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<document id="5CCABB026FE8F9A9A5D3CB1A20A947CF" ID="10.11646/zootaxa.4139.4.2" ID-DOI="10.11646/zootaxa.4139.4.2" ID-GBIF-Dataset="10fb08e8-a300-4132-a234-e7278fc643ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="264222" ID-ZooBank="247A52DC-530C-46FB-A985-ED8E0444CF09" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469185573468" checkinUser="plazi" docAuthor="Palero, Ferran, Genis-Armero, Rebeca, Hall, Michael R. &amp; Clark, Paul F." docDate="2016" docId="038C6004FFBEFF83FF34FF3CFEE467C6" docLanguage="en" docName="zootaxa.4139.4.2.pdf" docOrigin="Zootaxa 4139 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scyllarides squammosus H. Milne Edwards 1837" docType="treatment" docVersion="8" lastPageNumber="495" masterDocId="FFB5187CFFB8FF8DFFA3FFE1FF906577" masterDocTitle="DNA barcoding the phyllosoma of Scyllarides squammosus (H. Milne Edwards, 1837) (Decapoda: Achelata: Scyllaridae)" masterLastPageNumber="498" masterPageNumber="481" pageNumber="487" updateTime="1698365587425" updateUser="plazi">
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<mods:title id="41EFCEDD06BDCEDE445981878FD198F4">DNA barcoding the phyllosoma of Scyllarides squammosus (H. Milne Edwards, 1837) (Decapoda: Achelata: Scyllaridae)</mods:title>
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<mods:namePart id="C6032A8AAB9BFA77D4DBD7449A66B88F">Palero, Ferran</mods:namePart>
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<mods:namePart id="8D0CD885D9237379D228B30003D2863B">Genis-Armero, Rebeca</mods:namePart>
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<mods:namePart id="8714968737BAFC87D19DFBCADB55C222">Hall, Michael R.</mods:namePart>
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<mods:namePart id="07BB6F0F7E0F0F0AC277C58F724DBFA7">Clark, Paul F.</mods:namePart>
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<mods:title id="C1E80393F4693DB2AB35CC6B8A262E05">Zootaxa</mods:title>
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<mods:date id="D974F8BC0FD36971F9492A277CF86FE2">2016</mods:date>
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<mods:number id="288BD7B364EFFFF71BEB00FDBF3BD9F5">4139</mods:number>
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<treatment id="038C6004FFBEFF83FF34FF3CFEE467C6" ID-DOI="http://doi.org/10.5281/zenodo.5626505" ID-GBIF-Taxon="123199418" ID-Zenodo-Dep="5626505" LSID="urn:lsid:plazi:treatment:038C6004FFBEFF83FF34FF3CFEE467C6" httpUri="http://treatment.plazi.org/id/038C6004FFBEFF83FF34FF3CFEE467C6" lastPageId="14" lastPageNumber="495" pageId="6" pageNumber="487">
<subSubSection id="C33F8299FFBEFF8BFF34FF3CFE6A644A" pageId="6" pageNumber="487" type="nomenclature">
<paragraph id="8B9AD112FFBEFF8BFF34FF3CFC986580" blockId="6.[151,776,221,247]" box="[151,776,221,247]" pageId="6" pageNumber="487">
<heading id="D0D2667EFFBEFF8BFF34FF3CFC986580" bold="true" box="[151,776,221,247]" fontSize="11" level="1" pageId="6" pageNumber="487" reason="1">
<taxonomicName id="4C25AA91FFBEFF8BFF34FF3CFC986580" ID-CoL="854SS" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[151,776,221,247]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="487" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFBEFF8BFF34FF3CFC986580" bold="true" box="[151,776,221,247]" pageId="6" pageNumber="487">
<emphasis id="B9510D00FFBEFF8BFF34FF3CFE2E6580" bold="true" box="[151,446,221,247]" italics="true" pageId="6" pageNumber="487">Scyllarides squammosus</emphasis>
(H. Milne Edwards, 1837)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9AD112FFBEFF8BFF34FEC5FE6A644A" blockId="6.[151,1436,292,677]" box="[151,506,292,317]" pageId="6" pageNumber="487">Stage VI (NHMUK 2015.3279)</paragraph>
</subSubSection>
<subSubSection id="C33F8299FFBEFF83FF64FEA6FEE467C6" lastPageId="14" lastPageNumber="495" pageId="6" pageNumber="487" type="description">
<paragraph id="8B9AD112FFBEFF8BFF64FEA6FC636428" blockId="6.[151,1436,292,677]" box="[199,1011,327,352]" pageId="6" pageNumber="487">
<emphasis id="B9510D00FFBEFF8BFF64FEA6FEE66417" bold="true" box="[199,374,327,352]" pageId="6" pageNumber="487">Measurements</emphasis>
(
<figureCitation id="131ECD97FFBEFF8BFE26FEA6FE586417" box="[389,456,327,352]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="6" pageNumber="487">Fig. 2</figureCitation>
A). TL =
<quantity id="4CDD7CF7FFBEFF8BFD90FEA9FD096428" box="[563,665,328,352]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="6" pageNumber="487" unit="mm" value="11.7">11.7 mm</quantity>
; CW =
<quantity id="4CDD7CF7FFBEFF8BFD4CFEA6FCD86428" box="[751,840,327,352]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="6" pageNumber="487" unit="mm" value="6.3">6.3 mm</quantity>
; CL = 9.0 mm.
</paragraph>
<paragraph id="8B9AD112FFBEFF8BFF64FE8DFF7564DF" blockId="6.[151,1436,292,677]" pageId="6" pageNumber="487">
<emphasis id="B9510D00FFBEFF8BFF64FE8DFEED64F2" bold="true" box="[199,381,364,389]" pageId="6" pageNumber="487">Cephalic shield</emphasis>
(
<figureCitation id="131ECD97FFBEFF8BFE2FFE8DFE5F64F3" box="[396,463,364,389]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="6" pageNumber="487">Fig. 2</figureCitation>
A). Spindle-shaped, longer than wide (CL/CW = 1.42), maximum width located to mid length.
</paragraph>
<paragraph id="8B9AD112FFBEFF8BFF64FE55FF7D6762" blockId="6.[151,1436,292,677]" pageId="6" pageNumber="487">
<emphasis id="B9510D00FFBEFF8BFF64FE55FED364BA" bold="true" box="[199,323,436,461]" pageId="6" pageNumber="487">Antennule</emphasis>
(
<figureCitation id="131ECD97FFBEFF8BFEF7FE55FE0A64BB" box="[340,410,436,461]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="6" pageNumber="487">Fig. 2</figureCitation>
B). Peduncle 3-segmented, each segment with similar length. Distal segment biflagellated, accesory flagellum half the length of primary flagellum. Primary flagellum with 7 rows of aesthetascs on inner margin.
</paragraph>
<paragraph id="8B9AD112FFBEFF8BFF64FDFEFEE9672A" blockId="6.[151,1436,292,677]" pageId="6" pageNumber="487">
<emphasis id="B9510D00FFBEFF8BFF64FDFEFEBE674F" bold="true" box="[199,302,543,568]" pageId="6" pageNumber="487">Antenna</emphasis>
(
<figureCitation id="131ECD97FFBEFF8BFEE1FDFEFE1C674F" box="[322,396,543,568]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="6" pageNumber="487">Fig. 2</figureCitation>
B). Biramous, endopod 2-segmented. The inner ramus reaching the anterior end of first antennular segment.
</paragraph>
<paragraph id="8B9AD112FFBEFF8BFF64FD86FBF067D3" blockId="6.[151,1436,292,677]" pageId="6" pageNumber="487">
<emphasis id="B9510D00FFBEFF8BFF64FD86FEA967F7" bold="true" box="[199,313,615,640]" pageId="6" pageNumber="487">Maxillule</emphasis>
(
<figureCitation id="131ECD97FFBEFF8BFEEFFD86FE0467F7" box="[332,404,615,640]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="6" pageNumber="487">Fig. 2</figureCitation>
C). Uniramous, palp (endopod) absent. Coxal endite with 4 setae (three strong terminal cuspidate setae). Basal endite with 4 setae (three long strong terminal cuspidate setae).
</paragraph>
<caption id="DF5A819AFFBEFF8BFF34F845FAC162AE" httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="6" pageNumber="487" targetBox="[154,1425,737,1926]" targetPageId="6">
<paragraph id="8B9AD112FFBEFF8BFF34F845FAC162AE" blockId="6.[151,1435,1956,2009]" pageId="6" pageNumber="487">
<emphasis id="B9510D00FFBEFF8BFF34F845FE8362CE" bold="true" box="[151,275,1956,1978]" pageId="6" pageNumber="487">FIGURE 2.</emphasis>
<taxonomicName id="4C25AA91FFBEFF8BFEB9F845FD8262CD" box="[282,530,1956,1978]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="487" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFBEFF8BFEB9F845FD8262CD" box="[282,530,1956,1978]" italics="true" pageId="6" pageNumber="487">Scyllarides squammosus</emphasis>
</taxonomicName>
, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5th pereiopod. A scale bar = 1 mm; B = 0.5 mm; CE = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B9AD112FFBFFF8AFF64FF76FDAE65A2" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FF76FEB365C7" bold="true" box="[199,291,151,176]" pageId="7" pageNumber="488">Maxilla</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFE97FF76FEE965C7" box="[308,377,151,176]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="7" pageNumber="488">Fig. 2</figureCitation>
D). Rudimentary bud with small elongation at distal end. Triangular shape. Endites, endopod and scaphognathite not differentiated.
</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF64FF3EFBC1658F" blockId="7.[151,1437,151,464]" box="[199,1105,223,248]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FF3EFE15658F" bold="true" box="[199,389,223,248]" pageId="7" pageNumber="488">First maxilliped</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFE37FF3EFE48658F" box="[404,472,223,248]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="7" pageNumber="488">Fig. 2</figureCitation>
D). Undeveloped, present as small conical protuberance.
</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF64FEE5FF4A6437" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FEE5FE02646A" bold="true" box="[199,402,260,285]" pageId="7" pageNumber="488">Third maxilliped</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFE01FEE5FE76646B" box="[418,486,260,285]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="7" pageNumber="488">Fig. 2</figureCitation>
A). Uniramous. Five segmented (ischio-merus fused to basis). Without ventral coxal spine.
</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF64FEADFB5A64DA" blockId="7.[151,1437,151,464]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FEADFEDB6412" bold="true" box="[199,331,332,357]" pageId="7" pageNumber="488">Pereiopods</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFEF9FEADFE0D6413" box="[346,413,332,357]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="7" pageNumber="488">Fig. 2</figureCitation>
A). Biramous, without coxal or subexopodal spines. Exopods of pereiopods 14 flagellated distally with ~1214 annulations, each annulation bears a pair of plumose natatory setae. Pereiopod 5 underdeveloped, small single segment with conical shape. Distal end nearly reaching the telson.
</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF64FE56FCF864A7" blockId="7.[151,1437,151,464]" box="[199,872,439,464]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FE56FE9A64A7" bold="true" box="[199,266,439,464]" pageId="7" pageNumber="488">Pleon</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFEBAFE56FECC64A7" box="[281,348,439,464]" captionStart="FIGURE 2" captionStartId="6.[151,250,1956,1978]" captionTargetBox="[154,1425,737,1926]" captionTargetId="figure@6.[151,1435,729,1935]" captionTargetPageId="6" captionText="FIGURE 2. Scyllarides squammosus, State VI. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. A scale bar = 1 mm; B = 0.5 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264224/files/figure.png" pageId="7" pageNumber="488">Fig. 2</figureCitation>
E). Unsegmented. Pleopods or uropods absent.
</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF34FDE1FD93676F" blockId="7.[151,1437,511,752]" box="[151,515,511,536]" pageId="7" pageNumber="488">Stage VII (NHMUK 2015.3280)</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF64FDC5FB91674B" blockId="7.[151,1437,511,752]" box="[199,1025,548,573]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FDC5FEE6674A" bold="true" box="[199,374,548,573]" pageId="7" pageNumber="488">Measurements</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFE26FDC5FE57674B" box="[389,455,548,573]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="7" pageNumber="488">Fig. 3</figureCitation>
A). TL =
<quantity id="4CDD7CF7FFBFFF8AFD90FDC5FD0A674B" box="[563,666,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="7" pageNumber="488" unit="mm" value="14.9">14.9 mm</quantity>
; CW =
<quantity id="4CDD7CF7FFBFFF8AFD53FDC5FCD9674B" box="[752,841,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="7" pageNumber="488" unit="mm" value="8.2">8.2 mm</quantity>
; CL = 12.0 mm.
</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF64FDA6FF7567F2" blockId="7.[151,1437,511,752]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FDA6FE106717" bold="true" box="[199,384,583,608]" pageId="7" pageNumber="488">Cephalic shield</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFE32FDA6FE466717" box="[401,470,583,608]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="7" pageNumber="488">Fig. 3</figureCitation>
A). Pear-shaped, longer than wide (CL/CW = 1.46), maximum width located to mid length.
</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF64FD6EFBE967DF" blockId="7.[151,1437,511,752]" box="[199,1145,655,680]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FD6EFED367DF" bold="true" box="[199,323,655,680]" pageId="7" pageNumber="488">Antennule</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFEF0FD6EFE0567DF" box="[339,405,655,680]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="7" pageNumber="488">Fig. 3</figureCitation>
B). Primary flagellum with 8 rows of aesthetascs on inner margin.
</paragraph>
<paragraph id="8B9AD112FFBFFF8AFF64FD55FBC96787" blockId="7.[151,1437,511,752]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF64FD55FEBE67BA" bold="true" box="[199,302,692,717]" pageId="7" pageNumber="488">Antenna</emphasis>
(
<figureCitation id="131ECD97FFBFFF8AFE9DFD55FE1267BB" box="[318,386,692,717]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="7" pageNumber="488">Fig. 3</figureCitation>
B). Longer than first antennular segment. Endopod 2-segmented, reaching the middle of the second antennular segment. Exopod retracts and widens, becoming triangular-shaped.
</paragraph>
<caption id="DF5A819AFFBFFF8AFF34F86EFAF862B3" httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="7" pageNumber="488" targetBox="[154,1425,812,1904]" targetPageId="7">
<paragraph id="8B9AD112FFBFFF8AFF34F86EFAF862B3" blockId="7.[151,1435,1935,1988]" pageId="7" pageNumber="488">
<emphasis id="B9510D00FFBFFF8AFF34F86EFE8262D3" bold="true" box="[151,274,1935,1957]" pageId="7" pageNumber="488">FIGURE 3.</emphasis>
<taxonomicName id="4C25AA91FFBFFF8AFEBBF86EFDC462D2" authority="Stage" authorityName="Stage" box="[280,596,1935,1957]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="488" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFBFFF8AFEBBF86EFD9F62D2" box="[280,527,1935,1957]" italics="true" pageId="7" pageNumber="488">Scyllarides squammosus</emphasis>
, Stage
</taxonomicName>
VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9AD112FFB0FF85FF64FF76FB8365A2" blockId="8.[151,1437,151,464]" pageId="8" pageNumber="489">
<emphasis id="B9510D00FFB0FF85FF64FF76FEAA65C7" bold="true" box="[199,314,151,176]" pageId="8" pageNumber="489">Maxillula</emphasis>
(
<figureCitation id="131ECD97FFB0FF85FEEAFF76FE1A65C7" box="[329,394,151,176]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="8" pageNumber="489">Fig. 3</figureCitation>
C). Coxal endite with 6 setae, 4 located in apical position and
<quantity id="4CDD7CF7FFB0FF85FB98FF79FBF565C7" box="[1083,1125,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="489" unit="in" value="2.0">2 in</quantity>
basal position. Basal endite with 6 setae (3 terminal, elongated, with strong structure and serrated laterally).
</paragraph>
<paragraph id="8B9AD112FFB0FF85FF64FF3EFADD658F" blockId="8.[151,1437,151,464]" box="[199,1357,223,248]" pageId="8" pageNumber="489">
<emphasis id="B9510D00FFB0FF85FF64FF3EFEB3658F" bold="true" box="[199,291,223,248]" pageId="8" pageNumber="489">Maxilla</emphasis>
(
<figureCitation id="131ECD97FFB0FF85FE91FF3EFEE4658F" box="[306,372,223,248]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="8" pageNumber="489">Fig. 3</figureCitation>
D). Small tapered elongation becomes less evident, reduced in length. Triangular shape.
</paragraph>
<paragraph id="8B9AD112FFB0FF85FF64FEE5FD19646A" blockId="8.[151,1437,151,464]" box="[199,649,260,285]" pageId="8" pageNumber="489">
<emphasis id="B9510D00FFB0FF85FF64FEE5FE15646A" bold="true" box="[199,389,260,285]" pageId="8" pageNumber="489">First maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB0FF85FE37FEE5FE47646B" box="[404,471,260,285]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="8" pageNumber="489">Fig. 3</figureCitation>
D). Unchanged.
</paragraph>
<paragraph id="8B9AD112FFB0FF85FF64FEC6FD066437" blockId="8.[151,1437,151,464]" box="[199,662,295,320]" pageId="8" pageNumber="489">
<emphasis id="B9510D00FFB0FF85FF64FEC6FE026437" bold="true" box="[199,402,295,320]" pageId="8" pageNumber="489">Third maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB0FF85FE02FEC6FE736437" box="[417,483,295,320]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="8" pageNumber="489">Fig. 3</figureCitation>
A). Unchanged.
</paragraph>
<paragraph id="8B9AD112FFB0FF85FF64FEADFB9A64FF" blockId="8.[151,1437,151,464]" pageId="8" pageNumber="489">
<emphasis id="B9510D00FFB0FF85FF64FEADFEDB6412" bold="true" box="[199,331,332,357]" pageId="8" pageNumber="489">Pereiopods</emphasis>
(
<figureCitation id="131ECD97FFB0FF85FEFEFEADFE336413" box="[349,419,332,357]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="8" pageNumber="489">Fig. 3</figureCitation>
E). Exopods of pereiopods 14 distally flagellated with 1517 annulations. Pereiopod 5 underdeveloped, with conical base at thorax. Distal end reaches beyond telson.
</paragraph>
<paragraph id="8B9AD112FFB0FF85FF64FE75FE5564A7" blockId="8.[151,1437,151,464]" pageId="8" pageNumber="489">
<emphasis id="B9510D00FFB0FF85FF64FE75FE9A64DA" bold="true" box="[199,266,404,429]" pageId="8" pageNumber="489">Pleon</emphasis>
(
<figureCitation id="131ECD97FFB0FF85FEBEFE75FEF364DB" box="[285,355,404,429]" captionStart="FIGURE 3" captionStartId="7.[151,250,1935,1957]" captionTargetBox="[154,1425,812,1904]" captionTargetId="figure@7.[151,1435,805,1914]" captionTargetPageId="7" captionText="FIGURE 3. Scyllarides squammosus, Stage VII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 5 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264225/files/figure.png" pageId="8" pageNumber="489">Fig. 3</figureCitation>
E). Pleon non-segmented, uropods present as small uniramous buds. Telson more developed, longer than previous stage.
</paragraph>
<paragraph id="8B9AD112FFB0FF85FF34FDE1FD9D676F" blockId="8.[151,1437,511,645]" box="[151,525,511,536]" pageId="8" pageNumber="489">Stage VIII (NHMUK 2015.3 281)</paragraph>
<paragraph id="8B9AD112FFB0FF85FF64FDC5FF7567F2" blockId="8.[151,1437,511,645]" pageId="8" pageNumber="489">
<emphasis id="B9510D00FFB0FF85FF64FDC5FEE6674A" bold="true" box="[199,374,548,573]" pageId="8" pageNumber="489">Measurements</emphasis>
(
<figureCitation id="131ECD97FFB0FF85FE26FDC5FE58674B" box="[389,456,548,573]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="8" pageNumber="489">Fig. 4</figureCitation>
A). TL =
<quantity id="4CDD7CF7FFB0FF85FD90FDC5FD0A674B" box="[563,666,548,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="8" pageNumber="489" unit="mm" value="19.5">19.5 mm</quantity>
; CW =
<quantity id="4CDD7CF7FFB0FF85FD53FDC5FCC7674B" box="[752,855,548,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="8" pageNumber="489" unit="mm" value="10.6">10.6 mm</quantity>
; CL =
<quantity id="4CDD7CF7FFB0FF85FC07FDC5FB9C674B" box="[932,1036,548,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="8" pageNumber="489" unit="mm" value="15.5">15.5 mm</quantity>
.
<emphasis id="B9510D00FFB0FF85FF64FDA6FE106717" bold="true" box="[199,384,583,608]" pageId="8" pageNumber="489">Cephalic shield</emphasis>
(
<figureCitation id="131ECD97FFB0FF85FE32FDA6FE476717" box="[401,471,583,608]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="8" pageNumber="489">Fig. 4</figureCitation>
A). Pear-shaped, longer than wide (CL/CW = 1.46), maximum width located to mid length.
</paragraph>
<caption id="DF5A819AFFB0FF85FF34F87DFAE462A7" httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="8" pageNumber="489" targetBox="[152,1427,686,1917]" targetPageId="8">
<paragraph id="8B9AD112FFB0FF85FF34F87DFAE462A7" blockId="8.[151,1436,1948,2000]" pageId="8" pageNumber="489">
<emphasis id="B9510D00FFB0FF85FF34F87DFE8362C6" bold="true" box="[151,275,1948,1970]" pageId="8" pageNumber="489">FIGURE 4.</emphasis>
<taxonomicName id="4C25AA91FFB0FF85FEB8F87DFDC962C5" authority="Stage" authorityName="Stage" box="[283,601,1948,1970]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="489" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFB0FF85FEB8F87DFD8362C5" box="[283,531,1948,1970]" italics="true" pageId="8" pageNumber="489">Scyllarides squammosus</emphasis>
, Stage
</taxonomicName>
VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5th pereiopod. Scale bar: A = 10 mm; B = 1 mm; CE = 0.1 mm.
</paragraph>
</caption>
<caption id="DF5A819AFFB1FF84FF34F960FAE463C2" httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="9" pageNumber="490" targetBox="[163,1422,204,1638]" targetPageId="9">
<paragraph id="8B9AD112FFB1FF84FF34F960FAE463C2" blockId="9.[151,1435,1665,1717]" pageId="9" pageNumber="490">
<emphasis id="B9510D00FFB1FF84FF34F960FE8263E1" bold="true" box="[151,274,1665,1687]" pageId="9" pageNumber="490">FIGURE 5.</emphasis>
<taxonomicName id="4C25AA91FFB1FF84FEBAF960FDC663E0" authority="Stage" authorityName="Stage" box="[281,598,1665,1687]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFB1FF84FEBAF960FD8163E0" box="[281,529,1665,1687]" italics="true" pageId="9" pageNumber="490">Scyllarides squammosus</emphasis>
, Stage
</taxonomicName>
IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9AD112FFB1FF84FF64F93CFDF1626D" blockId="9.[151,1437,1757,1998]" pageId="9" pageNumber="490">
<emphasis id="B9510D00FFB1FF84FF64F93CFED36381" bold="true" box="[199,323,1757,1782]" pageId="9" pageNumber="490">Antennule</emphasis>
(
<figureCitation id="131ECD97FFB1FF84FEF1F93CFE056381" box="[338,405,1757,1782]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="9" pageNumber="490">Fig. 4</figureCitation>
B). Accessary flagellum reaches middle length of primary flagellum. Primary flagellum with 89 rows of aesthetascs on inner margin.
</paragraph>
<paragraph id="8B9AD112FFB1FF84FF64F8C4FA9B6216" blockId="9.[151,1437,1757,1998]" pageId="9" pageNumber="490">
<emphasis id="B9510D00FFB1FF84FF64F8C4FEBE6249" bold="true" box="[199,302,1829,1854]" pageId="9" pageNumber="490">Antenna</emphasis>
(
<figureCitation id="131ECD97FFB1FF84FEE4F8C4FE056249" box="[327,405,1829,1854]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="9" pageNumber="490">Fig. 4</figureCitation>
B). Reaches proximal end of third antennular segment. Endopod now 3-segmented. Significant increase in length of second segment. Triangular outer ramus becomes shorter and stouter.
</paragraph>
<paragraph id="8B9AD112FFB1FF84FF64F88CFDE762DD" blockId="9.[151,1437,1757,1998]" pageId="9" pageNumber="490">
<emphasis id="B9510D00FFB1FF84FF64F88CFEA962F1" bold="true" box="[199,313,1901,1926]" pageId="9" pageNumber="490">Maxillule</emphasis>
(
<figureCitation id="131ECD97FFB1FF84FEE4F88CFE1C62F1" box="[327,396,1901,1926]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="9" pageNumber="490">Fig. 4</figureCitation>
C). Coxal endite with 6 setae, 4 located at distal portion and 2 located in basal position. Basal endite with 7 setae (3 terminal, elongated).
</paragraph>
<paragraph id="8B9AD112FFB1FF84FF64F854FB9F62B9" blockId="9.[151,1437,1757,1998]" box="[199,1039,1973,1998]" pageId="9" pageNumber="490">
<emphasis id="B9510D00FFB1FF84FF64F854FEB362B9" bold="true" box="[199,291,1973,1998]" pageId="9" pageNumber="490">Maxilla</emphasis>
(
<figureCitation id="131ECD97FFB1FF84FE91F854FEE662B9" box="[306,374,1973,1998]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="9" pageNumber="490">Fig. 4</figureCitation>
D). Still underdeveloped, but wider than in previous stages.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FF76FE1265A2" blockId="10.[151,1436,151,320]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FF76FE1865C7" bold="true" box="[199,392,151,176]" pageId="10" pageNumber="491">First maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE39FF76FE7165C7" box="[410,481,151,176]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="10" pageNumber="491">Fig. 4</figureCitation>
D). Present as a minute protuberance. Diameter of conical base significantly larger than previous stages.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FF3EFD36646A" blockId="10.[151,1436,151,320]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FF3EFEDB658F" bold="true" box="[199,331,223,248]" pageId="10" pageNumber="491">Pereiopods</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEFEFF3EFE34658F" box="[349,420,223,248]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="10" pageNumber="491">Fig. 4</figureCitation>
E). Exopods of pereiopods 14 distally flagellated with 1618 annulations. Pereiopod 5 now 2-segmented, and reaching beyond telson.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FEC6FC416437" blockId="10.[151,1436,151,320]" box="[199,977,295,320]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FEC6FE9A6437" bold="true" box="[199,266,295,320]" pageId="10" pageNumber="491">Pleon</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEBAFEC6FECC6437" box="[281,348,295,320]" captionStart="FIGURE 4" captionStartId="8.[151,250,1948,1970]" captionTargetBox="[152,1427,686,1917]" captionTargetId="figure@8.[151,1436,677,1926]" captionTargetPageId="8" captionText="FIGURE 4. Scyllarides squammosus, Stage VIII. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B = 1 mm; C E = 0.1 mm." httpUri="https://zenodo.org/record/264226/files/figure.png" pageId="10" pageNumber="491">Fig. 4</figureCitation>
E). Somite divisions incomplete. Uropods are biramous.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF34FE91FE6A64FF" blockId="10.[151,1437,367,861]" box="[151,506,367,392]" pageId="10" pageNumber="491">Stage IX (NHMUK 2015.3282)</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FE75FB8064DB" blockId="10.[151,1437,367,861]" box="[199,1040,404,429]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FE75FEE664DA" bold="true" box="[199,374,404,429]" pageId="10" pageNumber="491">Measurements</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE26FE75FE5764DA" box="[389,455,404,429]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
A). TL =
<quantity id="4CDD7CF7FFB2FF87FD90FE75FD0A64DB" box="[563,666,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="10" pageNumber="491" unit="mm" value="23.6">23.6 mm</quantity>
; CW =
<quantity id="4CDD7CF7FFB2FF87FD53FE75FCC764DB" box="[752,855,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="10" pageNumber="491" unit="mm" value="13.8">13.8 mm</quantity>
; CL = 19.0 mm.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FE56FAFF64A7" blockId="10.[151,1437,367,861]" box="[199,1391,439,464]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FE56FEEE64A7" bold="true" box="[199,382,439,464]" pageId="10" pageNumber="491">Cephalic shield</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE2EFE56FE5F64A7" box="[397,463,439,464]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
A). Pear-shaped, longer than wide (CL/CW = 1.37). Slightly narrower than thorax.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FE3DFB346482" blockId="10.[151,1437,367,861]" box="[199,1188,476,501]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FE3DFED36482" bold="true" box="[199,323,476,501]" pageId="10" pageNumber="491">Antennule</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEF0FE3DFE066482" box="[339,406,476,501]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
B). Primary flagellum with 910 rows of aesthetascs on inner margin.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FE1EFF6F674A" blockId="10.[151,1437,367,861]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FE1EFEBE676F" bold="true" box="[199,302,511,536]" pageId="10" pageNumber="491">Antenna</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE9FFE1EFEEF676F" box="[316,383,511,536]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
B). Exceeds second antennular segment. Four segments remain but the exopod now reduced to small tip.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FDA6FE4467F2" blockId="10.[151,1437,367,861]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FDA6FEA96717" bold="true" box="[199,313,583,608]" pageId="10" pageNumber="491">Maxillule</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEEAFDA6FE1D6717" box="[329,397,583,608]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
C). Coxal endite with 7 setae (3 longer setae in distal portion). Basal endite with 9 setae (3 terminal, long and serrated).
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FD6EFAFA67DF" blockId="10.[151,1437,367,861]" box="[199,1386,655,680]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FD6EFEB367DF" bold="true" box="[199,291,655,680]" pageId="10" pageNumber="491">Maxilla</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE91FD6EFEE467DF" box="[306,372,655,680]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
D). Similar to previous stages, but scaphognathite acquires rudimentary rectangular shape.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FD55FA0367BA" blockId="10.[151,1437,367,861]" box="[199,1427,692,717]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FD55FE1567BA" bold="true" box="[199,389,692,717]" pageId="10" pageNumber="491">First maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE37FD55FE4767BA" box="[404,471,692,717]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
D). Adopting more tubular form with length significantly larger than previous stages.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FD36FE326662" blockId="10.[151,1437,367,861]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FD36FEDB6787" bold="true" box="[199,331,727,752]" pageId="10" pageNumber="491">Pereiopods</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEFFFD36FE306787" box="[348,416,727,752]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
A, E). Exopods of pereiopods 14 distally flagellated. Pereiopod 5 now 3-segmented and reaching beyond telson.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FCFEFBBE662A" blockId="10.[151,1437,367,861]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FCFEFE9A664F" bold="true" box="[199,266,799,824]" pageId="10" pageNumber="491">Pleon</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEBDFCFEFEF5664F" box="[286,357,799,824]" captionStart="FIGURE 5" captionStartId="9.[151,250,1665,1687]" captionTargetBox="[163,1422,204,1638]" captionTargetId="figure@9.[151,1435,193,1644]" captionTargetPageId="9" captionText="FIGURE 5. Scyllarides squammosus, Stage IX. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264227/files/figure.png" pageId="10" pageNumber="491">Fig. 5</figureCitation>
E). Divisions of somites becomes evident. Pleopods present, underdeveloped and biramous. Uropods biramous, length of internal lobe half than outer lobe. Telson unchanged.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF34FC66FE6166E8" blockId="10.[151,1437,902,1611]" box="[151,497,902,927]" pageId="10" pageNumber="491">Stage X (NHMUK 2015.3283)</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FC4BFB8066B5" blockId="10.[151,1437,902,1611]" box="[199,1040,938,963]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FC4BFEE666B4" bold="true" box="[199,374,938,963]" pageId="10" pageNumber="491">Measurements</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE26FC4BFE5866B4" box="[389,456,938,963]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
A). TL = 31.0 mm; CW =
<quantity id="4CDD7CF7FFB2FF87FD53FC4BFCC766B5" box="[752,855,938,963]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="10" pageNumber="491" unit="mm" value="18.5">18.5 mm</quantity>
; CL = 24.0 mm.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FC2FFF73617C" blockId="10.[151,1437,902,1611]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FC2FFEEE6690" bold="true" box="[199,382,974,999]" pageId="10" pageNumber="491">Cephalic shield</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE2EFC2FFE416690" box="[397,465,974,999]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
A). Pear-shaped, longer than wide (CL/CW = 1.29), but relatively wider than previous stages.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FBF7FD266124" blockId="10.[151,1437,902,1611]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FBF7FED36158" bold="true" box="[199,323,1046,1071]" pageId="10" pageNumber="491">Antennule</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEF6FBF7FE0C6158" box="[341,412,1046,1071]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
B). Accessory flagellum longer than half length of primary flagellum. Primary flagellum with 1012 rows of aesthetascs on inner margin.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FBBFFBC561EC" blockId="10.[151,1437,902,1611]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FBBFFEBE6100" bold="true" box="[199,302,1118,1143]" pageId="10" pageNumber="491">Antenna</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE9EFBBFFE126100" box="[317,386,1118,1143]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
B). Five-segmented and reaching beyond third antennular segment. Second antennal segment develops a lateral extension (lateral process) while first segment becomes cylindrical.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FB47FADD61C8" blockId="10.[151,1437,902,1611]" box="[199,1357,1190,1215]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FB47FEA961C8" bold="true" box="[199,313,1190,1215]" pageId="10" pageNumber="491">Maxillule</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEEBFB47FE1B61C8" box="[328,395,1190,1215]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
C). Coxal endite with 13 setae. Basal endite with 9 setae (3 long, strong and serrated).
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FB2BFAD46194" blockId="10.[151,1437,902,1611]" box="[199,1348,1226,1251]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FB2BFEB36194" bold="true" box="[199,291,1226,1251]" pageId="10" pageNumber="491">Maxilla</emphasis>
. (
<figureCitation id="131ECD97FFB2FF87FE9AFB2BFEEC6194" box="[313,380,1226,1251]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
D). Scaphognathite slightly differentiated, without setae, flattened and reduced in size.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FB0FFD756070" blockId="10.[151,1437,902,1611]" box="[199,741,1262,1287]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FB0FFE156070" bold="true" box="[199,389,1262,1287]" pageId="10" pageNumber="491">First maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE37FB0FFE486070" box="[404,472,1262,1287]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
D). Like previous stage.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FAF3FAE8605D" blockId="10.[151,1437,902,1611]" box="[199,1400,1298,1323]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FAF3FE31605C" bold="true" box="[199,417,1298,1323]" pageId="10" pageNumber="491">Second maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE12FAF3FE64605C" box="[433,500,1298,1323]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
A). Five-segmented, with rudimentary and unarmed minute exopod bud present.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FAD7FCFA6005" blockId="10.[151,1437,902,1611]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FAD7FE056038" bold="true" box="[199,405,1334,1359]" pageId="10" pageNumber="491">Third maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FE0BFAD7FE7E6038" box="[424,494,1334,1359]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
A, F). Uniramous. Five segmented (ischio-merus fused to basis). Without ventral coxal spine. Distal part of propodus and dactylus densely setose.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FA9FFB0D60A8" blockId="10.[151,1437,902,1611]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FA9FFEDB60E0" bold="true" box="[199,331,1406,1431]" pageId="10" pageNumber="491">Pereiopods</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEC2FA9FFE3B60E0" box="[353,427,1406,1431]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
E, G). Pereiopods 14 biramous, without coxal or subexopodal spines. Exopods of pereiopods 14 flagellated distally with 1719 annulations, each annulation bears a pair of plumose natatory setae. Pereiopod 5 with four-segmented endopod, without setae. Distal end reaches beyond telson.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64FA0BFB26633C" blockId="10.[151,1437,902,1611]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64FA0BFE9A6374" bold="true" box="[199,266,1514,1539]" pageId="10" pageNumber="491">Pleon</emphasis>
(
<figureCitation id="131ECD97FFB2FF87FEB8FA0BFEF06374" box="[283,352,1514,1539]" captionStart="FIGURE 6" captionStartId="11.[151,250,1412,1434]" captionTargetBox="[160,1430,181,1382]" captionTargetId="figure@11.[151,1435,179,1391]" captionTargetPageId="11" captionText="FIGURE 6. Scyllarides squammosus, Stage X. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5 th pereiopod; F, 3 rd maxilliped; G, 5 th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm." httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="10" pageNumber="491">Fig. 6</figureCitation>
E). Five-segmented (five somites plus telson, somite 1 not differentiated). Pleonites 25 with a pair of biramous pleopods, unsegmented and unarmed. Uropods biramous and incipiently segmented, but not outreaching posterior margin of telson. Telson rounded posteriorly, without processes or teeth.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF34F997FE6A63F9" blockId="10.[151,1437,1653,2038]" box="[151,506,1653,1678]" pageId="10" pageNumber="491">Stage XI (NHMUK 2015.3284)</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64F978FB8063C6" blockId="10.[151,1437,1653,2038]" box="[199,1040,1689,1714]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64F978FEE663C5" bold="true" box="[199,374,1689,1714]" pageId="10" pageNumber="491">Measurements</emphasis>
(Fig. 7A). TL = 39.0 mm; CW =
<quantity id="4CDD7CF7FFB2FF87FD53F97BFCC763C6" box="[752,855,1690,1714]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="10" pageNumber="491" unit="mm" value="23.1">23.1 mm</quantity>
; CL = 29.0 mm.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64F95CFF75638D" blockId="10.[151,1437,1653,2038]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64F95CFE1063A1" bold="true" box="[199,384,1725,1750]" pageId="10" pageNumber="491">Cephalic shield</emphasis>
(Fig. 7A). Pear-shaped, longer than wide (CL/CW = 1.26), maximum width located to mid length.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64F8E4FB216269" blockId="10.[151,1437,1653,2038]" box="[199,1201,1797,1822]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64F8E4FED36269" bold="true" box="[199,323,1797,1822]" pageId="10" pageNumber="491">Antennule</emphasis>
(Fig. 7B). Primary flagellum with 1214 rows of aesthetascs on inner margin.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64F8C8FE656212" blockId="10.[151,1437,1653,2038]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64F8C8FEBE6235" bold="true" box="[199,302,1833,1858]" pageId="10" pageNumber="491">Antenna</emphasis>
(Fig. 7B). Almost equal in length to antennule. Lateral expansion on the second segment widens while segment 4 gets narrower.
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64F890FD2362D9" blockId="10.[151,1437,1653,2038]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64F890FEA962FD" bold="true" box="[199,313,1905,1930]" pageId="10" pageNumber="491">Maxillule</emphasis>
(Fig. 7C). Coxal endite with 14 setae (2 terminal setae considerably longer and stronger). Basal endite with 9 setae (3 long, strong and serrated).
</paragraph>
<paragraph id="8B9AD112FFB2FF87FF64F858FD266281" blockId="10.[151,1437,1653,2038]" pageId="10" pageNumber="491">
<emphasis id="B9510D00FFB2FF87FF64F858FEB362A5" bold="true" box="[199,291,1977,2002]" pageId="10" pageNumber="491">Maxilla</emphasis>
(Fig. 7D). Similar to previous stage, but scaphognathite more differentiated, without setae, flattened and slightly expanded anteriorly and posteriorly.
</paragraph>
<caption id="DF5A819AFFB3FF86FF34FA65FCB460A0" httpUri="https://zenodo.org/record/264228/files/figure.png" pageId="11" pageNumber="492" targetBox="[160,1430,181,1382]" targetPageId="11">
<paragraph id="8B9AD112FFB3FF86FF34FA65FCB460A0" blockId="11.[151,1436,1412,1495]" pageId="11" pageNumber="492">
<emphasis id="B9510D00FFB3FF86FF34FA65FE8360EE" bold="true" box="[151,275,1412,1434]" pageId="11" pageNumber="492">FIGURE 6.</emphasis>
<taxonomicName id="4C25AA91FFB3FF86FEB9FA65FD0260ED" authority="Stage X. A" authorityName="Stage X. A" box="[282,658,1412,1434]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="492" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFB3FF86FEB9FA65FD8960ED" box="[282,537,1412,1434]" italics="true" pageId="11" pageNumber="492">Scyllarides squammosus,</emphasis>
Stage X. A
</taxonomicName>
, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and 5th pereiopod; F, 3rd maxilliped; G, 5th segment and dactyl of 1 st pereiopod. Scale bar: A = 10 mm; B, E = 1 mm; C, D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9AD112FFB3FF86FF64FA1FFD476360" blockId="11.[151,1436,1534,1775]" box="[199,727,1534,1559]" pageId="11" pageNumber="492">
<emphasis id="B9510D00FFB3FF86FF64FA1FFE156360" bold="true" box="[199,389,1534,1559]" pageId="11" pageNumber="492">First maxilliped</emphasis>
(Fig. 7D). Further developed.
</paragraph>
<paragraph id="8B9AD112FFB3FF86FF64F9C3FD0263F4" blockId="11.[151,1436,1534,1775]" pageId="11" pageNumber="492">
<emphasis id="B9510D00FFB3FF86FF64F9C3FEDB634C" bold="true" box="[199,331,1570,1595]" pageId="11" pageNumber="492">Pereiopods</emphasis>
(Fig. 7E). Biramous, without coxal or subexopodal spines. Exopods of pereiopods 14 flagellated distally with&gt;18 annulations, each annulation bears a pair of plumose natatory setae. Pereiopod 5 with 5 segments present, segment 2 with rudimentary exopod.
</paragraph>
<paragraph id="8B9AD112FFB3FF86FF64F96FFD3D6398" blockId="11.[151,1436,1534,1775]" pageId="11" pageNumber="492">
<emphasis id="B9510D00FFB3FF86FF64F96FFE9A63D0" bold="true" box="[199,266,1678,1703]" pageId="11" pageNumber="492">Pleon</emphasis>
(Fig. 7E, F). Segmented, with six pleonites plus telson. Pleonites 25 with a pair of biramous and unsegmented pleopods each, without appendix interna. Presence of two dorsal spines on pleonites 34. Biramous uropods outreaching posterior margin of telson.
</paragraph>
<paragraph id="8B9AD112FFB3FF86FF34F8FEFD936240" blockId="11.[151,1436,1822,2027]" box="[151,515,1822,1847]" pageId="11" pageNumber="492">Stage XII (NHMUK 2015.3285)</paragraph>
<paragraph id="8B9AD112FFB3FF86FF64F8A3FB98622D" blockId="11.[151,1436,1822,2027]" box="[199,1032,1858,1883]" pageId="11" pageNumber="492">
<emphasis id="B9510D00FFB3FF86FF64F8A3FEE6622C" bold="true" box="[199,374,1858,1883]" pageId="11" pageNumber="492">Measurements</emphasis>
(
<figureCitation id="131ECD97FFB3FF86FE26F8A3FE57622C" box="[389,455,1858,1883]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="11" pageNumber="492">Fig. 8</figureCitation>
A). TL = 51.0 mm; CW =
<quantity id="4CDD7CF7FFB3FF86FD53F8A3FCC7622D" box="[752,855,1858,1883]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="11" pageNumber="492" unit="mm" value="26.5">26.5 mm</quantity>
; CL = 36.0 mm
</paragraph>
<paragraph id="8B9AD112FFB3FF86FF64F887FEAF62D4" blockId="11.[151,1436,1822,2027]" pageId="11" pageNumber="492">
<emphasis id="B9510D00FFB3FF86FF64F887FE116208" bold="true" box="[199,385,1894,1919]" pageId="11" pageNumber="492">Cephalic shield</emphasis>
(
<figureCitation id="131ECD97FFB3FF86FE37F887FE496208" box="[404,473,1894,1919]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="11" pageNumber="492">Fig. 8</figureCitation>
A). Pear-shaped, longer than wide (CL/CW = 1.35), but proportionally longer than previous stage.
</paragraph>
<paragraph id="8B9AD112FFB3FF86FF64F84FFD26629C" blockId="11.[151,1436,1822,2027]" pageId="11" pageNumber="492">
<emphasis id="B9510D00FFB3FF86FF64F84FFED362B0" bold="true" box="[199,323,1966,1991]" pageId="11" pageNumber="492">Antennule</emphasis>
(
<figureCitation id="131ECD97FFB3FF86FEF7F84FFE0762B0" box="[340,407,1966,1991]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="11" pageNumber="492">Fig. 8</figureCitation>
B). Accessory flagellum extended, equal in length to primary flagellum. Primary flagellum with 1315 rows of aesthetascs on inner margin.
</paragraph>
<paragraph id="8B9AD112FFB4FF81FF34FA4CFEDD6377" pageId="12" pageNumber="493">
<table id="F92523B2FFB40072FF34FA4DFA0B6377" box="[151,1435,1452,1536]" gridcols="3" gridrows="3" pageId="12" pageNumber="493">
<tr id="3515D350FFB40072FF34FA4DFA0B60B4" box="[151,1435,1452,1475]" gridrow="0" pageId="12" pageNumber="493">
<th id="76C4BA2CFFB40072FF34FA4DFD8860B4" box="[151,536,1452,1475]" gridcol="0" gridrow="0" pageId="12" pageNumber="493">
<emphasis id="B9510D00FFB4FF81FF34FA4CFE8360B5" bold="true" box="[151,275,1453,1475]" pageId="12" pageNumber="493">FIGURE 7.</emphasis>
<taxonomicName id="4C25AA91FFB4FF81FEB9FA4DFD8360B4" box="[282,531,1452,1475]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="493" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFB4FF81FEB9FA4DFD8360B4" box="[282,531,1452,1475]" italics="true" pageId="12" pageNumber="493">Scyllarides squammosus</emphasis>
</taxonomicName>
,
</th>
<th id="76C4BA2CFFB40072FDBCFA4DFCB260B4" box="[543,802,1452,1475]" gridcol="1" gridrow="0" pageId="12" pageNumber="493">Stage X. A, ventral view;</th>
<th id="76C4BA2CFFB40072FC8AFA4DFA0B60B4" box="[809,1435,1452,1475]" gridcol="2" gridrow="0" pageId="12" pageNumber="493">B, details of antennule and antenna; C, detail of maxillule; D,</th>
</tr>
<tr id="3515D350FFB40072FF34FA2AFA0B6096" box="[151,1435,1483,1505]" gridrow="1" pageId="12" pageNumber="493">
<th id="76C4BA2CFFB40072FF34FA2AFD886096" box="[151,536,1483,1505]" gridcol="0" gridrow="1" pageId="12" pageNumber="493">maxilla and first maxilliped; E, detail</th>
<td id="76C4BA2CFFB40072FDBCFA2AFCB26096" box="[543,802,1483,1505]" gridcol="1" gridrow="1" pageId="12" pageNumber="493">of abdomen and detail of</td>
<td id="76C4BA2CFFB40072FC8AFA2AFA0B6096" box="[809,1435,1483,1505]" gridcol="2" gridrow="1" pageId="12" pageNumber="493">5th pereiopod; F, dorsal view of abdomen. Scale bar: A = 10</td>
</tr>
<tr id="3515D350FFB40072FF34FA0BFA0B6377" box="[151,1435,1514,1536]" gridrow="2" pageId="12" pageNumber="493" rowspan-1="1" rowspan-2="1">
<th id="76C4BA2CFFB40072FF34FA0BFD886377" box="[151,536,1514,1536]" gridcol="0" gridrow="2" pageId="12" pageNumber="493">mm; BF = 1mm.</th>
</tr>
</table>
</paragraph>
<paragraph id="8B9AD112FFB4FF81FF64F9CDFB3A631E" blockId="12.[151,1437,1580,2038]" pageId="12" pageNumber="493">
<emphasis id="B9510D00FFB4FF81FF64F9CDFEBE6332" bold="true" box="[199,302,1580,1605]" pageId="12" pageNumber="493">Antenna</emphasis>
(
<figureCitation id="131ECD97FFB4FF81FE9DF9CDFE126332" box="[318,386,1580,1605]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
B). Longer than antennule. Segments fully developed. Lateral expansion on second segment directed anteriorly (more developed than previous stage). Last antennal segment leaf-shaped.
</paragraph>
<paragraph id="8B9AD112FFB4FF81FF64F994FD5063C6" blockId="12.[151,1437,1580,2038]" pageId="12" pageNumber="493">
<emphasis id="B9510D00FFB4FF81FF64F994FEA963F9" bold="true" box="[199,313,1653,1678]" pageId="12" pageNumber="493">Maxillule</emphasis>
(
<figureCitation id="131ECD97FFB4FF81FEE8F994FE0163FA" box="[331,401,1653,1678]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
C). Coxal endite with 14 setae (2 terminal setae considerably longer and stronger). Basal endite with 11 setae (3 long, strong and serrated).
</paragraph>
<paragraph id="8B9AD112FFB4FF81FF64F95CFE4F638E" blockId="12.[151,1437,1580,2038]" pageId="12" pageNumber="493">
<emphasis id="B9510D00FFB4FF81FF64F95CFEB363A1" bold="true" box="[199,291,1725,1750]" pageId="12" pageNumber="493">Maxilla</emphasis>
(
<figureCitation id="131ECD97FFB4FF81FE9FF95CFE1763A2" box="[316,391,1725,1750]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
D). Fully developed, scaphognathite considerably expanded anteriorly and posteriorly. Endopod becomes triangular.
</paragraph>
<paragraph id="8B9AD112FFB4FF81FF64F8E4FC5F6269" blockId="12.[151,1437,1580,2038]" box="[199,975,1797,1822]" pageId="12" pageNumber="493">
<emphasis id="B9510D00FFB4FF81FF64F8E4FE156269" bold="true" box="[199,389,1797,1822]" pageId="12" pageNumber="493">First maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB4FF81FE37F8E4FE47626A" box="[404,471,1797,1822]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
D). Fully developed and markedly biramous.
</paragraph>
<paragraph id="8B9AD112FFB4FF81FF64F8C9FCB26236" blockId="12.[151,1437,1580,2038]" box="[199,802,1832,1857]" pageId="12" pageNumber="493">
<emphasis id="B9510D00FFB4FF81FF64F8C9FE316236" bold="true" box="[199,417,1832,1857]" pageId="12" pageNumber="493">Second maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB4FF81FE12F8C9FE636236" box="[433,499,1832,1857]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
A). Exopod bud elongated.
</paragraph>
<paragraph id="8B9AD112FFB4FF81FF64F8ACFEB362FE" blockId="12.[151,1437,1580,2038]" pageId="12" pageNumber="493">
<emphasis id="B9510D00FFB4FF81FF64F8ACFE066211" bold="true" box="[199,406,1869,1894]" pageId="12" pageNumber="493">Third maxilliped</emphasis>
(
<figureCitation id="131ECD97FFB4FF81FE09F8ACFE606212" box="[426,496,1869,1894]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
A). Gill buds present, with one pleurobranch, one arthrobranch and epipod with podobranch.
</paragraph>
<paragraph id="8B9AD112FFB4FF81FF64F874FA896281" blockId="12.[151,1437,1580,2038]" pageId="12" pageNumber="493">
<emphasis id="B9510D00FFB4FF81FF64F874FEDB62D9" bold="true" box="[199,331,1941,1966]" pageId="12" pageNumber="493">Pereiopods</emphasis>
(
<figureCitation id="131ECD97FFB4FF81FEFDF874FE3462DA" box="[350,420,1941,1966]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
E). Pereiopod 1 with one pleurobranch, one arthrobranch and epipod with podobranch. Pereiopods 24 with two pleurobranchs, one arthrobranch and epipod with podobranch. Pereiopod 5 with one pleurobranch. Pereiopod 5 now 6-segmented. Basis with exopod bud and ischium and merus are fused.
</paragraph>
<caption id="DF5A819AFFB5FF80FF34FAA5FD5760E0" httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="13" pageNumber="494" targetBox="[158,1426,200,1317]" targetPageId="13">
<paragraph id="8B9AD112FFB5FF80FF34FAA5FD5760E0" blockId="13.[151,1436,1348,1431]" pageId="13" pageNumber="494">
<emphasis id="B9510D00FFB5FF80FF34FAA5FE86602E" bold="true" box="[151,278,1348,1370]" pageId="13" pageNumber="494">FIGURE 8.</emphasis>
<taxonomicName id="4C25AA91FFB5FF80FE82FAA5FDF6602D" authority="Stage" authorityName="Stage" box="[289,614,1348,1370]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="494" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFB5FF80FE82FAA5FD8D602D" box="[289,541,1348,1370]" italics="true" pageId="13" pageNumber="494">Scyllarides squammosus</emphasis>
, Stage
</taxonomicName>
XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; BF = 1mm.
</paragraph>
</caption>
<paragraph id="8B9AD112FFB5FF80FF64FA5FFEC66334" blockId="13.[151,1437,1470,1897]" pageId="13" pageNumber="494">
<emphasis id="B9510D00FFB5FF80FF64FA5FFE9A60A0" bold="true" box="[199,266,1470,1495]" pageId="13" pageNumber="494">Pleon</emphasis>
(
<figureCitation id="131ECD97FFB5FF80FEBAFA5FFECA60A0" box="[281,346,1470,1495]" captionStart="FIGURE 8" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1426,200,1317]" captionTargetId="figure@13.[151,1435,193,1326]" captionTargetPageId="13" captionText="FIGURE 8. Scyllarides squammosus, Stage XII-Final. A, ventral view; B, details of antennule and antenna; C, detail of maxillule; D, maxilla and first maxilliped; E, detail of abdomen and detail of 5 th pereiopod; F, dorsal view of abdomen; G, detail of pleopods. Scale bar: A = 10 mm; B F = 1 mm." httpUri="https://zenodo.org/record/264229/files/figure.png" pageId="13" pageNumber="494">Fig. 8</figureCitation>
E, F). Fully segmented with 4 pairs of segmented pleopods. Posterolateral margin of the pleura of somites 2, 3, 4 and 6 ended with acute spine. Dorsally, somites 4 and 5 bear a strong median spine. Pleopods biramous, segmented and without setae, endopod with appendix interna. Biramous uropods outreaching posterior margin of telson.
</paragraph>
<paragraph id="8B9AD112FFB5FF80FF64F9AFFD21621F" blockId="13.[151,1437,1470,1897]" pageId="13" pageNumber="494">
Morphometric analyses showed that total length of the larvae follows an exponential growth in late stages of development (see
<tableCitation id="C6A7E4A9FFB5FF80FED2F993FE5E63FC" box="[369,462,1650,1675]" captionStart="TABLE 2" captionStartId="14.[151,239,152,175]" captionTargetBox="[159,1265,202,564]" captionTargetPageId="14" captionText="TABLE 2. Larval development of Scyllarides squammosus phyllosoma." httpUri="http://table.plazi.org/id/DF5A819AFFB6FF83FF34FF79FC0965D8" pageId="13" pageNumber="494" tableUuid="DF5A819AFFB6FF83FF34FF79FC0965D8">Table 2</tableCitation>
). The ratio of cephalic width over the total length increased during phyllosoma development up to a maximum value of CW/TL = 0.61 at around
<quantity id="4CDD7CF7FFB5FF80FCDFF976FC5C63D8" box="[892,972,1687,1711]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="494" unit="mm" value="30.0">30 mm</quantity>
TL (see
<figureCitation id="131ECD97FFB5FF80FB92F977FBE363D8" box="[1073,1139,1686,1711]" captionStart="FIGURE 9" captionStartId="14.[151,250,1345,1367]" captionTargetBox="[328,1259,742,1320]" captionTargetId="figure@14.[327,1259,742,1323]" captionTargetPageId="14" captionText="FIGURE 9. Relationship between CW / TL ratio and TL using all Scyllarides squammosus larvae sampled in this study (n = 34, including stages VI and VII) and those present in the previous literature." httpUri="https://zenodo.org/record/264230/files/figure.png" pageId="13" pageNumber="494">Fig. 9</figureCitation>
) and then decreased again in the sub-final and final stages (CW/ TL = 0.43 + 0.011 TL0.00018 TL2; r2 = 0.999). The relative growth of CL and CW during larval development in phyllosoma larvae assigned to
<taxonomicName id="4C25AA91FFB5FF80FC02F900FB23638E" box="[929,1203,1761,1786]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="494" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFB5FF80FC02F900FB23638E" box="[929,1203,1761,1786]" italics="true" pageId="13" pageNumber="494">Scyllarides squammosus</emphasis>
</taxonomicName>
was found to follow a power law equation (CW = 0.84 CL0.96), with a high correlation coefficient (r2 = 0.996). However, the confidence interval for the exponent parameter comprised the value b = 1 (0.911.01), which indicates that the growth of CW does not show a significantly allometric pattern.
</paragraph>
<caption id="DF5A819AFFB6FF83FF34FF79FC0965D8" ID-Table-UUID="DF5A819AFFB6FF83FF34FF79FC0965D8" box="[151,921,151,175]" httpUri="http://table.plazi.org/id/DF5A819AFFB6FF83FF34FF79FC0965D8" pageId="14" pageNumber="495" targetBox="[159,1265,202,564]" targetIsTable="true" targetPageId="14">
<paragraph id="8B9AD112FFB6FF83FF34FF79FC0965D8" blockId="14.[151,921,151,175]" box="[151,921,151,175]" pageId="14" pageNumber="495">
<emphasis id="B9510D00FFB6FF83FF34FF79FE9965D8" bold="true" box="[151,265,152,175]" pageId="14" pageNumber="495">TABLE 2.</emphasis>
Larval development of
<taxonomicName id="4C25AA91FFB6FF83FDA9FF76FC9F65D8" box="[522,783,151,175]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="495" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="B9510D00FFB6FF83FDA9FF76FC9F65D8" box="[522,783,151,175]" italics="true" pageId="14" pageNumber="495">Scyllarides squammosus</emphasis>
</taxonomicName>
phyllosoma.
</paragraph>
</caption>
<paragraph id="8B9AD112FFB6FF83FF3CFF2BFB2E6747" pageId="14" pageNumber="495">
<table id="F92523B2FFB60072FF3CFF2BFB616743" box="[159,1265,202,564]" gridcols="4" gridrows="7" pageId="14" pageNumber="495">
<tr id="3515D350FFB60072FF3CFF2BFB616588" box="[159,1265,202,255]" gridrow="0" pageId="14" pageNumber="495">
<th id="76C4BA2CFFB60072FF3CFF2BFF6F6588" box="[159,255,202,255]" gridcol="0" gridrow="0" pageId="14" pageNumber="495">Stage</th>
<th id="76C4BA2CFFB60072FE53FF2BFD926588" box="[496,514,202,255]" gridcol="1" gridrow="0" pageId="14" pageNumber="495">N</th>
<th id="76C4BA2CFFB60072FCE2FF2BFC246588" box="[833,948,202,255]" gridcol="2" gridrow="0" pageId="14" pageNumber="495">TL Range (mm)</th>
<th id="76C4BA2CFFB60072FB31FF2BFB616588" box="[1170,1265,202,255]" gridcol="3" gridrow="0" pageId="14" pageNumber="495">TL Mean (mm)</th>
</tr>
<tr id="3515D350FFB60072FF3CFEF3FB61645F" box="[159,1265,274,296]" gridrow="1" pageId="14" pageNumber="495">
<th id="76C4BA2CFFB60072FF3CFEF3FF6F645F" box="[159,255,274,296]" gridcol="0" gridrow="1" pageId="14" pageNumber="495">VI</th>
<td id="76C4BA2CFFB60072FE53FEF3FD92645F" box="[496,514,274,296]" gridcol="1" gridrow="1" pageId="14" pageNumber="495">1</td>
<td id="76C4BA2CFFB60072FCE2FEF3FC24645F" box="[833,948,274,296]" gridcol="2" gridrow="1" pageId="14" pageNumber="495">11.7</td>
<td id="76C4BA2CFFB60072FB31FEF3FB61645F" box="[1170,1265,274,296]" gridcol="3" gridrow="1" pageId="14" pageNumber="495">11.7</td>
</tr>
<tr id="3515D350FFB60072FF3CFEDDFB616409" box="[159,1265,316,382]" gridrow="2" pageId="14" pageNumber="495">
<th id="76C4BA2CFFB60072FF3CFEDDFF6F6409" box="[159,255,316,382]" gridcol="0" gridrow="2" pageId="14" pageNumber="495">VII VIII</th>
<td id="76C4BA2CFFB60072FE53FEDDFD926409" box="[496,514,316,382]" gridcol="1" gridrow="2" pageId="14" pageNumber="495">1 3</td>
<td id="76C4BA2CFFB60072FCE2FEDDFC246409" box="[833,948,316,382]" gridcol="2" gridrow="2" pageId="14" pageNumber="495">14.9 19.522.0</td>
<td id="76C4BA2CFFB60072FB31FEDDFB616409" box="[1170,1265,316,382]" gridcol="3" gridrow="2" pageId="14" pageNumber="495">14.9 19.8</td>
</tr>
<tr id="3515D350FFB60072FF3CFE73FB6164DB" box="[159,1265,402,428]" gridrow="3" pageId="14" pageNumber="495">
<th id="76C4BA2CFFB60072FF3CFE73FF6F64DB" box="[159,255,402,428]" gridcol="0" gridrow="3" pageId="14" pageNumber="495">IX</th>
<td id="76C4BA2CFFB60072FE53FE73FD9264DB" box="[496,514,402,428]" gridcol="1" gridrow="3" pageId="14" pageNumber="495">5</td>
<td id="76C4BA2CFFB60072FCE2FE73FC2464DB" box="[833,948,402,428]" gridcol="2" gridrow="3" pageId="14" pageNumber="495">22.326.8</td>
<td id="76C4BA2CFFB60072FB31FE73FB6164DB" box="[1170,1265,402,428]" gridcol="3" gridrow="3" pageId="14" pageNumber="495">24.6</td>
</tr>
<tr id="3515D350FFB60072FF3CFE5EFB6164AE" box="[159,1265,447,473]" gridrow="4" pageId="14" pageNumber="495">
<th id="76C4BA2CFFB60072FF3CFE5EFF6F64AE" box="[159,255,447,473]" gridcol="0" gridrow="4" pageId="14" pageNumber="495">X</th>
<td id="76C4BA2CFFB60072FE53FE5EFD9264AE" box="[496,514,447,473]" gridcol="1" gridrow="4" pageId="14" pageNumber="495">7</td>
<td id="76C4BA2CFFB60072FCE2FE5EFC2464AE" box="[833,948,447,473]" gridcol="2" gridrow="4" pageId="14" pageNumber="495">29.035.6</td>
<td id="76C4BA2CFFB60072FB31FE5EFB6164AE" box="[1170,1265,447,473]" gridcol="3" gridrow="4" pageId="14" pageNumber="495">31.4</td>
</tr>
<tr id="3515D350FFB60072FF3CFE0DFB616771" box="[159,1265,492,518]" gridrow="5" pageId="14" pageNumber="495">
<th id="76C4BA2CFFB60072FF3CFE0DFF6F6771" box="[159,255,492,518]" gridcol="0" gridrow="5" pageId="14" pageNumber="495">XI</th>
<td id="76C4BA2CFFB60072FE53FE0DFD926771" box="[496,514,492,518]" gridcol="1" gridrow="5" pageId="14" pageNumber="495">8</td>
<td id="76C4BA2CFFB60072FCE2FE0DFC246771" box="[833,948,492,518]" gridcol="2" gridrow="5" pageId="14" pageNumber="495">37.040.0</td>
<td id="76C4BA2CFFB60072FB31FE0DFB616771" box="[1170,1265,492,518]" gridcol="3" gridrow="5" pageId="14" pageNumber="495">38.7</td>
</tr>
<tr id="3515D350FFB60072FF3CFDFBFB616743" box="[159,1265,538,564]" gridrow="6" pageId="14" pageNumber="495">
<th id="76C4BA2CFFB60072FF3CFDFBFF6F6743" box="[159,255,538,564]" gridcol="0" gridrow="6" pageId="14" pageNumber="495">XII-Final</th>
<td id="76C4BA2CFFB60072FE53FDFBFD926743" box="[496,514,538,564]" gridcol="1" gridrow="6" pageId="14" pageNumber="495">9</td>
<td id="76C4BA2CFFB60072FCE2FDFBFC246743" box="[833,948,538,564]" gridcol="2" gridrow="6" pageId="14" pageNumber="495">47.051.0</td>
<td id="76C4BA2CFFB60072FB31FDFBFB616743" box="[1170,1265,538,564]" gridcol="3" gridrow="6" pageId="14" pageNumber="495">49.3</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9AD112FFB6FF83FF34FDB4FEE467C6" blockId="14.[151,1436,597,689]" pageId="14" pageNumber="495">Stage = Stage of development; N = Sample size; Range = range of lengths per stage; Mean = Average total length per stage. Total length (TL) measured from the anterior margin of the cephalic shield between the eyes to the posterior margin of the telson.</paragraph>
</subSubSection>
</treatment>
</document>