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<document id="C74EF48579E8015AA2D967898FA225C8" ID-DOI="10.11646/zootaxa.5062.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5808035" ID-ZooBank="2B468045-816B-4F29-AE0D-BDC63C503265" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636374593637" checkinUser="plazi" docAuthor="Alipanah, Helen, Yakovlev, Roman, Falsafi, Hossein, Witt, Thomas &amp; Saldaitis, Aidas" docDate="2021" docId="9A3E9F2B6508FFA4FF07FACFFB4055F2" docLanguage="en" docName="zootaxa.5062.1.1.pdf" docOrigin="Zootaxa 5062 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5062.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Cossulus kopetdaghensis Yakovlev &amp; Alipanah, sp. n." docType="treatment" docVersion="3" lastPageNumber="38" masterDocId="6607E753652CFF81FF90FFFFFFCB5423" masterDocTitle="5062" masterLastPageNumber="100" masterPageNumber="1" pageNumber="37" updateTime="1699421341510" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:namePart id="9E1B76160BE6BE007B1DE042DBC6D5D7">Alipanah, Helen</mods:namePart>
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<mods:namePart id="0D37C1E36C0F08F8EDB54429236E1C65">Yakovlev, Roman</mods:namePart>
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<mods:namePart id="8FECB947DB5F2F93F91B33944955E8E7">Falsafi, Hossein</mods:namePart>
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<mods:namePart id="9A6D14E22D6653ADC33AA26E3A96FE4B">Witt, Thomas</mods:namePart>
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<mods:namePart id="F4D8011119E4D33E83DAB16A7AA2C686">Saldaitis, Aidas</mods:namePart>
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<treatment id="9A3E9F2B6508FFA4FF07FACFFB4055F2" ID-DOI="http://doi.org/10.5281/zenodo.5808145" ID-Zenodo-Dep="5808145" LSID="urn:lsid:plazi:treatment:9A3E9F2B6508FFA4FF07FACFFB4055F2" httpUri="http://treatment.plazi.org/id/9A3E9F2B6508FFA4FF07FACFFB4055F2" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<subSubSection id="5A8D7DB66508FFA5FF07FACFFCE25168" box="[151,809,1328,1355]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="12282E3D6508FFA5FF07FACFFCE25168" blockId="36.[151,809,1328,1391]" box="[151,809,1328,1355]" pageId="36" pageNumber="37">
<heading id="496099516508FFA5FF07FACFFCE25168" bold="true" box="[151,809,1328,1355]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<emphasis id="20E3F22F6508FFA5FF07FACFFCE25168" bold="true" box="[151,809,1328,1355]" pageId="36" pageNumber="37">
<taxonomicName id="D59755BE6508FFA5FF07FACFFD115169" ID-CoL="B8QM2" authority="Yakovlev &amp; Alipanah" authorityName="Yakovlev &amp; Alipanah" box="[151,730,1328,1355]" class="Insecta" family="Cossidae" genus="Cossulus" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kopetdaghensis" status="sp. nov.">
<emphasis id="20E3F22F6508FFA5FF07FACFFE0D5168" bold="true" box="[151,454,1328,1355]" italics="true" pageId="36" pageNumber="37">Cossulus kopetdaghensis</emphasis>
Yakovlev &amp; Alipanah
</taxonomicName>
,
<taxonomicNameLabel id="3BD04F546508FFA5FD77FACEFCE25168" box="[743,809,1329,1355]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5A8D7DB66508FFA5FF07FAABFEF2514C" box="[151,313,1364,1391]" pageId="36" pageNumber="37" type="description">
<paragraph id="12282E3D6508FFA5FF07FAABFEF2514C" blockId="36.[151,809,1328,1391]" box="[151,313,1364,1391]" pageId="36" pageNumber="37">
(
<figureCitation id="8AAC32B86508FFA5FF0FFAABFEFB514C" box="[159,304,1364,1391]" captionStart="FIGURE 12" captionStartId="41.[151,250,738,762]" captionTargetBox="[151,1436,181,714]" captionTargetId="figure-18@41.[151,1436,181,714]" captionTargetPageId="41" captionText="FIGURE 12. Cossulus kopetdaghensis Yakovlev &amp; Alipanah, sp. n., holotype (scale bar= 10 mm). A) Adult male, upperside. B) Male genitalia, main body and phallus." figureDoi="http://doi.org/10.5281/zenodo.5808071" httpUri="https://zenodo.org/record/5808071/files/figure.png" pageId="36" pageNumber="37">Figs. 12A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5A8D7DB66508FFA5FF07FA63FF3651DD" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="12282E3D6508FFA5FF07FA63FF3651DD" blockId="36.[151,1437,1436,2039]" pageId="36" pageNumber="37">
<emphasis id="20E3F22F6508FFA5FF07FA63FEB45195" bold="true" box="[151,383,1436,1462]" pageId="36" pageNumber="37">Material examined.</emphasis>
<materialsCitation id="A2FF24606508FFA5FE15FA62FF3651DD" collectingDate="2000-06-27" collectingDateMax="2000-06-28" collectingDateMin="2000-06-27" collectionCode="MWM" collectorName="Gazko &amp; No." country="Iran" county="Kopet Dagh" location="Barjovein Mt." municipality="Bargovein Mt." pageId="36" pageNumber="37" specimenCode="MWM 34.604" specimenCount="4" specimenCount-male="4" stateProvince="Khorasan-e Razavi" typeStatus="holotype">
<emphasis id="20E3F22F6508FFA5FE15FA62FE145194" bold="true" box="[389,479,1437,1463]" pageId="36" pageNumber="37">
<collectionCode id="7486B6F86508FFA5FE15FA62FE115194" box="[389,474,1437,1463]" pageId="36" pageNumber="37">MWM</collectionCode>
:
</emphasis>
<typeStatus id="CD2C909F6508FFA5FE75FA63FD9B5195" box="[485,592,1436,1462]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
,
<specimenCount id="0491E5B46508FFA5FDC9FA62FD4F5194" box="[601,644,1437,1463]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
,
<collectingCountry id="6A806EAD6508FFA5FD1FFA62FD745194" box="[655,703,1437,1463]" name="Iran" pageId="36" pageNumber="37">Iran</collectingCountry>
,
<collectingRegion id="D053E0DF6508FFA5FD59FA63FC075194" box="[713,972,1436,1463]" country="Iran" name="Khorasan-e Razavi" pageId="36" pageNumber="37">Razavi Khorâsân Prov.</collectingRegion>
:
<collectingCounty id="FB4956B16508FFA5FC44FA62FB955195" box="[980,1118,1436,1463]" pageId="36" pageNumber="37">Kopet Dagh</collectingCounty>
,
<collectingMunicipality id="F24CB4476508FFA5FBF8FA63FAC15194" box="[1128,1290,1436,1463]" pageId="36" pageNumber="37">Bargovein Mt.</collectingMunicipality>
(=
<location id="174878E66508FFA5FABEFA63FF0951F8" LSID="urn:lsid:plazi:treatment:9A3E9F2B6508FFA4FF07FACFFB4055F2:174878E66508FFA5FABEFA63FF0951F8" country="Iran" county="Kopet Dagh" municipality="Bargovein Mt." name="Barjovein Mt." pageId="36" pageNumber="37" stateProvince="Khorasan-e Razavi">Barjovein Mt.</location>
),
<location id="174878E66508FFA5FF47FA3FFE5A51F9" LSID="urn:lsid:plazi:treatment:9A3E9F2B6508FFA4FF07FACFFB4055F2:174878E66508FFA5FF47FA3FFE5A51F9" box="[215,401,1472,1498]" country="Iran" county="Kopet Dagh" municipality="Bargovein Mt." name="Shahr-e Kohneh" pageId="36" pageNumber="37" stateProvince="Khorasan-e Razavi">Shahr-e Kohneh</location>
,
<date id="662908FD6508FFA5FE0CFA3FFD8351F9" box="[412,584,1472,1498]" pageId="36" pageNumber="37" value="2000-06-27" valueMax="2000-06-28" valueMin="2000-06-27">
<collectingDate id="766DF1156508FFA5FE0CFA3FFD8351F9" box="[412,584,1472,1498]" pageId="36" pageNumber="37" value="2000-06-27" valueMax="2000-06-28" valueMin="2000-06-27">27.–28.vi.2000</collectingDate>
</date>
, leg.
<collectorName id="BF624BEB6508FFA5FD14FA3FFD0651F9" box="[644,717,1472,1498]" pageId="36" pageNumber="37">Gazkó</collectorName>
(gen. prep.
<collectorName id="BF624BEB6508FFA5FCC5FA3EFCB451F8" box="[853,895,1473,1499]" pageId="36" pageNumber="37">No.</collectorName>
<specimenCode id="423186466508FFA5FC16FA3EFBE751F8" box="[902,1068,1472,1499]" collectionCode="MWM" pageId="36" pageNumber="37">MWM 34.604</specimenCode>
);
<typeStatus id="CD2C909F6508FFA5FBAFFA3EFB6651F8" box="[1087,1197,1473,1499]" pageId="36" pageNumber="37" type="paratype">paratypes</typeStatus>
,
<specimenCount id="0491E5B46508FFA5FB29FA3EFB3051F8" box="[1209,1275,1473,1499]" pageId="36" pageNumber="37" type="male">3 ♂♂</specimenCount>
, same locality and data.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5A8D7DB66508FFA5FF57F9F7FDEF52AC" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="12282E3D6508FFA5FF57F9F7FDEF52AC" blockId="36.[151,1437,1436,2039]" pageId="36" pageNumber="37">
<emphasis id="20E3F22F6508FFA5FF57F9F7FE895201" bold="true" box="[199,322,1544,1570]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
From all the studied species of
<taxonomicName id="D59755BE6508FFA5FD30F9F7FCCE5201" authorityName="Staudinger" authorityYear="1887" box="[672,773,1544,1570]" class="Insecta" family="Cossidae" genus="Cossulus" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="20E3F22F6508FFA5FD30F9F7FCCE5201" box="[672,773,1544,1570]" italics="true" pageId="36" pageNumber="37">Cossulus</emphasis>
</taxonomicName>
, which was recently revised by
<bibRefCitation id="760653CC6508FFA5FBF6F9F7FA495200" author="Yakovlev, R. V. &amp; Witt, Th." box="[1126,1410,1544,1571]" pageId="36" pageNumber="37" pagination="1" refId="ref82632" refString="Yakovlev, R. V. &amp; Witt, Th. (2017 a) World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa, 4311 (1), 062 - 080. https: // doi. org / 10.11646 / zootaxa. 4311.1.4" type="journal article" year="2017">
<taxonomicName id="D59755BE6508FFA5FBF6F9F7FB1B5201" box="[1126,1232,1544,1570]" class="Insecta" family="Cossidae" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="99" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Yakovlev">Yakovlev</taxonomicName>
&amp; Witt (2017a)
</bibRefCitation>
, it externally differs well by the specific reticulated pattern of the forewing and the pale brown portion on its surface in the center (
<figureCitation id="8AAC32B86508FFA5FE83F9AFFEB05248" box="[275,379,1616,1643]" captionStart="FIGURE 12" captionStartId="41.[151,250,738,762]" captionTargetBox="[151,1436,181,714]" captionTargetId="figure-18@41.[151,1436,181,714]" captionTargetPageId="41" captionText="FIGURE 12. Cossulus kopetdaghensis Yakovlev &amp; Alipanah, sp. n., holotype (scale bar= 10 mm). A) Adult male, upperside. B) Male genitalia, main body and phallus." figureDoi="http://doi.org/10.5281/zenodo.5808071" httpUri="https://zenodo.org/record/5808071/files/figure.png" pageId="36" pageNumber="37">Fig. 12A</figureCitation>
). Additionally, the newly described species is differentiated from the other species of the genus by having a short uncus (
<figureCitation id="8AAC32B86508FFA5FE20F98BFDDC52AC" box="[432,535,1652,1679]" captionStart="FIGURE 12" captionStartId="41.[151,250,738,762]" captionTargetBox="[151,1436,181,714]" captionTargetId="figure-18@41.[151,1436,181,714]" captionTargetPageId="41" captionText="FIGURE 12. Cossulus kopetdaghensis Yakovlev &amp; Alipanah, sp. n., holotype (scale bar= 10 mm). A) Adult male, upperside. B) Male genitalia, main body and phallus." figureDoi="http://doi.org/10.5281/zenodo.5808071" httpUri="https://zenodo.org/record/5808071/files/figure.png" pageId="36" pageNumber="37">Fig. 12B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5A8D7DB66508FFA4FF57F967FE5A5546" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="12282E3D6508FFA5FF57F967FAEE52D9" blockId="36.[151,1437,1436,2039]" pageId="36" pageNumber="37">
<emphasis id="20E3F22F6508FFA5FF57F967FE925291" bold="true" box="[199,345,1688,1714]" pageId="36" pageNumber="37">Description.</emphasis>
Male (
<figureCitation id="8AAC32B86508FFA5FE3BF967FDDE5290" box="[427,533,1688,1715]" captionStart="FIGURE 12" captionStartId="41.[151,250,738,762]" captionTargetBox="[151,1436,181,714]" captionTargetId="figure-18@41.[151,1436,181,714]" captionTargetPageId="41" captionText="FIGURE 12. Cossulus kopetdaghensis Yakovlev &amp; Alipanah, sp. n., holotype (scale bar= 10 mm). A) Adult male, upperside. B) Male genitalia, main body and phallus." figureDoi="http://doi.org/10.5281/zenodo.5808071" httpUri="https://zenodo.org/record/5808071/files/figure.png" pageId="36" pageNumber="37">Fig. 12A</figureCitation>
).
<emphasis id="20E3F22F6508FFA5FDBAF967FDAC5291" box="[554,615,1688,1714]" italics="true" pageId="36" pageNumber="37">Head</emphasis>
: Vertex and frons with yellow scales; labial palpus covered with similar scales, with length slightly more than the half length of horizontal diameter of compound eye; antenna unipectinate, with length 0.4 times the length of forewing, pecten process shorter than the antennal segment diameter.
</paragraph>
<paragraph id="12282E3D6508FFA5FF57F8FBFBFF538D" blockId="36.[151,1437,1436,2039]" pageId="36" pageNumber="37">
<emphasis id="20E3F22F6508FFA5FF57F8FBFED3533D" box="[199,280,1796,1822]" italics="true" pageId="36" pageNumber="37">Thorax</emphasis>
: Densely covered with yellow-brown scales; tegula and patagium yellow-brown. Wingspan
<quantity id="D56F83D86508FFA5FAB4F8FBFA56533C" box="[1316,1437,1796,1823]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="36" pageNumber="37" unit="mm" value="37.0" valueMax="39.0" valueMin="35.0">35–39 mm</quantity>
(n = 4). Forewing (
<figureCitation id="8AAC32B86508FFA5FEFDF8D7FE1E5360" box="[365,469,1832,1859]" captionStart="FIGURE 12" captionStartId="41.[151,250,738,762]" captionTargetBox="[151,1436,181,714]" captionTargetId="figure-18@41.[151,1436,181,714]" captionTargetPageId="41" captionText="FIGURE 12. Cossulus kopetdaghensis Yakovlev &amp; Alipanah, sp. n., holotype (scale bar= 10 mm). A) Adult male, upperside. B) Male genitalia, main body and phallus." figureDoi="http://doi.org/10.5281/zenodo.5808071" httpUri="https://zenodo.org/record/5808071/files/figure.png" pageId="36" pageNumber="37">Fig. 12A</figureCitation>
). Elongate, with obliquely rounded termen, apex rounded; upperside yellowish-brown with pale brown streakes along the costal margin, pale brown pattern of thin streakes throughout the wing area, and a pale brown portion from discal area to the apex, fringes mottled (pale brown at veins and yellow between the veins). Hindwing upperside pale brown, without pattern, fringes pale yellow, unicolorous.
</paragraph>
<paragraph id="12282E3D6508FFA4FF57F847FC475562" blockId="36.[151,1437,1436,2039]" lastBlockId="37.[151,1437,151,465]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<emphasis id="20E3F22F6508FFA5FF57F847FEFA53F1" box="[199,305,1976,2002]" italics="true" pageId="36" pageNumber="37">Abdomen</emphasis>
: Densely covered with yellow-brown scales. Genitalia (
<figureCitation id="8AAC32B86508FFA5FC3EF847FBD853F0" box="[942,1043,1976,2003]" captionStart="FIGURE 12" captionStartId="41.[151,250,738,762]" captionTargetBox="[151,1436,181,714]" captionTargetId="figure-18@41.[151,1436,181,714]" captionTargetPageId="41" captionText="FIGURE 12. Cossulus kopetdaghensis Yakovlev &amp; Alipanah, sp. n., holotype (scale bar= 10 mm). A) Adult male, upperside. B) Male genitalia, main body and phallus." figureDoi="http://doi.org/10.5281/zenodo.5808071" httpUri="https://zenodo.org/record/5808071/files/figure.png" pageId="36" pageNumber="37">Fig. 12B</figureCitation>
) typical for the genus in structure, with phallus being thin, almost straight, equal to the length of valva, having the same width along its length. Bulbus ejaculatorius positioned dorso-apically, length of its entrance one-third of the length of phallus, vesica without cornuti; saccus developed; juxta thin, with long lateral processes (equal to the transtilla processes) diluted at right angle; transtilla process bent uncinately, with sharp apex; valva sclerotized basally, membranous distally, with lanceolate apex and poorly developed crest on costal margin; gnathos robust, densely covered with small spines, gnathos arms long and thick; uncus short, wide, with rounded apex.
</paragraph>
<paragraph id="12282E3D6509FFA4FF57FEB4FE5A5546" blockId="37.[151,1437,151,465]" box="[199,401,331,357]" pageId="37" pageNumber="38">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="5A8D7DB66509FFA4FF57FE91FD6B558E" pageId="37" pageNumber="38" type="distribution">
<paragraph id="12282E3D6509FFA4FF57FE91FE5755AA" blockId="37.[151,1437,151,465]" box="[199,412,366,393]" pageId="37" pageNumber="38">
<emphasis id="20E3F22F6509FFA4FF57FE91FEAA55AB" bold="true" box="[199,353,366,392]" pageId="37" pageNumber="38">Distribution.</emphasis>
<collectingCountry id="6A806EAD6509FFA4FEF7FE90FE5355AA" box="[359,408,367,393]" name="Iran" pageId="37" pageNumber="38">Iran</collectingCountry>
.
</paragraph>
<paragraph id="12282E3D6509FFA4FF57FE6DFD6B558E" blockId="37.[151,1437,151,465]" box="[199,672,402,429]" pageId="37" pageNumber="38">
<emphasis id="20E3F22F6509FFA4FF57FE6DFE70558E" bold="true" box="[199,443,402,429]" pageId="37" pageNumber="38">
Distribution in
<collectingCountry id="6A806EAD6509FFA4FEEFFE6CFE7D558E" box="[383,438,403,429]" name="Iran" pageId="37" pageNumber="38">Iran</collectingCountry>
.
</emphasis>
NE Iran (
<figureCitation id="8AAC32B86509FFA4FDBAFE6CFD58558E" box="[554,659,403,429]" captionStart="FIGURE 29" captionStartId="82.[151,250,1943,1967]" captionTargetBox="[190,1395,183,1916]" captionTargetId="figure-22@82.[189,1399,181,1919]" captionTargetPageId="82" captionText="FIGURE 29. Distribution of Cossidae species in Iran. A) Cossulus herzi (Alpheraky). B) Cossulus intractatus (Staudinger). C) Cossulus irani (Daniel). D) Cossulus kopetdaghensis Yakovlev &amp; Alipanah, sp. n. E) Cossulus lignosus (Brandt). F) Cossulus mollis (Christoph)." figureDoi="http://doi.org/10.5281/zenodo.5808119" httpUri="https://zenodo.org/record/5808119/files/figure.png" pageId="37" pageNumber="38">Fig. 29D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5A8D7DB66509FFA4FF57FE48FB4055F2" box="[199,1163,439,465]" pageId="37" pageNumber="38" type="etymology">
<paragraph id="12282E3D6509FFA4FF57FE48FB4055F2" blockId="37.[151,1437,151,465]" box="[199,1163,439,465]" pageId="37" pageNumber="38">
<emphasis id="20E3F22F6509FFA4FF57FE48FE8555F2" bold="true" box="[199,334,439,465]" pageId="37" pageNumber="38">Etymology.</emphasis>
This species is named after the collecting area in Kopet Dagh Mountains.
</paragraph>
</subSubSection>
</treatment>
</document>
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