="243">
<emphasis id="B93DEAADBE43FFF9FF3FFD8AFE06A140" bold="true" box="[199,390,546,573]" pageId="10" pageNumber="243">Natural history.</emphasis>
<bibRefCitation id="EFD84B4EBE43FFF9FE76FD8AFDA7A141" author="Raven, R. J." box="[398,551,547,573]" pageId="10" pageNumber="243" pagination="357 - 362" refId="ref5736" refString="Raven, R. J. (1987) A new mygalomorph spider genus from Mexico (Nemesiinae, Nemesiidae, Arachnida). Journal of Arachnology, 14, 357 - 362." type="journal article" year="1987">Raven (1987)</bibRefCitation>
did not report any natural history data about the specimens collected from the
<typeStatus id="54F2881DBE43FFF9FF6FFDEEFF47A11D" box="[151,199,583,609]" pageId="10" pageNumber="243">type</typeStatus>
locality. However,
<bibRefCitation id="EFD84B4EBE43FFF9FE5AFDEEFC28A11D" author="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R." box="[418,936,583,609]" pageId="10" pageNumber="243" pagination="81" refId="ref5872" refString="Valdez-Mondragon, A. &amp; Cortez-Roldan, M. R. (2016) On the trapdoor spiders of Mexico: description of the first new species of the spider genus Aptostichus from Mexico and the description of the female of Eucteniza zapatista (Araneae, Mygalomorphae, Euctenizidae). ZooKeys, 641, 81. https: // doi. org / 10.3897 / zookeys. 641.10521" type="journal article" year="2016">Valdez-Mondragón and Cortez-Roldán (2016)</bibRefCitation>
mentioned that the specimen from Cueva Li Nita was collected inside the cave, in a temperate forest at
<quantity id="4CB19B5ABE43FFF9FCCCFDC2FC0AA1F9" box="[820,906,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.919" pageId="10" pageNumber="243" unit="m" value="1919.0">1919 m</quantity>
of elevation. Although the male specimen was collected within the cave, it does not present any troglomorphism or adaptation to cave life.
</paragraph>
</subSubSection>
<subSubSection id="C3536534BE43FFF9FF3FFD1BFC91A1B1" box="[199,785,690,717]" pageId="10" pageNumber="243" type="distribution">
<paragraph id="8BF636BFBE43FFF9FF3FFD1BFC91A1B1" blockId="10.[151,1437,151,717]" box="[199,785,690,717]" pageId="10" pageNumber="243">
<emphasis id="B93DEAADBE43FFF9FF3FFD1BFEE1A1B0" bold="true" box="[199,353,690,716]" pageId="10" pageNumber="243">Distribution.</emphasis>
<collectingCountry id="F35E762FBE43FFF9FE9FFD1AFE5AA1B1" box="[359,474,691,717]" name="Mexico" pageId="10" pageNumber="243">MEXICO</collectingCountry>
:
<collectingRegion id="498DF85DBE43FFF9FE1DFD1AFDC2A1B1" box="[485,578,691,717]" country="Mexico" name="Chiapas" pageId="10" pageNumber="243">Chiapas</collectingRegion>
,
<collectingRegion id="498DF85DBE43FFF9FDB5FD1AFD21A1B1" box="[589,673,691,717]" country="Mexico" name="Oaxaca" pageId="10" pageNumber="243">Oaxaca</collectingRegion>
(
<figureCitation id="13722A3ABE43FFF9FD48FD1AFC84A1B1" box="[688,772,690,717]" captionStart="FIGURE 45" captionStartId="10.[151,250,1382,1406]" captionTargetBox="[318,1270,741,1359]" captionTargetId="figure-264@10.[318,1270,741,1359]" captionTargetPageId="10" captionText="FIGURE 45. Known records of Mexentypesa hidalguensis sp. nov. (red circle) and M. chiapas (red diamonds)." figureDoi="http://doi.org/10.5281/zenodo.11233910" httpUri="https://zenodo.org/record/11233910/files/figure.png" pageId="10" pageNumber="243">Fig. 45</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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