247 lines
30 KiB
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247 lines
30 KiB
XML
<document id="DC44D7C74E95DBBB9B5FCFD6901FA847" ID-CLB-Dataset="21986" ID-DOI="10.5852/ejt.2017.361" ID-GBIF-Dataset="db3fd15a-2d90-4f3f-87f8-10b8953e26a5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3836508" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589998651298" checkinUser="carolina" docAuthor="Fend, Steven V., Rodriguez, Pilar, Achurra, Ainara & Erséus, Christer" docDate="2017" docId="03BC8649FF8DFF81FD81FDF5DD7BB298" docLanguage="en" docName="ejt-2017-361.pdf.imf" docOrigin="European Journal of Taxonomy 361" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Guestphalinus Michaelsen 1933" docType="treatment" docVersion="7" lastPageNumber="19" masterDocId="FF85FE31FF9FFF93FFD7FFD9DE4DB573" masterDocTitle="On Kincaidiana Altman, 1936 and Guestphalinus Michaelsen, 1933 (Annelida, Clitellata, Lumbriculidae), with the descriptions of three new species" masterLastPageNumber="46" masterPageNumber="1" pageNumber="19" updateTime="1698840559482" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="57F9DE82893E93303030531039320AC6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2A11273393E1C802289CC45B79E2F065">
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<mods:title id="0C619DBF8C568C5E6BE6D7ADA0842DDE">On Kincaidiana Altman, 1936 and Guestphalinus Michaelsen, 1933 (Annelida, Clitellata, Lumbriculidae), with the descriptions of three new species</mods:title>
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<mods:namePart id="46716DB30A370808C9539A4B629C8EC0">Fend, Steven V.</mods:namePart>
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<mods:affiliation id="ED76D46A6AAB87A145096FBA6A1B7A60">US Geological Survey, 345 Middlefield Rd., Menlo Park, CA, 94025, USA. & urn: lsid: zoobank. org: author: E 0 C 611 FA- 9619 - 4 DFD- 9701 - 026417181232 & Corresponding author: svfend @ usgs. gov</mods:affiliation>
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<mods:nameIdentifier id="09325A240D9E66F8A3053B0DABF5FFAD" type="email">svfend@usgs.gov</mods:nameIdentifier>
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<mods:name id="94E950A7F266071EAC0D3E90EF47802A" type="personal">
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<mods:roleTerm id="4E60B7F7BD6E52CD9C2F731A3F58D3F2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F97897E0BB999A555BF9222003DC4B1C">Rodriguez, Pilar</mods:namePart>
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<mods:nameIdentifier id="FB356CBA92AA95510C3B887D3CDCF55C" type="LSID">C4600D2B-17E2-4CD0-AE75-B79BA9DD35D6</mods:nameIdentifier>
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<mods:affiliation id="ADCC08ED9B29BF9BC63FBAF2D414911A">University of the Basque Country, UPV / EHU, Sarriena s / n, Leioa 48940, Spain. & Email: pilar. rodriguez @ ehu. eus & urn: lsid: zoobank. org: author: C 4600 D 2 B- 17 E 2 - 4 CD 0 - AE 75 - B 79 BA 9 DD 35 D 6</mods:affiliation>
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<mods:nameIdentifier id="F2BFAF222F4891483D462FA548BD135F" type="email">pilar.rodriguez@ehu.eus</mods:nameIdentifier>
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</mods:name>
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<mods:name id="DA988FFCC4A459FE2773D46ED2C1FF6E" type="personal">
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<mods:roleTerm id="C7629094AB7AA3CD8D50966C5452EDA5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DEEFEAD617820A2569026334063A64B2">Achurra, Ainara</mods:namePart>
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<mods:nameIdentifier id="9349BDDDD2E5253DCADD1E90EF95243C" type="LSID">105572EB-B070-4BE4-93D2-50FA50847C9E</mods:nameIdentifier>
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<mods:affiliation id="D4EFBAA328171FBDF7426C48A8B68A36">University of the Basque Country, UPV / EHU, Sarriena s / n, Leioa 48940, Spain. & Email: ainara. achurra @ ehu. eus & urn: lsid: zoobank. org: author: 105572 EB-B 070 - 4 BE 4 - 93 D 2 - 50 FA 50847 C 9 E</mods:affiliation>
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<mods:nameIdentifier id="52CF1A7D01D4531A04B5A5F3919E2810" type="email">ainara.achurra@ehu.eus</mods:nameIdentifier>
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</mods:name>
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<mods:name id="D9360118191D93D21F610C19BE6481D4" type="personal">
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<mods:roleTerm id="0251EA77D9AA4973131F33403A76B41A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E36AD38A8AD953624AB7887FD4C283E7">Erséus, Christer</mods:namePart>
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<mods:nameIdentifier id="C755BCCBE46CDEA68BCF32FC7FB2543A" type="LSID">D98F606A-B273-4F50-95F5-C35F17B12C85</mods:nameIdentifier>
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<mods:affiliation id="167D7A892A42D074CD8D82CAC5BBEB8F">University of Gothenburg, SE- 405 30 Göteborg, Sweden. & Email: christer. erseus @ bioenv. gu. se & urn: lsid: zoobank. org: author: D 98 F 606 A-B 273 - 4 F 50 - 95 F 5 - C 35 F 17 B 12 C 85</mods:affiliation>
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<mods:nameIdentifier id="167E0EE4DF57126746C8ADD81688D9A2" type="email">christer.erseus@bioenv.gu.se</mods:nameIdentifier>
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<mods:typeOfResource id="8BAF88385A7A09872436D4313C99C534">text</mods:typeOfResource>
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<mods:titleInfo id="1A0BCD2B8FD4D543E951F18CD445BE03">
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<mods:title id="D93D3E58115FBBD394176076A16D3BF9">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="D58CD81C664DEDBBEF98BFF7840F17ED">
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<mods:date id="3FA2F5F9B8ABD120A1CBBB66C9453DA8">2017</mods:date>
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<mods:detail id="F7D76446483EB7F9DBD214BD07234FB1" type="pubDate">
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<mods:number id="1C80EAE1D533F9A5659F87EB04758474">2017-10-24</mods:number>
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<mods:detail id="A7A8526574ED50D654B0462A4D149869" type="issue">
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<mods:number id="327CDD63D947E82C5EEAD8EA66DB52A8">361</mods:number>
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<mods:start id="777581733894D89AB0592AD9A07EB9C7">1</mods:start>
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<mods:end id="97AFFB62C24E32A531A940367098FCD2">46</mods:end>
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</mods:extent>
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<mods:classification id="00969CA0C9228B4EDC912F29BDAA2343">journal article</mods:classification>
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<mods:identifier id="5B731D1C06B13D7B73031225EB3F07B0" type="CLB-Dataset">21986</mods:identifier>
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<mods:identifier id="4E90D22AA1A71545324C841265A5B637" type="DOI">10.5852/ejt.2017.361</mods:identifier>
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<mods:identifier id="6F19EE48722F3DFFACDF39C464EBE329" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="AA4BE2930BA5B8864445E7FCC2D36BB9" type="Zenodo-Dep">3836508</mods:identifier>
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<treatment id="03BC8649FF8DFF81FD81FDF5DD7BB298" ID-DOI="http://doi.org/10.5281/zenodo.3851842" ID-GBIF-Taxon="164243723" ID-Zenodo-Dep="3851842" LSID="urn:lsid:plazi:treatment:03BC8649FF8DFF81FD81FDF5DD7BB298" httpUri="http://treatment.plazi.org/id/03BC8649FF8DFF81FD81FDF5DD7BB298" lastPageNumber="19" pageId="18" pageNumber="19">
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<subSubSection id="C30F64D4FF8DFF81FD81FDF5DD93B734" box="[598,990,556,583]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8BAA375FFF8DFF81FD81FDF5DD93B734" blockId="18.[598,990,556,583]" box="[598,990,556,583]" pageId="18" pageNumber="19">
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<heading id="D0E28033FF8DFF81FD81FDF5DD93B734" box="[598,990,556,583]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
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<taxonomicName id="4C154CDCFF8DFF81FD81FDF5DD93B734" ID-CoL="84NWF" authority="Michaelsen, 1933" authorityName="Michaelsen" authorityYear="1933" box="[598,990,556,583]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
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<emphasis id="B961EB4DFF8DFF81FD81FDF5DD4BB734" bold="true" box="[598,774,556,583]" italics="true" pageId="18" pageNumber="19">Guestphalinus</emphasis>
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<bibRefCitation id="EF844AAEFF8DFF81FCDAFDF4DD93B734" author="Michaelsen W." box="[781,990,557,583]" pageId="18" pageNumber="19" pagination="1 - 19" refId="ref24546" refString="Michaelsen W. 1933. Uber Hohlen Oligochaten. Mitteilungen uber Hohlen- und Karstforschung, Zeitschrift des Hauptverbandes Deutscher Hohlenforscher 1: 1 - 19." type="journal article" year="1933">Michaelsen, 1933</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F64D4FF8DFF81FF6AFDB6DD79B7C4" pageId="18" pageNumber="19" type="type_taxon">
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<paragraph id="8BAA375FFF8DFF81FF6AFDB6DF15B7FB" blockId="18.[189,344,622,649]" box="[189,344,622,649]" pageId="18" pageNumber="19">
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<heading id="D0E28033FF8DFF81FF6AFDB6DF15B7FB" bold="true" box="[189,344,622,649]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
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<emphasis id="B961EB4DFF8DFF81FF6AFDB6DF15B7FB" bold="true" box="[189,344,622,649]" pageId="18" pageNumber="19">
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<typeStatus id="54AE89FDFF8DFF81FF6AFDB6DEB7B7FA" box="[189,250,623,649]" pageId="18" pageNumber="19">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAA375FFF8DFF81FF6AFD44DD79B7C4" blockId="18.[189,820,669,696]" box="[189,820,669,696]" pageId="18" pageNumber="19">
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<taxonomicName id="4C154CDCFF8DFF81FF6AFD44DD7DB7C4" authority="Michaelsen, 1933" authorityName="Michaelsen" authorityYear="1933" box="[189,816,669,696]" class="Clitellata" family="Dorydrilidae" genus="Dorydrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="wiardi" subGenus="Guestphalinus">
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<emphasis id="B961EB4DFF8DFF81FF6AFD44DF73B7C4" box="[189,318,669,695]" italics="true" pageId="18" pageNumber="19">Dorydrilus</emphasis>
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(
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<emphasis id="B961EB4DFF8DFF81FE98FD44DFB5B7C4" box="[335,504,669,695]" italics="true" pageId="18" pageNumber="19">Guestphalinus</emphasis>
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)
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<emphasis id="B961EB4DFF8DFF81FDDEFD44DC18B7C4" box="[521,597,669,695]" italics="true" pageId="18" pageNumber="19">wiardi</emphasis>
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<bibRefCitation id="EF844AAEFF8DFF81FD8BFD44DD7DB7C4" author="Michaelsen W." box="[604,816,669,695]" pageId="18" pageNumber="19" pagination="1 - 19" refId="ref24546" refString="Michaelsen W. 1933. Uber Hohlen Oligochaten. Mitteilungen uber Hohlen- und Karstforschung, Zeitschrift des Hauptverbandes Deutscher Hohlenforscher 1: 1 - 19." type="journal article" year="1933">Michaelsen, 1933</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F64D4FF8DFF81FF6AFD06DCEFB1C7" pageId="18" pageNumber="19" type="discussion">
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<paragraph id="8BAA375FFF8DFF81FF6AFD06DFC4B78A" blockId="18.[189,393,735,761]" box="[189,393,735,761]" pageId="18" pageNumber="19">
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<heading id="D0E28033FF8DFF81FF6AFD06DFC4B78A" bold="true" box="[189,393,735,761]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
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<emphasis id="B961EB4DFF8DFF81FF6AFD06DFC4B78A" bold="true" box="[189,393,735,761]" pageId="18" pageNumber="19">Included species</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAA375FFF8DFF81FF6AFCD7DCE9B65B" blockId="18.[189,839,782,879]" box="[189,676,782,808]" pageId="18" pageNumber="19">
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<taxonomicName id="4C154CDCFF8DFF81FF6AFCD7DCE9B65B" authority="(Michaelsen, 1933)" baseAuthorityName="Michaelsen" baseAuthorityYear="1933" box="[189,676,782,808]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="wiardi">
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<emphasis id="B961EB4DFF8DFF81FF6AFCD7DFF7B65B" box="[189,442,782,808]" italics="true" pageId="18" pageNumber="19">Guestphalinus wiardi</emphasis>
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(
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<bibRefCitation id="EF844AAEFF8DFF81FE1EFCD7DCD1B65B" author="Michaelsen W." box="[457,668,782,808]" pageId="18" pageNumber="19" pagination="1 - 19" refId="ref24546" refString="Michaelsen W. 1933. Uber Hohlen Oligochaten. Mitteilungen uber Hohlen- und Karstforschung, Zeitschrift des Hauptverbandes Deutscher Hohlenforscher 1: 1 - 19." type="journal article" year="1933">Michaelsen, 1933</bibRefCitation>
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)
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</taxonomicName>
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</paragraph>
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<paragraph id="8BAA375FFF8DFF81FF6AFCE8DCA2B63F" blockId="18.[189,839,782,879]" box="[189,751,817,844]" pageId="18" pageNumber="19">
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<taxonomicName id="4C154CDCFF8DFF81FF6AFCE8DCC0B638" authority="Fend & Rodriguez" authorityName="Fend & Rodriguez & Achurra & Erséus" authorityYear="2017" box="[189,653,817,843]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="exilis" status="sp. nov.">
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<emphasis id="B961EB4DFF8DFF81FF6AFCE8DFE1B638" box="[189,428,817,843]" italics="true" pageId="18" pageNumber="19">Guestphalinus exilis</emphasis>
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Fend & Rodriguez
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</taxonomicName>
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<taxonomicNameLabel id="A2525636FF8DFF81FD42FCEBDCA2B63F" box="[661,751,818,844]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BAA375FFF8DFF81FF6AFC8CDD0AB61C" blockId="18.[189,839,782,879]" box="[189,839,853,879]" pageId="18" pageNumber="19">
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<taxonomicName id="4C154CDCFF8DFF81FF6AFC8CDCA8B61C" authority="Fend & Rodriguez" authorityName="Fend & Rodriguez & Achurra & Erséus" authorityYear="2017" box="[189,741,853,879]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="elephantinus" status="sp. nov.">
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<emphasis id="B961EB4DFF8DFF81FF6AFC8CDC49B61C" box="[189,516,853,879]" italics="true" pageId="18" pageNumber="19">Guestphalinus elephantinus</emphasis>
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Fend & Rodriguez
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</taxonomicName>
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<taxonomicNameLabel id="A2525636FF8DFF81FD3AFC8CDD0AB61C" box="[749,839,853,879]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BAA375FFF8DFF81FF6AFC4EDD99B6C2" blockId="18.[189,980,918,945]" box="[189,980,918,945]" pageId="18" pageNumber="19">
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<emphasis id="B961EB4DFF8DFF81FF6AFC4EDFE2B6C3" bold="true" box="[189,431,918,945]" pageId="18" pageNumber="19">Emended diagnosis</emphasis>
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(modified from
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<bibRefCitation id="EF844AAEFF8DFF81FDA7FC4FDD77B6C3" author="Michaelsen W." box="[624,826,918,944]" pageId="18" pageNumber="19" pagination="1 - 19" refId="ref24546" refString="Michaelsen W. 1933. Uber Hohlen Oligochaten. Mitteilungen uber Hohlen- und Karstforschung, Zeitschrift des Hauptverbandes Deutscher Hohlenforscher 1: 1 - 19." type="journal article" year="1933">Michaelsen 1933</bibRefCitation>
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;
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<bibRefCitation id="EF844AAEFF8DFF81FC9EFC4FDD86B6C3" author="Cook D. G." box="[841,971,918,944]" pageId="18" pageNumber="19" pagination="200 - 285" refId="ref22982" refString="Cook D. G. 1971. Family Lumbriculidae. In: Brinkhurst R. O. & Jamieson B. G. M. (eds) Aquatic Oligochaeta of the World: 200 - 285. University of Toronto Press, Toronto." type="book chapter" year="1971">Cook 1971</bibRefCitation>
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)
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</paragraph>
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<paragraph id="8BAA375FFF8DFF81FF6AFC1CDCEFB1C7" blockId="18.[189,1399,965,1204]" pageId="18" pageNumber="19">
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Medium-sized to large worms with a filiform proboscis. Chaetae two per bundle. Paired testes in VIII and IX, one pair of ovaries in X. Male pores paired in IX, near 9/10. Spermathecal pores paired in IX, anterior to the male pores. Petiolate copulatory glands (= Pubertätsdrüsen in
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<bibRefCitation id="EF844AAEFF8DFF81FC31FBD5DAFDB155" author="Michaelsen W." box="[998,1200,1036,1062]" pageId="18" pageNumber="19" pagination="1 - 19" refId="ref24546" refString="Michaelsen W. 1933. Uber Hohlen Oligochaten. Mitteilungen uber Hohlen- und Karstforschung, Zeitschrift des Hauptverbandes Deutscher Hohlenforscher 1: 1 - 19." type="journal article" year="1933">Michaelsen 1933</bibRefCitation>
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) associated with male and/or spermathecal pores. Male duct semiprosoporous. Penes absent. Atria elongate, cylindrical to club-shaped, ental part loosely covered with multicellular, pyriform prostate glands. Anterior vasa deferentia form a loop in the pre-atrial segment before entering the atrial segment. Paired spermathecae in the atrial segment, anterior to the atria.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F64D4FF8DFF81FF6AFB02DD63B0FC" pageId="18" pageNumber="19" type="distribution">
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<paragraph id="8BAA375FFF8DFF81FF6AFB02DF1BB186" blockId="18.[189,342,1243,1269]" box="[189,342,1243,1269]" pageId="18" pageNumber="19">
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<heading id="D0E28033FF8DFF81FF6AFB02DF1BB186" bold="true" box="[189,342,1243,1269]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
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<emphasis id="B961EB4DFF8DFF81FF6AFB02DF1BB186" bold="true" box="[189,342,1243,1269]" pageId="18" pageNumber="19">Distribution</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAA375FFF8DFF81FF6AFAD2DD63B0FC" blockId="18.[189,1399,1290,1423]" pageId="18" pageNumber="19">
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Europe and northwestern
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<collectingCountry id="F30277CFFF8DFF81FE26FAD2DC61B056" box="[497,556,1291,1317]" name="United States of America" pageId="18" pageNumber="19">USA</collectingCountry>
|
|
(present study).
|
|
<taxonomicName id="4C154CDCFF8DFF81FD21FAD3DDB8B057" baseAuthorityName="Michaelsen" baseAuthorityYear="1933" box="[758,1013,1290,1316]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="species" species="wiardi">
|
|
<emphasis id="B961EB4DFF8DFF81FD21FAD3DDB8B057" box="[758,1013,1290,1316]" italics="true" pageId="18" pageNumber="19">Guestphalinus wiardi</emphasis>
|
|
</taxonomicName>
|
|
is rarely reported, but is known from subterranean or spring habitats in
|
|
<collectingCountry id="F30277CFFF8DFF81FD79FAF7DD56B03B" box="[686,795,1326,1352]" name="Germany" pageId="18" pageNumber="19">Germany</collectingCountry>
|
|
(
|
|
<bibRefCitation id="EF844AAEFF8DFF81FCE5FAF7DA4CB03B" author="Michaelsen W." box="[818,1025,1326,1352]" pageId="18" pageNumber="19" pagination="1 - 19" refId="ref24546" refString="Michaelsen W. 1933. Uber Hohlen Oligochaten. Mitteilungen uber Hohlen- und Karstforschung, Zeitschrift des Hauptverbandes Deutscher Hohlenforscher 1: 1 - 19." type="journal article" year="1933">Michaelsen 1933</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EF844AAEFF8DFF81FBC1FAF7DABFB03B" author="Griepenburg W." box="[1046,1266,1326,1352]" pageId="18" pageNumber="19" pagination="73 - 116" refId="ref23879" refString="Griepenburg W. 1941. Ein Beitrag zur Kenntnis der Wurmfauna der westfalischen Hohlen. Decheniana 100 B: 73 - 116." type="journal article" year="1941">Griepenburg 1941</bibRefCitation>
|
|
),
|
|
<collectingCountry id="F30277CFFF8DFF81FAC7FAF7DB3AB03B" box="[1296,1399,1326,1352]" name="Slovenia" pageId="18" pageNumber="19">Slovenia</collectingCountry>
|
|
(
|
|
<bibRefCitation id="EF844AAEFF8DFF81FF10FA88DF1EB01F" author="Hrabe S." box="[199,339,1361,1388]" pageId="18" pageNumber="19" pagination="39 - 50" refId="ref24075" refString="Hrabe S. 1973. On a collection of Oligochaeta from various parts of Yugoslavia. Biloski vestnik Ljubljana 21: 39 - 50." type="journal article" year="1973">Hrabě 1973</bibRefCitation>
|
|
) and
|
|
<collectingCountry id="F30277CFFF8DFF81FE4EFA88DF82B018" box="[409,463,1361,1387]" name="Italy" pageId="18" pageNumber="19">Italy</collectingCountry>
|
|
(new material used for the present description, see below), with other records from mountain streams in Crimea (
|
|
<bibRefCitation id="EF844AAEFF8DFF81FD8DFAACDD50B0FC" author="Dembitsky E. B." box="[602,797,1397,1423]" pageId="18" pageNumber="19" pagination="12 - 16" refId="ref23145" refString="Dembitsky E. B. 1987. Novyj rod vod nyh maloscetinkovyh cervej v faune Sovetskogo Sojuza. In: Kacalova O. L. & Parele E. A. (eds) Vodnye Maloscetinkovye Cervi. Materialy Sestogo Vsesojuznogo Simpoziuma, Salaspils 27 - 30 Aprelja 1987, Riga: 12 - 16. [In Russian: A new genus of oligochaetes in fauna of the Soviet Union. In: Kacalova O. L. & Parele E. A. (eds) Aquatic Oligochaeta, Materials of the 6 th Symposium, Salaspils, 27 - 30 April 1987]." type="book chapter" year="1987">Dembitsky 1987</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30F64D4FF8DFF81FF6AFA6EDD7BB298" pageId="18" pageNumber="19" type="discussion">
|
|
<paragraph id="8BAA375FFF8DFF81FF6AFA6EDF62B0A2" blockId="18.[189,303,1463,1489]" box="[189,303,1463,1489]" pageId="18" pageNumber="19">
|
|
<heading id="D0E28033FF8DFF81FF6AFA6EDF62B0A2" bold="true" box="[189,303,1463,1489]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
|
|
<emphasis id="B961EB4DFF8DFF81FF6AFA6EDF62B0A2" bold="true" box="[189,303,1463,1489]" pageId="18" pageNumber="19">Remarks</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="8BAA375FFF8DFF81FF6AFA3FDDF0B24C" blockId="18.[189,1399,1509,1855]" pageId="18" pageNumber="19">
|
|
Among the lumbriculids with a filiform proboscis,
|
|
<taxonomicName id="4C154CDCFF8DFF81FCB9FA3CDA5AB08C" authorityName="Michaelsen" authorityYear="1933" box="[878,1047,1509,1535]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
|
|
<emphasis id="B961EB4DFF8DFF81FCB9FA3CDA5AB08C" box="[878,1047,1509,1535]" italics="true" pageId="18" pageNumber="19">Guestphalinus</emphasis>
|
|
</taxonomicName>
|
|
is distinguished from the semiprosoporous species of
|
|
<taxonomicName id="4C154CDCFF8DFF81FDC1F9D0DCE1B350" authorityName="Hoffmeister" authorityYear="1843" box="[534,684,1545,1571]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
|
|
<emphasis id="B961EB4DFF8DFF81FDC1F9D0DCE1B350" box="[534,684,1545,1571]" italics="true" pageId="18" pageNumber="19">Rhynchelmis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C154CDCFF8DFF81FD68F9D0DAE7B350" authority="Smith, 1900" authorityName="Smith" authorityYear="1900" box="[703,1194,1545,1571]" class="Clitellata" family="Lumbriculidae" genus="Eclipidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="subGenus" subGenus="Premnodrilus">
|
|
<emphasis id="B961EB4DFF8DFF81FD68F9D0DD07B350" box="[703,842,1545,1571]" italics="true" pageId="18" pageNumber="19">Eclipidrilus</emphasis>
|
|
(
|
|
<emphasis id="B961EB4DFF8DFF81FC88F9D0DDB2B350" box="[863,1023,1545,1571]" italics="true" pageId="18" pageNumber="19">Premnodrilus</emphasis>
|
|
) Smith, 1900
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C154CDCFF8DFF81FB3CF9D0DFA5B335" authority="Fend & Rodriguez, 2003" authorityName="Fend & Rodriguez" authorityYear="2003" class="Clitellata" family="Lumbriculidae" genus="Eremidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
|
|
<emphasis id="B961EB4DFF8DFF81FB3CF9D0DB3BB350" box="[1259,1398,1545,1571]" italics="true" pageId="18" pageNumber="19">Eremidrilus</emphasis>
|
|
<bibRefCitation id="EF844AAEFF8DFF81FF6AF9F5DFA5B335" author="Fend S. V. & Rodriguez P." box="[189,488,1580,1606]" pageId="18" pageNumber="19" pagination="515 - 542" refId="ref23707" refString="Fend S. V. & Rodriguez P. 2003. Eremidrilus n. gen. (Annelida, Clitellata, Lumbriculidae) and new species from California, U. S. A. Canadian Journal of Zoology 81: 515 - 542. https: // doi. org / 10.1139 / z 02 - 235" type="journal article" year="2003">Fend & Rodriguez, 2003</bibRefCitation>
|
|
</taxonomicName>
|
|
by the location of the male pores in IX (instead of X), the spermathecae in the atrial segment, the anterior vasa deferentia entering the pre-atrial segment, and the morphology of the elongate spermathecae. In addition to being prosoporous, the two
|
|
<taxonomicName id="4C154CDCFF8DFF81FB85F9AADAAEB3FE" authorityName="Altman" authorityYear="1936" box="[1106,1251,1651,1677]" class="Clitellata" family="Lumbriculidae" genus="Kincaidiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
|
|
<emphasis id="B961EB4DFF8DFF81FB85F9AADAAEB3FE" box="[1106,1251,1651,1677]" italics="true" pageId="18" pageNumber="19">Kincaidiana</emphasis>
|
|
</taxonomicName>
|
|
species are easily distinguished from
|
|
<taxonomicName id="4C154CDCFF8DFF81FE20F94EDCEDB3C2" authorityName="Michaelsen" authorityYear="1933" box="[503,672,1687,1713]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
|
|
<emphasis id="B961EB4DFF8DFF81FE20F94EDCEDB3C2" box="[503,672,1687,1713]" italics="true" pageId="18" pageNumber="19">Guestphalinus</emphasis>
|
|
</taxonomicName>
|
|
by the presence of spermathecae also in the first two postatrial segments, and by a different
|
|
<typeStatus id="54AE89FDFF8DFF81FDB6F962DCD9B3A6" box="[609,660,1723,1749]" pageId="18" pageNumber="19">type</typeStatus>
|
|
of modified chaetae in several pre-clitellar segments.
|
|
<taxonomicName id="4C154CDCFF8DFF81FAF5F963DB3AB3A7" box="[1314,1399,1722,1748]" class="Clitellata" family="Lumbriculidae" genus="Uktena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
|
|
<emphasis id="B961EB4DFF8DFF81FAF5F963DB3AB3A7" box="[1314,1399,1722,1748]" italics="true" pageId="18" pageNumber="19">Uktena</emphasis>
|
|
</taxonomicName>
|
|
is distinguished from other proboscis-bearing lumbriculids by having atria and spermathecae in VIII (rather than IX), in addition to characters unique within the family: spermatophores, a spermathecal copulatory organ, and multiple genital chaetae (
|
|
<bibRefCitation id="EF844AAEFF8DFF81FD39F8FDDDE0B24C" author="Fend S. V. & Rodriguez P. & Lenat D. R." box="[750,941,1828,1855]" pageId="18" pageNumber="19" pagination="411 - 424" refId="ref23765" refString="Fend S. V., Rodriguez P. & Lenat D. R. 2015. Uktena riparia n. gen., n. sp. (Annelida, Clitellata, Lumbriculidae), a new spermatophore-producing oligochaete. Zootaxa 3994 (3): 411 - 424. https: // doi. org / 10.11646 / zootaxa. 3994.3.5" type="journal article" year="2015">
|
|
Fend
|
|
<emphasis id="B961EB4DFF8DFF81FCE7F8FCDD27B24C" box="[816,874,1829,1855]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
|
2015
|
|
</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BAA375FFF8DFF81FF6AF8BFDD7BB298" blockId="18.[189,1399,1894,2027]" pageId="18" pageNumber="19">
|
|
<taxonomicName id="4C154CDCFF8DFF81FF6AF8BFDF2BB2F3" authorityName="Michaelsen" authorityYear="1933" box="[189,358,1894,1920]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
|
|
<emphasis id="B961EB4DFF8DFF81FF6AF8BFDF2BB2F3" box="[189,358,1894,1920]" italics="true" pageId="18" pageNumber="19">Guestphalinus</emphasis>
|
|
</taxonomicName>
|
|
was originally described as a subgenus of
|
|
<taxonomicName id="4C154CDCFF8DFF81FCBEF8BFDB3DB2F2" authority="Piguet, 1913" authorityName="Piguet" authorityYear="1913" box="[873,1392,1894,1921]" class="Clitellata" family="Dorydrilidae" genus="Dorydrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="18" pageNumber="19" phylum="Annelida" rank="genus">
|
|
<emphasis id="B961EB4DFF8DFF81FCBEF8BFDDA7B2F3" box="[873,1002,1894,1920]" italics="true" pageId="18" pageNumber="19">Dorydrilus</emphasis>
|
|
Piguet, 1913 (
|
|
<bibRefCitation id="EF844AAEFF8DFF81FB4CF8BFDB2BB2F3" author="Michaelsen W." box="[1179,1382,1894,1920]" pageId="18" pageNumber="19" pagination="1 - 19" refId="ref24546" refString="Michaelsen W. 1933. Uber Hohlen Oligochaten. Mitteilungen uber Hohlen- und Karstforschung, Zeitschrift des Hauptverbandes Deutscher Hohlenforscher 1: 1 - 19." type="journal article" year="1933">Michaelsen 1933</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
, but was later elevated to generic status (
|
|
<bibRefCitation id="EF844AAEFF8DFF81FD7DF853DD7AB2D7" author="Hrabe S." box="[682,823,1930,1956]" pageId="18" pageNumber="19" pagination="3 - 18" refId="ref24011" refString="Hrabe S. 1936. Uber Dorydrilus (Piguetia) mirabilis n. subgen., n. sp. aus einem Sodbrunnen in der Umgebung von Basel sowie uber Dorydrilus (Dorydrilus) michaelseni Pig. und Bichaeta sanguinea Bret. Spisy vydavane prirodovedeckou fakultou Masarykovy University [Aus dem Zoologischen Institut der Masaryk-Universitat in Brno] 227: 3 - 18." type="journal article" year="1936">Hrabě 1936</bibRefCitation>
|
|
, although spelled
|
|
<emphasis id="B961EB4DFF8DFF81FBCFF853DA8CB2D7" box="[1048,1217,1930,1956]" italics="true" pageId="18" pageNumber="19">Questphalinus</emphasis>
|
|
in that paper). This decision, although considered provisional in the 1936 paper, has been maintained in subsequent literature (
|
|
<bibRefCitation id="EF844AAEFF8DFF81FEEFF808DFF7B298" author="Cook D. G." box="[312,442,2001,2027]" pageId="18" pageNumber="19" pagination="200 - 285" refId="ref22982" refString="Cook D. G. 1971. Family Lumbriculidae. In: Brinkhurst R. O. & Jamieson B. G. M. (eds) Aquatic Oligochaeta of the World: 200 - 285. University of Toronto Press, Toronto." type="book chapter" year="1971">Cook 1971</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EF844AAEFF8DFF81FE1EF808DC1EB298" author="Hrabe S." box="[457,595,2001,2027]" pageId="18" pageNumber="19" pagination="39 - 50" refId="ref24075" refString="Hrabe S. 1973. On a collection of Oligochaeta from various parts of Yugoslavia. Biloski vestnik Ljubljana 21: 39 - 50." type="journal article" year="1973">Hrabě 1973</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EF844AAEFF8DFF81FDB5F808DD68B298" author="Dembitsky E. B." box="[610,805,2001,2027]" pageId="18" pageNumber="19" pagination="12 - 16" refId="ref23145" refString="Dembitsky E. B. 1987. Novyj rod vod nyh maloscetinkovyh cervej v faune Sovetskogo Sojuza. In: Kacalova O. L. & Parele E. A. (eds) Vodnye Maloscetinkovye Cervi. Materialy Sestogo Vsesojuznogo Simpoziuma, Salaspils 27 - 30 Aprelja 1987, Riga: 12 - 16. [In Russian: A new genus of oligochaetes in fauna of the Soviet Union. In: Kacalova O. L. & Parele E. A. (eds) Aquatic Oligochaeta, Materials of the 6 th Symposium, Salaspils, 27 - 30 April 1987]." type="book chapter" year="1987">Dembitsky 1987</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |