128 lines
15 KiB
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128 lines
15 KiB
XML
<document id="F0783B844C47D0D883088AF8151D2896" ID-DOI="10.11646/zootaxa.1745.1.3" ID-ISSN="11755334" ID-Zenodo-Dep="5108342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626382332205" checkinUser="felipe" docAuthor="Sabu, Thomas K., Abhitha, Prabhakar & Zhao, Danyang" docDate="2008" docId="03B97607FFFA647C47D62C47FA03FC25" docLanguage="en" docName="zt01745p046.pdf" docOrigin="Zootaxa 1745 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Helluodes Westwood" docType="key" docVersion="3" lastPageNumber="35" masterDocId="FF800E7FFFFF647947412D70FF96FF81" masterDocTitle="A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae)" masterLastPageNumber="46" masterPageNumber="30" pageNumber="35" updateTime="1699011263822" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="50C55C5D95E21AD1D66E4349D47FC150">
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<mods:title id="6EB1A3C4939B5CC3F17B9B88486E0F32">A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae)</mods:title>
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<mods:name id="23887E0B3C7F9AC987C0DC2B5D8E8EE4" type="personal">
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<mods:roleTerm id="087033239BCA078FD3B1EDD318D4BFE3">Author</mods:roleTerm>
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<mods:namePart id="B21A9C607721B9DAF243AC9AB071A36B">Sabu, Thomas K.</mods:namePart>
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<mods:affiliation id="269FF101F18015B6D2A4E4397E62F51D">Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="14183C52BA20DBFC14B3F0E67CE675E1" type="email">sabukthomas@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="382A5868953995EBAEACC213FCF286CE">Author</mods:roleTerm>
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<mods:namePart id="744332950B39B5DEDD25FBFB5E733824">Abhitha, Prabhakar</mods:namePart>
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<mods:affiliation id="BB08412CFD4C13A102CCB7C7C8CA9FBD">Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com</mods:affiliation>
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<mods:roleTerm id="E3BE32026F91CCDC22C28DDDEBA1D346">Author</mods:roleTerm>
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<mods:namePart id="80A40C7BD01D08F1EA209639DA9E2B33">Zhao, Danyang</mods:namePart>
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<mods:title id="6E00BE9F4895EC3FB2433DFC731B099B">Zootaxa</mods:title>
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<mods:part id="F1C5685F834B133987173B2688D037E7">
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<mods:date id="408B8003726EBA340849FF9BDD88DE3F">2008</mods:date>
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<mods:number id="4661A429B8E8E91E41250BE1B626BA4A">2008-04-09</mods:number>
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<mods:number id="D06E284590A7E4519EEA56C29D975112">1745</mods:number>
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<mods:url id="4C127B3D72AFDF38B3E57FD1139A9CFF">https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3</mods:url>
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<mods:identifier id="B20738D30A970CB091A04083272AADEF" type="DOI">10.11646/zootaxa.1745.1.3</mods:identifier>
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<mods:identifier id="A6C0E8274D9A671158461A78D4CCC2C0" type="ISSN">11755334</mods:identifier>
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<treatment id="03B97607FFFA647C47D62C47FA03FC25" LSID="urn:lsid:plazi:treatment:03B97607FFFA647C47D62C47FA03FC25" httpUri="http://treatment.plazi.org/id/03B97607FFFA647C47D62C47FA03FC25" lastPageNumber="35" pageId="5" pageNumber="35">
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<subSubSection id="C30A949AFFFA647C47D62C47FD08FED0" box="[151,670,311,337]" pageId="5" pageNumber="35" type="nomenclature">
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<paragraph id="8BAFC711FFFA647C47D62C47FD08FED0" blockId="5.[151,670,311,337]" box="[151,670,311,337]" pageId="5" pageNumber="35">
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<heading id="D0E7707DFFFA647C47D62C47FD08FED0" bold="true" box="[151,670,311,337]" fontSize="11" level="1" pageId="5" pageNumber="35" reason="1">
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<emphasis id="B9641B03FFFA647C47D62C47FD08FED0" bold="true" box="[151,670,311,337]" pageId="5" pageNumber="35">
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Key to the species of
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<taxonomicName id="4C10BC92FFFA647C46DD2C47FD08FED0" ID-CoL="8KX6Q" authority="Westwood" authorityName="Westwood" box="[412,670,311,337]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="B9641B03FFFA647C46DD2C47FD80FED0" bold="true" box="[412,534,311,337]" italics="true" pageId="5" pageNumber="35">Helluodes</emphasis>
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Westwood
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30A949AFFFA647C47D62CF2FA03FC25" pageId="5" pageNumber="35" type="key">
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<keyStep id="30E4DFB4FFFA647C47D62CF2FA02FD35" pageId="5" pageNumber="35">
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<paragraph id="8BAFC711FFFA647C47D62CF2FA03FD95" blockId="5.[151,1437,386,932]" pageId="5" pageNumber="35">
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<keyLead id="30E16424FFFA647C47D62CF2FA03FD95" pageId="5" pageNumber="35">
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1. Anterior margin of labrum with a distinct median emargination between rounded anterior angles (
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<figureCitation id="132BDB94FFFA647C42772CF2FA1BFE1D" box="[1334,1421,386,412]" captionStart="FIGURES 6–14" captionStartId="4.[151,269,1435,1459]" captionTargetBox="[175,1403,493,1408]" captionTargetId="figure-287@4.[151,1436,474,1411]" captionTargetPageId="4" captionText="FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="5" pageNumber="35">Fig. 12</figureCitation>
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); anterior angles of pronotum expanded, rounded and weakly acute; pronotum distinctly sinuated anterior to posterior angles; posterior angles of pronotum strongly reflexed and sharp at apex (
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<figureCitation id="132BDB94FFFA647C43E62CA2FB69FE6D" box="[1191,1279,466,492]" captionStart="FIGURES 6–14" captionStartId="4.[151,269,1435,1459]" captionTargetBox="[175,1403,493,1408]" captionTargetId="figure-287@4.[151,1436,474,1411]" captionTargetPageId="4" captionText="FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="5" pageNumber="35">Fig. 14</figureCitation>
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); ligula slen- der, widest at middle; TLA = 25–27.5
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<emphasis id="B9641B03FFFA647C45C52C8AFBE1FD92" box="[644,1143,506,531]" italics="true" pageId="5" pageNumber="35">....................................................................</emphasis>
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<taxonomicName id="4C10BC92FFFA647C433A2C8AFA03FD95" authority="Chaudoir." authorityName="Chaudoir." box="[1147,1429,506,532]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="westwoodi">
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<emphasis id="B9641B03FFFA647C433A2C8AFA8CFD92" box="[1147,1306,506,531]" italics="true" pageId="5" pageNumber="35">H. westwoodi</emphasis>
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Chaudoir.
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BAFC711FFFA647C47D62F52FA02FD35" blockId="5.[151,1437,386,932]" pageId="5" pageNumber="35">
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<keyLead id="30E16424FFFA647C47D62F52FA02FD35" pageId="5" pageNumber="35">
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- Anterior margin of labrum nearly straight or slightly emarginated between rounded anterior angles; anterior angles of pronotum expanded and rounded; pronotum weakly sinuated anterior to posterior angles; posterior angles of pronotum weakly reflexed and blunt at apex (
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<figureCitation id="132BDB94FFFA647C44822F02FBD7FD0D" box="[963,1089,626,652]" captionStart="FIGURES 6–14" captionStartId="4.[151,269,1435,1459]" captionTargetBox="[175,1403,493,1408]" captionTargetId="figure-287@4.[151,1436,474,1411]" captionTargetPageId="4" captionText="FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="5" pageNumber="35">Figs. 8, 11</figureCitation>
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); ligula slender and narrower towards apex................................................................................................................................................2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30E4DFB4FFFA647C47D62FB2FA03FC25" pageId="5" pageNumber="35">
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<paragraph id="8BAFC711FFFA647C47D62FB2FA03FCAD" blockId="5.[151,1437,386,932]" pageId="5" pageNumber="35">
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<keyLead id="30E16424FFFA647C47D62FB2FA03FCAD" pageId="5" pageNumber="35">
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2 Anterior margin of labrum distinctly crenulate with 4 distinct indentations between anterior angles (
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<figureCitation id="132BDB94FFFA647C422D2FB2FF4EFC85" captionStart="FIGURES 6–14" captionStartId="4.[151,269,1435,1459]" captionTargetBox="[175,1403,493,1408]" captionTargetId="figure-287@4.[151,1436,474,1411]" captionTargetPageId="4" captionText="FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="5" pageNumber="35">Fig. 6</figureCitation>
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); maxillary palpomere 4 truncate and broadly dilated from middle to apex (
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<figureCitation id="132BDB94FFFA647C430D2F9AFB02FC85" box="[1100,1172,746,772]" captionStart="FIGURES 6–14" captionStartId="4.[151,269,1435,1459]" captionTargetBox="[175,1403,493,1408]" captionTargetId="figure-287@4.[151,1436,474,1411]" captionTargetPageId="4" captionText="FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="5" pageNumber="35">Fig. 7</figureCitation>
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); TLA = 24–26.2 ...... ..................................................................................................................................
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<taxonomicName id="4C10BC92FFFA647C43D32E62FAC4FCAA" ID-CoL="3KFT4" authority="Sabu & Abhitha & Zhao, 2008" authorityName="Sabu & Abhitha & Zhao" authorityYear="2008" box="[1170,1362,786,811]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
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<emphasis id="B9641B03FFFA647C43D32E62FAC4FCAA" box="[1170,1362,786,811]" italics="true" pageId="5" pageNumber="35">H. devagiriensis</emphasis>
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</taxonomicName>
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<emphasis id="B9641B03FFFA647C421B2E62FA03FCAD" bold="true" box="[1370,1429,786,812]" pageId="5" pageNumber="35">
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<taxonomicNameLabel id="A257A678FFFA647C421B2E62FA03FCAD" box="[1370,1429,786,812]" pageId="5" pageNumber="35" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8BAFC711FFFA647C47D62E4AFA03FC25" blockId="5.[151,1437,386,932]" pageId="5" pageNumber="35">
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<keyLead id="30E16424FFFA647C47D62E4AFA03FC25" pageId="5" pageNumber="35">
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- Anterior margin of labrum without crenulation and slightly emarginate between anterior angles (
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<figureCitation id="132BDB94FFFA647C42022E4AFA1AFCD5" box="[1347,1420,826,852]" captionStart="FIGURES 6–14" captionStartId="4.[151,269,1435,1459]" captionTargetBox="[175,1403,493,1408]" captionTargetId="figure-287@4.[151,1436,474,1411]" captionTargetPageId="4" captionText="FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="5" pageNumber="35">Fig. 9</figureCitation>
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); maxillary palpomere 4 roundly narrowed at apex and roundly dilated from middle to apex (
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<figureCitation id="132BDB94FFFA647C43B42E12FADAFCFD" box="[1269,1356,866,892]" captionStart="FIGURES 6–14" captionStartId="4.[151,269,1435,1459]" captionTargetBox="[175,1403,493,1408]" captionTargetId="figure-287@4.[151,1436,474,1411]" captionTargetPageId="4" captionText="FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="5" pageNumber="35">Fig. 10</figureCitation>
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); TLA = 24.5
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<emphasis id="B9641B03FFFA647C465C2EFAFBC9FC22" box="[285,1119,906,931]" italics="true" pageId="5" pageNumber="35">..................................................................................................................</emphasis>
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<taxonomicName id="4C10BC92FFFA647C43212EFAFA03FC25" ID-CoL="3KFT5" authority="Westwood." authorityName="Westwood." box="[1120,1429,906,932]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="taprobanae">
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<emphasis id="B9641B03FFFA647C43212EFAFA9CFC22" box="[1120,1290,906,931]" italics="true" pageId="5" pageNumber="35">H.taprobanae</emphasis>
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Westwood.
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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