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<mods:title id="8F5E37D98628B9E15F57C94B34CA324B">Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species</mods:title>
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<mods:namePart id="1274C5942B64B01701C817DC20CCE447">Guéorguiev, Borislav</mods:namePart>
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<mods:namePart id="D8CA042D29B329ABB351DEEE398B1A02">Liang, Hongbin</mods:namePart>
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<treatment id="03BC5E5B296AFF94FF4BFEDAE86CFF04" ID-DOI="http://doi.org/10.5281/zenodo.4480017" ID-GBIF-Taxon="177139737" ID-Zenodo-Dep="4480017" LSID="urn:lsid:plazi:treatment:03BC5E5B296AFF94FF4BFEDAE86CFF04" httpUri="http://treatment.plazi.org/id/03BC5E5B296AFF94FF4BFEDAE86CFF04" lastPageId="43" lastPageNumber="43" pageId="42" pageNumber="43">
<subSubSection id="C30FBCC6296AFF95FF4BFEDAEDA5FE75" box="[151,395,366,393]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8BAAEF4D296AFF95FF4BFEDAEDA5FE75" blockId="42.[151,395,366,393]" box="[151,395,366,393]" pageId="42" pageNumber="43">
<heading id="D0E25821296AFF95FF4BFEDAEDA5FE75" bold="true" box="[151,395,366,393]" fontSize="11" level="2" pageId="42" pageNumber="43" reason="4">
<taxonomicName id="4C1594CE296AFF95FF4BFEDAEDA5FE75" ID-CoL="8KXMQ" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[151,395,366,393]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FF4BFEDAEDA5FE75" bold="true" box="[151,395,366,393]" pageId="42" pageNumber="43">
<emphasis id="B961335F296AFF95FF4BFEDAECCCFE74" bold="true" box="[151,226,366,393]" italics="true" pageId="42" pageNumber="43">Oodes</emphasis>
Bonelli, 1810
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296AFF95FF4BFE07EDE5FE31" box="[151,459,435,460]" pageId="42" pageNumber="43" type="reference_group">
<paragraph id="8BAAEF4D296AFF95FF4BFE07EDE5FE31" blockId="42.[151,822,435,490]" box="[151,459,435,460]" pageId="42" pageNumber="43">
<taxonomicName id="4C1594CE296AFF95FF4BFE07ED42FE36" ID-CoL="8KXMQ" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[151,364,435,459]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FF4BFE07ECF6FE36" box="[151,216,435,459]" italics="true" pageId="42" pageNumber="43">Oodes</emphasis>
Bonelli, 1810
</taxonomicName>
: tab. syn.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296AFF95FF4BFE66EF18FE17" box="[151,822,466,490]" pageId="42" pageNumber="43" type="type_taxon">
<paragraph id="8BAAEF4D296AFF95FF4BFE66EF18FE17" blockId="42.[151,822,435,490]" box="[151,822,466,490]" pageId="42" pageNumber="43">
<typeStatus id="54AE51EF296AFF95FF4BFE66ECE7FE17" box="[151,201,466,490]" pageId="42" pageNumber="43">Type</typeStatus>
species:
<taxonomicName id="4C1594CE296AFF95FEFAFE66EEB1FE17" ID-CoL="QWW7" authority="Fabricius, 1792" authorityName="Fabricius" authorityYear="1792" box="[294,671,466,490]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="helopioides">
<emphasis id="B961335F296AFF95FEFAFE66EDD7FE17" box="[294,505,466,490]" italics="true" pageId="42" pageNumber="43">Carabus helopioides</emphasis>
<bibRefCitation id="EF8492BC296AFF95FE23FE66EEB1FE17" author="Fabricius, J. C." box="[511,671,466,490]" pageId="42" pageNumber="43" refId="ref61063" refString="Fabricius, J. C. (1792) Entomologia systematica emendata et aucta, secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tomus I. Pars I. C. G. Proft, Hafniae, xx + 330 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1792">Fabricius, 1792</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296AFF95FF4BFDBBEFBEFDBB" pageId="42" pageNumber="43" type="reference_group">
<paragraph id="8BAAEF4D296AFF95FF4BFDBBEFBEFDBB" blockId="42.[151,1436,527,582]" pageId="42" pageNumber="43">
<emphasis id="B961335F296AFF95FF4BFDBBED23FDDA" bold="true" box="[151,269,527,551]" pageId="42" pageNumber="43">References</emphasis>
.
<bibRefCitation id="EF8492BC296AFF95FEC0FDBBEE3CFDDA" author="Ganglbauer, L." box="[284,530,527,551]" pageId="42" pageNumber="43" refId="ref61349" refString="Ganglbauer, L. (1891 a) Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. Erster Band. Familienreihe Caraboidea. C. Gerold's Sohn, Wien, iii + 557 pp. https: // doi. org / 10.5962 / bhl. title. 8764" type="book" year="1891">Ganglbauer 1891a: 383</bibRefCitation>
;
<bibRefCitation id="EF8492BC296AFF95FDC3FDBBEED4FDDA" author="Peringuey, L." box="[543,762,527,551]" pageId="42" pageNumber="43" pagination="99 - 623" refId="ref63904" refString="Peringuey, L. (1896) Descriptive catalogue of the Coleoptera of South Africa. Part II. Cicindelidae supplement. Carabidae. The Transactions of the South African Philosophical Society, 7, 99 - 623 + i-xiv." type="journal article" year="1896">Péringuey 1896: 529</bibRefCitation>
;
<bibRefCitation id="EF8492BC296AFF95FCDBFDA4EF9FFDDA" author="Porta, A." box="[775,945,527,551]" pageId="42" pageNumber="43" refId="ref63945" refString="Porta, A. (1923) Fauna Coleopterorum Italica. Vol. I. Adephaga. Cicindelidae, Carabidae, Haliplidae, Hygrobiidae, Dyticidae, Gyrinidae, Rhisodidae, Paussidae. Stabilimento Tipographico Piacentino, Piacenza, 285 pp." type="book" year="1923">Porta 1923: 213</bibRefCitation>
;
<bibRefCitation id="EF8492BC296AFF95FC61FDBBE8B6FDDA" author="Andrewes, H. E." box="[957,1176,527,551]" pageId="42" pageNumber="43" pagination="203 - 208" refId="ref59739" refString="Andrewes, H. E. (1940) Keys to some Indian genera of Carabidae (Col.). X. The genus Oodes. The Proceedings of the Royal Entomological Society of London, Series B, 9, 203 - 208. https: // doi. org / 10.1111 / j. 1365 - 3113.1940. tb 00329. x" type="journal article" year="1940">Andrewes 1940: 203</bibRefCitation>
;
<bibRefCitation id="EF8492BC296AFF95FB79FDBBE9B7FDDA" author="Jeannel, R." box="[1189,1433,527,551]" pageId="42" pageNumber="43" pagination="572 - 1173" refId="ref62241" refString="Jeannel, R. (1942) Coleopteres carabiques. Deuxieme Partie. In: Faune de France. 40. Librairie de la Faculte des Sciences, Paris, pp. 572 - 1173." type="book chapter" year="1942">Jeannel 1942: 980981</bibRefCitation>
;
<bibRefCitation id="EF8492BC296AFF95FF1BFD9AED98FDBB" author="Jeannel, R." box="[199,438,558,582]" pageId="42" pageNumber="43" pagination="767 - 1146" refId="ref62277" refString="Jeannel, R. (1949) ColeiopteIres carabiques de la reigion Malgache. Troisieme Partie. Faune de l'Empire Francais. IX. Office de la Recherche scientifique coloniale, Paris, 767 - 1146." type="book chapter" year="1949">Jeannel 1949: 833834</bibRefCitation>
;
<bibRefCitation id="EF8492BC296AFF95FE1CFD9AEEAEFDBB" author="Habu, A." box="[448,640,558,582]" pageId="42" pageNumber="43" pagination="49 - 73" refId="ref61741" refString="Habu, A. (1956) On the genera and species of the Oodini (Coleoptera, Carabidae) from Japan. The Bulletin of the National Institute of Agricultural Sciences, Series C, 6, 49 - 73." type="journal article" year="1956">Habu 1956: 8081</bibRefCitation>
;
<bibRefCitation id="EF8492BC296AFF95FD56FD9AEFA5FDBB" author="Bousquet, Y." box="[650,907,558,582]" pageId="42" pageNumber="43" pagination="443 - 537" refId="ref60231" refString="Bousquet, Y. (1996) Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae). The Canadian Entomologist, 128, 443 - 537. https: // doi. org / 10.4039 / Ent 128443 - 3" type="journal article" year="1996">Bousquet 1996: 469471</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296AFF95FF4BFDC0EFAFFC77" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8BAAEF4D296AFF95FF4BFDC0EFE2FD72" blockId="42.[151,972,628,655]" box="[151,972,628,655]" pageId="42" pageNumber="43">
<emphasis id="B961335F296AFF95FF4BFDC0ED24FD73" bold="true" box="[151,266,628,654]" pageId="42" pageNumber="43">Diagnosis</emphasis>
. The species of this genus share the following character states:
</paragraph>
<paragraph id="8BAAEF4D296AFF95FF4BFD08EE68FD2B" blockId="42.[151,1122,700,907]" box="[151,582,700,726]" pageId="42" pageNumber="43">
(1)
<collectingRegion id="49D121AF296AFF95FF1BFD08ED08FD2B" box="[199,294,700,726]" country="Myanmar" name="Chin" pageId="42" pageNumber="43">Mentum</collectingRegion>
tooth with apex rounded;
</paragraph>
<paragraph id="8BAAEF4D296AFF95FF4BFD54E84CFD06" blockId="42.[151,1122,700,907]" box="[151,1122,736,763]" pageId="42" pageNumber="43">(2) Submentum with medial setiferous punctures, without lateral setiferous punctures;</paragraph>
<paragraph id="8BAAEF4D296AFF95FF4BFCB0EF1DFCE3" blockId="42.[151,1122,700,907]" box="[151,819,772,799]" pageId="42" pageNumber="43">(3) Pronotum without setiferous punctures at basal margin;</paragraph>
<paragraph id="8BAAEF4D296AFF95FF4BFC9CEE92FCBE" blockId="42.[151,1122,700,907]" box="[151,700,808,835]" pageId="42" pageNumber="43">(4) Abdominal ventrite 3 with ambulatory setae;</paragraph>
<paragraph id="8BAAEF4D296AFF95FF4BFCF8EF12FC9B" blockId="42.[151,1122,700,907]" box="[151,828,844,871]" pageId="42" pageNumber="43">(5) Male protarsomere 3 rectangular, with subparallel sides;</paragraph>
<paragraph id="8BAAEF4D296AFF95FF4BFCC4EFAFFC77" blockId="42.[151,1122,700,907]" box="[151,897,880,907]" pageId="42" pageNumber="43">(6) Basal bulb of median lobe closed dorsally, with apical aileron.</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296AFF95FF1BFC0CEF99F862" pageId="42" pageNumber="43" type="description">
<paragraph id="8BAAEF4D296AFF95FF1BFC0CE962F9EF" blockId="42.[151,1437,952,2023]" pageId="42" pageNumber="43">
<emphasis id="B961335F296AFF95FF1BFC0CED7FFC2F" bold="true" box="[199,337,952,978]" pageId="42" pageNumber="43">Description</emphasis>
. As reported by
<bibRefCitation id="EF8492BC296AFF95FDC5FC0DEF23FC2F" author="Bousquet, Y." box="[537,781,952,979]" pageId="42" pageNumber="43" pagination="443 - 537" refId="ref60231" refString="Bousquet, Y. (1996) Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae). The Canadian Entomologist, 128, 443 - 537. https: // doi. org / 10.4039 / Ent 128443 - 3" type="journal article" year="1996">Bousquet (1996: 468</bibRefCitation>
, 470471) for the Nearctic representatives of the genera
<taxonomicName id="4C1594CE296AFF95FF4BFC68EDBAFC0B" authority="LeConte" authorityName="LeConte" authorityYear="1853" box="[151,404,988,1014]" class="Insecta" family="Carabidae" genus="Lachnocrepis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FF4BFC68ED00FC0B" box="[151,302,988,1014]" italics="true" pageId="42" pageNumber="43">Lachnocrepis</emphasis>
LeConte
</taxonomicName>
and
<taxonomicName id="4C1594CE296AFF95FE14FC68EE45FC0B" authority="Bonelli" authorityName="Bonelli" box="[456,619,988,1014]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FE14FC68EE3EFC0B" box="[456,528,988,1014]" italics="true" pageId="42" pageNumber="43">Oodes</emphasis>
Bonelli
</taxonomicName>
, with the following specifications (see also “Diagnosis”).
<emphasis id="B961335F296AFF95FB37FC6FEDB9FBE7" bold="true" italics="true" pageId="42" pageNumber="43">Microsculpture and micropunctuation</emphasis>
. Dorsal surface of head, pronotum and elytra with regular micropunctuation and isodiametric meshes well impressed which together form “punctate rosettes” microreticulation; micropunctuation usually, larger, denser and more regular on head than on pronotum and elytra; ventral surface without regular micropunctuation, with reduced sculpticells.
<emphasis id="B961335F296AFF95FE1BFBDFEE64FB7B" bold="true" box="[455,586,1131,1158]" italics="true" pageId="42" pageNumber="43">Chaetotaxy</emphasis>
. Elytral interval 3 with two discal setiferous punctures situated closer to stria 2, first situated in medial third, second in posterior third. Basal gonocoxite with lateroapical setae, without medioapical setae.Apical gonocoxite with or without one or two very small dorsolateral ensiform setae; dorsomedial ensiform seta present, located near base; nematiform setae present, situated equidistant from middle and apex, rarely absent.
<emphasis id="B961335F296AFF95FF33FB4FED19FAEB" bold="true" box="[239,311,1275,1302]" italics="true" pageId="42" pageNumber="43">Elytra</emphasis>
. Stria 7 as distinct as stria 6 or markedly reduced (in
<taxonomicName id="4C1594CE296AFF95FC78FB48E830FAEB" authorityName="A. Villa &amp; G. B. Villa" authorityYear="1833" box="[932,1054,1276,1303]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F296AFF95FC78FB48EF96FAEB" box="[932,952,1276,1302]" italics="true" pageId="42" pageNumber="43">O</emphasis>
.
<emphasis id="B961335F296AFF95FC14FB48E830FAEB" box="[968,1054,1276,1302]" italics="true" pageId="42" pageNumber="43">gracilis</emphasis>
</taxonomicName>
). Granulation in marginal furrow continuous or discontinuous.
<emphasis id="B961335F296AFF95FE3AFA94EE32FAC7" bold="true" box="[486,540,1312,1338]" italics="true" pageId="42" pageNumber="43">Legs</emphasis>
. Metacoxal basal sulcus well-developed, extending to lateral one-third to onefourth of coxa, rarely reduced, ending at medial fifth. Male mesotibia not or only moderately modified, with apical swelling. Meso- and metatarsomeres 14 (or 24) with or without dense and long pubescence on ventral side.
<emphasis id="B961335F296AFF95FF4BFA3FED10FA5B" bold="true" box="[151,318,1419,1446]" italics="true" pageId="42" pageNumber="43">Male genitalia</emphasis>
. Ostium elongate, reaching or not reaching basal bulb. Internal sac with or without sclerite.
<emphasis id="B961335F296AFF95FA99FA3FECD2FA37" bold="true" italics="true" pageId="42" pageNumber="43">Female genitalia</emphasis>
. Apical gonocoxite stout, sub-triangular.
<collectingRegion id="49D121AF296AFF95FD1BFA05EF26FA36" box="[711,776,1457,1483]" country="Turkey" name="Bursa" pageId="42" pageNumber="43">Bursa</collectingRegion>
copulatrix expanded distally or not expanded; spermatheca long, undifferentiated (seminal canal), twisted or coiled apically (see also
<bibRefCitation id="EF8492BC296AFF95FC2BFA61E8CCFA12" author="Bousquet, Y." box="[1015,1250,1492,1519]" pageId="42" pageNumber="43" pagination="443 - 537" refId="ref60231" refString="Bousquet, Y. (1996) Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae). The Canadian Entomologist, 128, 443 - 537. https: // doi. org / 10.4039 / Ent 128443 - 3" type="journal article" year="1996">Bousquet 1996: 532</bibRefCitation>
, figs 130, 132); spermathecal gland with atrium, connected near apex of seminal canal; common oviduct with villous canal.
</paragraph>
<paragraph id="8BAAEF4D296AFF95FF1BF9A8EF99F862" blockId="42.[151,1437,952,2023]" pageId="42" pageNumber="43">
<emphasis id="B961335F296AFF95FF1BF9A8EE04F9CB" bold="true" box="[199,554,1564,1590]" pageId="42" pageNumber="43">Monophyly and relationships</emphasis>
. The taxa of
<taxonomicName id="4C1594CE296AFF95FD14F9A8EF3EF9CB" box="[712,784,1564,1590]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FD14F9A8EF3EF9CB" box="[712,784,1564,1590]" italics="true" pageId="42" pageNumber="43">Oodes</emphasis>
</taxonomicName>
possess a single synapomorphy, the submentum without lateral setiferous punctures, suggesting that the genus may constitute a clade. In addition, all members of the genus have two other derived features, the diagnostic states (3) and (5), which are supposedly homoplastic (see “Monophyly and relationships” under
<taxonomicName id="4C1594CE296AFF95FD9BF93CEEE0F95F" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" box="[583,718,1672,1698]" class="Insecta" family="Carabidae" genus="Sundaoodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FD9BF93CEEE0F95F" box="[583,718,1672,1698]" italics="true" pageId="42" pageNumber="43">Sundaoodes</emphasis>
</taxonomicName>
). The representatives of
<taxonomicName id="4C1594CE296AFF95FC3AF93CE800F95F" box="[998,1070,1672,1698]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FC3AF93CE800F95F" box="[998,1070,1672,1698]" italics="true" pageId="42" pageNumber="43">Oodes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1594CE296AFF95FBB9F93CE8C2F95F" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" box="[1125,1260,1672,1698]" class="Insecta" family="Carabidae" genus="Sundaoodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FBB9F93CE8C2F95F" box="[1125,1260,1672,1698]" italics="true" pageId="42" pageNumber="43">Sundaoodes</emphasis>
</taxonomicName>
share character states (1) and (3)(6). However, the members of the two genera differ in: (1) chaetotaxy of submentum; (2) structure of basal bulb of median lobe; (3) shape of apical gonocoxite; (4) stage of differentiation of spermatheca; and (5) connection of spermathecal gland to spermatheca.
<taxonomicName id="4C1594CE296AFF95FD0DF940EF37F8F3" box="[721,793,1780,1806]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FD0DF940EF37F8F3" box="[721,793,1780,1806]" italics="true" pageId="42" pageNumber="43">Oodes</emphasis>
</taxonomicName>
is likely the adelphotaxon of
<emphasis id="B961335F296AFF95FBB5F940E8C1F8F3" box="[1129,1263,1780,1806]" italics="true" pageId="42" pageNumber="43">Pseudoodes</emphasis>
+
<taxonomicName id="4C1594CE296AFF95FAD2F940E9BBF8F3" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" box="[1294,1429,1780,1806]" class="Insecta" family="Carabidae" genus="Sundaoodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FAD2F940E9BBF8F3" box="[1294,1429,1780,1806]" italics="true" pageId="42" pageNumber="43">Sundaoodes</emphasis>
</taxonomicName>
. The spermathecae in three genera are long, with spermathecal gland connected near apical third (with the exception of being connected near the middle in a few taxa of
<taxonomicName id="4C1594CE296AFF95FD33F888EF5BF8AB" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" box="[751,885,1852,1878]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FD33F888EF5BF8AB" box="[751,885,1852,1878]" italics="true" pageId="42" pageNumber="43">Pseudoodes</emphasis>
</taxonomicName>
). However, the spermatheca in
<taxonomicName id="4C1594CE296AFF95FB3CF888E906F8AB" box="[1248,1320,1852,1878]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FB3CF888E906F8AB" box="[1248,1320,1852,1878]" italics="true" pageId="42" pageNumber="43">Oodes</emphasis>
</taxonomicName>
species is undifferentiated while those of species from the other two genera are more or less differentiated to a seminal canal and a receptaculum (with reversed state in several taxa of
<taxonomicName id="4C1594CE296AFF95FCFDF830EF89F863" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" box="[801,935,1924,1950]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FCFDF830EF89F863" box="[801,935,1924,1950]" italics="true" pageId="42" pageNumber="43">Pseudoodes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296AFF94FF1BF81CE86CFF04" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BAAEF4D296AFF94FF1BF81CE86CFF04" blockId="42.[151,1437,952,2023]" lastBlockId="43.[151,1436,151,249]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
The species from the nominotypical subgenus and those from
<taxonomicName id="4C1594CE296AFF95FC40F81CE8B3F83F" authority="LeConte" authorityName="LeConte" authorityYear="1853" box="[924,1181,1960,1986]" class="Insecta" family="Carabidae" genus="Lachnocrepis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296AFF95FC40F81CE81DF83F" box="[924,1075,1960,1986]" italics="true" pageId="42" pageNumber="43">Lachnocrepis</emphasis>
LeConte
</taxonomicName>
share character states 16 which is a good evidence of a close relationship between the two groups and for their common origin. Likely
<taxonomicName id="4C1594CE296BFF94FF4BFF23ED00FF4C" authorityName="LeConte" authorityYear="1853" box="[151,302,151,177]" class="Insecta" family="Carabidae" genus="Lachnocrepis" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296BFF94FF4BFF23ED00FF4C" box="[151,302,151,177]" italics="true" pageId="43" pageNumber="44">Lachnocrepis</emphasis>
</taxonomicName>
arose within
<taxonomicName id="4C1594CE296BFF94FE1AFF23EE20FF4C" box="[454,526,151,177]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B961335F296BFF94FE1AFF23EE20FF4C" box="[454,526,151,177]" italics="true" pageId="43" pageNumber="44">Oodes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2528E24296BFF94FDC8FF23EE1BFF4C" box="[532,565,151,177]" pageId="43" pageNumber="44" sensu="lato">s.l.</taxonomicNameLabel>
during an early stage of diversification. In view of existing common features, including one synapomophy between
<taxonomicName id="4C1594CE296BFF94FD91FF0FEECAFF28" authorityName="LeConte" authorityYear="1853" box="[589,740,187,213]" class="Insecta" family="Carabidae" genus="Lachnocrepis" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296BFF94FD91FF0FEECAFF28" box="[589,740,187,213]" italics="true" pageId="43" pageNumber="44">Lachnocrepis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1594CE296BFF94FCFEFF0FEF44FF28" box="[802,874,187,213]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296BFF94FCFEFF0FEF44FF28" box="[802,874,187,213]" italics="true" pageId="43" pageNumber="44">Oodes</emphasis>
</taxonomicName>
not shared by any other representative from the
<taxonomicName id="4C1594CE296BFF94FF4BFF6BECF1FF04" box="[151,223,223,249]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296BFF94FF4BFF6BECF1FF04" box="[151,223,223,249]" italics="true" pageId="43" pageNumber="44">Oodes</emphasis>
</taxonomicName>
generic group, we prefer to rank the former group as a subgenus of the latter.
</paragraph>
</subSubSection>
</treatment>
</document>