167 lines
18 KiB
XML
167 lines
18 KiB
XML
<document id="6EC26869A1820F07F76A5B2FD58E5B3B" ID-DOI="10.1206/3961.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5370399" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268866948" checkinUser="carolina" docAuthor="Sinclair, Bradley J. & Grimaldi, David A." docDate="2020" docId="03B94C5BFFC13C4B67CD3538FD8A7598" docLanguage="en" docName="N3961.pdf" docOrigin="American Museum Novitates 2020 (3961)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3961/3961.1/Cretaceous-Diversity-of-the-Relict-Genus-Alavesia-Waters-and-Arillo/10.1206/3961.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Alavesia angusta Sinclair & Grimaldi 2020, new species" docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="FF803423FFCC3C46662D3721FC617321" masterDocTitle="Cretaceous diversity of the relict genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="14" updateTime="1699205599378" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="F054C16F2DE0E316CCC1CE582F6CA68B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="18BA5431F81977D30660A1DE816CD726">
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<mods:title id="B24572BE7B7CBF9BA4B98002FEBEED4E">Cretaceous diversity of the relict genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae)</mods:title>
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<mods:name id="C9739D687C8ABCDD6E2D43E65F7D53BC" type="personal">
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<mods:roleTerm id="5D3FF68C2FEA901DE78F6D76E435C0A0">Author</mods:roleTerm>
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<mods:namePart id="1D6AF41EA94934FD0AC65E9AB7D60BEE">Sinclair, Bradley J.</mods:namePart>
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<mods:name id="06E586C9FB70E9EB61203744DD175BF9" type="personal">
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<mods:roleTerm id="F2B27FFB8359B4136A24AF37EB92FED7">Author</mods:roleTerm>
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<mods:namePart id="F4E00051D4AEAE07DDC66B85ECBEEB08">Grimaldi, David A.</mods:namePart>
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<mods:typeOfResource id="1F2E7CBCBB7EA691181AA558A8C59181">text</mods:typeOfResource>
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<mods:title id="750406E9124FA5E36F73DE5C7EB9048B">American Museum Novitates</mods:title>
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<mods:part id="82682B2F49753B463D784A7247E68C5C">
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<mods:date id="713504553A82EB9AD66BA450B4DD02F4">2020</mods:date>
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<mods:number id="84AF0B5E9AA5E979AA38607E64DDE7A0">2020-10-06</mods:number>
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<mods:detail id="49EEA37F96595E93A8FB69FB2241B32A" type="volume">
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<mods:number id="2B9A991D92DAC87447B9251E7705D2FE">2020</mods:number>
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<mods:detail id="A8E79C581E320E93B2DEB77A6C2065FC" type="issue">
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<mods:number id="A629DB87648B6030D7CFF8A9253A8B8A">3961</mods:number>
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<mods:url id="63DD637D9DF534A30865615B4B32AFE7">https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3961/3961.1/Cretaceous-Diversity-of-the-Relict-Genus-Alavesia-Waters-and-Arillo/10.1206/3961.1.full</mods:url>
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</mods:location>
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<mods:classification id="92E2C074F55B7ED26BF7FB30F28227CA">journal article</mods:classification>
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<mods:identifier id="5F441C5EEF6B742C31F732D2639CB5EA" type="DOI">10.1206/3961.1</mods:identifier>
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<mods:identifier id="79CCB7C9CA94FCA9896BB54C79CE1BB8" type="ISSN">0003-0082</mods:identifier>
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<treatment id="03B94C5BFFC13C4B67CD3538FD8A7598" ID-DOI="http://doi.org/10.5281/zenodo.5462028" ID-Zenodo-Dep="5462028" LSID="urn:lsid:plazi:treatment:03B94C5BFFC13C4B67CD3538FD8A7598" httpUri="http://treatment.plazi.org/id/03B94C5BFFC13C4B67CD3538FD8A7598" lastPageNumber="14" pageId="13" pageNumber="14">
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<subSubSection id="C30AAEC6FFC13C4B67CD3538FF267112" box="[480,839,537,564]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8BAFFD4DFFC13C4B67CD3538FF267112" blockId="13.[480,839,537,564]" box="[480,839,537,564]" pageId="13" pageNumber="14">
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<heading id="D0E74A21FFC13C4B67CD3538FF267112" box="[480,839,537,564]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
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<emphasis id="B964215FFFC13C4B67CD3538FED47112" bold="true" box="[480,693,537,564]" pageId="13" pageNumber="14">
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<taxonomicName id="4C1086CEFFC13C4B67CD3538FECC7115" ID-CoL="89WYD" authority="Sinclair & Grimaldi, 2020" authorityName="Sinclair & Grimaldi" authorityYear="2020" box="[480,685,537,564]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angusta" status="sp. nov.">
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<emphasis id="B964215FFFC13C4B67CD3538FECC7115" bold="true" box="[480,685,537,564]" italics="true" pageId="13" pageNumber="14">Alavesia angusta</emphasis>
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</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A2579C24FFC13C4B6490353BFF267112" box="[701,839,537,564]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AAEC6FFC13C4B64313570FF6C714A" box="[540,781,593,619]" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BAFFD4DFFC13C4B64313570FF6C714A" blockId="13.[540,781,593,619]" box="[540,781,593,619]" pageId="13" pageNumber="14">
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<heading id="D0E74A21FFC13C4B64313570FF6C714A" box="[540,781,593,619]" centered="true" fontSize="36" level="3" pageId="13" pageNumber="14" reason="7">
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<figureCitation id="132BE1C8FFC13C4B64313570FEFC714A" box="[540,669,593,619]" captionStart="FIGURE" captionStartId="6.[108,197,1285,1306]" captionTargetBox="[200,1125,232,1256]" captionTargetId="figure-108@6.[108,1212,224,1262]" captionTargetPageId="6" captionText="FIGURE. 1. Antennae of Burmese amber Alavesia species, lateral views. All are males except N, O, P. Some antennae were reversed in direction for optimal comparison; longicornuta and longistylata are slightly reconstructed to account for curvature of the stylus. All are to the same scale. Specimen numbers: A. USNM PAL 726867. B. AMNH Bu-494. C. AMNH Bu-496. D. USNM PAL 726868. E. AMNH Bu-KL2-7. F. AMNH Bu-1476. G. JZC Bu-556 (AMNH). H. USNM PAL 726872. K. USNM PAL 726869. L. AMNH Bu-KL7-18. M. JZC Bu-282. N. Undetermined species. O. A. zigrasi. P. A. zigrasi." figureDoi="http://doi.org/10.5281/zenodo.5370401" httpUri="https://zenodo.org/record/5370401/files/figure.png" pageId="13" pageNumber="14">Figures 1A</figureCitation>
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,
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<figureCitation id="132BE1C8FFC13C4B64863570FEAE714A" box="[683,719,593,619]" captionStart="FIGURE 2" captionStartId="7.[108,194,1456,1477]" captionTargetBox="[121,1197,229,1434]" captionTargetId="figure-45@7.[108,1212,224,1437]" captionTargetPageId="7" captionText="FIGURE 2. Wings of six species of Alavesia in Burmese amber. All are males, to the same scale. Specimens: A. USNM PAL 726867. B. AMNH Bu-494. C. AMNH Bu-496. D. USNM PAL 726868. E. AMNH Bu-1476. F. JZC Bu-556." figureDoi="http://doi.org/10.5281/zenodo.5370403" httpUri="https://zenodo.org/record/5370403/files/figure.png" pageId="13" pageNumber="14">2A</figureCitation>
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,
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<figureCitation id="132BE1C8FFC13C4B64F13570FF6C714A" box="[732,781,593,619]" captionStart="FIGURE 12" captionStartId="21.[108,194,1223,1244]" captionTargetBox="[108,1212,224,1202]" captionTargetId="figure-51@21.[625,1212,705,1208]" captionTargetPageId="21" captionText="FIGURE 12. Alavesia spp. in Burmese amber. A. A. angusta, USNM PAL 726867. B. Drawing of A. brevipennae, AMNH Bu-494. C. Photomicrograph of A. brevipennae, AMNH Bu-494. D. A. myanmarensis, AMNH Bu-496." figureDoi="http://doi.org/10.5281/zenodo.5370423" httpUri="https://zenodo.org/record/5370423/files/figure.png" pageId="13" pageNumber="14">12A</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AAEC6FFC13C4B668E35A8FD487035" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="8BAFFD4DFFC13C4B668E35A8FD487035" blockId="13.[108,1214,649,1721]" pageId="13" pageNumber="14">
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<materialsCitation id="3B78F710FFC13C4B668E35A8FD447035" collectionCode="R" longitude="-2.8" pageId="13" pageNumber="14" specimenCode="R4" specimenCount="1">
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DIAGNOSIS: Postpedicel slender, fusiform, L/
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<geoCoordinate id="EE249B8AFFC13C4B648135A8FF607182" box="[684,769,649,675]" degrees="4.47" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="555" value="-4.47">W 4.47</geoCoordinate>
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; wing slender, L/
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<geoCoordinate id="EE249B8AFFC13C4B65E535A8F86E7182" box="[968,1039,649,675]" degrees="2.8" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="5555" value="-2.8">W 2.8</geoCoordinate>
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; M
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<subScript id="1794FF08FFC13C4B621835B6F8327189" attach="left" box="[1077,1107,663,680]" fontSize="7" pageId="13" pageNumber="14">1+2</subScript>
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and
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<specimenCode id="DBB65536FFC13C4B62A135ABF8C97189" box="[1164,1192,650,680]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="14">
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R
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<subScript id="1794FF08FFC13C4B62B335B6F8C97189" attach="left" box="[1182,1192,663,680]" fontSize="7" pageId="13" pageNumber="14">4</subScript>
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</specimenCode>
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+5 parallel for entire lengths; crossvein bm-cu at midlength of cell dm; crossvein dm-m slanted; CuA straight, directed to wing base, in line with crossvein m-cu; cell dm extending well beyond apex of cell cua
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AAEC6FFC13C4B668F343EFFE77568" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BAFFD4DFFC13C4B668F343EFD647088" blockId="13.[108,1214,649,1721]" pageId="13" pageNumber="14">
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DESCRIPTION:
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<emphasis id="B964215FFFC13C4B6762343EFDF17018" box="[335,400,799,825]" italics="true" pageId="13" pageNumber="14">Male.</emphasis>
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<emphasis id="B964215FFFC13C4B67BB343FFE7C7019" bold="true" box="[406,541,798,824]" pageId="13" pageNumber="14">Coloration</emphasis>
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not preserved.
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<emphasis id="B964215FFFC13C4B64FF343EFF7B7018" bold="true" box="[722,794,799,825]" pageId="13" pageNumber="14">Head:</emphasis>
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Scape small, goblet-shaped; pedicel quadrate; postpedicel narrow, almost fusiform, L/
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<geoCoordinate id="EE249B8AFFC13C4B64963465FF70707F" box="[699,785,836,862]" degrees="4.47" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="555" value="-4.47">W 4.47</geoCoordinate>
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(figs. 1A, 12A); stylus short, length 0.46
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<emphasis id="B964215FFFC13C4B66B03448FCCF70A2" bold="true" box="[157,174,873,899]" pageId="13" pageNumber="14">×</emphasis>
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the postpedicel, with 2 small basal articles. Palpus light, extended slightly beyond level of proboscis.
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</paragraph>
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<paragraph id="8BAFFD4DFFC13C4B668A3495FCDE7742" blockId="13.[108,1214,649,1721]" pageId="13" pageNumber="14">
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<emphasis id="B964215FFFC13C4B668A3495FD6770EF" bold="true" box="[167,262,948,974]" pageId="13" pageNumber="14">Thorax:</emphasis>
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Chaetotaxy: acrostichals fine, short, in biserial rows; dorsocentrals similar to acrostichals, ending in pair of long, fine prescutellars; 2 short postpronotal setae; 4 short, fine anterior notopleural setae, 2 large posterior notopleural setae; 1 postsutural supraalar seta; 1 finer postalar seta; 3 pairs scutellar setae, apical pair longest, upright, crossed for ~0.5
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<emphasis id="B964215FFFC13C4B62473305F81A771F" bold="true" box="[1130,1147,1060,1086]" pageId="13" pageNumber="14">×</emphasis>
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their length.
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</paragraph>
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<paragraph id="8BAFFD4DFFC13C4B668E334EFFAB77F5" blockId="13.[108,1214,649,1721]" pageId="13" pageNumber="14">
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<emphasis id="B964215FFFC13C4B668E334EFC8077A8" bold="true" box="[163,225,1135,1161]" pageId="13" pageNumber="14">Legs:</emphasis>
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Forecoxa with many short setae on anterior surface. Forefemur with ventral row ~15 short, stiff setae. Tibiae with long, apical ventral seta. Hind tibia with dense dorsal row of setae, length about half width of tibia; posterior surface with fine, brushy setae.
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</paragraph>
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<paragraph id="8BAFFD4DFFC13C4B668833FFF8DF7648" blockId="13.[108,1214,649,1721]" pageId="13" pageNumber="14">
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<emphasis id="B964215FFFC13C4B668833FFFC8E77D9" bold="true" box="[165,239,1246,1272]" pageId="13" pageNumber="14">Wing:</emphasis>
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Length
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<quantity id="4CE850A8FFC13C4B677933FEFDD177D8" box="[340,432,1247,1273]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="14" unit="mm" value="1.4">1.4 mm</quantity>
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(fig. 2A); C terminating slightly beyond M
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<subScript id="1794FF08FFC13C4B659933CDFFB377DC" attach="left" box="[948,978,1260,1277]" fontSize="7" pageId="13" pageNumber="14">1+2</subScript>
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; Sc nearly reaching costal margin; R
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<subScript id="1794FF08FFC13C4B67063233FD287602" attach="left" box="[299,329,1298,1315]" fontSize="7" pageId="13" pageNumber="14">4+5</subScript>
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parallel to M
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<subScript id="1794FF08FFC13C4B67CA3233FE647602" attach="left" box="[487,517,1298,1315]" fontSize="7" pageId="13" pageNumber="14">1+2</subScript>
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for entire lengths; crossvein r-m slightly distal to middle of cell dm; apex of M
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<subScript id="1794FF08FFC13C4B67613216FD347669" attach="left" box="[332,341,1335,1352]" fontSize="7" pageId="13" pageNumber="14">4</subScript>
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slightly closer to M
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<subScript id="1794FF08FFC13C4B646E3216FE007669" attach="left" box="[579,609,1335,1352]" fontSize="7" pageId="13" pageNumber="14">1+2</subScript>
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than to CuA+CuP; CuA strongly recurved, in line with bm-m; apex of cell cua truncate, cell cua shorter than cell bm; CuA+CuP slightly curved.
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</paragraph>
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<paragraph id="8BAFFD4DFFC13C4B668B3255FFE77568" blockId="13.[108,1214,649,1721]" pageId="13" pageNumber="14">
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<emphasis id="B964215FFFC13C4B668B3255FD4976AF" bold="true" box="[166,296,1396,1422]" pageId="13" pageNumber="14">Abdomen:</emphasis>
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No longer than thorax; setae short, without differentiated posteromarginal setae. Segments 1–5 broad, other segments retracted into segment 5. Terminalia well preserved in
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<typeStatus id="54AB43EFFFC13C4B66A0329EFC9576F8" box="[141,244,1471,1497]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
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(not illustrated): Epandrium complete dorsally, U-shaped, with fringelike row of five long setae on ventral margin; cercus flat, broad, scalelike, meeting medially. Surstylus long, slightly curved (as in
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<taxonomicName id="4C1086CEFFC13C4B6755312BFD8C7502" authority="Sinclair & Grimaldi, 2020" authorityName="Sinclair & Grimaldi" authorityYear="2020" box="[376,493,1546,1572]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="magna" status="sp. nov.">
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<emphasis id="B964215FFFC13C4B6755312BFD8C7502" box="[376,493,1546,1572]" italics="true" pageId="13" pageNumber="14">A. magna</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2579C24FFC13C4B67D3312BFE3F7505" box="[510,606,1546,1572]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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); postgonites + phallus as for most other species: thicker at base, hook-shaped, bend in middle, phallus downturned.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AAEC6FFC13C4B66893174FFEE754F" box="[164,911,1620,1647]" pageId="13" pageNumber="14" type="materials_examined">
|
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<paragraph id="8BAFFD4DFFC13C4B66893174FFEE754F" blockId="13.[108,1214,649,1721]" box="[164,911,1620,1647]" pageId="13" pageNumber="14">
|
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<materialsCitation id="3B78F710FFC13C4B66893174FFEE754F" box="[164,911,1620,1647]" collectionCode="TYPE, MATERIAL, USNM" pageId="13" pageNumber="14" specimenCode="PAL 726867" specimenCount="1" typeStatus="holotype">
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<collectionCode id="ED016588FFC13C4B66893174FD00754F" box="[164,353,1620,1647]" pageId="13" pageNumber="14">TYPE MATERIAL</collectionCode>
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:
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<typeStatus id="54AB43EFFFC13C4B67413175FDBA754F" box="[364,475,1620,1646]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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, male,
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<collectionCode id="ED016588FFC13C4B64023175FEE3754F" box="[559,642,1620,1646]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="14" type="Museum">USNM</collectionCode>
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<specimenCode id="DBB65536FFC13C4B64A73175FF79754F" box="[650,792,1620,1646]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="14" type="Museum">PAL 726867</specimenCode>
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(
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<collectionCode id="ED016588FFC13C4B65053175FFE3754F" box="[808,898,1620,1646]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="14" type="Museum">USNM</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30AAEC6FFC13C4B668E3158FD8A7598" pageId="13" pageNumber="14" type="etymology">
|
||
<paragraph id="8BAFFD4DFFC13C4B668E3158FD8A7598" blockId="13.[108,1214,649,1721]" pageId="13" pageNumber="14">
|
||
ETYMOLOGY: The specific epithet is from the Latin
|
||
<emphasis id="B964215FFFC13C4B64DC315BFF3775B2" box="[753,854,1658,1683]" italics="true" pageId="13" pageNumber="14">angustus</emphasis>
|
||
(narrow, slender), in reference to the narrow wing and cell cua.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |