232 lines
27 KiB
XML
232 lines
27 KiB
XML
<document id="AA072D33CDD567AD2BD646AE17A4B3DC" ID-CLB-Dataset="8579" ID-DOI="10.11646/zootaxa.4850.1.1" ID-GBIF-Dataset="686337a2-1528-43d0-ae32-68ca023d8df3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4407072" ID-ZooBank="18AA0411-0E54-4922-84C7-608EAC68D281" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609428297648" checkinUser="plazi" docAuthor="Guéorguiev, Borislav & Liang, Hongbin" docDate="2020" docId="03BC5E5B2947FFB7FF4BF972ECD5F988" docLanguage="en" docName="zootaxa.4850.1.1.pdf" docOrigin="Zootaxa 4850 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Oodes Bonelli" docType="treatment" docVersion="10" lastPageNumber="9" masterDocId="FF8526232940FFBFFFDCFFB4EC2EFFFD" masterDocTitle="Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species" masterLastPageNumber="89" masterPageNumber="1" pageNumber="8" updateTime="1698852686469" updateUser="ExternalLinkService">
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<mods:mods id="3C116BDDE7001C39334EC567E75331C5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3CCBDF678FF4DBC3DFB857B216056567">
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<mods:title id="DEBA41D982114618C8D54663EB7B46F2">Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species</mods:title>
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<mods:name id="28219421ED60D6246EFE00BC7294BA29" type="personal">
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<mods:role id="1B4DDF52A9B3E47D33B97384D4CD1927">
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<mods:roleTerm id="6C9EB26A52753EC943A194719DC5F602">Author</mods:roleTerm>
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<mods:namePart id="0A6750E5993E16FDAC68D85E9A92B565">Guéorguiev, Borislav</mods:namePart>
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<mods:nameIdentifier id="71E60C9AC4879DD9C4CCF4DC05DEF8A6" type="ORCID">0000-0002-8532-0657</mods:nameIdentifier>
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<mods:nameIdentifier id="2BE0539EE115F4241FAD001216F96A0A" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:name id="D2B0913DA5F83E897EB1BBFBD76D6215" type="personal">
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<mods:roleTerm id="6DFB2B0CE49E0FC52ED9E1ADF9470D19">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0913A04366D835803E95F2FF88B9BD99">Liang, Hongbin</mods:namePart>
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<mods:nameIdentifier id="7278E115CF7D3B9CB9DE000C512368D1" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
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<mods:nameIdentifier id="E0025258E25DE9B84266D0D225D41AF9" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:title id="5C8C50E94F46694EBA26AC55E7A6B7AF">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="76E6E6DE0ADFA75C59A3876029A6E836">
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<mods:date id="727F6D5E0A6B5C166C15B1CA7ABFF459">2020</mods:date>
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<mods:number id="B37A0486A354D3B6E7BE761AFBCAA612">2020-09-08</mods:number>
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<mods:detail id="CEDEE5A7B4707DE1DB9470AA19EAA6C5" type="volume">
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<mods:number id="25B2099C2849D9DE7EE16E10E67C3CA7">4850</mods:number>
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<mods:number id="1BC6841AD0F0708D8EE5162A5DAE4D20">1</mods:number>
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<mods:classification id="D09C276BE6ED1E82A5969F1ACB8D7B82">journal article</mods:classification>
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<mods:identifier id="ED7F5DAB5A1ED370230A4E2946E569A0" type="CLB-Dataset">8579</mods:identifier>
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<mods:identifier id="2EFE7430F61593F2DA59AC312DBB8D38" type="DOI">10.11646/zootaxa.4850.1.1</mods:identifier>
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<mods:identifier id="D100C50795FCB6E4B7D9579270488527" type="GBIF-Dataset">686337a2-1528-43d0-ae32-68ca023d8df3</mods:identifier>
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<mods:identifier id="56A083985FDBF8A663F850AC716AD003" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03BC5E5B2947FFB7FF4BF972ECD5F988" ID-DOI="http://doi.org/10.5281/zenodo.4479973" ID-GBIF-Taxon="177139742" ID-Zenodo-Dep="4479973" LSID="urn:lsid:plazi:treatment:03BC5E5B2947FFB7FF4BF972ECD5F988" httpUri="http://treatment.plazi.org/id/03BC5E5B2947FFB7FF4BF972ECD5F988" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection id="C30FBCC62947FFB8FF4BF972EDB8F91D" box="[151,406,1734,1761]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8BAAEF4D2947FFB8FF4BF972EDB8F91D" blockId="7.[151,406,1734,1761]" box="[151,406,1734,1761]" pageId="7" pageNumber="8">
