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<document id="F50377C917E0760579A1EAA9F19F1A21" ID-DOI="10.5281/zenodo.293925" ID-GBIF-Dataset="2d814db2-7388-4590-9367-abc3389321cb" ID-ISSN="1175-5326" ID-Zenodo-Dep="293925" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460195177616" checkinUser="plazi" docAuthor="Jungfer, Karl-Heinz" docDate="2010" docId="6E478797FFA9FF88B0B6F8B4FDBB30EF" docLanguage="en" docName="zt02407p050.pdf" docOrigin="Zootaxa 2407" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Osteocephalus inframaculatus Boulenger 1882" docType="treatment" docVersion="8" lastPageNumber="38" masterDocId="927EFFEFFFA1FF83B021FFE7F8353013" masterDocTitle="The taxonomic status of some spiny-backed treefrogs, genus Osteocephalus (Amphibia: Anura: Hylidae)" masterLastPageNumber="50" masterPageNumber="28" pageNumber="36" updateTime="1698236544781" updateUser="plazi">
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<mods:title id="00FCD38DD547C8506B1FB2F19BD5D9D7">The taxonomic status of some spiny-backed treefrogs, genus Osteocephalus (Amphibia: Anura: Hylidae)</mods:title>
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<mods:namePart id="E7ABF630440D80C68815CFD2B80EEB1E">Jungfer, Karl-Heinz</mods:namePart>
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<mods:date id="1B97924117659495B621AE32D4EC8A25">2010</mods:date>
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<treatment id="6E478797FFA9FF88B0B6F8B4FDBB30EF" ID-DOI="http://doi.org/10.5281/zenodo.5669082" ID-GBIF-Taxon="119383756" ID-Zenodo-Dep="5669082" LSID="urn:lsid:plazi:treatment:6E478797FFA9FF88B0B6F8B4FDBB30EF" httpUri="http://treatment.plazi.org/id/6E478797FFA9FF88B0B6F8B4FDBB30EF" lastPageId="11" lastPageNumber="38" pageId="8" pageNumber="36">
<subSubSection id="AEF4650AFFA9FF8BB0B6F8B4F8DB3781" pageId="8" pageNumber="36" type="nomenclature">
<paragraph id="E6513681FFA9FF8BB0B6F8B4FB42377F" blockId="8.[151,887,1874,1938]" box="[151,887,1874,1901]" pageId="8" pageNumber="36">
<heading id="BD1981EDFFA9FF8BB0B6F8B4FB42377F" bold="true" box="[151,887,1874,1901]" fontSize="11" level="1" pageId="8" pageNumber="36" reason="1">
<emphasis id="D49AEA93FFA9FF8BB0B6F8B4FB42377F" bold="true" box="[151,887,1874,1901]" pageId="8" pageNumber="36">
<taxonomicName id="21EE4D02FFA9FF8BB0B6F8B4FADC377F" ID-CoL="6TBN5" authority="Boulenger, 1882" authorityName="Boulenger" authorityYear="1882" box="[151,745,1874,1901]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="36" phylum="Chordata" rank="species" species="inframaculatus">
<emphasis id="D49AEA93FFA9FF8BB0B6F8B4FA3D377E" bold="true" box="[151,520,1875,1901]" italics="true" pageId="8" pageNumber="36">Osteocephalus inframaculatus</emphasis>
(
<bibRefCitation id="827F4B70FFA9FF8BB237F8B5FAD4377F" author="Boulenger" box="[534,737,1874,1900]" pageId="8" pageNumber="47" refString="Boulenger, G. A. (1882) Catalogue of the Batrachia Salientia s. Ecaudata in the collection of the British Museum. London, 503 pp." type="book" year="1882">Boulenger, 1882</bibRefCitation>
)
</taxonomicName>
nov. comb.
</emphasis>
</heading>
</paragraph>
<paragraph id="E6513681FFA9FF8BB0B6F89FF8DB3781" blockId="8.[151,887,1874,1938]" box="[151,238,1912,1938]" pageId="8" pageNumber="36">
(
<figureCitation id="7ED52A04FFA9FF8BB0BEF89FF8D03781" box="[159,229,1912,1938]" captionStart="FIGURE 5" captionStartId="9.[151,255,1962,1986]" captionTargetBox="[307,1267,298,1916]" captionTargetId="figure@9.[305,1268,293,1927]" captionTargetPageId="9" captionText="FIGURE 5. Osteocephalus inframaculatus (holotype): a. dorsolateral, b. ventral view, c. posterior surface of thigh and cloacal area. Not to scale." httpUri="https://zenodo.org/record/293930/files/figure.png" pageId="8" pageNumber="36">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="AEF4650AFFA9FF89B0B6F824FCD031CF" lastPageId="10" lastPageNumber="38" pageId="8" pageNumber="36" type="reference_group">
<paragraph id="E6513681FFA9FF8AB0B6F824F9C030FC" blockId="8.[151,543,1987,2010]" lastBlockId="9.[151,552,152,239]" lastPageId="9" lastPageNumber="37" pageId="8" pageNumber="36">
<taxonomicName id="21EE4D02FFA9FF8BB0B6F824FA2A37C9" ID-CoL="3NCX3" authority="Boulenger, 1882" authorityName="Boulenger" authorityYear="1882" box="[151,543,1987,2010]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="8" pageNumber="36" phylum="Chordata" rank="species" species="inframaculata">
<emphasis id="D49AEA93FFA9FF8BB0B6F824F95D37C9" box="[151,360,1987,2010]" italics="true" pageId="8" pageNumber="36">Hyla inframaculata</emphasis>
<bibRefCitation id="827F4B70FFA9FF8BB14EF824FA2A37C9" author="Boulenger" box="[367,543,1987,2010]" pageId="8" pageNumber="47" refString="Boulenger, G. A. (1882) Catalogue of the Batrachia Salientia s. Ecaudata in the collection of the British Museum. London, 503 pp." type="book" year="1882">Boulenger, 1882</bibRefCitation>
</taxonomicName>
<taxonomicName id="21EE4D02FFA8FF8AB0B6FF7FF95D30BC" ID-CoL="3NCX3" box="[151,360,152,175]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="37" phylum="Chordata" rank="species" species="inframaculata">
<emphasis id="D49AEA93FFA8FF8AB0B6FF7FF95D30BC" box="[151,360,152,175]" italics="true" pageId="9" pageNumber="37">Hyla inframaculata</emphasis>
</taxonomicName>
<bibRefCitation id="827F4B70FFA8FF8AB1A2FF7FFA3930BC" author="Nieden" box="[387,524,152,175]" pageId="9" pageNumber="48" refString="Nieden, F. (1923) Das Tierreich. Anura I. de Gruyter, Berlin and Leipzig, 584 pp." type="book" year="1923">Nieden 1923</bibRefCitation>
<taxonomicName id="21EE4D02FFA8FF8AB0B6FF5FF95D30DC" ID-CoL="3NCX3" box="[151,360,184,207]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="37" phylum="Chordata" rank="species" species="inframaculata">
<emphasis id="D49AEA93FFA8FF8AB0B6FF5FF95D30DC" box="[151,360,184,207]" italics="true" pageId="9" pageNumber="37">Hyla inframaculata</emphasis>
</taxonomicName>
<bibRefCitation id="827F4B70FFA8FF8AB1A2FF5FFA1D30DC" author="Duellman" box="[387,552,184,207]" pageId="9" pageNumber="47" refString="Duellman, W. E. (1977) Liste der rezenten Amphibien und Reptilien. Hylidae, Centrolenidae, Pseudidae. Das Tierreich, 95, 1 - 225." type="journal article" year="1977">Duellman 1977</bibRefCitation>
<taxonomicName id="21EE4D02FFA8FF8AB0B6FF3FF95D30FC" ID-CoL="3NCX3" box="[151,360,216,239]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="37" phylum="Chordata" rank="species" species="inframaculata">
<emphasis id="D49AEA93FFA8FF8AB0B6FF3FF95D30FC" box="[151,360,216,239]" italics="true" pageId="9" pageNumber="37">Hyla inframaculata</emphasis>
</taxonomicName>
<bibRefCitation id="827F4B70FFA8FF8AB1A2FF3FF9C030FC" author="Frost" box="[387,501,216,239]" pageId="9" pageNumber="47" refString="Frost, D. R. (2009) Amphibian Species of the World: an Online Reference. Version 5.3 (12 February, 2009). American Museum of Natural History, New York, USA. Electronic Database accessible at http: // research. amnh. org / herpetology / amphibia /" type="journal article" year="2009">Frost 2009</bibRefCitation>
</paragraph>
<caption id="B2916609FFA8FF8AB0B6F84DF99C37F5" httpUri="https://zenodo.org/record/293930/files/figure.png" pageId="9" pageNumber="37" targetBox="[307,1267,298,1916]" targetPageId="9">
<paragraph id="E6513681FFA8FF8AB0B6F84DF99C37F5" blockId="9.[151,1436,1962,2022]" pageId="9" pageNumber="37">
<emphasis id="D49AEA93FFA8FF8AB0B6F84DF92E37D1" bold="true" box="[151,283,1962,1986]" pageId="9" pageNumber="37">FIGURE 5.</emphasis>
<taxonomicName id="21EE4D02FFA8FF8AB103F84CFA5C37D1" ID-CoL="6TBN5" box="[290,617,1963,1986]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="37" phylum="Chordata" rank="species" species="inframaculatus">
<emphasis id="D49AEA93FFA8FF8AB103F84CFA5C37D1" box="[290,617,1963,1986]" italics="true" pageId="9" pageNumber="37">Osteocephalus inframaculatus</emphasis>
</taxonomicName>
(holotype): a. dorsolateral, b. ventral view, c. posterior surface of thigh and cloacal area. Not to scale.
</paragraph>
</caption>
<paragraph id="E6513681FFABFF89B0B6FF7FFCD031CF" blockId="10.[151,1437,152,2019]" pageId="10" pageNumber="38">
This species, known only from the
<typeStatus id="39558823FFABFF89B265FF7FFA8430A1" box="[580,689,152,178]" pageId="10" pageNumber="38" type="holotype">holotype</typeStatus>
, BM 1947.2.13.10, has only appeared in species lists after its description. Its
<typeStatus id="39558823FFABFF89B16AFF5AF94830C4" box="[331,381,189,215]" pageId="10" pageNumber="38">type</typeStatus>
locality is “Santarem”, Estado do Para,
<collectingCountry id="9EF97611FFABFF89B373FF5AFBAB30C4" box="[850,926,189,215]" name="Brazil" pageId="10" pageNumber="38">Brazil</collectingCountry>
. Although there are few characters found in species of
<taxonomicName id="21EE4D02FFABFF89B132FF05F9F530E8" ID-CoL="63GQL" box="[275,448,226,251]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="D49AEA93FFABFF89B132FF05F9F530E8" box="[275,448,226,251]" italics="true" pageId="10" pageNumber="38">Osteocephalus</emphasis>
</taxonomicName>
that are not present in other hylid genera (
<bibRefCitation id="827F4B70FFABFF89B395FF05FCFC30EF" author="Jungfer" box="[948,1225,226,252]" pageId="10" pageNumber="47" refString="Jungfer, K. - H. &amp; Hodl, W. (2002) A new species of Osteocephalus from Ecuador and a redescription of O. leprieurii (Dumeril &amp; Bibron, 1841) (Anura: Hylidae). Amphibia-Reptilia, 23, 21 - 46." type="journal article" year="2002">Jungfer and Hödl 2002</bibRefCitation>
), the combination of angular vomers, a bifid ultimate subarticular tubercle of Finger IV and tuberculate dorsal skin characterize
<taxonomicName id="21EE4D02FFABFF89B0B6FECAF94B3155" ID-CoL="3NCX3" box="[151,382,301,326]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="inframaculata">
<emphasis id="D49AEA93FFABFF89B0B6FECAF94B3155" box="[151,382,301,326]" italics="true" pageId="10" pageNumber="38">Hyla inframaculata</emphasis>
</taxonomicName>
as a species of
<taxonomicName id="21EE4D02FFABFF89B216FECAFAD63155" ID-CoL="63GQL" box="[567,739,301,326]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="D49AEA93FFABFF89B216FECAFAD63155" box="[567,739,301,326]" italics="true" pageId="10" pageNumber="38">Osteocephalus</emphasis>
</taxonomicName>
. The tuberculate dorsal skin in a female specimen together with areolate skin on the flanks, non-uniform coloration of the posterior surfaces of the thighs and a row of tubercles on the tarsus indicate that the species is a member of the
<taxonomicName id="21EE4D02FFABFF89B3D7FE9FFC553182" ID-CoL="6TBMM" box="[1014,1120,376,401]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="buckleyi">
<emphasis id="D49AEA93FFABFF89B3D7FE9FFC553182" box="[1014,1120,376,401]" italics="true" pageId="10" pageNumber="38">buckleyi</emphasis>
</taxonomicName>
complex. With only one undissected specimen at hand, neither sex, identified by lack of nuptial excrescences, lack of vocal sac apertures and externally visible vocal sacs, nor adult size can be taken for completely granted.
</paragraph>
</subSubSection>
<subSubSection id="AEF4650AFFABFF88B0E7FE00FDBB30EF" lastPageId="11" lastPageNumber="39" pageId="10" pageNumber="38" type="description">
<paragraph id="E6513681FFABFF89B0E7FE00F9F8333F" blockId="10.[151,1437,152,2019]" pageId="10" pageNumber="38">
<emphasis id="D49AEA93FFABFF89B0E7FE00FA763212" bold="true" box="[198,579,487,513]" pageId="10" pageNumber="38">
Redescription of the
<typeStatus id="39558823FFABFF89B1EFFE00FA0B3212" box="[462,574,487,513]" pageId="10" pageNumber="38" type="holotype">holotype</typeStatus>
:
</emphasis>
Snout rounded in dorsal and lateral view, as long as the diameter of the orbit. Head slightly wider than long, without raised lateral edges of the frontoparietals. Canthus rostralis rounded, indistinct; loreal region slightly concave; interorbital space as broad as the upper eyelid. Dorsal surfaces of head and canthus rostralis covered with small, irregular tubercles; larger tubercles on the orbits. Tympanum distinct, large, about 70% of the diameter of the eye. A heavy tuberculate supratympanic fold from the midlevel of the eye to a point at about 2 h (when tympanum is thought as a clock face), sloping towards the flanks to a point at about midlevel of the tympanum. Tongue circular, vomers angular, almost between the large choanae. Anteriorly, the choanae reach slightly beyond the anterior parts of the vomers, posteriorly it is vice versa.
</paragraph>
<paragraph id="E6513681FFABFF89B0E5FCDFFCCB341F" blockId="10.[151,1437,152,2019]" pageId="10" pageNumber="38">
Dorsum tuberculate. Two dorsal folds not mentioned in the original description, but shown in Plate XXIII (
<bibRefCitation id="827F4B70FFABFF89B0BEFCBAF9683364" author="Boulenger" box="[159,349,861,887]" pageId="10" pageNumber="47" refString="Boulenger, G. A. (1882) Catalogue of the Batrachia Salientia s. Ecaudata in the collection of the British Museum. London, 503 pp." type="book" year="1882">Boulenger 1882</bibRefCitation>
) and still present, are irregular and obviously artefacts of preservation. Anterior two thirds of the flanks areolate. Ventrally granulate. Cloacal opening at about midlevel of the thighs. Anal sheath absent. Circumanal tubercles present, low. Axillary membrane weak. Upper surfaces of limbs weakly tuberculate. A row of irregular tubercles on the outer ventrolateral edge of the lower arm and the tarsus. Hidden surfaces of limbs smooth. When bent forward, the tibiotarsal articulation reaches nearly the tip of the snout.
</paragraph>
<paragraph id="E6513681FFABFF89B0E5FBFFFC9034C3" blockId="10.[151,1437,152,2019]" pageId="10" pageNumber="38">
Finger discs rounded. Disc on Finger III about 70% the diameter of the tympanum. The relative lengths of the adpressed fingers is I &lt;II &lt;IV &lt;III. Finger webbing formula is I basal
<date id="92501041FFABFF89B41BFBA7FC533449" box="[1082,1126,1088,1114]" pageId="10" pageNumber="38">II 1</date>
½ — 2 2/3
<date id="92501041FFABFF89B4CCFBA7FD163449" box="[1261,1315,1088,1114]" pageId="10" pageNumber="38">III 3</date>
— 2+ IV. Webbing on the inner edge of Finger III reaches halfway between penultimate and ultimate subarticular tubercle. The distal subarticular tubercle on Finger IV is bifid. The relative length of adpressed toes is I &lt;II &lt;III ≤ V &lt;IV. The webbing formula is
<date id="92501041FFABFF89B270FB51FA4434C3" box="[593,625,1206,1232]" pageId="10" pageNumber="38">I 1</date>
+ — 2
<date id="92501041FFABFF89B299FB51FAD734C3" box="[696,738,1206,1232]" pageId="10" pageNumber="38">II 1</date>
+ — 2 ½
<date id="92501041FFABFF89B366FB51FB4F34C3" box="[839,890,1206,1232]" pageId="10" pageNumber="38">III 1</date>
1/3 — 2 1/3
<date id="92501041FFABFF89B3D9FB51FC1834C3" box="[1016,1069,1206,1232]" pageId="10" pageNumber="38">IV 2</date>
½ —
<date id="92501041FFABFF89B457FB51FC9034C3" box="[1142,1189,1206,1232]" pageId="10" pageNumber="38">1 V.</date>
</paragraph>
<paragraph id="E6513681FFABFF89B0E5FB39FA39350E" blockId="10.[151,1437,152,2019]" pageId="10" pageNumber="38">
Measurements of the
<typeStatus id="39558823FFABFF89B1E1FB39FA1234EB" box="[448,551,1246,1272]" pageId="10" pageNumber="38" type="holotype">holotype</typeStatus>
(in mm): SVL 41.9; HL 14.2; HW 14.5; TL 22.4; FL 27.6; ED 5.2; TD 3.7; FD 2.2; EN 4.6; IN 3.1; TE 2.2.
</paragraph>
<paragraph id="E6513681FFABFF89B0E5FACEF9D735A0" blockId="10.[151,1437,152,2019]" pageId="10" pageNumber="38">Coloration: Dorsally grayish brown, indistinctly marbled with dark brown; Flanks marbled white and brown; throat and chest with bold brown blotches on cream ground; posterior half of abdomen and ventral surfaces of thighs light brown; hidden surfaces of arms and legs boldly marbled with brown and cream. The tibiofibular bones are white.</paragraph>
<paragraph id="E6513681FFABFF89B0E5FA59FD4E36AB" blockId="10.[151,1437,152,2019]" pageId="10" pageNumber="38">
<emphasis id="D49AEA93FFABFF89B0E5FA59F94C35CB" bold="true" box="[196,377,1470,1496]" pageId="10" pageNumber="38">Comparisons:</emphasis>
The only known female is much smaller than both female
<taxonomicName id="21EE4D02FFABFF89B46DFA59FC8F35C4" box="[1100,1210,1470,1495]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="festae">
<emphasis id="D49AEA93FFABFF89B46DFA59FC8F35C4" box="[1100,1210,1470,1495]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B46DFA59FC5035C4" box="[1100,1125,1470,1495]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. festae
</emphasis>
</taxonomicName>
(
<quantity id="21169B64FFABFF89B4EBFA59FD0E35CB" box="[1226,1339,1470,1496]" metricMagnitude="-2" metricUnit="m" metricValue="7.89" pageId="10" pageNumber="38" unit="mm" value="78.9">78.9 mm</quantity>
) and
<taxonomicName id="21EE4D02FFABFF89B55EFA59F8CE35EE" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="cabrerai">
<emphasis id="D49AEA93FFABFF89B55EFA59F8CE35EE" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B55EFA59FDAD35C4" box="[1407,1432,1470,1495]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. cabrerai
</emphasis>
</taxonomicName>
(
<quantity id="21169B64FFABFF89B12BFA04F94D35EE" box="[266,376,1507,1533]" metricMagnitude="-2" metricUnit="m" metricValue="7.18" pageId="10" pageNumber="38" unit="mm" value="71.8">71.8 mm</quantity>
), and smaller than female
<taxonomicName id="21EE4D02FFABFF89B28CFA03FB0635EE" box="[685,819,1508,1533]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="buckleyi">
<emphasis id="D49AEA93FFABFF89B28CFA03FB0635EE" box="[685,819,1508,1533]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B28CFA03FAF035EE" box="[685,709,1508,1533]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. buckleyi
</emphasis>
</taxonomicName>
(
<quantity id="21169B64FFABFF89B362FA04FB8735EE" box="[835,946,1507,1533]" metricMagnitude="-2" metricUnit="m" metricValue="5.41" pageId="10" pageNumber="38" unit="mm" value="54.1">54.1 mm</quantity>
). Other differences to
<taxonomicName id="21EE4D02FFABFF89B497FA03FD0B35EE" box="[1206,1342,1508,1533]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="cabrerai">
<emphasis id="D49AEA93FFABFF89B497FA03FD0B35EE" box="[1206,1342,1508,1533]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B497FA03FCFB35EE" box="[1206,1230,1508,1533]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. cabrerai
</emphasis>
</taxonomicName>
include the outer edge of Finger IV, which is tuberculate in its proximal half and smooth in the distal half (with an irregular fringe in
<taxonomicName id="21EE4D02FFABFF89B151F9C9F9CC3654" box="[368,505,1582,1607]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="cabrerai">
<emphasis id="D49AEA93FFABFF89B151F9C9F9CC3654" box="[368,505,1582,1607]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B151F9C9F9BD3654" box="[368,392,1582,1607]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. cabrerai
</emphasis>
</taxonomicName>
) and there are no tubercles on the lower jaw (present in
<taxonomicName id="21EE4D02FFABFF89B4B4F9C9FD283654" box="[1173,1309,1582,1607]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="cabrerai">
<emphasis id="D49AEA93FFABFF89B4B4F9C9FD283654" box="[1173,1309,1582,1607]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B4B4F9C9FC983654" box="[1173,1197,1582,1607]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. cabrerai
</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="21EE4D02FFABFF89B0F8F9B3F971367E" box="[217,324,1620,1645]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="festae">
<emphasis id="D49AEA93FFABFF89B0F8F9B3F971367E" box="[217,324,1620,1645]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B0F8F9B3F8C7367E" box="[217,242,1620,1645]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. festae
</emphasis>
</taxonomicName>
in the coloration of the posterior surfaces of thighs in preservative, which are tan with light short vermiculation (uniform brown). It differs from
<taxonomicName id="21EE4D02FFABFF89B322F99EFBBD3681" box="[771,904,1657,1682]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="buckleyi">
<emphasis id="D49AEA93FFABFF89B322F99EFBBD3681" box="[771,904,1657,1682]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B322F99EFB2E3681" box="[771,795,1657,1682]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. buckleyi
</emphasis>
</taxonomicName>
,
<taxonomicName id="21EE4D02FFABFF89B3B6F99EFC2A3681" box="[919,1055,1657,1682]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="cabrerai">
<emphasis id="D49AEA93FFABFF89B3B6F99EFC2A3681" box="[919,1055,1657,1682]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B3B6F99EFB9A3681" box="[919,943,1657,1682]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. cabrerai
</emphasis>
</taxonomicName>
and
<taxonomicName id="21EE4D02FFABFF89B479F99EFCF53681" box="[1112,1216,1657,1682]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="festae">
<emphasis id="D49AEA93FFABFF89B479F99EFCF53681" box="[1112,1216,1657,1682]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B479F99EFC453681" box="[1112,1136,1657,1682]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. festae
</emphasis>
</taxonomicName>
in having a longer snout. The snout is round in dorsal view, while
<taxonomicName id="21EE4D02FFABFF89B2E3F979FB7D36A4" box="[706,840,1694,1719]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="buckleyi">
<emphasis id="D49AEA93FFABFF89B2E3F979FB7D36A4" box="[706,840,1694,1719]" italics="true" pageId="10" pageNumber="38">
<collectingCountry id="9EF97611FFABFF89B2E3F979FAEF36A4" box="[706,730,1694,1719]" name="Iceland" pageId="10" pageNumber="38">O</collectingCountry>
. buckleyi
</emphasis>
</taxonomicName>
usually has a bluntly rounded to truncate snout.
</paragraph>
<paragraph id="E6513681FFABFF88B0E5F923FDBB30EF" blockId="10.[151,1437,152,2019]" lastBlockId="11.[151,1437,152,252]" lastPageId="11" lastPageNumber="39" pageId="10" pageNumber="38">
<taxonomicName id="21EE4D02FFABFF89B0E5F923FA1E36CE" box="[196,555,1732,1757]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="inframaculatus">
<emphasis id="D49AEA93FFABFF89B0E5F923FA1E36CE" box="[196,555,1732,1757]" italics="true" pageId="10" pageNumber="38">Osteocephalus inframaculatus</emphasis>
</taxonomicName>
may be diagnosed as (1) a medium-sized species with sexual dimorphism unknown; (2) skin on dorsum of female tuberculate; (3) anterior two thirds of the flanks areolate; (4) canthus rostralis indistinct, rounded, curved inward; (5) frontoparietal ridges not visible through skin; (6) dentigerous processes of vomers angular; (7) tuberculate supratympanic fold from the midlevel of the eye sloping posterior to the tympanum towards the flanks up to about midlevel of the tympanum; (8) web on inner edge of third finger reaching halfway between penultimate and ultimate subarticular tubercle; (9) distal subarticular tubercle on Finger IV bifid; (10) dorsum grayish brown, indistinctly marbled with dark brown; (11) ventrally anterior half with large brown blotches on cream ground, posterior half light brown; (12) several light markings on the upper lip, the largest one posteroventral to the midlevel of the eye to the tympanum; (13) flanks marbled white and brown; (14) position of vocal sacs unknown; (15) juvenile coloration unknown; (16) tadpole habitat and labial tooth row formula unknown; (17) color of tibiofibular bones white in preservative.
</paragraph>
</subSubSection>
</treatment>
</document>