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<document id="AF8610687243153A4F87E46FFAD05426" ID-CLB-Dataset="31742" ID-DOI="10.11646/zootaxa.4338.2.3" ID-GBIF-Dataset="f1edbdd7-dc18-4c34-b236-fb45c9401096" ID-ISSN="1175-5326" ID-Zenodo-Dep="1036109" ID-ZooBank="BE839A35-8A0D-4949-907D-0DDE68A77356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1508915434225" checkinUser="plazi" docAuthor="Grismado, Cristian J. &amp; Carrión, Nicolás López" docDate="2017" docId="03C6927AFFC2FF825FDDC209FDD7F83B" docLanguage="en" docName="zootaxa.4338.2.3.pdf" docOrigin="Zootaxa 4338 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Almafuerte Grismado &amp; Carrión 2017, gen. nov." docType="treatment" docVersion="7" lastPageNumber="267" masterDocId="FFFFEA02FFC3FF865F4AC402FFDEFF8D" masterDocTitle="Description of Almafuerte, a new genus of ground spiders from South America (Araneae, Gnaphosidae)" masterLastPageNumber="291" masterPageNumber="263" pageNumber="264" updateTime="1698470190957" updateUser="ExternalLinkService">
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<mods:title id="B88B44654A64F7B9970A56A3F28E76FB">Description of Almafuerte, a new genus of ground spiders from South America (Araneae, Gnaphosidae)</mods:title>
</mods:titleInfo>
<mods:name id="269A36D1D7669C6A386812DC37A36F1C" type="personal">
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<mods:namePart id="517E0F08F03E6097D21F4297C4024A42">Grismado, Cristian J.</mods:namePart>
</mods:name>
<mods:name id="E2398B48EDBEBA625126A3EB6335AF30" type="personal">
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<mods:namePart id="FEC47F483000921AD5D79E6805E87A26">Carrión, Nicolás López</mods:namePart>
</mods:name>
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<mods:title id="46E7B095A420AEF4A8EE9E86A32455E3">Zootaxa</mods:title>
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<mods:part id="9EA31A3E043D0FAB0CED63C4A912DE9E">
<mods:date id="4D5FF12BAB240B094AA2BA7F6FE12D62">2017</mods:date>
<mods:detail id="54FEB07503E672E043907FAEE1711AC0" type="pubDate">
<mods:number id="9F646E5208BC020243B73F0EFE836130">2017-10-25</mods:number>
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<mods:classification id="2DE24CA5DDC7CFB82408A03B808F9A1F">journal article</mods:classification>
<mods:identifier id="16038B1494C6FBD79A5C80C4CC623127" type="CLB-Dataset">31742</mods:identifier>
<mods:identifier id="53AC7307064ACA0EB39CBB6F0D6AD3E6" type="DOI">10.11646/zootaxa.4338.2.3</mods:identifier>
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<mods:identifier id="2EB1D19A4099AB6C554860FCA13754A3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C6927AFFC2FF825FDDC209FDD7F83B" ID-DOI="http://doi.org/10.5281/zenodo.6033614" ID-GBIF-Taxon="135470733" ID-Zenodo-Dep="6033614" LSID="urn:lsid:plazi:treatment:03C6927AFFC2FF825FDDC209FDD7F83B" httpUri="http://treatment.plazi.org/id/03C6927AFFC2FF825FDDC209FDD7F83B" lastPageId="4" lastPageNumber="267" pageId="1" pageNumber="264">
<subSubSection id="C37570E7FFC2FF875FDDC209FE42F9AB" box="[151,412,1547,1574]" pageId="1" pageNumber="264" type="nomenclature">
<paragraph id="8BD0236CFFC2FF875FDDC209FE42F9AB" blockId="1.[151,412,1547,1574]" box="[151,412,1547,1574]" pageId="1" pageNumber="264">
<heading id="D0989400FFC2FF875FDDC209FE42F9AB" bold="true" box="[151,412,1547,1574]" fontSize="11" level="1" pageId="1" pageNumber="264" reason="1">
<emphasis id="B91BFF7EFFC2FF875FDDC209FE42F9AB" bold="true" box="[151,412,1547,1574]" pageId="1" pageNumber="264">
<taxonomicName id="4C6F58EFFFC2FF875FDDC209FEFEF9A8" ID-CoL="SPZ" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[151,288,1547,1573]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91BFF7EFFC2FF875FDDC209FEFEF9A8" bold="true" box="[151,288,1547,1573]" italics="true" pageId="1" pageNumber="264">Almafuerte</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2284205FFC2FF875E64C20EFE42F9AB" box="[302,412,1548,1574]" pageId="1" pageNumber="264" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37570E7FFC2FF875FDDC250FD19F9E7" box="[151,711,1618,1643]" pageId="1" pageNumber="264" type="materials_examined">
<paragraph id="8BD0236CFFC2FF875FDDC250FD19F9E7" blockId="1.[151,1437,1618,2003]" box="[151,711,1618,1643]" pageId="1" pageNumber="264">
<emphasis id="B91BFF7EFFC2FF875FDDC250FEECF9E6" bold="true" box="[151,306,1618,1643]" pageId="1" pageNumber="264">Type species.</emphasis>
<taxonomicName id="4C6F58EFFFC2FF875E73C250FDBCF9E6" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[313,610,1618,1643]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="peripampasica" status="sp. nov.">
<emphasis id="B91BFF7EFFC2FF875E73C250FDBCF9E6" box="[313,610,1618,1643]" italics="true" pageId="1" pageNumber="264">Almafuerte peripampasica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2284205FFC2FF875D3BC251FD19F9E7" box="[625,711,1619,1642]" pageId="1" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C37570E7FFC2FF875F8DC274FA97F95A" pageId="1" pageNumber="264" type="etymology">
<paragraph id="8BD0236CFFC2FF875F8DC274FA97F95A" blockId="1.[151,1437,1618,2003]" pageId="1" pageNumber="264">
<emphasis id="B91BFF7EFFC2FF875F8DC274FE90F902" bold="true" box="[199,334,1654,1679]" pageId="1" pageNumber="264">Etymology.</emphasis>
“
<taxonomicName id="4C6F58EFFFC2FF875E15C274FE3AF902" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[351,484,1654,1679]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">Almafuerte</taxonomicName>
” is the pseudonymous of Pedro Bonifacio Palacios (San Justo 1854- La Plata 1917), an influential
<collectingCountry id="F37863FCFFC2FF875E70C298FE77F93E" box="[314,425,1690,1715]" name="Argentina" pageId="1" pageNumber="264">Argentine</collectingCountry>
poet and teacher; the name is also shared with a famous
<collectingCountry id="F37863FCFFC2FF875B1CC298FB18F93E" box="[1110,1222,1690,1715]" name="Argentina" pageId="1" pageNumber="264">Argentine</collectingCountry>
heavy-metal band leaded by Ricardo Iorio.
<taxonomicName id="4C6F58EFFFC2FF875EE4C2BCFDEFF95A" box="[430,561,1726,1751]" class="Clitellata" family="Glossoscolecidae" genus="Alma" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="264" phylum="Annelida" rank="species" species="fuerte">
<emphasis id="B91BFF7EFFC2FF875EE4C2BCFDEFF95A" box="[430,561,1726,1751]" italics="true" pageId="1" pageNumber="264">Alma fuerte</emphasis>
</taxonomicName>
means “strong soul” in Spanish, and is considered feminine in gender.
</paragraph>
</subSubSection>
<subSubSection id="C37570E7FFC2FF845F8DC2E0FE9AFF75" lastPageId="2" lastPageNumber="265" pageId="1" pageNumber="264" type="diagnosis">
<paragraph id="8BD0236CFFC2FF845F8DC2E0FE9AFF75" blockId="1.[151,1437,1618,2003]" lastBlockId="2.[151,1437,151,248]" lastPageId="2" lastPageNumber="265" pageId="1" pageNumber="264">
<emphasis id="B91BFF7EFFC2FF875F8DC2E0FE9FF976" bold="true" box="[199,321,1762,1787]" pageId="1" pageNumber="264">Diagnosis.</emphasis>
<taxonomicName id="4C6F58EFFFC2FF875E03C2E0FE18F976" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[329,454,1762,1787]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91BFF7EFFC2FF875E03C2E0FE18F976" box="[329,454,1762,1787]" italics="true" pageId="1" pageNumber="264">Almafuerte</emphasis>
</taxonomicName>
species share with the genera assigned to the subfamily Echeminae (and with those of the “
<taxonomicName id="4C6F58EFFFC2FF875F84C304FEE9F892" authorityName="Simon" authorityYear="1878" box="[206,311,1798,1823]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus">Echemus</taxonomicName>
group” by
<bibRefCitation id="EFFE5E9DFFC2FF875E88C304FDADF892" author="Murphy" box="[450,627,1798,1823]" pageId="1" pageNumber="264" refString="Murphy, J. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambridgeshire 1, i - xii, 1 - 92; 2, i-ii, 93 - 605." type="journal article" year="2007">Murphy [2007]</bibRefCitation>
) the uniform brown coloration, toothed cheliceral promargin, dentate palpal claws in females, and well developed scutum on the dorsum males’ abdomen. The whole somatic morphology and, especially, the eye pattern are reminiscent to the members of the partially sympatric genera
<taxonomicName id="4C6F58EFFFC2FF875FDDC370FEEAF806" authorityName="Vellard" authorityYear="1924" box="[151,308,1906,1931]" class="Arachnida" family="Gnaphosidae" genus="Apodrassodes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B91BFF7EFFC2FF875FDDC370FEEAF806" box="[151,308,1906,1931]" italics="true" pageId="1" pageNumber="264">Apodrassodes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F58EFFFC2FF875E38C370FE3CF806" authorityName="Platnick &amp; Shadab" authorityYear="1984" box="[370,482,1906,1931]" class="Arachnida" family="Gnaphosidae" genus="Apopyllus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B91BFF7EFFC2FF875E38C370FE3CF806" box="[370,482,1906,1931]" italics="true" pageId="1" pageNumber="264">Apopyllus</emphasis>
</taxonomicName>
. They differ from all of them by having three teeth on the cheliceral retromargin (
<figureCitation id="13543FE9FFC2FF875FD5C394FEFBF822" box="[159,293,1942,1967]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1917]" captionTargetBox="[151,1436,841,1874]" captionTargetId="figure@3.[151,1436,841,1874]" captionTargetPageId="3" captionText="FIGURE 2. Almafuerte peripampasica sp. nov., female (MACN-Ar 30442). a, left chelicera, apical view; b, same, posterior view, c, same, detail of cheliceral fang and tooth; d, detail of venom outlet; e, left palp, retrolateral view; f, detail of dorsal cuticle of palpal tibia, showing the row of trichobothria; g, detail of trichobothrial socket; h, tarsal claw, prolateral view; i, detail of tarsal chemosensory seta." httpUri="https://zenodo.org/record/1036113/files/figure.png" pageId="1" pageNumber="264">Figs. 2 b, c</figureCitation>
), by the protrusion on the retrolateral side of the tegulum (RTP,
<figureCitation id="13543FE9FFC2FF875B5EC394FB6DF822" box="[1044,1203,1942,1967]" captionStart="FIGURE 6" captionStartId="7.[151,250,1375,1397]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure@7.[151,1436,193,1354]" captionTargetPageId="7" captionText="FIGURE 6. Almafuerte peripampasica sp. nov., male (MACN-Ar 30443). a, left palp, ventral view; b, same, closer view of the copulatory bulb, c, same, detail of embolus tip; d, left palp, retrolateral view; e, same, closer view of RTA; f, retrolateral tegular process (RTP), ventral view." httpUri="https://zenodo.org/record/1036121/files/figure.png" pageId="1" pageNumber="264">Figs. 6 a, b, f</figureCitation>
) and the absence of median apophysis, and by the shape of the internal female genitalia: two large, rounded primary spermathecae, secondary spermathecae small, anteriorly located; copulatory ducts with thickened walls, especially around the insertion area of the secondary spermathecae, which makes it appear to have bilobular primary spermathecae. (
<figureCitation id="13543FE9FFC1FF845FD5C4DDFEE9FF75" box="[159,311,223,248]" captionStart="FIGURE 5" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[292,1294,517,1921]" captionTargetId="figure@6.[292,1294,517,1921]" captionTargetPageId="6" captionText="FIGURE 5. Almafuerte peripampasica sp. nov., female (MACN-Ar 30442). a, epigynum, ventral view; b, same, detail of copulatory openings, c, same, detail of anterior hood; d, vulva, dorsal view; e, same, closer view; f, same, detail of the left secondary spermathecae." httpUri="https://zenodo.org/record/1036119/files/figure.png" pageId="2" pageNumber="265">Figs. 5 d, e, f</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1073E4FFC1FF845FDDC2F5FB14F8A6" httpUri="https://zenodo.org/record/1036111/files/figure.png" pageId="2" pageNumber="265" targetBox="[169,1418,301,1761]" targetPageId="2">
<paragraph id="8BD0236CFFC1FF845FDDC2F5FB14F8A6" blockId="2.[151,1436,1783,1835]" pageId="2" pageNumber="265">
<emphasis id="B91BFF7EFFC1FF845FDDC2F5FEC9F880" bold="true" box="[151,279,1783,1805]" pageId="2" pageNumber="265">FIGURE 1.</emphasis>
<taxonomicName id="4C6F58EFFFC1FF845E6BC2F5FDECF880" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[289,562,1783,1805]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="peripampasica" status="sp. nov.">
<emphasis id="B91BFF7EFFC1FF845E6BC2F5FDECF880" box="[289,562,1783,1805]" italics="true" pageId="2" pageNumber="265">Almafuerte peripampasica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2284205FFC1FF845D76C2F5FD4FF880" box="[572,657,1783,1805]" pageId="2" pageNumber="265" rank="species">
<emphasis id="B91BFF7EFFC1FF845D76C2F5FD4FF880" bold="true" box="[572,657,1783,1805]" pageId="2" pageNumber="265">sp. nov.</emphasis>
</taxonomicNameLabel>
, female (MACN-Ar 30442).
<emphasis id="B91BFF7EFFC1FF845C81C2F5FC06F880" bold="true" box="[971,984,1783,1805]" pageId="2" pageNumber="265">a</emphasis>
, carapace, dorsal view;
<emphasis id="B91BFF7EFFC1FF845B96C2F5FB34F880" bold="true" box="[1244,1258,1783,1805]" pageId="2" pageNumber="265">b</emphasis>
, detail of ocular region,
<emphasis id="B91BFF7EFFC1FF845FAEC317FF31F8A6" bold="true" box="[228,239,1813,1835]" pageId="2" pageNumber="265">c</emphasis>
, cephalothorax, ventral view;
<emphasis id="B91BFF7EFFC1FF845D69C317FDEFF8A6" bold="true" box="[547,561,1813,1835]" pageId="2" pageNumber="265">d</emphasis>
, mouthparts, ventral view;
<emphasis id="B91BFF7EFFC1FF845C02C317FC8DF8A6" bold="true" box="[840,851,1813,1835]" pageId="2" pageNumber="265">e</emphasis>
, feathery scales on carapace surface.
</paragraph>
</caption>
<subSubSection id="C37570E7FFC1FF825F8DC351FDD7F83B" lastPageId="4" lastPageNumber="267" pageId="2" pageNumber="265" type="description">
<paragraph id="8BD0236CFFC1FF845F8DC351FA43F85A" blockId="2.[151,1437,1875,2008]" pageId="2" pageNumber="265">
<emphasis id="B91BFF7EFFC1FF845F8DC351FE87F8E1" bold="true" box="[199,345,1875,1900]" pageId="2" pageNumber="265">Description.</emphasis>
Medium-size spiders (total length
<quantity id="4C978E89FFC1FF845DA6C351FC8FF8E6" box="[748,849,1875,1900]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="2" pageNumber="265" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">6−9 mm</quantity>
). Carapace oval in dorsal view, rounded anteriorly, truncated posteriorly, widest behind coxae II, slightly narrowed at the level of the palps, coloration uniform chestnut brown, with needle-like setae, most abundant on margins, longer in ocular and clypeal areas; feathery scales also present (
<figureCitation id="13543FE9FFC1FF845E33C3BDFE14F85A" box="[377,458,1983,2008]" captionStart="FIGURE 1" captionStartId="2.[151,250,1783,1805]" captionTargetBox="[169,1418,301,1761]" captionTargetId="figure@2.[169,1418,301,1761]" captionTargetPageId="2" captionText="FIGURE 1. Almafuerte peripampasica sp. nov., female (MACN-Ar 30442). a, carapace, dorsal view; b, detail of ocular region, c, cephalothorax, ventral view; d, mouthparts, ventral view; e, feathery scales on carapace surface." httpUri="https://zenodo.org/record/1036111/files/figure.png" pageId="2" pageNumber="265">Fig. 1e</figureCitation>
), scattered on the dorsal surface of carapace, and most abundant on the cephalic area.
</paragraph>
<paragraph id="8BD0236CFFC0FF825FDDC495FB30F88E" blockId="3.[151,1437,151,789]" lastBlockId="4.[151,1436,1373,1974]" lastPageId="4" lastPageNumber="267" pageId="3" pageNumber="266">
Cephalic area slightly elevated; thoracic groove short, deep. In front view, anterior eye row slightly procurved, posterior eye row strongly procurved; in dorsal view, anterior row nearly straight, posterior row slightly procurved; AME circular, dark, PME irregularly oval, light, ALE and PLE oval, light; eyes subequal in size; AME separated by about their diameter, by nearly their radius from ALE; PME separated by roughly their radius, farther from PLE; lateral eyes of each side separated by almost their diameter; MOQ longer than wide. Clypeus higher than AME diameter. Chelicerae usually with three promarginal teeth and three retromarginal teeth (
<figureCitation id="13543FE9FFC0FF855B3EC54EFB2DFEE9" box="[1140,1267,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1895,1917]" captionTargetBox="[151,1436,841,1874]" captionTargetId="figure@3.[151,1436,841,1874]" captionTargetPageId="3" captionText="FIGURE 2. Almafuerte peripampasica sp. nov., female (MACN-Ar 30442). a, left chelicera, apical view; b, same, posterior view, c, same, detail of cheliceral fang and tooth; d, detail of venom outlet; e, left palp, retrolateral view; f, detail of dorsal cuticle of palpal tibia, showing the row of trichobothria; g, detail of trichobothrial socket; h, tarsal claw, prolateral view; i, detail of tarsal chemosensory seta." httpUri="https://zenodo.org/record/1036113/files/figure.png" pageId="3" pageNumber="266">Figs. 2 b, c</figureCitation>
). Endites long, obliquely depressed; labium elongated, with rounded tip; sternum long, rebordered, with extensions to coxae and between them, with needle-like setae uniform on all surface. Leg formula 4123. Femora and patellae light brown; tarsi and almost all metatarsi (except the basal portion) with thick scopula, without preening combs; tarsi with two dentate claws and claw tufts; trochanters slightly notched; all segments with very long setae, and numerous feathery scales, especially on the distal segments. Leg spination: Legs III–IV more spinose than legs I–II. Regular pattern are nearly uniform in the forelegs: tibiae I and II of males with usually three pairs of ventral macrosetae (2- 2-2), females without macrosetae or with a single one in ventral retrolateral position; metatarsi I and II of both sexes with ventral macrosetae (2-
<quantity id="4C978E89FFC0FF855D46C66EFDE9FD08" box="[524,567,620,645]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="266" unit="in" value="2.0">2 in</quantity>
males, 2/1-0-0 in females), prolateral or retrolateral surfaces with spines only in males. Abdomen brownish gray with three pairs of apodemes; covered with needle-like setae and feathery scales, especially abundant on the posterior area. Males with orange-brown anterior scutum. Six long spinnerets, ALS sclerotized, separated at base by their width. Male palpal tibia with retrolateral apophysis bifid (
<figureCitation id="13543FE9FFC0FF855A5CC6D5FA53FD7D" box="[1302,1421,727,752]" captionStart="FIGURE 6" captionStartId="7.[151,250,1375,1397]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure@7.[151,1436,193,1354]" captionTargetPageId="7" captionText="FIGURE 6. Almafuerte peripampasica sp. nov., male (MACN-Ar 30443). a, left palp, ventral view; b, same, closer view of the copulatory bulb, c, same, detail of embolus tip; d, left palp, retrolateral view; e, same, closer view of RTA; f, retrolateral tegular process (RTP), ventral view." httpUri="https://zenodo.org/record/1036121/files/figure.png" pageId="3" pageNumber="266">Fig. 6 d, e</figureCitation>
), with a dorsal branch usually flattened, with serrated margin (except in
<taxonomicName id="4C6F58EFFFC0FF855CE4C6FEFBF0FC99" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[942,1070,764,788]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="vigorosa">
<emphasis id="B91BFF7EFFC0FF855CE4C6FEFBF0FC99" box="[942,1070,764,788]" italics="true" pageId="3" pageNumber="266">A. vigorosa</emphasis>
</taxonomicName>
, which is hook-shaped), and the ventral branch ending nearly parallel to the dorsal one; ventrally oriented at its base, curving then to dorsal (in
<taxonomicName id="4C6F58EFFFC7FF825ACEC15CFF14FA17" authorityName="Gerschman &amp; Schiapelli" authorityYear="1948" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="giaii">
<emphasis id="B91BFF7EFFC7FF825ACEC15CFF14FA17" italics="true" pageId="4" pageNumber="267">A. giaii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F58EFFFC7FF825E49C181FE17FA17" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[259,457,1411,1434]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="peripampasica">
<emphasis id="B91BFF7EFFC7FF825E49C181FE17FA17" box="[259,457,1411,1434]" italics="true" pageId="4" pageNumber="267">A. peripampasica</emphasis>
</taxonomicName>
), directly oriented nearly parallel to the dorsal one in all its length (in
<emphasis id="B91BFF7EFFC7FF825BA5C181FF32FA33" italics="true" pageId="4" pageNumber="267">
<taxonomicName id="4C6F58EFFFC7FF825BA5C181FAA9FA16" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[1263,1399,1410,1435]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="goloboffi">A. goloboffi</taxonomicName>
,
<taxonomicName id="4C6F58EFFFC7FF825ACEC181FF36FA33" authorityName="Grismado &amp; Carrión" authorityYear="2017" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="remota">A. remota</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C6F58EFFFC7FF825E6FC1A4FE5DFA33" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[293,387,1446,1470]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="facon">
<emphasis id="B91BFF7EFFC7FF825E6FC1A4FE5DFA33" box="[293,387,1446,1470]" italics="true" pageId="4" pageNumber="267">A. facon</emphasis>
</taxonomicName>
), or reduced, wide, almost not protruding (in
<taxonomicName id="4C6F58EFFFC7FF825CC4C1A4FBD1FA33" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[910,1039,1446,1470]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="vigorosa">
<emphasis id="B91BFF7EFFC7FF825CC4C1A4FBD1FA33" box="[910,1039,1446,1470]" italics="true" pageId="4" pageNumber="267">A. vigorosa</emphasis>
</taxonomicName>
). Palp (
<figureCitation id="13543FE9FFC7FF825B20C1A7FB6FFA33" box="[1130,1201,1445,1470]" captionStart="FIGURE 6" captionStartId="7.[151,250,1375,1397]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure@7.[151,1436,193,1354]" captionTargetPageId="7" captionText="FIGURE 6. Almafuerte peripampasica sp. nov., male (MACN-Ar 30443). a, left palp, ventral view; b, same, closer view of the copulatory bulb, c, same, detail of embolus tip; d, left palp, retrolateral view; e, same, closer view of RTA; f, retrolateral tegular process (RTP), ventral view." httpUri="https://zenodo.org/record/1036121/files/figure.png" pageId="4" pageNumber="267">Fig. 6</figureCitation>
): Embolus about the same size as the tegulum, with prolateral basal origin, apically directed; conductor apical, membranous, more or less parallel to the embolus tip; median apophysis absent; retrolateral tegular protrusion nearly rounded, laterally directed (except in
<taxonomicName id="4C6F58EFFFC7FF825EC8C211FDD6F9A7" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[386,520,1555,1578]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="vigorosa">
<emphasis id="B91BFF7EFFC7FF825EC8C211FDD6F9A7" box="[386,520,1555,1578]" italics="true" pageId="4" pageNumber="267">A. vigorosa</emphasis>
</taxonomicName>
, where it point to apical), and central part of tegulum ventrally protruding, sometimes as a whole bulging area (especially in
<taxonomicName id="4C6F58EFFFC7FF825D82C234FC50F9C3" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[712,910,1590,1614]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="peripampasica">
<emphasis id="B91BFF7EFFC7FF825D82C234FC50F9C3" box="[712,910,1590,1614]" italics="true" pageId="4" pageNumber="267">A. peripampasica</emphasis>
</taxonomicName>
) or as a digitiform projection (especially in
<taxonomicName id="4C6F58EFFFC7FF825ACEC234FF22F9FE" authorityName="Grismado &amp; Carrión" authorityYear="2017" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="goloboffi">
<emphasis id="B91BFF7EFFC7FF825ACEC234FF22F9FE" italics="true" pageId="4" pageNumber="267">A. goloboffi</emphasis>
</taxonomicName>
). Epigynum (
<figureCitation id="13543FE9FFC7FF825ED7C258FDFBF9FF" box="[413,549,1626,1651]" captionStart="FIGURE 5" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[292,1294,517,1921]" captionTargetId="figure@6.[292,1294,517,1921]" captionTargetPageId="6" captionText="FIGURE 5. Almafuerte peripampasica sp. nov., female (MACN-Ar 30442). a, epigynum, ventral view; b, same, detail of copulatory openings, c, same, detail of anterior hood; d, vulva, dorsal view; e, same, closer view; f, same, detail of the left secondary spermathecae." httpUri="https://zenodo.org/record/1036119/files/figure.png" pageId="4" pageNumber="267">Figs. 5 a −c</figureCitation>
): with anteroventral hood with variable shape; a transversaly folded, weakly sclerotized area between the copulatory openings and the hood; copulatory openings oriented ventrally (
<emphasis id="B91BFF7EFFC7FF825ACEC27CFE43F936" italics="true" pageId="4" pageNumber="267">
<taxonomicName id="4C6F58EFFFC7FF825ACEC27CFE9EF937" authorityName="Grismado &amp; Carrión" authorityYear="2017" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="peripampasica">A. peripampasica</taxonomicName>
,
<taxonomicName id="4C6F58EFFFC7FF825E01C2A1FE43F936" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[331,413,1698,1723]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="kuru">A. kuru</taxonomicName>
</emphasis>
) sometimes in a transversal epigynal groove, or laterally (
<emphasis id="B91BFF7EFFC7FF825B61C2A1FA55F936" box="[1067,1419,1698,1723]" italics="true" pageId="4" pageNumber="267">
<taxonomicName id="4C6F58EFFFC7FF825B61C2A1FB42F937" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[1067,1180,1699,1722]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="remota">A. remota</taxonomicName>
,
<taxonomicName id="4C6F58EFFFC7FF825BEDC2A1FB22F937" authorityName="Gerschman &amp; Schiapelli" authorityYear="1948" box="[1191,1276,1699,1722]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="giaii">A. giaii</taxonomicName>
,
<taxonomicName id="4C6F58EFFFC7FF825A4DC2A1FA55F936" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[1287,1419,1698,1723]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="goloboffi">A. goloboffi</taxonomicName>
</emphasis>
). Internal genitalia (
<figureCitation id="13543FE9FFC7FF825E22C2C7FE3FF953" box="[360,481,1733,1758]" captionStart="FIGURE 5" captionStartId="6.[151,250,1942,1964]" captionTargetBox="[292,1294,517,1921]" captionTargetId="figure@6.[292,1294,517,1921]" captionTargetPageId="6" captionText="FIGURE 5. Almafuerte peripampasica sp. nov., female (MACN-Ar 30442). a, epigynum, ventral view; b, same, detail of copulatory openings, c, same, detail of anterior hood; d, vulva, dorsal view; e, same, closer view; f, same, detail of the left secondary spermathecae." httpUri="https://zenodo.org/record/1036119/files/figure.png" pageId="4" pageNumber="267">Fig. 5 d −f</figureCitation>
): two large, rounded primary spermathecae, anterior secondary spermathecae small, copulatory ducts with thickened walls, especially near the insertion of the secondary spermathecae.
</paragraph>
<caption id="DF1073E4FFC0FF855FDDC365FE2AF854" httpUri="https://zenodo.org/record/1036113/files/figure.png" pageId="3" pageNumber="266" targetBox="[151,1436,841,1874]" targetPageId="3">
<paragraph id="8BD0236CFFC0FF855FDDC365FE2AF854" blockId="3.[151,1436,1895,2009]" pageId="3" pageNumber="266">
<emphasis id="B91BFF7EFFC0FF855FDDC365FECAF8F0" bold="true" box="[151,276,1895,1917]" pageId="3" pageNumber="266">FIGURE 2.</emphasis>
<taxonomicName id="4C6F58EFFFC0FF855E51C365FDF7F8F3" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[283,553,1895,1918]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="peripampasica" status="sp. nov.">
<emphasis id="B91BFF7EFFC0FF855E51C365FDF7F8F3" box="[283,553,1895,1918]" italics="true" pageId="3" pageNumber="266">Almafuerte peripampasica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2284205FFC0FF855D78C365FD5DF8F0" box="[562,643,1895,1917]" pageId="3" pageNumber="266" rank="species">
<emphasis id="B91BFF7EFFC0FF855D78C365FD5DF8F0" bold="true" box="[562,643,1895,1917]" pageId="3" pageNumber="266">sp. nov.</emphasis>
</taxonomicNameLabel>
, female (MACN-Ar 30442).
<emphasis id="B91BFF7EFFC0FF855CF9C365FC61F8F0" bold="true" box="[947,959,1895,1917]" pageId="3" pageNumber="266">a</emphasis>
, left chelicera, apical view;
<emphasis id="B91BFF7EFFC0FF855BAEC365FB2CF8F0" bold="true" box="[1252,1266,1895,1917]" pageId="3" pageNumber="266">b</emphasis>
, same, posterior view,
<emphasis id="B91BFF7EFFC0FF855F9FC384FF3EF811" bold="true" box="[213,224,1926,1948]" pageId="3" pageNumber="266">c</emphasis>
, same, detail of cheliceral fang and tooth;
<emphasis id="B91BFF7EFFC0FF855DEBC384FD71F811" bold="true" box="[673,687,1926,1948]" pageId="3" pageNumber="266">d</emphasis>
, detail of venom outlet;
<emphasis id="B91BFF7EFFC0FF855CF8C384FC63F811" bold="true" box="[946,957,1926,1948]" pageId="3" pageNumber="266">e</emphasis>
, left palp, retrolateral view;
<emphasis id="B91BFF7EFFC0FF855BA3C384FB2FF811" bold="true" box="[1257,1265,1926,1948]" pageId="3" pageNumber="266">f</emphasis>
, detail of dorsal cuticle of palpal tibia, showing the row of trichobothria;
<emphasis id="B91BFF7EFFC0FF855DA1C3A6FD29F837" bold="true" box="[747,759,1956,1978]" pageId="3" pageNumber="266">g</emphasis>
, detail of trichobothrial socket;
<emphasis id="B91BFF7EFFC0FF855B03C3A6FB89F837" bold="true" box="[1097,1111,1956,1978]" pageId="3" pageNumber="266">h</emphasis>
, tarsal claw, prolateral view;
<emphasis id="B91BFF7EFFC0FF855AC4C3A6FA4BF837" bold="true" box="[1422,1429,1956,1978]" pageId="3" pageNumber="266">i</emphasis>
, detail of tarsal chemosensory seta.
</paragraph>
</caption>
<caption id="DF1073E4FFC7FF825FDDC0C6FD9BFABB" httpUri="https://zenodo.org/record/1036115/files/figure.png" pageId="4" pageNumber="267" targetBox="[151,1436,193,1198]" targetPageId="4">
<paragraph id="8BD0236CFFC7FF825FDDC0C6FD9BFABB" blockId="4.[151,1436,1220,1334]" pageId="4" pageNumber="267">
<emphasis id="B91BFF7EFFC7FF825FDDC0C6FECDFB57" bold="true" box="[151,275,1220,1242]" pageId="4" pageNumber="267">FIGURE 3.</emphasis>
<taxonomicName id="4C6F58EFFFC7FF825E51C0C6FDF4FB56" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[283,554,1220,1243]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="peripampasica" status="sp. nov.">
<emphasis id="B91BFF7EFFC7FF825E51C0C6FDF4FB56" box="[283,554,1220,1243]" italics="true" pageId="4" pageNumber="267">Almafuerte peripampasica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2284205FFC7FF825D7AC0C6FD5FFB57" box="[560,641,1220,1242]" pageId="4" pageNumber="267" rank="species">
<emphasis id="B91BFF7EFFC7FF825D7AC0C6FD5FFB57" bold="true" box="[560,641,1220,1242]" pageId="4" pageNumber="267">sp. nov.</emphasis>
</taxonomicNameLabel>
, female (MACN-Ar 30442).
<emphasis id="B91BFF7EFFC7FF825CFAC0C6FC63FB57" bold="true" box="[944,957,1220,1242]" pageId="4" pageNumber="267">a</emphasis>
, left leg I, retrolateral view;
<emphasis id="B91BFF7EFFC7FF825BACC0C6FB2AFB57" bold="true" box="[1254,1268,1220,1242]" pageId="4" pageNumber="267">b</emphasis>
, detail of dorsal cuticle of tibia I, showing feathery scales, normal setae, and trichobothria;
<emphasis id="B91BFF7EFFC7FF825CEDC0E1FC6CFB74" bold="true" box="[935,946,1251,1273]" pageId="4" pageNumber="267">c</emphasis>
, same, detail of trichobothrial socket;
<emphasis id="B91BFF7EFFC7FF825A02C0E1FA88FB74" bold="true" box="[1352,1366,1251,1273]" pageId="4" pageNumber="267">d</emphasis>
, tarsal claws and claw tuft;
<emphasis id="B91BFF7EFFC7FF825E25C100FEA4FA95" bold="true" box="[367,378,1282,1304]" pageId="4" pageNumber="267">e</emphasis>
, tarsal organ I;
<emphasis id="B91BFF7EFFC7FF825D6BC100FDF7FA95" bold="true" box="[545,553,1282,1304]" pageId="4" pageNumber="267">f</emphasis>
, left leg IV, prolateral view;
<emphasis id="B91BFF7EFFC7FF825C13C100FCBBFA95" bold="true" box="[857,869,1282,1304]" pageId="4" pageNumber="267">g</emphasis>
, tarsus IV, showing scopula;
<emphasis id="B91BFF7EFFC7FF825BDDC100FB7BFA95" bold="true" box="[1175,1189,1282,1304]" pageId="4" pageNumber="267">h</emphasis>
, tarsal claws, prolateral view, showing claw tuft;
<emphasis id="B91BFF7EFFC7FF825EDCC122FE43FABB" bold="true" box="[406,413,1312,1334]" pageId="4" pageNumber="267">i</emphasis>
, tarsal organ IV.
</paragraph>
</caption>
<paragraph id="8BD0236CFFC7FF825F8DC30FFDD7F83B" blockId="4.[151,1436,1373,1974]" pageId="4" pageNumber="267">
<emphasis id="B91BFF7EFFC7FF825F8DC30FFD97F8AB" bold="true" box="[199,585,1805,1830]" pageId="4" pageNumber="267">Composition and distribution.</emphasis>
Seven species from central parts of Southern South America;
<taxonomicName id="4C6F58EFFFC7FF825ACEC30CFEE3F8C7" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="peripampasica" status="sp. nov.">
<emphasis id="B91BFF7EFFC7FF825ACEC30CFEE3F8C7" italics="true" pageId="4" pageNumber="267">A. peripampasica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2284205FFC7FF825E02C331FE57F8C7" box="[328,393,1843,1866]" pageId="4" pageNumber="267" rank="species">n. sp.</taxonomicNameLabel>
from Argentina and Uruguay,
<taxonomicName id="4C6F58EFFFC7FF825DBCC331FC89F8C7" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[758,855,1842,1866]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="facon" status="sp. nov.">
<emphasis id="B91BFF7EFFC7FF825DBCC331FC89F8C7" box="[758,855,1842,1866]" italics="true" pageId="4" pageNumber="267">A. facon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2284205FFC7FF825C23C331FC74F8C7" box="[873,938,1843,1866]" pageId="4" pageNumber="267" rank="species">n. sp.</taxonomicNameLabel>
from Cochabamba, Bolivia, and five only known from Argentina:
<taxonomicName id="4C6F58EFFFC7FF825EE5C354FCE5F8E3" authority="(Gerschman &amp; Schiapelli)" authorityName="Gerschman &amp; Schiapelli" box="[431,827,1877,1902]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="giaii">
<emphasis id="B91BFF7EFFC7FF825EE5C354FDDAF8E3" box="[431,516,1878,1902]" italics="true" pageId="4" pageNumber="267">A. giaii</emphasis>
(Gerschman &amp; Schiapelli)
</taxonomicName>
from Santa Fe,
<taxonomicName id="4C6F58EFFFC7FF825CB1C354FBB5F8E3" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[1019,1131,1878,1902]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="remota" status="sp. nov.">
<emphasis id="B91BFF7EFFC7FF825CB1C354FBB5F8E3" box="[1019,1131,1878,1902]" italics="true" pageId="4" pageNumber="267">A. remota</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2284205FFC7FF825B37C355FB60F8E3" box="[1149,1214,1879,1902]" pageId="4" pageNumber="267" rank="species">n. sp.</taxonomicNameLabel>
, from Santiago del Estero and Mendoza,
<emphasis id="B91BFF7EFFC7FF825ED8C379FE30F81E" box="[402,494,1914,1939]" italics="true" pageId="4" pageNumber="267">
<taxonomicName id="4C6F58EFFFC7FF825ED8C379FE34F81E" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[402,490,1914,1939]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="kuru" status="sp. nov.">A. kuru</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2284205FFC7FF825EBDC379FDE6F81F" box="[503,568,1915,1938]" pageId="4" pageNumber="267" rank="species">n. sp.</taxonomicNameLabel>
, from Santiago del Estero,
<taxonomicName id="4C6F58EFFFC7FF825C3FC379FC25F81E" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" box="[885,1019,1914,1939]" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="goloboffi" status="sp. nov.">
<emphasis id="B91BFF7EFFC7FF825C3FC379FC25F81E" box="[885,1019,1914,1939]" italics="true" pageId="4" pageNumber="267">A. goloboffi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2284205FFC7FF825B46C379FB94F81F" box="[1036,1098,1915,1938]" pageId="4" pageNumber="267" rank="species">n. sp.</taxonomicNameLabel>
from Salta and Jujuy, and
<taxonomicName id="4C6F58EFFFC7FF825ACEC379FF26F83B" authority="Grismado &amp; Carrión, 2017" authorityName="Grismado &amp; Carrión" authorityYear="2017" class="Arachnida" family="Gnaphosidae" genus="Almafuerte" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="267" phylum="Arthropoda" rank="species" species="vigorosa" status="sp. nov.">
<emphasis id="B91BFF7EFFC7FF825ACEC379FF26F83B" italics="true" pageId="4" pageNumber="267">A. vigorosa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2284205FFC7FF825E4CC39DFE9DF83B" box="[262,323,1951,1974]" pageId="4" pageNumber="267" rank="species">n. sp.</taxonomicNameLabel>
, from Corrientes.
</paragraph>
</subSubSection>
</treatment>
</document>
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