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<heading id="D0E258212947FFB8FF4BF972EDB8F91D" bold="true" box="[151,406,1734,1761]" fontSize="11" level="2" pageId="7" pageNumber="8" reason="4">
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<emphasis id="B961335F2947FFB8FF4BF972EDB8F91D" bold="true" box="[151,406,1734,1761]" pageId="7" pageNumber="8">
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<taxonomicName id="4C1594CE2947FFB8FF4BF972ECCCF91C" ID-CoL="8KXMQ" authority="Bonelli" authorityName="Bonelli" box="[151,226,1734,1761]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2947FFB8FF4BF972ECCCF91C" bold="true" box="[151,226,1734,1761]" italics="true" pageId="7" pageNumber="8">Oodes</emphasis>
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</taxonomicName>
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generic group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FBCC62947FFB7FF4BF8BAECD5F988" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="8BAAEF4D2947FFB8FF4BF8BAEFE2F8D4" blockId="7.[151,972,1806,1833]" box="[151,972,1806,1833]" pageId="7" pageNumber="8">
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<emphasis id="B961335F2947FFB8FF4BF8BAED24F8D5" bold="true" box="[151,266,1806,1832]" pageId="7" pageNumber="8">Diagnosis</emphasis>
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. The species of this group share the following character states:
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</paragraph>
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<paragraph id="8BAAEF4D2947FFB8FF4BF8E4EE40F897" blockId="7.[151,1436,1871,2042]" box="[151,622,1871,1898]" pageId="7" pageNumber="8">(1) Labrum with six setiferous punctures;</paragraph>
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<paragraph id="8BAAEF4D2947FFB8FF4BF8C0EFCAF870" blockId="7.[151,1436,1871,2042]" box="[151,996,1907,1934]" pageId="7" pageNumber="8">(2) Labrum not appreciably shortened, with flat or slightly convex surface;</paragraph>
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<paragraph id="8BAAEF4D2947FFB8FF4BF82CED5EF807" blockId="7.[151,1436,1871,2042]" pageId="7" pageNumber="8">(3) Pronotum with or without basal setiferous punctures. If present, the punctures are medially remote from the posterior angles; usually they are quite small (in many cases setae have fallen off and punctures themselves are hardly visible);</paragraph>
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<paragraph id="8BAAEF4D2948FFB7FF4BFF23EFAFFF4C" blockId="8.[151,1436,151,393]" box="[151,897,151,177]" pageId="8" pageNumber="9">(4) Elytral interval 3 with two setiferous pores, adjacent to stria 2;</paragraph>
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<paragraph id="8BAAEF4D2948FFB7FF4BFF0EEF34FF28" blockId="8.[151,1436,151,393]" box="[151,794,186,213]" pageId="8" pageNumber="9">(5) Elytral intervals 6 and 7 do not join the basal margin;</paragraph>
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<paragraph id="8BAAEF4D2948FFB7FF4BFF6BEE5DFEE0" blockId="8.[151,1436,151,393]" pageId="8" pageNumber="9">
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(6) Elytral interval
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<quantity id="4CED42A82948FFB7FEA5FF6BED88FF04" box="[377,422,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="9" unit="in" value="8.0">8 in</quantity>
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anterior half approximately as wide as intervals 1-7 (never significantly constricted, like species of
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<taxonomicName id="4C1594CE2948FFB7FEE0FEB7EE4BFEE0" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" box="[316,613,259,285]" class="Insecta" family="Carabidae" genus="Brachyodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FEE0FEB7EDEEFEE0" box="[316,448,259,285]" italics="true" pageId="8" pageNumber="9">Brachyodes</emphasis>
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<bibRefCitation id="EF8492BC2948FFB7FE1BFEB7EE4BFEE0" author="Jeannel, R." box="[455,613,259,285]" pageId="8" pageNumber="9" pagination="767 - 1146" refId="ref62277" refString="Jeannel, R. (1949) ColeiopteIres carabiques de la reigion Malgache. Troisieme Partie. Faune de l'Empire Francais. IX. Office de la Recherche scientifique coloniale, Paris, 767 - 1146." type="book chapter" year="1949">Jeannel, 1949</bibRefCitation>
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</taxonomicName>
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);
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</paragraph>
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<paragraph id="8BAAEF4D2948FFB7FF4BFE93E825FEBC" blockId="8.[151,1436,151,393]" box="[151,1035,295,321]" pageId="8" pageNumber="9">(7) Elytral interval 9 completely disappeared, transformed in marginal furrow;</paragraph>
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<paragraph id="8BAAEF4D2948FFB7FF4BFEFFEF35FE74" blockId="8.[151,1436,151,393]" pageId="8" pageNumber="9">
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(8) Apical part of median lobe of aedeagus tubular (not flattened dorsoventrally, as in members of
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<taxonomicName id="4C1594CE2948FFB7FAE6FEFFEDAEFE74" authority="Andrewes, 1922" authorityName="Andrewes" authorityYear="1922" class="Insecta" family="Carabidae" genus="Miltodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FAE6FEFFE9B2FE98" box="[1338,1436,331,357]" italics="true" pageId="8" pageNumber="9">Miltodes</emphasis>
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Andrewes, 1922
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</taxonomicName>
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, and
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<taxonomicName id="4C1594CE2948FFB7FE67FEDBEF20FE74" authority="Basilewsky, 1949" authorityName="Basilewsky" authorityYear="1949" box="[443,782,367,393]" class="Insecta" family="Carabidae" genus="Protopidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FE67FEDBEE10FE74" box="[443,574,367,393]" italics="true" pageId="8" pageNumber="9">Protopidius</emphasis>
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Basilewsky, 1949
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BAAEF4D2948FFB7FF1BFE03EED8FCA0" blockId="8.[151,1437,438,1653]" pageId="8" pageNumber="9">
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<emphasis id="B961335F2948FFB7FF1BFE03EE0DFE2D" bold="true" box="[199,547,438,465]" pageId="8" pageNumber="9">Monophyly and relationships</emphasis>
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. This group includes 48 species, 23 of them classified in the genera
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<taxonomicName id="4C1594CE2948FFB7FAD3FE03E9BBFE2C" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1295,1429,439,465]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FAD3FE03E9BBFE2C" box="[1295,1429,439,465]" italics="true" pageId="8" pageNumber="9">Pseudoodes</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1594CE2948FFB7FF4BFE6FED30FE08" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[151,286,475,501]" class="Insecta" family="Carabidae" genus="Sundaoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FF4BFE6FED30FE08" box="[151,286,475,501]" italics="true" pageId="8" pageNumber="9">Sundaoodes</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1594CE2948FFB7FEF1FE6FED5BFE08" box="[301,373,475,501]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FEF1FE6FED5BFE08" box="[301,373,475,501]" italics="true" pageId="8" pageNumber="9">Oodes</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C1594CE2948FFB7FE6EFE6FEE04FE08" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[434,554,475,501]" class="Insecta" family="Carabidae" genus="Nothoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FE6EFE6FEE04FE08" box="[434,554,475,501]" italics="true" pageId="8" pageNumber="9">Nothoodes</emphasis>
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</taxonomicName>
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, as well as 25
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<taxonomicName id="4C1594CE2948FFB7FD13FE6FEF39FE08" box="[719,791,475,501]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FD13FE6FEF39FE08" box="[719,791,475,501]" italics="true" pageId="8" pageNumber="9">Oodes</emphasis>
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</taxonomicName>
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species
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<emphasis id="B961335F2948FFB7FCAAFE6FE83FFE08" box="[886,1041,475,501]" italics="true" pageId="8" pageNumber="9">incertae sedis</emphasis>
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. The last category includes species still not taxonomically revised. The species from the above mentioned genera have a high degree of morphological similarity. Most of its diagnostic characters (1–2, 4, 6, and 8) are plesiomorphic, and the rest seem to be homoplastic characters shared with other unrelated
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<taxonomicName id="4C1594CE2948FFB7FD8EFDF3EE8DFD9C" authorityName=", Bousquet" authorityYear="1996" box="[594,675,583,609]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
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. Representatives from the first three genera have the basal bulb of median lobe closed dorsally and well-developed, elongate spermatheca with spermathecal gland connected near the apical third of the seminal canal, which suggests that they are more closely related to each other than to
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<taxonomicName id="4C1594CE2948FFB7FAC1FD3BE9BBFD54" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1309,1429,655,681]" class="Insecta" family="Carabidae" genus="Nothoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FAC1FD3BE9BBFD54" box="[1309,1429,655,681]" italics="true" pageId="8" pageNumber="9">Nothoodes</emphasis>
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</taxonomicName>
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. The relationships among the former three genera are difficult to assess. Despite sharing two derived features,
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<taxonomicName id="4C1594CE2948FFB7FA89FD07E9B3FD30" box="[1365,1437,691,717]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FA89FD07E9B3FD30" box="[1365,1437,691,717]" italics="true" pageId="8" pageNumber="9">Oodes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1594CE2948FFB7FF1AFD63ED63FD0C" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[198,333,727,753]" class="Insecta" family="Carabidae" genus="Sundaoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FF1AFD63ED63FD0C" box="[198,333,727,753]" italics="true" pageId="8" pageNumber="9">Sundaoodes</emphasis>
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</taxonomicName>
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are probably not adelphotaxa (see “Monophyly and relationships” under
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<taxonomicName id="4C1594CE2948FFB7FBA7FD63E92CFD0C" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1147,1282,727,753]" class="Insecta" family="Carabidae" genus="Sundaoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FBA7FD63E92CFD0C" box="[1147,1282,727,753]" italics="true" pageId="8" pageNumber="9">Sundaoodes</emphasis>
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</taxonomicName>
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) and together with
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<taxonomicName id="4C1594CE2948FFB7FF0FFD4FED77FCE8" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[211,345,763,789]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FF0FFD4FED77FCE8" box="[211,345,763,789]" italics="true" pageId="8" pageNumber="9">Pseudoodes</emphasis>
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</taxonomicName>
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could be separate lineages that arose from a common ancestor. Alternatively,
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<taxonomicName id="4C1594CE2948FFB7FB38FD4FE944FCE8" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1252,1386,763,789]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FB38FD4FE944FCE8" box="[1252,1386,763,789]" italics="true" pageId="8" pageNumber="9">Pseudoodes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1594CE2948FFB7FF4BFCABED30FCC4" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[151,286,799,825]" class="Insecta" family="Carabidae" genus="Sundaoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B961335F2948FFB7FF4BFCABED30FCC4" box="[151,286,799,825]" italics="true" pageId="8" pageNumber="9">Sundaoodes</emphasis>
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</taxonomicName>
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may be adelphotaxa (see “Monophyly and relationships” under
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<taxonomicName id="4C1594CE2948FFB7FC29FCABE855FCC4" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1013,1147,799,825]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FC29FCABE855FCC4" box="[1013,1147,799,825]" italics="true" pageId="8" pageNumber="9">Pseudoodes</emphasis>
|
||
</taxonomicName>
|
||
) and the clade formed by these two genera might be the adelphotaxon to
|
||
<taxonomicName id="4C1594CE2948FFB7FD74FCF7EEDEFCA0" box="[680,752,835,861]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FD74FCF7EEDEFCA0" box="[680,752,835,861]" italics="true" pageId="8" pageNumber="9">Oodes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAAEF4D2948FFB7FF1BFCD3EDCAFACC" blockId="8.[151,1437,438,1653]" pageId="8" pageNumber="9">
|
||
<emphasis id="B961335F2948FFB7FF1BFCD3ED24FC7C" bold="true" box="[199,266,871,897]" pageId="8" pageNumber="9">Notes</emphasis>
|
||
. The species from the above four genera can be distinguished from those of the genus
|
||
<taxonomicName id="4C1594CE2948FFB7FB0AFCD3ECFCFC58" authority="Chaudoir, 1882" authorityName="Chaudoir" authorityYear="1882" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FB0AFCD3E909FC7C" box="[1238,1319,871,897]" italics="true" pageId="8" pageNumber="9">Simous</emphasis>
|
||
<bibRefCitation id="EF8492BC2948FFB7FAF2FCD3ECFCFC58" author="Chaudoir, M. de" pageId="8" pageNumber="9" pagination="317 - 378" refId="ref60500" refString="Chaudoir, M. de (1882) Monographie des oodides 1 re partie. Annales de la Societe Entomologique de France, Series 6, 2, 317 - 378." type="journal article" year="1882">Chaudoir, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
, by the following characters: (1) labrum relatively long and not convex, with anterior angles moderately rounded; (2) mentum tooth nearly as long as wide, with apex rounded (most species), rarely truncate (most
|
||
<taxonomicName id="4C1594CE2948FFB7FF4BFC67ED33FC10" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[151,285,979,1005]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FF4BFC67ED33FC10" box="[151,285,979,1005]" italics="true" pageId="8" pageNumber="9">Pseudoodes</emphasis>
|
||
</taxonomicName>
|
||
spp.) or emarginate (
|
||
<taxonomicName id="4C1594CE2948FFB7FDFFFC67EF3EFC11" authorityName="Guéorguiev & Liang" authorityYear="2020" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[547,784,979,1005]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="B961335F2948FFB7FDFFFC67EF3EFC11" box="[547,784,979,1005]" italics="true" pageId="8" pageNumber="9">Pseudoodes vicarius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1594CE2948FFB7FCF8FC67EFE2FC10" authorityName="Jedlicka. Consists" authorityYear="1931" box="[804,972,979,1005]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis id="B961335F2948FFB7FCF8FC67EF1BFC11" box="[804,821,979,1004]" italics="true" pageId="8" pageNumber="9">P</emphasis>
|
||
.
|
||
<emphasis id="B961335F2948FFB7FC95FC67EFE2FC10" box="[841,972,979,1005]" italics="true" pageId="8" pageNumber="9">rambouseki</emphasis>
|
||
</taxonomicName>
|
||
); and (3) basal setiferous punctures of pronotum, if present, medially remote from posterior angles. The species of
|
||
<taxonomicName id="4C1594CE2948FFB7FBDBFC43E876FBEC" authorityName="Chaudoir" authorityYear="1882" box="[1031,1112,1015,1041]" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FBDBFC43E876FBEC" box="[1031,1112,1015,1041]" italics="true" pageId="8" pageNumber="9">Simous</emphasis>
|
||
</taxonomicName>
|
||
have a labrum two or more times shorter than the clypeus whereas the two structures have similar lengths in the species dealt with here. Also, the species of
|
||
<taxonomicName id="4C1594CE2948FFB7FEE7FB8BEDA2FBA4" authorityName="Chaudoir" authorityYear="1882" box="[315,396,1087,1113]" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FEE7FB8BEDA2FBA4" box="[315,396,1087,1113]" italics="true" pageId="8" pageNumber="9">Simous</emphasis>
|
||
</taxonomicName>
|
||
have a mentum tooth considerably shorter (considering distance from paramedial border to apex) and wider, with apex emarginate or truncate, as well as basal setiferous punctures of the pronotum situated near posterior angles. A few still undescribed species of this genus have lost their basal punctures of the pronotum, and are thus distinguishable from those of the generic group of
|
||
<taxonomicName id="4C1594CE2948FFB7FCB0FB1FEF9AFB38" box="[876,948,1195,1221]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FCB0FB1FEF9AFB38" box="[876,948,1195,1221]" italics="true" pageId="8" pageNumber="9">Oodes</emphasis>
|
||
</taxonomicName>
|
||
only by the first two features listed above. Moreover, all
|
||
<taxonomicName id="4C1594CE2948FFB7FEEBFB7BEDA6FB14" authorityName="Chaudoir" authorityYear="1882" box="[311,392,1231,1257]" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FEEBFB7BEDA6FB14" box="[311,392,1231,1257]" italics="true" pageId="8" pageNumber="9">Simous</emphasis>
|
||
</taxonomicName>
|
||
species studied have a submentum with one pair of setae. Among taxa considered, this state is present only in species of
|
||
<taxonomicName id="4C1594CE2948FFB7FE02FB47EE08FAF0" box="[478,550,1267,1293]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FE02FB47EE08FAF0" box="[478,550,1267,1293]" italics="true" pageId="8" pageNumber="9">Oodes</emphasis>
|
||
</taxonomicName>
|
||
, which are distinct however from
|
||
<taxonomicName id="4C1594CE2948FFB7FC73FB47E82EFAF0" authorityName="Chaudoir" authorityYear="1882" box="[943,1024,1267,1293]" class="Insecta" family="Carabidae" genus="Simous" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FC73FB47E82EFAF0" box="[943,1024,1267,1293]" italics="true" pageId="8" pageNumber="9">Simous</emphasis>
|
||
</taxonomicName>
|
||
by their rounded mentum tooth and proportionally longer labrum.
|
||
</paragraph>
|
||
<paragraph id="8BAAEF4D2948FFB7FF1BFA8FECD5F988" blockId="8.[151,1437,438,1653]" pageId="8" pageNumber="9">
|
||
Also, the species treated in this paper are distinct in character states 4 and 7 from those belonging to another group of
|
||
<collectingRegion id="49D121AF2948FFB7FF20FAEBED76FA84" box="[252,344,1375,1401]" country="Morocco" name="Oriental" pageId="8" pageNumber="9">Oriental</collectingRegion>
|
||
oodines that are currently classified in
|
||
<taxonomicName id="4C1594CE2948FFB7FCD4FAEBEF7EFA84" box="[776,848,1375,1401]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FCD4FAEBEF7EFA84" box="[776,848,1375,1401]" italics="true" pageId="8" pageNumber="9">Oodes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1594CE2948FFB7FC58FAEBE8D9FA84" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" box="[900,1271,1375,1401]" class="Insecta" family="Carabidae" genus="Pseudosphaerodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FC58FAEBE87BFA84" box="[900,1109,1375,1401]" italics="true" pageId="8" pageNumber="9">Pseudosphaerodes</emphasis>
|
||
<bibRefCitation id="EF8492BC2948FFB7FB87FAEBE8D9FA84" author="Jeannel, R." box="[1115,1271,1375,1401]" pageId="8" pageNumber="9" pagination="767 - 1146" refId="ref62277" refString="Jeannel, R. (1949) ColeiopteIres carabiques de la reigion Malgache. Troisieme Partie. Faune de l'Empire Francais. IX. Office de la Recherche scientifique coloniale, Paris, 767 - 1146." type="book chapter" year="1949">Jeannel, 1949</bibRefCitation>
|
||
</taxonomicName>
|
||
. The taxa from the latter group have the elytra without setiferous punctures in interval 3, and the interval 9 is not or incompletely transformed into a marginal furrow. Furthermore, the taxa from this still unrevised taxonomic group have three other characteristics: (1) basal bulb of the median lobe open dorsally (a trait present in species of
|
||
<taxonomicName id="4C1594CE2948FFB7FB70FA7FE90AFA18" authorityName="Guéorguiev & Liang" authorityYear="2020" box="[1196,1316,1483,1509]" class="Insecta" family="Carabidae" genus="Nothoodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961335F2948FFB7FB70FA7FE90AFA18" box="[1196,1316,1483,1509]" italics="true" pageId="8" pageNumber="9">Nothoodes</emphasis>
|
||
</taxonomicName>
|
||
); (2) male protarsomere 3 semicircular with the medial side rounded; and (3) interval 8 (and 9, if present) flat and situated laterally, in contrast with adjacent medial intervals which lie dorsally. Interval
|
||
<quantity id="4CED42A82948FFB7FBC0F9A7E867F9D0" box="[1052,1097,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="9" unit="in" value="8.0">8 in</quantity>
|
||
the species dealt with here is usually more convex than the medial intervals and occupies an intermediate position between the lateral and dorsal surfaces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |