treatments-xml/data/03/E0/87/03E0879F2472FFABFAD068B4FE20F94F.xml
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<document id="C5C0674CA43DDE5A437B4C2486DC296C" ID-DOI="10.11646/zootaxa.3948.2.3" ID-GBIF-Dataset="5465e49a-9ad0-4891-a32e-f633cc9a87e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="244094" ID-ZooBank="064A12F0-5742-4828-8D26-85AD7B226091" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461043916503" checkinUser="plazi" docAuthor="David, Patrick, Pauwels, Olivier S. G., Nguyen, Truong Quang &amp; Vogel, Gernot" docDate="2015" docId="03E0879F2472FFABFAD068B4FE20F94F" docLanguage="en" docName="zt03948p217.pdf" docOrigin="Zootaxa 3948 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parahelicops Bourret 1934" docType="treatment" docVersion="7" lastPageNumber="208" masterDocId="FFD9FFE72477FFAEFA476B6DFFE5FFF4" masterDocTitle="On the taxonomic status of the Thai endemic freshwater snake Parahelicops boonsongi, with the erection of a new genus (Squamata: Natricidae)" masterLastPageNumber="217" masterPageNumber="203" pageNumber="208" updateTime="1698576256631" updateUser="plazi">
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<mods:title id="5C7CF9A2A69DCC3C274C3D5013BA7B3F">On the taxonomic status of the Thai endemic freshwater snake Parahelicops boonsongi, with the erection of a new genus (Squamata: Natricidae)</mods:title>
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<mods:namePart id="4B39389371E3F8539EB6702F9F55563E">David, Patrick</mods:namePart>
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<mods:namePart id="1A30215EF0C9142B8C4F0E2DEE1888A7">Pauwels, Olivier S. G.</mods:namePart>
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<mods:namePart id="03F9D09A9A6EAE55B2F051E9FC916650">Nguyen, Truong Quang</mods:namePart>
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<mods:date id="EA545E706EEF43210B34573AD98D97C7">2015</mods:date>
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<taxonomicName id="4C494D0A2472FFABFAD068B4FE0DFC07" authority="Bourret, 1934" authorityName="Bourret" authorityYear="1934" box="[151,488,985,1011]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABFAD068B4FE0DFC07" bold="true" box="[151,488,985,1011]" pageId="5" pageNumber="208">
<emphasis id="B93DEA9B2472FFABFAD068B4FED0FC07" bold="true" box="[151,309,985,1011]" italics="true" pageId="5" pageNumber="208">Parahelicops</emphasis>
Bourret, 1934
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<paragraph id="8BF636892472FFABFAD06F4DFA8FFBC2" blockId="5.[151,1386,1056,1078]" box="[151,1386,1056,1078]" pageId="5" pageNumber="208">
<treatmentCitationGroup id="AB5911A72472FFABFAD06F4DFA8FFBC2" box="[151,1386,1056,1078]" pageId="5" pageNumber="208">
<taxonomicName id="4C494D0A2472FFABFAD06F4DFE14FBC2" authority="Bourret, 1934b: 170" authorityName="Bourret" authorityPageNumber="170" authorityYear="1934" box="[151,497,1056,1078]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABFAD06F4DFEFBFBC2" box="[151,286,1056,1078]" italics="true" pageId="5" pageNumber="208">Parahelicops</emphasis>
<treatmentCitation id="0AE810982472FFABFB636F4CFE14FBC2" author="Bourret" box="[292,497,1056,1078]" page="170" pageId="5" pageNumber="208" year="1934">
<bibRefCitation id="EFD84B782472FFABFB636F4CFE24FBC2" author="Bourret" box="[292,449,1056,1078]" pageId="5" pageNumber="208" refString="Bourret, R. (1934 b) Notes herpetologiques sur l'Indochine francaise. III. Ophidiens d'Annam et du Moyen Laos. Bulletin general de l'Instruction Publique, 13 e Annee (9, May), 167 - 176. [separate reprint: 3 - 12]" type="journal article" year="1934" yearSuffix="b">Bourret, 1934b</bibRefCitation>
: 170
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[separate: p. 6].—
<emphasis id="B93DEA9B2472FFABF8E96F4DFCD1FBC2" bold="true" box="[686,820,1056,1078]" pageId="5" pageNumber="208">
<typeStatus id="54F2882B2472FFABF8E96F4DFD06FBC2" box="[686,739,1056,1078]" pageId="5" pageNumber="208">Type</typeStatus>
species
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.
<taxonomicName id="4C494D0A2472FFABF9076F4DFB3DFBC2" authority="Bourret, 1934" authorityName="Bourret" authorityYear="1934" box="[832,1240,1056,1078]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="B93DEA9B2472FFABF9076F4DFBA0FBC2" box="[832,1093,1056,1078]" italics="true" pageId="5" pageNumber="208">Parahelicops annamensis</emphasis>
Bourret, 1934
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by monotypy.
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<emphasis id="B93DEA9B2472FFABFAD06F0EFEEFFB88" bold="true" box="[151,266,1123,1148]" pageId="5" pageNumber="208">Diagnosis</emphasis>
. A genus of medium-sized snakes of the family
<taxonomicName id="4C494D0A2472FFABF9696F0EFC46FB88" box="[814,931,1123,1148]" class="Reptilia" family="Natricidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="family">Natricidae</taxonomicName>
characterized by (1) body elongate, (2) head moderately distinct from the neck, (3) eye moderate in size with a round pupil, (4) maxillary teeth progressively increasing in size in a continuous series, last 2 moderately enlarged, not separated from anterior teeth by a diastema, (5) nostrils piercing distinctly dorsolaterally, (6) upper head scales complete as for typical colubroids but prefrontal single in most specimens, (7) internasals distinctly narrowed anteriorly, (8) all SL distinctly longer than high, (9) dorsal scales in 15 or 17 rows at midbody, moderately keeled at midbody, strongly keeled posteriorly, especially before vent, (10) caudal scales around the base of the tail very strongly keeled, (11) more than 110 SC, paired, (12) hemipenes short, thin, single, and spinose.
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Although
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<emphasis id="B93DEA9B2472FFABFB7B6EEEFD85FA6F" box="[316,608,1411,1435]" italics="true" pageId="5" pageNumber="208">Parahelicops annamensis</emphasis>
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usually has a single prefrontal, as in most species of
<taxonomicName id="4C494D0A2472FFABFEA66EEEFA9FFA6F" box="[1249,1402,1411,1435]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABFEA66EEEFA9FFA6F" box="[1249,1402,1411,1435]" italics="true" pageId="5" pageNumber="208">Opisthotropis</emphasis>
</taxonomicName>
, it differs from this latter genus, as defined above, by (1) maxillary teeth enlarged vs. subequal teeth in
<taxonomicName id="4C494D0A2472FFABFEBB6ECAFA70FA4B" box="[1276,1429,1447,1471]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABFEBB6ECAFA70FA4B" box="[1276,1429,1447,1471]" italics="true" pageId="5" pageNumber="208">Opisthotropis</emphasis>
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, (2) tail long vs. rather short or average in
<taxonomicName id="4C494D0A2472FFABF8236EA6FD18FA17" box="[612,765,1483,1507]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABF8236EA6FD18FA17" box="[612,765,1483,1507]" italics="true" pageId="5" pageNumber="208">Opisthotropis</emphasis>
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(see the descriptions), (3) head moderately distinct from the neck vs. barely distinct in
<taxonomicName id="4C494D0A2472FFABFBFC6E82FDB1F9F3" box="[443,596,1519,1543]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABFBFC6E82FDB1F9F3" box="[443,596,1519,1543]" italics="true" pageId="5" pageNumber="208">Opisthotropis</emphasis>
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, (4) eye large vs. small in
<taxonomicName id="4C494D0A2472FFABF93D6E82FBF6F9F3" box="[890,1043,1519,1543]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABF93D6E82FBF6F9F3" box="[890,1043,1519,1543]" italics="true" pageId="5" pageNumber="208">Opisthotropis</emphasis>
</taxonomicName>
, (5) nostrils directed dorsolaterally vs. directed upwards in
<taxonomicName id="4C494D0A2472FFABFB806D7EFD85F9DF" box="[455,608,1555,1579]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABFB806D7EFD85F9DF" box="[455,608,1555,1579]" italics="true" pageId="5" pageNumber="208">Opisthotropis</emphasis>
</taxonomicName>
, (6) SL much longer than high vs. distinctly higher than long in
<taxonomicName id="4C494D0A2472FFABFAD06D5AFED5F9BB" box="[151,304,1591,1615]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABFAD06D5AFED5F9BB" box="[151,304,1591,1615]" italics="true" pageId="5" pageNumber="208">Opisthotropis</emphasis>
</taxonomicName>
, and (7) dorsal pattern with two rows of large rusty blotches, vs. more or less uniform, or with crossbars in species of
<taxonomicName id="4C494D0A2472FFABFBDE6D36FDD7F987" box="[409,562,1627,1651]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABFBDE6D36FDD7F987" box="[409,562,1627,1651]" italics="true" pageId="5" pageNumber="208">Opisthotropis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF636892472FFABFA806D13FE20F94F" blockId="5.[151,1436,1123,1723]" pageId="5" pageNumber="208">
<emphasis id="B93DEA9B2472FFABFA806D13FED7F963" bold="true" box="[199,306,1662,1687]" pageId="5" pageNumber="208">Contents</emphasis>
<collectingCountry id="F35E76192472FFABFB056D12FE84F963" box="[322,353,1663,1687]" name="American Samoa" pageId="5" pageNumber="208">As</collectingCountry>
currently defined, the genus
<taxonomicName id="4C494D0A2472FFABF89F6D12FC88F963" box="[728,877,1663,1687]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B93DEA9B2472FFABF89F6D12FC88F963" box="[728,877,1663,1687]" italics="true" pageId="5" pageNumber="208">Parahelicops</emphasis>
</taxonomicName>
includes only a single species,
<taxonomicName id="4C494D0A2472FFABFF416D12FE24F94F" authority="Bourret, 1934" authorityName="Bourret" authorityYear="1934" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="208" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="B93DEA9B2472FFABFF416D12FEFEF94F" italics="true" pageId="5" pageNumber="208">Parahelicops annamensis</emphasis>
Bourret, 1934
</taxonomicName>
.
</paragraph>
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