400 lines
44 KiB
XML
400 lines
44 KiB
XML
<document ID-DOI="10.5281/zenodo.3382461" ID-GBIF-Dataset="fd03f669-c046-46cd-861a-6bf56b7fc989" ID-Handle="http://hdl.handle.net/2246/1099" ID-Zenodo-Dep="3382461" approvalRequired="468" approvalRequired_for_bibRefs="7" approvalRequired_for_document="1" approvalRequired_for_illustrations="16" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="18" approvalRequired_for_textStreams="406" approvalRequired_for_treatments="19" checkinTime="1567239673843" checkinUser="plazi" docAuthor="Sahni, Ashok" docDate="1972" docId="1A7187CFFFDC1765FE8AFCCEE7855DD3" docLanguage="en" docName="B147.6.pdf" docOrigin="Bulletin of the American Museum of Natural History 147 (6)" docStyle="DocumentStyle{}" docTitle="Cimolomys clarki Sahni 1972, new species" docType="treatment" docVersion="22" lastPageNumber="373" masterDocId="E648FFB7FFEA175DFFBAFFBEE2785A7A" masterDocTitle="The vertebrate Fauna of the Judith River formation, Montana" masterLastPageNumber="416" masterPageNumber="319" pageNumber="371" updateTime="1673867241423" updateUser="jeremy">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>The vertebrate Fauna of the Judith River formation, Montana</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Sahni, Ashok</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Bulletin of the American Museum of Natural History</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>1972</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>147</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="issue">
|
|
<mods:number>6</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>319</mods:start>
|
|
<mods:end>416</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">10.5281/zenodo.3382461</mods:identifier>
|
|
<mods:identifier type="GBIF-Dataset">fd03f669-c046-46cd-861a-6bf56b7fc989</mods:identifier>
|
|
<mods:identifier type="Handle">http://hdl.handle.net/2246/1099</mods:identifier>
|
|
<mods:identifier type="Zenodo-Dep">3382461</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4710502" ID-GBIF-Taxon="159397326" ID-Zenodo-Dep="4710502" LSID="urn:lsid:plazi:treatment:1A7187CFFFDC1765FE8AFCCEE7855DD3" httpUri="http://treatment.plazi.org/id/1A7187CFFFDC1765FE8AFCCEE7855DD3" lastPageId="56" lastPageNumber="373" pageId="54" pageNumber="371">
|
|
<subSubSection box="[304,864,879,915]" pageId="54" pageNumber="371" type="nomenclature">
|
|
<paragraph blockId="54.[158,1013,679,2777]" box="[304,864,879,915]" pageId="54" pageNumber="371">
|
|
<taxonomicName authority="Sahni, 1972" authorityName="Sahni" authorityYear="1972" box="[304,647,880,915]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="clarki" status="new species">
|
|
<emphasis bold="true" box="[304,647,880,915]" italics="true" pageId="54" pageNumber="371">Cimolomys clarki</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel box="[667,864,879,914]" pageId="54" pageNumber="371" rank="species">new species</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection box="[478,688,939,973]" pageId="54" pageNumber="371" type="description">
|
|
<paragraph blockId="54.[158,1013,679,2777]" box="[478,688,939,973]" pageId="54" pageNumber="371">Figure11A-J</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="54" pageNumber="371" type="etymology">
|
|
<paragraph blockId="54.[158,1013,679,2777]" pageId="54" pageNumber="371">ETYMOLOGY: For Captain William Clark, pioneer explorer of the Missouri River.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="54" pageNumber="371" type="materials_examined">
|
|
<paragraph blockId="54.[158,1013,679,2777]" box="[205,840,1092,1128]" pageId="54" pageNumber="371">HOLOTYPE: AMNH 77179, right P4.</paragraph>
|
|
<paragraph blockId="54.[158,1013,679,2777]" pageId="54" pageNumber="371">TYPE LOCALITY: Clayball Hill, Judith River Formation, Montana.</paragraph>
|
|
<paragraph blockId="54.[158,1013,679,2777]" pageId="54" pageNumber="371">KNOWN DISTRIBUTION: Chouteau County, Montana.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="54" pageNumber="371" type="diagnosis">
|
|
<paragraph blockId="54.[158,1013,679,2777]" pageId="54" pageNumber="371">
|
|
<emphasis box="[204,386,1324,1359]" italics="true" pageId="54" pageNumber="371">DIAGNOSIS</emphasis>
|
|
:
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[406,665,1323,1358]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[406,569,1323,1358]" italics="true" pageId="54" pageNumber="371">Cimolomys</emphasis>
|
|
clarki
|
|
</taxonomicName>
|
|
is smaller than, but similar to, its Maestrichtian descendant,
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[974,1001,1368,1402]" italics="true" pageId="54" pageNumber="371">C</emphasis>
|
|
.
|
|
<emphasis box="[162,280,1416,1451]" italics="true" pageId="54" pageNumber="371">gracilis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="56" lastPageNumber="373" pageId="54" pageNumber="371" type="description">
|
|
<paragraph blockId="54.[158,1013,679,2777]" pageId="54" pageNumber="371">
|
|
DESCRIPTION: P4; no complete P4 of
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[872,1010,1461,1495]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[872,898,1461,1495]" italics="true" pageId="54" pageNumber="371">C</emphasis>
|
|
.
|
|
<emphasis box="[923,1010,1461,1495]" italics="true" pageId="54" pageNumber="371">clarki</emphasis>
|
|
</taxonomicName>
|
|
has yet been found. A reconstruction of the premolar was made from AMNH 77179 and 77180 (fig. I1 A-D). P4 probably had eight or nine serrations arranged in a nearly symmetric arc. Ridges arising from the anterior two or three serrations are extremely short; those from the middle are well developed and extend anteroventrally to the level of the posterolabial ledge. The posterior two or three serrations are distinct, and are grooved labially. The posterolabial ledge
|
|
<emphasis box="[276,299,1968,2002]" italics="true" pageId="54" pageNumber="371">is</emphasis>
|
|
well developed and indicates the presence of three worn cuspules.
|
|
</paragraph>
|
|
<paragraph blockId="54.[158,1013,679,2777]" pageId="54" pageNumber="371">
|
|
The anterolabial lobe of the crown at the anterior root is poorly formed, as in
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" box="[833,1003,2105,2139]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[833,859,2105,2139]" italics="true" pageId="54" pageNumber="371">C</emphasis>
|
|
.
|
|
<emphasis box="[881,1003,2106,2139]" italics="true" pageId="54" pageNumber="371">gracilis</emphasis>
|
|
</taxonomicName>
|
|
; the anterobasal cavity is small and enamel covered except for the posterior wall. The anterior root is much larger than the posterior, and a low interradicular crest
|
|
<emphasis box="[701,725,2290,2323]" italics="true" pageId="54" pageNumber="371">is</emphasis>
|
|
present.
|
|
</paragraph>
|
|
<paragraph blockId="54.[158,1013,679,2777]" pageId="54" pageNumber="371">
|
|
Ml; this tooth is apparently identical in size and cusp formula with that of
|
|
<taxonomicName class="Mammalia" family="Ptilodontidae" genus="Mesodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="primaevus">
|
|
<emphasis italics="true" pageId="54" pageNumber="371">Mesodma primaevus</emphasis>
|
|
</taxonomicName>
|
|
, suggesting a P4/M1 length ratio of about 1.0±0.1, which is similar to that in the other species of this family (
|
|
<bibRefCitation author="CLEMENS, W. A." bookContentInfo="1 map" box="[551,834,2517,2552]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="54" pageNumber="371" pagination="1 - 105" part="48" refId="ref56384" refString="1963 b. Fossil mammals of the type Lance Formation, Wyoming. Pt. 1. Introduction and Multituberculata. Univ. California Publ. Geol. Sci., vol. 48, pp. 1 - 105, figs. 1 - 51, tables 1 - 10, 1 map." title="Fossil mammals of the type Lance Formation, Wyoming. Pt. 1. Introduction and Multituberculata" type="book chapter" year="1963">Clemens, 1963b</bibRefCitation>
|
|
, p. 80).
|
|
</paragraph>
|
|
<paragraph blockId="54.[158,1013,679,2777]" lastBlockId="54.[1078,1936,351,2780]" pageId="54" pageNumber="371">
|
|
M2; morphologically, these teeth bear close resemblance to M2s of
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" box="[562,720,2609,2642]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[562,588,2609,2642]" italics="true" pageId="54" pageNumber="371">C</emphasis>
|
|
.
|
|
<emphasis box="[606,720,2609,2641]" italics="true" pageId="54" pageNumber="371">gracilis</emphasis>
|
|
</taxonomicName>
|
|
except that they are slightly smaller. Cusp formula of AMNH 77212 is 4/2 (fig. 1 1E). External cusps are four in number, subcrescentic with deep intervening valleys. The posteroexternal cusp occupies the posterolabial portion of the crown and is linked to the internal row by a low ridge, in some instances bearing small cuspules. The internal row consists of two large cusps which are much higher than the external ones and separated from them by broad valleys. An anterior ridge usually joins the anterior cusps of the external and internal rows.
|
|
</paragraph>
|
|
<paragraph blockId="54.[1078,1936,351,2780]" pageId="54" pageNumber="371">
|
|
Upper incisor; a right upper incisor, AMNH 77266, is referred to
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[1420,1677,814,850]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[1420,1677,814,850]" italics="true" pageId="54" pageNumber="371">Cimolomys clarki</emphasis>
|
|
</taxonomicName>
|
|
on the basis of size and the presence of two distinct cusps (fig. 1 IJ).
|
|
</paragraph>
|
|
<paragraph blockId="54.[1078,1936,351,2780]" pageId="54" pageNumber="371">
|
|
P4; two specimens of P4, AMNH 77217 and 77218, both from Clambank Hollow, are referred to
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[1195,1330,998,1035]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[1195,1221,999,1034]" italics="true" pageId="54" pageNumber="371">C</emphasis>
|
|
.
|
|
<emphasis box="[1240,1330,998,1035]" italics="true" pageId="54" pageNumber="371">clarki</emphasis>
|
|
</taxonomicName>
|
|
. Cusp formula of AMNH 77217 is 2/5 or 6/2 (fig. 1 IH, I). Medial cusps gradually increase in height posteriorly and have prominent vertical striations on both their lingual and labial sides. The crown is low, and at its base enamel thickens and bulges on both external and internal sides. As is the case with
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" box="[1706,1873,1273,1310]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[1706,1732,1275,1310]" italics="true" pageId="54" pageNumber="371">C</emphasis>
|
|
.
|
|
<emphasis box="[1756,1873,1273,1310]" italics="true" pageId="54" pageNumber="371">gracilis</emphasis>
|
|
</taxonomicName>
|
|
, to which the teeth are similar, a distinct ridge connects the highest medial cusp to the basal cusps situated on the posterolabial corner of the crown. Lingual to the basal cusp, there is a depressed triangular area with two to three deep pits and grooves. Two external cusps in AMNH 77217 are situated on the anterolabial expansion of the crown; the posterior cusp is usually niuch larger than the anterior.
|
|
</paragraph>
|
|
<paragraph blockId="54.[1078,1936,351,2780]" pageId="54" pageNumber="371">The coronal shape of P4 in occlusal view is roughly rectangular, with a constriction on both the lingual and the labial sides of the crown. Roots of AMNH 77217 are broken but their sockets indicate a subrounded cross section, and a small interradicular crest is present.</paragraph>
|
|
<paragraph blockId="54.[1078,1936,351,2780]" pageId="54" pageNumber="371">
|
|
Ml; there are at least a dozen Mls in the collection from Clambank Hollow. They are similar to M1s of
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" box="[1430,1601,2104,2141]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="54" pageNumber="371" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[1430,1456,2106,2141]" italics="true" pageId="54" pageNumber="371">C</emphasis>
|
|
.
|
|
<emphasis box="[1486,1601,2104,2141]" italics="true" pageId="54" pageNumber="371">gracilis</emphasis>
|
|
</taxonomicName>
|
|
except that their dimensions are much smaller and they possess fewer medial cusps. Cusp formula of AMNH 77219 is 5/6/6 (fig. I1 G). External cusps are conical or pyramidal in shape. The anterior cusps are joined by a ridge to the anteromedial cusp. Other external cusps are well separated from each other by distinct valleys, especially the posterolabial cusp, in which the valley is wide and projects labially. Cusps ofthe medial row are crescentic with their apexes directed forward. Those on the internal row diminish in size anteriorly and are six in number. Anterolingual cusps are conical in shape, with circular bases, and extend to the secondmost anterior cusp of the medial row.
|
|
</paragraph>
|
|
<caption httpUri="https://zenodo.org/record/3382471/files/figure.png" pageId="55" pageNumber="372" targetBox="[336,2099,331,2525]" targetPageId="55">
|
|
<paragraph pageId="55" pageNumber="372">
|
|
FIG. 1 1.
|
|
<taxonomicName authority="Sahni, 1972" authorityName="Sahni" authorityYear="1972" box="[499,740,2543,2577]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="55" pageNumber="372" phylum="Chordata" rank="species" species="clarki" status="new species">
|
|
<emphasis box="[499,646,2544,2577]" italics="true" pageId="55" pageNumber="372">Cimolomys</emphasis>
|
|
clarki
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel box="[761,946,2544,2577]" pageId="55" pageNumber="372" rank="species">new species</taxonomicNameLabel>
|
|
. A. AMNH 77179, labial view of right P4. B. Lingual view. C. AMNH 77180, labial view of right P4. D. Lingual view. E. AMNH 77212, occlusal view of right M2. F. AMNH 77241, occlusal view of left M2. G. AMNH 77219, occlusal view of right M'. H. AMNH 77217, labial view of right P4.
|
|
<emphasis bold="true" box="[326,342,2668,2699]" pageId="55" pageNumber="372">I</emphasis>
|
|
. Lingual view. J. AMNH 77266, labial view of right upper incisor. K.?
|
|
<taxonomicName authority="(Lambe)" authorityName="Lambe" baseAuthorityName="Lambe" box="[1508,1951,2666,2699]" class="Mammalia" family="Ptilodontidae" genus="Mesodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="55" pageNumber="372" phylum="Chordata" rank="species" species="primaevus">
|
|
<emphasis box="[1508,1794,2667,2699]" italics="true" pageId="55" pageNumber="372">Mesodma primaevus</emphasis>
|
|
(Lambe)
|
|
</taxonomicName>
|
|
, AMNH 77178, proximal portion of right femur. L.
|
|
<taxonomicName authority="Sahni, 1972" authorityName="Sahni" authorityYear="1972" box="[1004,1273,2706,2740]" class="Mammalia" family="Taeniolabidae" genus="Cimexomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="55" pageNumber="372" phylum="Chordata" rank="species" species="magnus" status="new species">
|
|
<emphasis box="[1004,1273,2706,2740]" italics="true" pageId="55" pageNumber="372">Cimexomys magnus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel box="[1292,1474,2708,2739]" pageId="55" pageNumber="372" rank="species">new species</taxonomicNameLabel>
|
|
, AMNH 77112, lingual view of posterior fragment of left P4. M.
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" box="[771,913,2749,2780]" class="Mammalia" family="Cimolodontidae" genus="Cimolodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="55" pageNumber="376" phylum="Chordata" rank="genus">Cimolodon</taxonomicName>
|
|
<emphasis box="[926,966,2750,2780]" italics="true" pageId="55" pageNumber="372">sp</emphasis>
|
|
., AMNH 77267, labial view of anterior fragment of left P4. N. Lingual view. 0. Anuran, AMNH 8424, anterior view of vertebra. P. Posterior view. All x 10.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph blockId="56.[163,1021,357,2781]" pageId="56" pageNumber="373">The cross section of the anterior root is elongated, whereas that of the posterior is transversely rectangular. Two small accessory roots are present near the posterior root and on either side of the interradicular crest.</paragraph>
|
|
<paragraph blockId="56.[163,1021,357,2781]" pageId="56" pageNumber="373">M2; cusp formula of AMNH 77241 is 2/3/4 (fig. 11F). The internal row consists of four pyramidal cusps, which are well separated from each other by deep valleys. The anterointernal cusp is linked by a low ridge to the anteromedial cusp and also to the anteroexternal cusp, forming the anterior ridge which closely fits against the posterior side of M1. Medial cusps are well separated, and the most posterior one is crescentic and joined to the external cusps by a cuspidate ridge. Roots are equal in size, and an interradicular crest is feebly developed.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="56" pageNumber="373" type="discussion">
|
|
<paragraph blockId="56.[163,1021,357,2781]" pageId="56" pageNumber="373">
|
|
<emphasis box="[209,403,1142,1177]" italics="true" pageId="56" pageNumber="373">DISCUSSION</emphasis>
|
|
: Although five species have been assigned to the genus
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[572,739,1185,1222]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[572,739,1185,1222]" italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
, only three,
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[978,1005,1187,1221]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
. clarki
|
|
</taxonomicName>
|
|
, from the Campanian and
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" box="[764,925,1232,1269]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[764,790,1233,1268]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
.
|
|
<emphasis box="[811,925,1232,1269]" italics="true" pageId="56" pageNumber="373">gracilis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Lillegraven" authorityYear="1969" box="[166,346,1279,1315]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="trochuus">
|
|
<emphasis box="[166,193,1280,1315]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
.
|
|
<emphasis box="[220,346,1279,1315]" italics="true" pageId="56" pageNumber="373">trochuus</emphasis>
|
|
</taxonomicName>
|
|
, from the Maestrichtian, are validly included.
|
|
<taxonomicName authority="Lambe (1902)" authorityName="Lambe" authorityYear="1902" box="[347,909,1322,1361]" class="Mammalia" family="Ptilodontidae" genus="Ptilodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="primaevus">
|
|
<emphasis box="[347,641,1322,1361]" italics="true" pageId="56" pageNumber="373">Ptilodus primaevus</emphasis>
|
|
<bibRefCitation author="LAMBE, L. M." box="[660,909,1323,1360]" journalOrPublisher="Contrib. Canadian Paleont." pageId="56" pageNumber="373" pagination="25 - 81" part="3" refId="ref60224" refString="LAMBE, L. M. 1902. New genera and species from the Belly River Series (mid-Cretaceous). Contrib. Canadian Paleont., vol. 3, pp. 25 - 81, figs. 1 - 24, pls. 1 - 2 1." title="New genera and species from the Belly River Series (mid-Cretaceous)." type="journal article" year="1902">Lambe (1902)</bibRefCitation>
|
|
</taxonomicName>
|
|
from the Oldman Formation was referred to
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
by
|
|
<bibRefCitation author="SIMPSON, G. G." bookContentInfo="171 pp." box="[331,613,1416,1454]" journalOrPublisher="Mem. Peabody Mus. Nat. Hist." pageId="56" pageNumber="373" part="3" refId="ref63592" refString="1929. American Mesozoic Mammalia. Mem. Peabody Mus. Nat. Hist., vol. 3, pt. 1, xv + 171 pp., figs. 1 - 62, pls. 1 - 32." title="American Mesozoic Mammalia." type="book" year="1929">Simpson (1929)</bibRefCitation>
|
|
, and then later transferred to
|
|
<taxonomicName box="[349,496,1464,1499]" class="Mammalia" family="Ptilodontidae" genus="Mesodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[349,496,1464,1499]" italics="true" pageId="56" pageNumber="373">Mesodma</emphasis>
|
|
</taxonomicName>
|
|
by
|
|
<bibRefCitation author="CLEMENS, W. A." bookContentInfo="1 map" box="[584,893,1462,1499]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="56" pageNumber="373" pagination="1 - 105" part="48" refId="ref56384" refString="1963 b. Fossil mammals of the type Lance Formation, Wyoming. Pt. 1. Introduction and Multituberculata. Univ. California Publ. Geol. Sci., vol. 48, pp. 1 - 105, figs. 1 - 51, tables 1 - 10, 1 map." title="Fossil mammals of the type Lance Formation, Wyoming. Pt. 1. Introduction and Multituberculata" type="book chapter" year="1963">Clemens (1963b)</bibRefCitation>
|
|
.
|
|
<taxonomicName authority="Russell (1937)" authorityName="Russell" authorityYear="1937" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="major">
|
|
<emphasis italics="true" pageId="56" pageNumber="373">Cimolomys major</emphasis>
|
|
<bibRefCitation author="RUSSELL, L. S." box="[363,607,1508,1546]" journalOrPublisher="Ibid." pageId="56" pageNumber="373" pagination="75 - 80" part="30" refId="ref63105" refString="1937. New and interesting mammalian fossils from western Canada. Ibid., vol. 30, pp. 75 - 80, p 1. 1." title="New and interesting mammalian fossils from western Canada" type="journal article" year="1937">Russell (1937)</bibRefCitation>
|
|
</taxonomicName>
|
|
is herein transferred to
|
|
<taxonomicName authorityName="Cope" authorityYear="1882" box="[169,374,1557,1592]" class="Mammalia" family="Taeniolabididae" genus="Meniscoessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[169,374,1557,1592]" italics="true" pageId="56" pageNumber="373">Meniscoessus</emphasis>
|
|
</taxonomicName>
|
|
. All teeth of
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[611,879,1554,1591]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[611,879,1554,1591]" italics="true" pageId="56" pageNumber="373">Cimolomys clarki</emphasis>
|
|
</taxonomicName>
|
|
, except P4, are approximately 70 per cent of the size of their counterparts in
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" box="[546,709,1646,1682]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[546,573,1646,1681]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
.
|
|
<emphasis box="[592,709,1646,1682]" italics="true" pageId="56" pageNumber="373">gracilis</emphasis>
|
|
</taxonomicName>
|
|
. P4, on the other hand, is larger; the average length of the two specimens of
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[406,543,1740,1775]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[406,432,1740,1775]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
.
|
|
<emphasis box="[455,543,1740,1775]" italics="true" pageId="56" pageNumber="373">clarki</emphasis>
|
|
</taxonomicName>
|
|
is 4.0 mm. in comparison with the average length of the specimens of
|
|
<taxonomicName authority="(Clemens, 1963 b, p. 81)" authorityName="Clemens" authorityPageNumber="81" authorityYear="1963" baseAuthorityName="Clemens" baseAuthorityPageNumber="81" baseAuthorityYear="1963" box="[168,790,1829,1867]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[168,194,1831,1866]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
.
|
|
<emphasis box="[219,333,1830,1867]" italics="true" pageId="56" pageNumber="373">gracilis</emphasis>
|
|
(
|
|
<bibRefCitation author="CLEMENS, W. A." bookContentInfo="1 map" box="[367,657,1829,1867]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="56" pageNumber="373" pagination="1 - 105" part="48" refId="ref56384" refString="1963 b. Fossil mammals of the type Lance Formation, Wyoming. Pt. 1. Introduction and Multituberculata. Univ. California Publ. Geol. Sci., vol. 48, pp. 1 - 105, figs. 1 - 51, tables 1 - 10, 1 map." title="Fossil mammals of the type Lance Formation, Wyoming. Pt. 1. Introduction and Multituberculata" type="book chapter" year="1963">Clemens, 1963b</bibRefCitation>
|
|
, p. 81)
|
|
</taxonomicName>
|
|
of 3.0 mm. This observation can be explained as the result of the progressive tendency toward reduction of size of the shearing premolars relative to the other teeth in species of the family. Morphologically, however, the teeth in the two species are similar and it seems reasonable to derive
|
|
<taxonomicName baseAuthorityName="Clemens" baseAuthorityYear="1963" box="[169,456,2151,2189]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[169,456,2151,2189]" italics="true" pageId="56" pageNumber="373">Cimolomys gracilis</emphasis>
|
|
</taxonomicName>
|
|
from
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[563,699,2152,2187]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[563,589,2152,2187]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
.
|
|
<emphasis box="[609,699,2152,2187]" italics="true" pageId="56" pageNumber="373">clarki</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph blockId="56.[163,1021,357,2781]" lastBlockId="56.[1082,1937,355,1961]" pageId="56" pageNumber="373">
|
|
Absence of associated teeth in mandibles and maxillae from the Lance or Judith River formations has resulted in casual reference of teeth to specific taxa. All jaws previously described as belonging to
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[399,563,2381,2418]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[399,563,2381,2418]" italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
have been removed from the genus; thus, in the Judith River and Oldman formations, mandibles referred to
|
|
<taxonomicName authority="(Lambe, 1902)" authorityName="Lambe" authorityYear="1902" baseAuthorityName="Lambe" baseAuthorityYear="1902" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="primaevus">
|
|
<emphasis italics="true" pageId="56" pageNumber="373">Cimolomys primaevus</emphasis>
|
|
(
|
|
<bibRefCitation author="LAMBE, L. M." box="[472,717,2518,2554]" journalOrPublisher="Contrib. Canadian Paleont." pageId="56" pageNumber="373" pagination="25 - 81" part="3" refId="ref60224" refString="LAMBE, L. M. 1902. New genera and species from the Belly River Series (mid-Cretaceous). Contrib. Canadian Paleont., vol. 3, pp. 25 - 81, figs. 1 - 24, pls. 1 - 2 1." title="New genera and species from the Belly River Series (mid-Cretaceous)." type="journal article" year="1902">Lambe, 1902</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authority="Russell (1937)" authorityName="Russell" authorityYear="1937" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="major">
|
|
<emphasis italics="true" pageId="56" pageNumber="373">Cimolomys major</emphasis>
|
|
<bibRefCitation author="RUSSELL, L. S." box="[284,545,2564,2600]" journalOrPublisher="Ibid." pageId="56" pageNumber="373" pagination="75 - 80" part="30" refId="ref63105" refString="1937. New and interesting mammalian fossils from western Canada. Ibid., vol. 30, pp. 75 - 80, p 1. 1." title="New and interesting mammalian fossils from western Canada" type="journal article" year="1937">Russell (1937)</bibRefCitation>
|
|
</taxonomicName>
|
|
are placed in different genera. Also, with the removal of
|
|
<taxonomicName box="[783,930,2609,2644]" class="Mammalia" family="Ptilodontidae" genus="Mesodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[783,930,2609,2644]" italics="true" pageId="56" pageNumber="373">Mesodma</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Marsh" authorityYear="1889" box="[167,325,2655,2690]" class="Mammalia" family="Cimolodontidae" genus="Cimolodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="376" phylum="Chordata" rank="genus">Cimolodon</taxonomicName>
|
|
by
|
|
<bibRefCitation author="CLEMENS, W. A." bookContentInfo="1 map" box="[396,687,2653,2689]" journalOrPublisher="Univ. California Publ. Geol. Sci." pageId="56" pageNumber="373" pagination="1 - 105" part="48" refId="ref56384" refString="1963 b. Fossil mammals of the type Lance Formation, Wyoming. Pt. 1. Introduction and Multituberculata. Univ. California Publ. Geol. Sci., vol. 48, pp. 1 - 105, figs. 1 - 51, tables 1 - 10, 1 map." title="Fossil mammals of the type Lance Formation, Wyoming. Pt. 1. Introduction and Multituberculata" type="book chapter" year="1963">Clemens (1963b)</bibRefCitation>
|
|
from the group of multituberculates that had been included by
|
|
<bibRefCitation author="SIMPSON, G. G." bookContentInfo="171 pp." box="[169,439,2743,2780]" journalOrPublisher="Mem. Peabody Mus. Nat. Hist." pageId="56" pageNumber="373" part="3" refId="ref63592" refString="1929. American Mesozoic Mammalia. Mem. Peabody Mus. Nat. Hist., vol. 3, pt. 1, xv + 171 pp., figs. 1 - 62, pls. 1 - 32." title="American Mesozoic Mammalia." type="book" year="1929">Simpson (1929)</bibRefCitation>
|
|
in
|
|
<taxonomicName authority="Sahni, 1972" authorityName="Sahni" authorityYear="1972" box="[500,667,2743,2778]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus" sensu="lato" status="sensu lato">
|
|
<emphasis box="[500,667,2743,2778]" italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<emphasis box="[687,838,2743,2779]" italics="true" pageId="56" pageNumber="373">
|
|
<taxonomicNameLabel box="[687,838,2743,2779]" pageId="56" pageNumber="373" sensu="lato">sensu lato</taxonomicNameLabel>
|
|
</emphasis>
|
|
, there are now no jaws that can be referred to
|
|
<taxonomicName authority="Sahni, 1972" authorityName="Sahni" authorityYear="1972" box="[1755,1925,358,393]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus" sensu="stricto" status="sensu stricto">
|
|
<emphasis box="[1755,1925,358,393]" italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<emphasis box="[1083,1288,406,442]" italics="true" pageId="56" pageNumber="373">
|
|
<taxonomicNameLabel box="[1083,1288,406,442]" pageId="56" pageNumber="373" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
|
</emphasis>
|
|
. The method used by Clemens (1 963b, p. 80) for distinguishing
|
|
<taxonomicName authority="Sahni, 1972" authorityName="Sahni" authorityYear="1972" box="[1668,1837,450,485]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus" sensu="stricto" status="sensu stricto">
|
|
<emphasis box="[1668,1837,450,485]" italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<emphasis italics="true" pageId="56" pageNumber="373">
|
|
<taxonomicNameLabel pageId="56" pageNumber="373" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
|
</emphasis>
|
|
was to eliminate teeth referable to
|
|
<taxonomicName authority=", Cimolodon" authorityName="Cimolodon" box="[1087,1437,543,579]" class="Mammalia" family="Ptilodontidae" genus="Mesodma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1087,1239,543,578]" italics="true" pageId="56" pageNumber="373">Mesodma</emphasis>
|
|
, Cimolodon
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName authorityName="Cope" authorityYear="1882" box="[1571,1773,544,579]" class="Mammalia" family="Taeniolabididae" genus="Meniscoessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1571,1773,544,579]" italics="true" pageId="56" pageNumber="373">Meniscoessus</emphasis>
|
|
</taxonomicName>
|
|
and to assign the few remaining teeth to
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[1690,1858,588,624]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1690,1858,588,624]" italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
. He corroborated his findings by computing ratios for various upper and lower teeth. Material from the Judith River Formation tends to confirm Clemens's identification.
|
|
</paragraph>
|
|
<paragraph blockId="56.[1082,1937,355,1961]" pageId="56" pageNumber="373">
|
|
<taxonomicName authority="Russell" authorityName="Russell" authorityYear="1937" box="[1128,1530,821,856]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="major">
|
|
<emphasis box="[1128,1394,821,856]" italics="true" pageId="56" pageNumber="373">Cimolomys major</emphasis>
|
|
<bibRefCitation author="RUSSELL, L. S." box="[1408,1646,821,856]" journalOrPublisher="Ibid." pageId="56" pageNumber="373" pagination="75 - 80" part="30" refId="ref63105" refString="1937. New and interesting mammalian fossils from western Canada. Ibid., vol. 30, pp. 75 - 80, p 1. 1." title="New and interesting mammalian fossils from western Canada" type="journal article" year="1937">Russell (1937</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<bibRefCitation author="RUSSELL, L. S." box="[1675,1753,820,855]" journalOrPublisher="Bull. Natl. Mus. Canada" pageId="56" pageNumber="373" pagination="110 - 119" part="126" refId="ref63168" refString="1952. Cretaceous mammals of Alberta. Bull. Natl. Mus. Canada, vol. 126, pp. 110 - 119, pls. 1, 2." title="Cretaceous mammals of Alberta" type="journal article" year="1952">1952</bibRefCitation>
|
|
) from the Oldman Formation is in reality closer to
|
|
<taxonomicName authorityName="Cope" authorityYear="1882" class="Mammalia" family="Taeniolabididae" genus="Meniscoessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="56" pageNumber="373">Meniscoessus</emphasis>
|
|
</taxonomicName>
|
|
than to
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[1340,1504,912,947]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1340,1504,912,947]" italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<emphasis box="[1615,1639,913,948]" italics="true" pageId="56" pageNumber="373">is</emphasis>
|
|
discussed later. Only the reasons for excluding it from
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
will be mentioned here. The type specimen is NMC 8743, a broken mandible with M1-M2, and the root of the incisor. Later a P4 was obtained at another locality. The root of this P4
|
|
<emphasis box="[1087,1111,1190,1225]" italics="true" pageId="56" pageNumber="373">is</emphasis>
|
|
the same size as that expected for the alveolus of the type specimen. The teeth are large and the combined length of P4-M2 is 17.2 mm., nearly twice that of
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[1323,1460,1327,1362]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="clarki">
|
|
<emphasis box="[1323,1349,1327,1362]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
.
|
|
<emphasis box="[1370,1460,1327,1362]" italics="true" pageId="56" pageNumber="373">clarki</emphasis>
|
|
</taxonomicName>
|
|
. They are almost identical in size and morphology to teeth of the smallest species of
|
|
<taxonomicName authorityName="Cope" authorityYear="1882" box="[1259,1463,1418,1453]" class="Mammalia" family="Taeniolabididae" genus="Meniscoessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1259,1463,1418,1453]" italics="true" pageId="56" pageNumber="373">Meniscoessus</emphasis>
|
|
</taxonomicName>
|
|
, found in the Judith River Formation. All species of
|
|
<taxonomicName authorityName="Cope" authorityYear="1882" box="[1619,1820,1464,1499]" class="Mammalia" family="Taeniolabididae" genus="Meniscoessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1619,1820,1464,1499]" italics="true" pageId="56" pageNumber="373">Meniscoessus</emphasis>
|
|
</taxonomicName>
|
|
have characteristic grooves on the cusps, especially on the molars. In some cases similar grooves are present on the incisor and P4. Such grooves are present on M1 and M2 of Russell's type specimen NMC 8743. The structure of
|
|
<emphasis bold="true" box="[1604,1625,1695,1731]" pageId="56" pageNumber="373">P</emphasis>
|
|
4 of
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[1695,1858,1694,1731]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1695,1858,1694,1731]" italics="true" pageId="56" pageNumber="373">Cimolomys</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Cope" authorityYear="1882" box="[1091,1292,1742,1777]" class="Mammalia" family="Taeniolabididae" genus="Meniscoessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1091,1292,1742,1777]" italics="true" pageId="56" pageNumber="373">Meniscoessus</emphasis>
|
|
</taxonomicName>
|
|
is also different; in the Campanian
|
|
<taxonomicName authorityName="Cope" authorityYear="1882" box="[1089,1291,1788,1824]" class="Mammalia" family="Taeniolabididae" genus="Meniscoessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="genus">
|
|
<emphasis box="[1089,1291,1788,1824]" italics="true" pageId="56" pageNumber="373">Meniscoessus</emphasis>
|
|
</taxonomicName>
|
|
and NMC 8820 (P4 referred by
|
|
<bibRefCitation author="RUSSELL, L. S." box="[1090,1327,1833,1869]" journalOrPublisher="Bull. Natl. Mus. Canada" pageId="56" pageNumber="373" pagination="110 - 119" part="126" refId="ref63168" refString="1952. Cretaceous mammals of Alberta. Bull. Natl. Mus. Canada, vol. 126, pp. 110 - 119, pls. 1, 2." title="Cretaceous mammals of Alberta" type="journal article" year="1952">Russell, 1952</bibRefCitation>
|
|
), 1 1 serrations are present instead of the usual eight or nine found in P4 of
|
|
<emphasis italics="true" pageId="56" pageNumber="373">
|
|
Cimilomys
|
|
<taxonomicName authorityName="Sahni" authorityYear="1972" box="[1186,1273,1925,1960]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="clarki">clarki</taxonomicName>
|
|
</emphasis>
|
|
and
|
|
<taxonomicName baseAuthorityName="Clemens" baseAuthorityYear="1963" box="[1364,1528,1926,1961]" class="Mammalia" family="Cimolomyidae" genus="Cimolomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="56" pageNumber="373" phylum="Chordata" rank="species" species="gracilis">
|
|
<emphasis box="[1364,1390,1926,1961]" italics="true" pageId="56" pageNumber="373">C</emphasis>
|
|
.
|
|
<emphasis box="[1409,1528,1926,1961]" italics="true" pageId="56" pageNumber="373">gracilis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |