175 lines
19 KiB
XML
175 lines
19 KiB
XML
<document id="3D379398A486012B613CB3E3CB60A889" ID-DOI="10.11646/zootaxa.1745.1.3" ID-ISSN="11755334" ID-Zenodo-Dep="5108342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626382332205" checkinUser="felipe" docAuthor="Sabu, Thomas K., Abhitha, Prabhakar & Zhao, Danyang" docDate="2008" docId="03B97607FFFC647C47D62853FAAEFF5B" docLanguage="en" docName="zt01745p046.pdf" docOrigin="Zootaxa 1745 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Helluodes Westwood 1847" docType="treatment" docVersion="4" lastPageNumber="35" masterDocId="FF800E7FFFFF647947412D70FF96FF81" masterDocTitle="A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae)" masterLastPageNumber="46" masterPageNumber="30" pageNumber="33" updateTime="1699011263822" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B67A93F4C953BBC582F6DFB69F8DC20B">
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<mods:title id="5868785FB3E80133345F0F66E0EBB611">A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="BDB4BAD6230EB1915B88EC689FE03ABF" type="personal">
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<mods:role id="695024F9B03F53F25B6D840A68965663">
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<mods:roleTerm id="6DFEF521E88C7E5B30F9A44252936C80">Author</mods:roleTerm>
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<mods:namePart id="70663D755C2EC778C9059BB1774147D0">Sabu, Thomas K.</mods:namePart>
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<mods:affiliation id="D84627C53BFA4F5ABD059A49E7C2416B">Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="5CA4DA6CF8E6AA5D18FFB0E97DCDAF21" type="email">sabukthomas@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="CE6140BA02DD8987B8BB18AAE0911798" type="personal">
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<mods:roleTerm id="CE48B614FB6404C9D79BBB83F4D141D6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1528E4EC0E758AEB06DAD7C2DD7F457B">Abhitha, Prabhakar</mods:namePart>
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<mods:affiliation id="829344490C0E94D1F71A9917044B6E68">Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com</mods:affiliation>
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</mods:name>
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<mods:name id="F57445B77C172F011CD6720BC3776759" type="personal">
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<mods:role id="8D21FC960AA6D8CF87AA186420967CCB">
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<mods:roleTerm id="F5870289314CBCE7E8A7A09E61E7C28B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="90F7F48177938AD77424D718FC818B7E">Zhao, Danyang</mods:namePart>
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<mods:typeOfResource id="5C2B6B23234C87337A4BE3D4638F5072">text</mods:typeOfResource>
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<mods:titleInfo id="F60F8BB5D9F0BC4AAAE2063C7DF05452">
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<mods:title id="F3EB5C002D2B0ADFBA3EB27EF025C196">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A184359D44AD92988225756AA38B0884">
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<mods:date id="50781BE44203CF307F135D45D07E6137">2008</mods:date>
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<mods:detail id="8AF8FCCB116C340B23CAB5B2812F3B2C" type="pubDate">
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<mods:number id="3F551312D2751B6A8A4F2B47F79BA54A">2008-04-09</mods:number>
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</mods:detail>
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<mods:detail id="AF189AD1A236C1D529E871BE3362C373" type="volume">
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<mods:number id="1F656379C3046573628565F436F7D48E">1745</mods:number>
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</mods:detail>
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<mods:detail id="77B2688AFEA909709143E71060DBF842" type="issue">
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<mods:number id="D32CC075E98C8DC66CEAEC774AF0D7C5">1</mods:number>
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<mods:start id="511A625922DCE93DD5F8216D7D44CB96">30</mods:start>
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<mods:end id="0F2E539B28C7786353CA2F0285C79CFA">46</mods:end>
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<mods:url id="30D6574388AB5AF8841F13CD36F56F63">https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3</mods:url>
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<mods:classification id="D1DB5503A8B37CF0E01F895E218EEF91">journal article</mods:classification>
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<mods:identifier id="61277914EE75C0C06655C9711F5EBB6B" type="DOI">10.11646/zootaxa.1745.1.3</mods:identifier>
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<mods:identifier id="5A9124D54E9BFF90646ABBA26C15F3BE" type="ISSN">11755334</mods:identifier>
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<treatment id="03B97607FFFC647C47D62853FAAEFF5B" ID-DOI="http://doi.org/10.5281/zenodo.5108818" ID-Zenodo-Dep="5108818" LSID="urn:lsid:plazi:treatment:03B97607FFFC647C47D62853FAAEFF5B" httpUri="http://treatment.plazi.org/id/03B97607FFFC647C47D62853FAAEFF5B" lastPageId="5" lastPageNumber="35" pageId="3" pageNumber="33">
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<subSubSection id="C30A949AFFFC647A47D62853FE74FABC" box="[151,482,1315,1341]" pageId="3" pageNumber="33" type="nomenclature">
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<paragraph id="8BAFC711FFFC647A47D62853FE74FABC" blockId="3.[151,482,1315,1341]" box="[151,482,1315,1341]" pageId="3" pageNumber="33">
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<heading id="D0E7707DFFFC647A47D62853FE74FABC" bold="true" box="[151,482,1315,1341]" fontSize="11" level="1" pageId="3" pageNumber="33" reason="1">
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<taxonomicName id="4C10BC92FFFC647A47D62853FE74FABC" ID-CoL="8KX6Q" authority="Westwood, 1847" authorityName="Westwood" authorityYear="1847" box="[151,482,1315,1341]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="genus">
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<emphasis id="B9641B03FFFC647A47D62853FE74FABC" bold="true" box="[151,482,1315,1341]" pageId="3" pageNumber="33">
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<emphasis id="B9641B03FFFC647A47D62853FE87FABC" bold="true" box="[151,273,1315,1341]" italics="true" pageId="3" pageNumber="33">Helluodes</emphasis>
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<bibRefCitation id="EF81BAE0FFFC647A46592853FE74FABC" author="Westwood, J. O." box="[280,482,1315,1341]" pageId="3" pageNumber="33" pagination="278 - 280" refId="ref10481" refString="Westwood, J. O. (1847) Descriptions of two new genera of carabideous insects. Transactions of the Entomological Society of London, Series I, vol IV, 278 - 280." type="journal article" year="1847">Westwood, 1847</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30A949AFFFC647A47D62813FD87FA41" pageId="3" pageNumber="33" type="reference_group">
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<paragraph id="8BAFC711FFFC647A47D62813FD74FAFB" blockId="3.[151,738,1379,1402]" box="[151,738,1379,1402]" pageId="3" pageNumber="33">
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<treatmentCitationGroup id="AB00E03FFFFC647A47D62813FD74FAFB" box="[151,738,1379,1402]" pageId="3" pageNumber="33">
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<treatmentCitation id="0AB1E100FFFC647A47D62813FE63FAFB" author="Westwood, J. O." box="[151,501,1379,1402]" page="279" pageId="3" pageNumber="33" year="1847">
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<taxonomicName id="4C10BC92FFFC647A47D62813FE63FAFB" ID-CoL="8KX6Q" authority="Westwood, 1847: 279" authorityName="Westwood" authorityPageNumber="279" authorityYear="1847" box="[151,501,1379,1402]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="genus">
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<emphasis id="B9641B03FFFC647A47D62813FE95FAFB" box="[151,259,1379,1402]" italics="true" pageId="3" pageNumber="33">Helluodes</emphasis>
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<bibRefCitation id="EF81BAE0FFFC647A464B2813FE63FAFB" author="Westwood, J. O." box="[266,501,1379,1402]" pageId="3" pageNumber="33" pagination="278 - 280" refId="ref10481" refString="Westwood, J. O. (1847) Descriptions of two new genera of carabideous insects. Transactions of the Entomological Society of London, Series I, vol IV, 278 - 280." type="journal article" year="1847">Westwood, 1847: 279</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="0AB1E100FFFC647A45412813FD48FAFB" author="Chaudoir, M." box="[512,734,1379,1402]" page="302" pageId="3" pageNumber="33" year="1862">
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<bibRefCitation id="EF81BAE0FFFC647A45412813FD48FAFB" author="Chaudoir, M." box="[512,734,1379,1402]" pageId="3" pageNumber="33" pagination="275 - 320" refId="ref9253" refString="Chaudoir, M. (1862) Mat e riaux pour servir l'etude des Cicind eletes et Carabiques 3 - e partie. Extraits du Bulletin de la Societe Imperiale des Naturalist de Moscou, 35 (4), 275 - 320." type="journal article" year="1862">Chaudoir, 1862: 302</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BAFC711FFFC647A47D628D5FD87FA41" blockId="3.[151,1436,1445,2035]" box="[151,529,1445,1472]" pageId="3" pageNumber="33">
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<emphasis id="B9641B03FFFC647A47D628D5FF75FA3E" bold="true" box="[151,227,1445,1471]" pageId="3" pageNumber="33">Color.</emphasis>
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Black or brownish black.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A949AFFFC647D478428BDFF42F85A" lastPageId="4" lastPageNumber="34" pageId="3" pageNumber="33" type="description">
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<paragraph id="8BAFC711FFFC647A478428BDFB70F872" blockId="3.[151,1436,1445,2035]" pageId="3" pageNumber="33">
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<emphasis id="B9641B03FFFC647A478428BDFE99FA66" bold="true" box="[197,271,1485,1511]" pageId="3" pageNumber="33">Head.</emphasis>
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Large, massive and protracted, narrower than prothorax, surface strongly wrinkled, moderately setose and punctate. Labrum subrectangular, as wide as long, emarginate or truncate between anterior angles. Clypeus subtruncate, with 6–12 setae; frontal foveae deeply depressed and distinct; anterior supraorbital seta situated at about middle of eye near inner margin, posterior supraorbital seta situated near posterior margin of eye. Genae convex, large and densely setose behind eye. Mandibles protracted, acute, medially smooth, apex symmetrically acuminate. Maxillae with elongate and massive lacinia, galeae curved; maxillary palpi with palpomere 2 longest, palpomere 4 shortest, inflexed medially and medial surface with distinct, deep puncture, apex rounded or truncate. Mentum wide, strongly emarginate, with bifid median tooth. Ligula slender, elongate and tapered towards apex, with a dorsal horn-like process ahead of tip, with 2–3 elongate apical setae. Paraglossae membranous, narrow and distinctly shorter than ligula. Palpomere 2 of labial palpi with dense and long setae medially. Antennae narrow, elongate, pilose from basal antennomere, with apical ring of setae; antennomeres 1-5 cylindrical; scape longest, elongate; antennomere 2 more than half the length of 3; antennomere 3 slightly longer than 4
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<superScript id="7C656A59FFFC647A46B02ADEFE6BF83D" attach="none" box="[497,509,1966,1980]" fontSize="6" pageId="3" pageNumber="33">th</superScript>
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, antennomeres 4 and 5 shorter; antennomeres 5–11 flattened, with a glabrous impunctate stripe along middle of dorsal and ventral surfaces, laterally chagreened and setose.
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</paragraph>
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<paragraph id="8BAFC711FFFB647D47842DE7FB1AFE23" blockId="4.[151,1436,151,418]" pageId="4" pageNumber="34">
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<emphasis id="B9641B03FFFB647D47842DE7FEDDFF30" bold="true" box="[197,331,151,177]" pageId="4" pageNumber="34">Pronotum.</emphasis>
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Large, cordiform, widest at anterior third, with distinctly explanated and reflexed lateral borders; apical margin with or without emargination; anterior angles rounded and extended anteriorly beyond arc of emargination; lateral margins reflexed with marginal setae at base and slightly before middle; disc weakly convex with distinct median line, median line extended neither to apex nor base; anterior and posterior transverse sulci faint; basal foveae deep and wide extended forward to anterior one-third as distinct lateral furrows; lateral margins distinctly or weakly sinuate in front of posterior angles; base narrower than apex, slightly sinuate in middle and distinctly oblique laterally; densely or sparsely punctate and pilose.
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</paragraph>
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<caption id="DF6F9799FFFB647D47D628EBFDE8FA54" ID-DOI="http://doi.org/10.5281/zenodo.5108350" ID-Zenodo-Dep="5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="4" pageNumber="34" startId="4.[151,269,1435,1459]" targetBox="[175,1403,493,1408]" targetPageId="4">
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<paragraph id="8BAFC711FFFB647D47D628EBFDE8FA54" blockId="4.[151,1436,1435,1493]" pageId="4" pageNumber="34">
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<emphasis id="B9641B03FFFB647D47D628EBFEC7FA32" bold="true" box="[151,337,1435,1459]" pageId="4" pageNumber="34">FIGURES 6–14.</emphasis>
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Labrum, maxillary palpi and pronotum: 6–8
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<taxonomicName id="4C10BC92FFFB647D447E28EBFBD4FA33" authority="Sabu & Abhitha & Zhao, 2008" authorityName="Sabu & Abhitha & Zhao" authorityYear="2008" box="[831,1090,1435,1458]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
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<emphasis id="B9641B03FFFB647D447E28EBFBD4FA33" box="[831,1090,1435,1458]" italics="true" pageId="4" pageNumber="34">Helluodes devagiriensis</emphasis>
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</taxonomicName>
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<emphasis id="B9641B03FFFB647D430B28EBFBE9FA32" bold="true" box="[1098,1151,1435,1459]" pageId="4" pageNumber="34">
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<taxonomicNameLabel id="A257A678FFFB647D430B28EBFBE9FA32" box="[1098,1151,1435,1459]" pageId="4" pageNumber="34" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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; 9–11
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<taxonomicName id="4C10BC92FFFB647D438A28EBFF2BFA54" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="taprobanae">
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<emphasis id="B9641B03FFFB647D438A28EBFF2BFA54" italics="true" pageId="4" pageNumber="34">Helluodes taprobanae</emphasis>
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</taxonomicName>
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; 12–14
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<emphasis id="B9641B03FFFB647D465528CEFE69FA54" box="[276,511,1470,1493]" italics="true" pageId="4" pageNumber="34">
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<taxonomicName id="4C10BC92FFFB647D465528CEFE6DFA54" box="[276,507,1470,1493]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="westwoodi">Helluodes westwoodi</taxonomicName>
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.
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</emphasis>
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Not scaled.
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</paragraph>
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</caption>
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<paragraph id="8BAFC711FFFB647D47842B78FDF5F9F2" blockId="4.[151,1436,1544,2011]" pageId="4" pageNumber="34">
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<emphasis id="B9641B03FFFB647D47842B78FE8AF9A3" bold="true" box="[197,284,1544,1570]" pageId="4" pageNumber="34">Elytra.</emphasis>
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Convex posteriorly, widest medially, densely punctate and setose; margins faintly subserrate; apices obliquely truncate with wide or narrow membranous margins, sinuate near outer angles; striae entire; intervals variably punctate and convex.
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</paragraph>
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<paragraph id="8BAFC711FFFB647D47842BF0FD8CF91A" blockId="4.[151,1436,1544,2011]" box="[197,538,1664,1691]" pageId="4" pageNumber="34">
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<emphasis id="B9641B03FFFB647D47842BF0FEC2F91B" bold="true" box="[197,340,1664,1690]" pageId="4" pageNumber="34">Hindwings.</emphasis>
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Well developed.
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</paragraph>
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<paragraph id="8BAFC711FFFB647D47842BD8FBBAF942" blockId="4.[151,1436,1544,2011]" box="[197,1068,1704,1731]" pageId="4" pageNumber="34">
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<emphasis id="B9641B03FFFB647D47842BD8FEB6F943" bold="true" box="[197,288,1704,1730]" pageId="4" pageNumber="34">Venter.</emphasis>
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Punctate-pubescent, except posterior area of prosternum glabrous.
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</paragraph>
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<paragraph id="8BAFC711FFFB647D47842BA0FAD0F96A" blockId="4.[151,1436,1544,2011]" box="[197,1350,1744,1771]" pageId="4" pageNumber="34">
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<emphasis id="B9641B03FFFB647D47842BA0FE91F96B" bold="true" box="[197,263,1744,1770]" pageId="4" pageNumber="34">Legs.</emphasis>
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Short, front tibiae emarginate at apex, dorsal surface of tarsi pubescent or not; claws simple.
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</paragraph>
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<paragraph id="8BAFC711FFFB647D47842B88FAFDF8BA" blockId="4.[151,1436,1544,2011]" pageId="4" pageNumber="34">
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<emphasis id="B9641B03FFFB647D47842B88FED6F893" bold="true" box="[197,320,1784,1810]" pageId="4" pageNumber="34">Genitalia.</emphasis>
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Male aedeagus well sclerotised; median lobe with a dorsal opening, apex narrow, tapered and rounded; left paramere well developed, lower surface almost straight. Styli of female ovipositor glabrous.
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</paragraph>
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<paragraph id="8BAFC711FFFB647D47842A38FF42F85A" blockId="4.[151,1436,1544,2011]" pageId="4" pageNumber="34">
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<emphasis id="B9641B03FFFB647D47842A38FE2BF8E3" bold="true" box="[197,445,1864,1890]" pageId="4" pageNumber="34">Sexual dimorphism</emphasis>
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. Male with abdominal sternite 2 with one and sternite 3 with two oval, median depressions and a cluster of long filamentous setae; female without such depressions or setae. Male protarsus with dense pads of adhesive squamose setae on ventral surface of three basal tarsomeres; female without such pads.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A949AFFFA647C47842DE7FAAEFF5B" pageId="5" pageNumber="35" type="type_taxon">
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<paragraph id="8BAFC711FFFA647C47842DE7FC01FF33" blockId="5.[197,1336,151,218]" box="[197,919,151,178]" pageId="5" pageNumber="35">
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<materialsCitation id="3B78CD4CFFFA647C47842DE7FC05FF33" box="[197,915,151,178]" country="Sri Lanka" location="Sri Lanka" pageId="5" pageNumber="35" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9641B03FFFA647C47842DE7FEF1FF30" bold="true" box="[197,359,151,177]" pageId="5" pageNumber="35">
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<typeStatus id="54AB79B3FFFA647C47842DE7FE94FF30" box="[197,258,151,177]" pageId="5" pageNumber="35">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C10BC92FFFA647C462E2DE8FD60FF33" ID-CoL="3KFT5" authority="Westwood" authorityName="Westwood" box="[367,758,152,178]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="taprobanae">
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<emphasis id="B9641B03FFFA647C462E2DE8FDE5FF30" box="[367,627,152,177]" italics="true" pageId="5" pageNumber="35">Helluodes taprobanae</emphasis>
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Westwood
|
||
</taxonomicName>
|
||
of
|
||
<collectingCountry id="F3078781FFFA647C445D2DE8FC05FF33" box="[796,915,152,178]" name="Sri Lanka" pageId="5" pageNumber="35">Sri Lanka</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAFC711FFFA647C47842DCFFAAEFF5B" blockId="5.[197,1336,151,218]" box="[197,1336,191,218]" pageId="5" pageNumber="35">
|
||
<emphasis id="B9641B03FFFA647C47842DCFFE6CFF58" bold="true" box="[197,506,191,217]" pageId="5" pageNumber="35">Geographic distribution.</emphasis>
|
||
Species in this genus are distributed in
|
||
<collectingCountry id="F3078781FFFA647C448D2DB0FBA8FF5B" box="[972,1086,192,218]" name="Sri Lanka" pageId="5" pageNumber="35">Sri Lanka</collectingCountry>
|
||
and southwest
|
||
<collectingCountry id="F3078781FFFA647C43B52DB0FAA2FF5B" box="[1268,1332,192,218]" name="India" pageId="5" pageNumber="35">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |