484 lines
70 KiB
XML
484 lines
70 KiB
XML
<document id="E076E784E2671E70663123AD02F90FB4" ID-CLB-Dataset="7426" ID-DOI="10.11646/zootaxa.4930.1.1" ID-GBIF-Dataset="ac692e2d-3f49-42b8-8855-02f81512a3c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4544896" ID-ZooBank="97110C21-173C-4552-96AC-4B5DC987FF1C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1613556916485" checkinUser="plazi" docAuthor="Blake, James A." docDate="2021" docId="03C9912CFFF6FF9E01A717E8FB12FDDC" docLanguage="en" docName="zootaxa.4930.1.1.pdf" docOrigin="Zootaxa 4930 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scoloplos papillatus Blake 2021, new species" docType="treatment" docUuid="DF1B8B75-38BE-4841-8452-13DCF90DD7AB" docUuidSource="ZooBank" docVersion="7" lastPageNumber="42" masterDocId="FFF0E954FFD2FFB70130126AFF83FFBD" masterDocTitle="New species and records of Orbiniidae (Annelida, Polychaeta) from continental shelf and slope depths of the Western North Atlantic Ocean" masterLastPageNumber="123" masterPageNumber="1" pageNumber="37" updateTime="1698929903993" updateUser="ExternalLinkService">
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<mods:titleInfo id="8A42075461BD930EA415D344AAAFDB5B">
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<mods:title id="E0685A8D89E961745C60BA7C58903BE2">New species and records of Orbiniidae (Annelida, Polychaeta) from continental shelf and slope depths of the Western North Atlantic Ocean</mods:title>
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<mods:name id="374F30E823315BF9D9B9742D4620026A" type="personal">
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<mods:namePart id="B70B6C5BFA0874F06DF2C48AF0C4AFE7">Blake, James A.</mods:namePart>
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<mods:nameIdentifier id="9E9B4563DC078B939A51E8D5D1142BC6" type="email">jablake9@gmail.com</mods:nameIdentifier>
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<mods:title id="CD5A6C98081C2B74608BDF011A8304FE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0244E5EF521E3473A4DF2563368D10C1">
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<mods:date id="028F100A986332542C379F312F9232DB">2021</mods:date>
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<mods:number id="702D9A4A5A93A2A7419F359A40880CAA">2021-02-17</mods:number>
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<mods:number id="35FAB0CB0ABAA3E1B6B91A5E87954296">4930</mods:number>
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<mods:identifier id="591B7D9E5730EBD67A736EC5AF66F0E4" type="CLB-Dataset">7426</mods:identifier>
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<mods:identifier id="CFDC2123728BBC966798C087E584340F" type="DOI">10.11646/zootaxa.4930.1.1</mods:identifier>
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<treatment id="03C9912CFFF6FF9E01A717E8FB12FDDC" ID-DOI="http://doi.org/10.5281/zenodo.4678527" ID-GBIF-Taxon="180947682" ID-Zenodo-Dep="4678527" LSID="urn:lsid:zoobank.org:act:DF1B8B75-38BE-4841-8452-13DCF90DD7AB" httpUri="http://treatment.plazi.org/id/03C9912CFFF6FF9E01A717E8FB12FDDC" lastPageId="41" lastPageNumber="42" pageId="36" pageNumber="37">
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<subSubSection id="C37A73B1FFF6FF9301A717E8FDA3FA21" box="[151,544,1410,1437]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BDF203AFFF6FF9301A717E8FDA3FA21" blockId="36.[151,977,1410,1509]" box="[151,544,1410,1437]" pageId="36" pageNumber="37">
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<heading id="D0979756FFF6FF9301A717E8FDA3FA21" bold="true" box="[151,544,1410,1437]" fontSize="11" level="2" pageId="36" pageNumber="37" reason="4">
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<emphasis id="B914FC28FFF6FF9301A717E8FDA3FA21" bold="true" box="[151,544,1410,1437]" pageId="36" pageNumber="37">
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<taxonomicName id="4C605BB9FFF6FF9301A717E8FE0AFA20" ID-CoL="9BDHL" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[151,393,1410,1437]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
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<emphasis id="B914FC28FFF6FF9301A717E8FE0AFA20" bold="true" box="[151,393,1410,1437]" italics="true" pageId="36" pageNumber="37">Scoloplos papillatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2274153FFF6FF9300A117E9FDA3FA21" box="[401,544,1410,1437]" pageId="36" pageNumber="37" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37A73B1FFF6FF9301A717CDFC52FA58" pageId="36" pageNumber="37" type="description">
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<paragraph id="8BDF203AFFF6FF9301A717CDFEBBFA7C" blockId="36.[151,977,1410,1509]" box="[151,312,1446,1473]" pageId="36" pageNumber="37">
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<figureCitation id="135B3CBFFFF6FF9301A717CDFEBBFA7C" box="[151,312,1446,1473]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStartId-0="38.[151,250,1943,1967]" captionStartId-1="39.[151,250,1862,1886]" captionStartId-2="40.[151,250,1886,1910]" captionTargetBox-0="[193,1395,186,1911]" captionTargetBox-1="[161,1423,185,1824]" captionTargetBox-2="[217,1372,189,1855]" captionTargetId-0="figure-19@38.[184,1403,181,1919]" captionTargetId-1="figure-15@39.[151,1436,181,1838]" captionTargetId-2="figure-19@40.[209,1377,181,1861]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionTargetPageId-2="40" captionText-0="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." captionText-1="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." captionText-2="FIGURE 17. Scoloplos papillatus n. sp. A, anterior end of holotype, dorsolateral view; B, anterior end of paratype ventrolateral view; C, furcate notoseta and camerated capillaries, abdominal setiger; D, anterior abdominal segments dorsal view with pigmented interramal swellings denoted with arrows; E, middle abdominal segments, lateral view; F, posterior end and pygidium. A, D–F, holotype, (USNM 1621017); B–C, G paratype, (USNM 1621018). A–F stained with Shirlastain A." figureDoi-0="http://doi.org/10.5281/zenodo.4544931" figureDoi-1="http://doi.org/10.5281/zenodo.4544933" figureDoi-2="http://doi.org/10.5281/zenodo.4544935" httpUri-0="https://zenodo.org/record/4544931/files/figure.png" httpUri-1="https://zenodo.org/record/4544933/files/figure.png" httpUri-2="https://zenodo.org/record/4544935/files/figure.png" pageId="36" pageNumber="37">Figures 15–17</figureCitation>
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</paragraph>
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<paragraph id="8BDF203AFFF6FF9301A717A0FC52FA58" blockId="36.[151,977,1410,1509]" box="[151,977,1482,1509]" pageId="36" pageNumber="37">
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urn:lsid:zoobank.org:act:
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</subSubSection>
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<subSubSection id="C37A73B1FFF6FF9301A71466FD71F92E" pageId="36" pageNumber="37" type="materials_examined">
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<paragraph id="8BDF203AFFF6FF9301A71466FD71F92E" blockId="36.[151,1437,1548,2043]" pageId="36" pageNumber="37">
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<emphasis id="B914FC28FFF6FF9301A71466FEF9F99B" bold="true" box="[151,378,1548,1574]" pageId="36" pageNumber="37">Material examined</emphasis>
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. (
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<emphasis id="B914FC28FFF6FF9300A21466FDAAF99B" box="[402,553,1548,1574]" italics="true" pageId="36" pageNumber="37">
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<specimenCount id="9D66EBB3FFF6FF9300A21466FDAAF99B" box="[402,553,1548,1574]" pageId="36" pageNumber="37" type="generic">36 specimens</specimenCount>
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</emphasis>
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)
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<materialsCitation id="3B082A67FFF6FF93030B1466FDBAF9D2" ID-GBIF-Occurrence="3069842878" collectingDate="1992-08" collectionCode="USNM" collectorName="J. A. Blake " country="United States of America" elevation="812" latitude="35.618" location="Cape Hatteras" longLatPrecision="11" longitude="-74.76867" pageId="36" pageNumber="37" specimenCode="USNM 1621017" specimenCount="1" stateProvince="Cape Hatteras" typeStatus="holotype">
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<emphasis id="B914FC28FFF6FF93030B1466FEB5F9F7" bold="true" pageId="36" pageNumber="37">
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||
Southeastern
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<collectingCountry id="F37760AAFFF6FF9303EF1466FC9AF99B" box="[735,793,1548,1574]" name="United States of America" pageId="36" pageNumber="37">USA</collectingCountry>
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, off
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||
<collectingRegion id="49A4EED8FFF6FF93027F1466FC7CF99B" box="[847,1023,1548,1574]" country="United States of America" name="Cape Hatteras" pageId="36" pageNumber="37">Cape Hatteras</collectingRegion>
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,
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<collectingRegion id="49A4EED8FFF6FF93053C1466FB45F99B" box="[1036,1222,1548,1574]" country="United States of America" name="North Carolina" pageId="36" pageNumber="37">North Carolina</collectingRegion>
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, MMS
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<collectingRegion id="49A4EED8FFF6FF9304101466FEB3F9F7" country="United States of America" name="Cape Hatteras Survey" pageId="36" pageNumber="37">Cape Hatteras Survey</collectingRegion>
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||
,
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||
</emphasis>
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||
coll.
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||
<collectorName id="269545ECFFF6FF93004B145AFE75F9F7" box="[379,502,1584,1610]" pageId="36" pageNumber="37">J.A. Blake</collectorName>
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,
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||
<collectorName id="269545ECFFF6FF930335145AFD52F9F7" box="[517,721,1584,1610]" pageId="36" pageNumber="37">Chief Scientist, R</collectorName>
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/
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||
<collectorName id="269545ECFFF6FF9303EA145AFC23F9F7" box="[730,928,1584,1610]" pageId="36" pageNumber="37">
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||
V
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||
<emphasis id="B914FC28FFF6FF9303C7145AFCEAF9F7" box="[759,873,1584,1610]" italics="true" pageId="36" pageNumber="37">Endeavor.</emphasis>
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||
<emphasis id="B914FC28FFF6FF930242145AFC23F9F7" bold="true" box="[882,928,1584,1610]" pageId="36" pageNumber="37">Sta.</emphasis>
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||
</collectorName>
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||
CH-3:
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||
<date id="FFDE06FAFFF6FF9302CC145AFBF2F9F7" box="[1020,1137,1584,1610]" pageId="36" pageNumber="37" value="1992-08">
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||
<collectingDate id="EF9AFF12FFF6FF9302CC145AFBF2F9F7" box="[1020,1137,1584,1610]" pageId="36" pageNumber="37" value="1992-08">Aug 1992</collectingDate>
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||
</date>
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||
,
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<geoCoordinate id="EE5446FDFFF6FF93054F145BFA80F9F1" box="[1151,1283,1585,1612]" degrees="35" direction="north" minutes="37.08" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="35.618">35°37.08′N</geoCoordinate>
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||
,
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<geoCoordinate id="EE5446FDFFF6FF930421145BFA1BF9F1" box="[1297,1432,1585,1612]" degrees="74" direction="west" minutes="46.12" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="-74.76867">74°46.12′W</geoCoordinate>
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,
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<quantity id="4C988DDFFFF6FF9301A7143EFF66F9D2" box="[151,229,1620,1647]" metricMagnitude="2" metricUnit="m" metricValue="8.12" pageId="36" pageNumber="37" unit="m" value="812.0">
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<elevation id="004DC709FFF6FF9301A7143EFF66F9D2" box="[151,229,1620,1647]" metricMagnitude="2" metricUnit="m" metricValue="8.12" pageId="36" pageNumber="37" unit="m" value="812.0">812 m</elevation>
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</quantity>
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,
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<emphasis id="B914FC28FFF6FF9301C5143EFEDFF9D3" bold="true" box="[245,348,1620,1646]" pageId="36" pageNumber="37">
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<typeStatus id="54DB9E98FFF6FF9301C5143EFEDFF9D3" box="[245,348,1620,1646]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
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</emphasis>
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(
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<specimenCode id="DBC68841FFF6FF93005F143EFDB2F9D2" box="[367,561,1620,1647]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="36" pageNumber="37" type="Museum">USNM 1621017</specimenCode>
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)
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</materialsCitation>
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<materialsCitation id="3B082A67FFF6FF930374143EFC74F9D2" ID-GBIF-Occurrence="3069842721" box="[580,1015,1620,1647]" collectionCode="USNM" country="United States of America" pageId="36" pageNumber="37" specimenCode="USNM 1621018" specimenCount="17" stateProvince="Cape Hatteras" typeStatus="paratype">
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and
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<specimenCount id="9D66EBB3FFF6FF930348143EFC95F9D3" box="[632,790,1620,1646]" pageId="36" pageNumber="37" type="generic" typeStatus="paratypes">
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17
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<emphasis id="B914FC28FFF6FF930390143EFC95F9D3" bold="true" box="[672,790,1620,1646]" pageId="36" pageNumber="37">paratypes</emphasis>
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</specimenCount>
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(
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||
<specimenCode id="DBC68841FFF6FF930219143EFC6CF9D2" box="[809,1007,1620,1647]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="36" pageNumber="37" type="Museum">USNM 1621018</specimenCode>
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)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B082A67FFF6FF930535143EFD6CF92E" ID-GBIF-Occurrence="3069842775" collectingDate="1992-09" collectionCode="USNM" country="United States of America" elevation="620" latitude="35.472668" longLatPrecision="11" longitude="-74.68767" pageId="36" pageNumber="37" specimenCode="USNM 1621019" specimenCount="18" stateProvince="Cape Hatteras" typeStatus="paratype">
|
||
<emphasis id="B914FC28FFF6FF930535143EFBF9F9D3" bold="true" box="[1029,1146,1620,1646]" pageId="36" pageNumber="37">Sta. SA-9</emphasis>
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||
:
|
||
<date id="FFDE06FAFFF6FF9305BD143EFA86F9D3" box="[1165,1285,1620,1646]" pageId="36" pageNumber="37" value="1992-09">
|
||
<collectingDate id="EF9AFF12FFF6FF9305BD143EFA86F9D3" box="[1165,1285,1620,1646]" pageId="36" pageNumber="37" value="1992-09">Sep. 1992</collectingDate>
|
||
</date>
|
||
,
|
||
<geoCoordinate id="EE5446FDFFF6FF930425143FFA1BF9CD" box="[1301,1432,1621,1648]" degrees="35" direction="north" minutes="28.36" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="35.472668">35°28.36′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5446FDFFF6FF9301A71413FE9DF929" box="[151,286,1657,1684]" degrees="74" direction="west" minutes="41.26" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="-74.68767">74°41.26′W</geoCoordinate>
|
||
,
|
||
<quantity id="4C988DDFFFF6FF9300191412FEF1F92E" box="[297,370,1656,1683]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="36" pageNumber="37" unit="m" value="620.0">
|
||
<elevation id="004DC709FFF6FF9300191412FEF1F92E" box="[297,370,1656,1683]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="36" pageNumber="37" unit="m" value="620.0">620 m</elevation>
|
||
</quantity>
|
||
,
|
||
<specimenCount id="9D66EBB3FFF6FF93004E1412FD94F92F" box="[382,535,1656,1682]" pageId="36" pageNumber="37" type="generic" typeStatus="paratypes">
|
||
18
|
||
<emphasis id="B914FC28FFF6FF9300911412FD94F92F" bold="true" box="[417,535,1656,1682]" pageId="36" pageNumber="37">paratypes</emphasis>
|
||
</specimenCount>
|
||
(
|
||
<specimenCode id="DBC68841FFF6FF9303161412FD64F92E" box="[550,743,1656,1683]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="36" pageNumber="37" type="Museum">USNM 1621019</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A73B1FFF6FF9201F714F6FBFBFAC4" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="8BDF203AFFF6FF9201F714F6FE41FF68" blockId="36.[151,1437,1548,2043]" lastBlockId="37.[151,1437,151,2013]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
|
||
<emphasis id="B914FC28FFF6FF9301F714F6FED2F90B" bold="true" box="[199,337,1692,1718]" pageId="36" pageNumber="37">Description</emphasis>
|
||
. A large, elongate species; thoracic segments dorsoventrally flattened, often with middle thoracic segments swollen dorsally, likely from location of internal retracted pharynx; abdominal segments ventrally round- ed, dorsally flattened with elevated parapodia. Shallow ventral groove present along entire body on most specimens; dorsal grooves and ridges absent.
|
||
<typeStatus id="54DB9E98FFF6FF9303201562FDFBF89F" box="[528,632,1800,1826]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
complete, coiled, with 182 setigers,
|
||
<quantity id="4C988DDFFFF6FF9305211562FBDCF89E" box="[1041,1119,1800,1827]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="36" pageNumber="37" unit="mm" value="45.0">45 mm</quantity>
|
||
long,
|
||
<quantity id="4C988DDFFFF6FF9305951562FB79F89E" box="[1189,1274,1800,1827]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="36" pageNumber="37" unit="mm" value="1.8">1.8 mm</quantity>
|
||
across thorax; with 17 thoracic setigers and branchiae from setiger 14 (
|
||
<figureCitation id="135B3CBFFFF6FF9302251546FCFCF8FB" box="[789,895,1836,1862]" captionStart="FIGURE 17" captionStartId="40.[151,250,1886,1910]" captionTargetBox="[217,1372,189,1855]" captionTargetId="figure-19@40.[209,1377,181,1861]" captionTargetPageId="40" captionText="FIGURE 17. Scoloplos papillatus n. sp. A, anterior end of holotype, dorsolateral view; B, anterior end of paratype ventrolateral view; C, furcate notoseta and camerated capillaries, abdominal setiger; D, anterior abdominal segments dorsal view with pigmented interramal swellings denoted with arrows; E, middle abdominal segments, lateral view; F, posterior end and pygidium. A, D–F, holotype, (USNM 1621017); B–C, G paratype, (USNM 1621018). A–F stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.4544935" httpUri="https://zenodo.org/record/4544935/files/figure.png" pageId="36" pageNumber="37">Fig. 17A</figureCitation>
|
||
). Large incomplete
|
||
<typeStatus id="54DB9E98FFF6FF9305501547FB43F8FA" box="[1120,1216,1837,1863]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||
(USNM 1621018), with 180 setigers,
|
||
<quantity id="4C988DDFFFF6FF930054153AFE30F8D6" box="[356,435,1872,1899]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="36" pageNumber="37" unit="mm" value="37.0">37 mm</quantity>
|
||
long,
|
||
<quantity id="4C988DDFFFF6FF9300C9153AFDCCF8D6" box="[505,591,1872,1899]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="36" pageNumber="37" unit="mm" value="2.1">2.1 mm</quantity>
|
||
wide across thorax; with 18 thoracic setigers and branchiae from setiger 14 (
|
||
<figureCitation id="135B3CBFFFF6FF930190151EFE84F833" box="[160,263,1908,1934]" captionStart="FIGURE 17" captionStartId="40.[151,250,1886,1910]" captionTargetBox="[217,1372,189,1855]" captionTargetId="figure-19@40.[209,1377,181,1861]" captionTargetPageId="40" captionText="FIGURE 17. Scoloplos papillatus n. sp. A, anterior end of holotype, dorsolateral view; B, anterior end of paratype ventrolateral view; C, furcate notoseta and camerated capillaries, abdominal setiger; D, anterior abdominal segments dorsal view with pigmented interramal swellings denoted with arrows; E, middle abdominal segments, lateral view; F, posterior end and pygidium. A, D–F, holotype, (USNM 1621017); B–C, G paratype, (USNM 1621018). A–F stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.4544935" httpUri="https://zenodo.org/record/4544935/files/figure.png" pageId="36" pageNumber="37">Fig. 17B</figureCitation>
|
||
). Transition from thorax to abdomen abrupt, denoted by appearance of elongate and enlarged neuropodia, simple at first, becoming longer and with apex split over successive segments. Thoracic segments short, about five times wider than long. Color in alcohol light tan; dorsum of abdominal segments between branchiae with 1–3 medial dorsal brown pigment spots in intersegmental swelling (
|
||
<figureCitation id="135B3CBFFFF6FF930268158AFC38F847" box="[856,955,2016,2042]" captionStart="FIGURE 17" captionStartId="40.[151,250,1886,1910]" captionTargetBox="[217,1372,189,1855]" captionTargetId="figure-19@40.[209,1377,181,1861]" captionTargetPageId="40" captionText="FIGURE 17. Scoloplos papillatus n. sp. A, anterior end of holotype, dorsolateral view; B, anterior end of paratype ventrolateral view; C, furcate notoseta and camerated capillaries, abdominal setiger; D, anterior abdominal segments dorsal view with pigmented interramal swellings denoted with arrows; E, middle abdominal segments, lateral view; F, posterior end and pygidium. A, D–F, holotype, (USNM 1621017); B–C, G paratype, (USNM 1621018). A–F stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.4544935" httpUri="https://zenodo.org/record/4544935/files/figure.png" pageId="36" pageNumber="37">Fig. 17D</figureCitation>
|
||
arrows), this distinctive pigment conspicu- ous on large specimens; large subpodial flanges of middle and posterior abdominal neuropodia often with darkly pigmented internal glands.
|
||
</paragraph>
|
||
<paragraph id="8BDF203AFFF7FF9201F712B5FC51FE34" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
|
||
Pre-setiger region relatively short, slightly wider than long, triangular, overall as long as first two setigers (
|
||
<figureCitation id="135B3CBFFFF7FF9201901368FE82FEA0" box="[160,257,258,285]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig.15A</figureCitation>
|
||
). Prostomium conical, tapering to pointed tip; nuchal organs vertical curved slits on posterior lateral margin; eyespots absent. Peristomium slightly longer than first setiger, with a single smooth annular ring dorsally (
|
||
<figureCitation id="135B3CBFFFF7FF92045F134DFF49FED8" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig. 15A</figureCitation>
|
||
), surrounding mouth ventrally, forming upper and lower lips; upper lip in two parts each with numerous narrow lobes (
|
||
<figureCitation id="135B3CBFFFF7FF9201D21305FEB2FE34" box="[226,305,366,393]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig. 15</figureCitation>
|
||
B–C); lower lip with an equivalent number of narrow lobes.
|
||
</paragraph>
|
||
<paragraph id="8BDF203AFFF7FF9201F713F9FE0CFCAB" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
|
||
Anterior thoracic notopodia with digitiform postsetal lobe, short at first (
|
||
<figureCitation id="135B3CBFFFF7FF9202C313F9FBD8FE10" box="[1011,1115,402,429]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig. 15D</figureCitation>
|
||
), then becoming longer, narrow in posterior thoracic segments (
|
||
<figureCitation id="135B3CBFFFF7FF92031913DDFD0DFE6C" box="[553,654,438,465]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig. 15E</figureCitation>
|
||
); each notopodial lobe arising from a thick base. Thoracic neuropodia initially with a shorter, thick postsetal lobe arising from a broadly rounded base (
|
||
<figureCitation id="135B3CBFFFF7FF92051213B1FB09FE48" box="[1058,1162,474,501]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig. 15D</figureCitation>
|
||
); posterior thoracic neuropodia developing short postsetal lobe and a short subpodial papilla ventral to neuropodium (
|
||
<figureCitation id="135B3CBFFFF7FF9205821395FA94FDA4" box="[1202,1303,510,537]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig. 15E</figureCitation>
|
||
). Transition to abdominal segments abrupt, with fewer noto- and neurosetae. Abdominal notopodial postsetal lobes becoming narrow and elongate along rest of body (
|
||
<figureCitation id="135B3CBFFFF7FF920366102DFD27FDDF" box="[598,676,583,610]" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Fig. 16</figureCitation>
|
||
A–C); abdominal neuropodia narrow, divided apically into two parts separated by notch; medial lobe long, pointed; ventral lobe shorter, rounded apically, directed laterally, appearing as ventral cirrus (
|
||
<figureCitation id="135B3CBFFFF7FF9200B810E5FE5AFD17" box="[392,473,655,682]" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Fig. 16</figureCitation>
|
||
A–C). Interramal cirrus absent. Each abdominal neuropodium with narrow subpodial flange (
|
||
<figureCitation id="135B3CBFFFF7FF9201DC10D9FEC6FD70" box="[236,325,691,717]" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Figs. 16</figureCitation>
|
||
B–C, 17E); with one subpodial papilla on anterior abdominal segments (
|
||
<figureCitation id="135B3CBFFFF7FF92055A10D9FB58FD70" box="[1130,1243,691,717]" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Figs. 16A</figureCitation>
|
||
,
|
||
<figureCitation id="135B3CBFFFF7FF9205D510D9FA9BFD70" box="[1253,1304,691,717]" captionStart="FIGURE 17" captionStartId="40.[151,250,1886,1910]" captionTargetBox="[217,1372,189,1855]" captionTargetId="figure-19@40.[209,1377,181,1861]" captionTargetPageId="40" captionText="FIGURE 17. Scoloplos papillatus n. sp. A, anterior end of holotype, dorsolateral view; B, anterior end of paratype ventrolateral view; C, furcate notoseta and camerated capillaries, abdominal setiger; D, anterior abdominal segments dorsal view with pigmented interramal swellings denoted with arrows; E, middle abdominal segments, lateral view; F, posterior end and pygidium. A, D–F, holotype, (USNM 1621017); B–C, G paratype, (USNM 1621018). A–F stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.4544935" httpUri="https://zenodo.org/record/4544935/files/figure.png" pageId="37" pageNumber="38">17C</figureCitation>
|
||
). Subpodial flanges of middle and posterior abdominal setigers with numerous internal glands; these darkly pigmented on many specimens (
|
||
<figureCitation id="135B3CBFFFF7FF92002B1091FE01FCAB" box="[283,386,763,790]" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Fig. 16C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDF203AFFF7FF9201F71175FE4BFC74" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
|
||
Branchiae from posterior thoracic setiger 13 or 14 (
|
||
<figureCitation id="135B3CBFFFF7FF9203CE1175FCE5FC84" box="[766,870,798,825]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig. 15A</figureCitation>
|
||
); branchia short, narrow at first, becoming leaflike, longer, and full size by first abdominal segment; each anterior branchiae broad, tapering to narrow papillate tip (
|
||
<figureCitation id="135B3CBFFFF7FF92045F1129FF48FC3F" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Fig. 16A</figureCitation>
|
||
); a few anterior abdominal branchiae bifid, with a second apical papilla (
|
||
<figureCitation id="135B3CBFFFF7FF920530110DFBE0FC3C" box="[1024,1123,871,897]" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Fig. 16A</figureCitation>
|
||
inset). Branchiae of middle and posterior setigers becoming longer, triangulate, rarely asymmetrical (
|
||
<figureCitation id="135B3CBFFFF7FF9202F711E1FB95FC1B" box="[967,1046,907,934]" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Fig. 16</figureCitation>
|
||
B–C). Each branchia ciliated along lateral and medial margins.
|
||
</paragraph>
|
||
<paragraph id="8BDF203AFFF7FF9201F711B9FDB6FA8C" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
|
||
Thoracic notosetae numerous thick, long, camerated capillaries arranged in 4–5 rows. Thoracic neurosetae arranged in about five rows with similar appearing camerated capillaries and 5–10 uncini alternating with capillaries in second row; uncini occurring over first 7–10 setigers, becoming reduced to 3–4 uncini in lower part of setal fascicle; then either absent or not observed in later thoracic setigers. Individual uncini with shafts smooth on convex side, tapering to narrow, rounded tip; concave side of shaft flattened, bearing paired transverse knobs or barbs along most of shaft (
|
||
<figureCitation id="135B3CBFFFF7FF92000E16EDFE0DFB1C" box="[318,398,1158,1185]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Fig. 15</figureCitation>
|
||
F–G). Abdominal notosetae thin, camerated capillaries and 1–2 furcate setae. Furcate setae with unequal tynes, each tyne with blunted tip and apical notch; shaft with rows of transverse barbs or ridges; tynes with a row of thin needles extending medially (
|
||
<figureCitation id="135B3CBFFFF7FF92038016A5FCA0FB54" box="[688,803,1230,1257]" captionStart="FIGURE 15" captionStartId="38.[151,250,1943,1967]" captionTargetBox="[193,1395,186,1911]" captionTargetId="figure-19@38.[184,1403,181,1919]" captionTargetPageId="38" captionText="FIGURE 15. Scoloplos papillatus n. sp. Paratype (USNM 1621019):A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae." figureDoi="http://doi.org/10.5281/zenodo.4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="37" pageNumber="38">Figs. 15H</figureCitation>
|
||
,
|
||
<figureCitation id="135B3CBFFFF7FF92021F16A5FCE0FB54" box="[815,867,1231,1257]" captionStart="FIGURE 17" captionStartId="40.[151,250,1886,1910]" captionTargetBox="[217,1372,189,1855]" captionTargetId="figure-19@40.[209,1377,181,1861]" captionTargetPageId="40" captionText="FIGURE 17. Scoloplos papillatus n. sp. A, anterior end of holotype, dorsolateral view; B, anterior end of paratype ventrolateral view; C, furcate notoseta and camerated capillaries, abdominal setiger; D, anterior abdominal segments dorsal view with pigmented interramal swellings denoted with arrows; E, middle abdominal segments, lateral view; F, posterior end and pygidium. A, D–F, holotype, (USNM 1621017); B–C, G paratype, (USNM 1621018). A–F stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.4544935" httpUri="https://zenodo.org/record/4544935/files/figure.png" pageId="37" pageNumber="38">17G</figureCitation>
|
||
). Abdominal neurosetae with up to 4–5 thin capillaries, each with short barbs along one edge and 1–2 curved aciculae, sometimes protruding, with rounded tip (
|
||
<figureCitation id="135B3CBFFFF7FF92045F1699FF45FA8C" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Fig. 16A</figureCitation>
|
||
insets). Flail setae not observed.
|
||
</paragraph>
|
||
<paragraph id="8BDF203AFFF7FF9201F71751FBFBFAC4" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
|
||
Pygidium short, with about ten narrow dorsal lobes, two large lateral lobes, and a single large ventral lobe surrounding anal opening; with two short dorsal anal cirri arising dorsally (
|
||
<figureCitation id="135B3CBFFFF7FF92028E1735FBB3FAC4" box="[958,1072,1375,1401]" captionStart="FIGURE 16" captionStartId="39.[151,250,1862,1886]" captionTargetBox="[161,1423,185,1824]" captionTargetId="figure-15@39.[151,1436,181,1838]" captionTargetPageId="39" captionText="FIGURE 16. Scoloplos papillatus n. sp. A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017)." figureDoi="http://doi.org/10.5281/zenodo.4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="37" pageNumber="38">Figs. 16D</figureCitation>
|
||
,
|
||
<figureCitation id="135B3CBFFFF7FF92050C1735FBE8FAC4" box="[1084,1131,1375,1401]" captionStart="FIGURE 17" captionStartId="40.[151,250,1886,1910]" captionTargetBox="[217,1372,189,1855]" captionTargetId="figure-19@40.[209,1377,181,1861]" captionTargetPageId="40" captionText="FIGURE 17. Scoloplos papillatus n. sp. A, anterior end of holotype, dorsolateral view; B, anterior end of paratype ventrolateral view; C, furcate notoseta and camerated capillaries, abdominal setiger; D, anterior abdominal segments dorsal view with pigmented interramal swellings denoted with arrows; E, middle abdominal segments, lateral view; F, posterior end and pygidium. A, D–F, holotype, (USNM 1621017); B–C, G paratype, (USNM 1621018). A–F stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.4544935" httpUri="https://zenodo.org/record/4544935/files/figure.png" pageId="37" pageNumber="38">17F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A73B1FFF7FF9E01F717E9FD5DFF68" lastPageId="41" lastPageNumber="42" pageId="37" pageNumber="38" type="discussion">
|
||
<paragraph id="8BDF203AFFF7FF9201F717E9FD32F8F0" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
|
||
<emphasis id="B914FC28FFF7FF9201F717E9FEB8FA20" bold="true" box="[199,315,1411,1437]" pageId="37" pageNumber="38">Remarks.</emphasis>
|
||
Among species of
|
||
<taxonomicName id="4C605BB9FFF7FF92032E17E9FD08FA20" authorityName="de Blainville" authorityYear="1828" box="[542,651,1411,1437]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="genus">
|
||
<emphasis id="B914FC28FFF7FF92032E17E9FD08FA20" box="[542,651,1411,1437]" italics="true" pageId="37" pageNumber="38">Scoloplos</emphasis>
|
||
</taxonomicName>
|
||
along the
|
||
<collectingCountry id="F37760AAFFF7FF92023817E9FCB9FA20" box="[776,826,1411,1437]" name="United States of America" pageId="37" pageNumber="38">U.S.</collectingCountry>
|
||
Atlantic coast having a reduced number of uncini in thoracic neuropodia,
|
||
<taxonomicName id="4C605BB9FFF7FF9200B617CDFD91FA7C" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[390,530,1447,1473]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9200B617CDFD91FA7C" box="[390,530,1447,1473]" italics="true" pageId="37" pageNumber="38">S. papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF92032A17CDFDD9FA7C" box="[538,602,1447,1473]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF92032A17CDFDD1FA7C" bold="true" box="[538,594,1447,1473]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from upper slope depths is most similar to
|
||
<taxonomicName id="4C605BB9FFF7FF92057517CDFB33FA7C" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[1093,1200,1447,1473]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="verrilli" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF92057517CDFB33FA7C" box="[1093,1200,1447,1473]" italics="true" pageId="37" pageNumber="38">S. verrilli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B914FC28FFF7FF92058717CDFB74FA7C" bold="true" box="[1207,1271,1447,1473]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A2274153FFF7FF92058717CDFB74FA7C" box="[1207,1271,1447,1473]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, (see below) a common species in New
|
||
<collectingCountry id="F37760AAFFF7FF92008217A1FD8CFA58" box="[434,527,1483,1509]" name="United Kingdom" pageId="37" pageNumber="38">England</collectingCountry>
|
||
near-shore shelf sediments. The two species are similar in size (up to
|
||
<quantity id="4C988DDFFFF7FF92041317A0FA1EFA58" box="[1315,1437,1482,1509]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="37" pageNumber="38" unit="mm" value="43.5" valueMax="45.0" valueMin="42.0">42–45 mm</quantity>
|
||
long) and number of setigers (about 180–200).
|
||
<taxonomicName id="4C605BB9FFF7FF9203821785FCBCF9B4" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[690,831,1519,1545]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9203821785FCBCF9B4" box="[690,831,1519,1545]" italics="true" pageId="37" pageNumber="38">S. papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF9202781785FC0BF9B4" box="[840,904,1519,1545]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF9202781785FC03F9B4" bold="true" box="[840,896,1519,1545]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has 17–18 thoracic setigers whereas
|
||
<taxonomicName id="4C605BB9FFF7FF9204011785FA1FF9B4" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[1329,1436,1519,1545]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="verrilli" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9204011785FA1FF9B4" box="[1329,1436,1519,1545]" italics="true" pageId="37" pageNumber="38">S. verrilli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF9201A71479FF55F990" box="[151,214,1555,1581]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF9201A71479FF4CF990" bold="true" box="[151,207,1555,1581]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has 16–17 thoracic setigers; both species have branchiae from setigers 13–14. Obvious differences are in the development of extra podial lobes in posterior thoracic setigers:
|
||
<taxonomicName id="4C605BB9FFF7FF920258145DFC77F9EC" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[872,1012,1591,1617]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF920258145DFC77F9EC" box="[872,1012,1591,1617]" italics="true" pageId="37" pageNumber="38">S. papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF9202CB145DFBB9F9EC" box="[1019,1082,1591,1617]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF9202CB145DFBB0F9EC" bold="true" box="[1019,1075,1591,1617]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
develops both a subpodial lobe and subpodial papillae ventral to the neuropodium, whereas
|
||
<taxonomicName id="4C605BB9FFF7FF9202051431FC1DF9C8" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[821,926,1627,1653]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="verrilli" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9202051431FC1DF9C8" box="[821,926,1627,1653]" italics="true" pageId="37" pageNumber="38">S. verrilli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF9202941431FC61F9C8" box="[932,994,1627,1653]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF9202941431FC58F9C8" bold="true" box="[932,987,1627,1653]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
develops a second postsetal lobe on the neuropodium and a subpodial lobe. There also appear to be differences in the nature of the upper lip of the mouth: in
|
||
<taxonomicName id="4C605BB9FFF7FF9201A714C9FEA7F900" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[151,292,1699,1725]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9201A714C9FEA7F900" box="[151,292,1699,1725]" italics="true" pageId="37" pageNumber="38">S. papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF92001B14C9FEEAF900" box="[299,361,1699,1725]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF92001B14C9FEE0F900" bold="true" box="[299,355,1699,1725]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, the upper lip is divided into two parts, each of which has numerous narrow lobes; in
|
||
<taxonomicName id="4C605BB9FFF7FF92040214C9FA1FF900" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[1330,1436,1699,1725]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="verrilli" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF92040214C9FA1FF900" box="[1330,1436,1699,1725]" italics="true" pageId="37" pageNumber="38">S. verrilli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF9201A714ADFF56F95C" box="[151,213,1735,1761]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF9201A714ADFF4CF95C" bold="true" box="[151,207,1735,1761]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, the upper lobe is formed into two large, relatively smooth lobes that do not appear to be further divided. The most conspicuous difference between the two species, however, is that
|
||
<taxonomicName id="4C605BB9FFF7FF9202F11481FBCCF8B8" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[961,1103,1771,1797]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9202F11481FBCCF8B8" box="[961,1103,1771,1797]" italics="true" pageId="37" pageNumber="38">S. papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF9205671481FB1BF8B8" box="[1111,1176,1771,1797]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF9205671481FB13F8B8" bold="true" box="[1111,1168,1771,1797]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has 1–3 medial brown pigment spots in dorsal intersegmental swelling in abdominal segments, whereas in
|
||
<taxonomicName id="4C605BB9FFF7FF9205631565FB3CF894" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[1107,1215,1807,1833]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="verrilli" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9205631565FB3CF894" box="[1107,1215,1807,1833]" italics="true" pageId="37" pageNumber="38">S. verrilli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF9205F71565FA84F894" box="[1223,1287,1807,1833]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF9205F71565FA83F894" bold="true" box="[1223,1280,1807,1833]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
the intersegmental areas are neither swollen nor pigmented.
|
||
</paragraph>
|
||
<paragraph id="8BDF203AFFF7FF9E01F7153DFD5DFF68" blockId="37.[151,1437,151,2013]" lastBlockId="41.[151,1437,151,609]" lastPageId="41" lastPageNumber="42" pageId="37" pageNumber="38">
|
||
<taxonomicName id="4C605BB9FFF7FF9201F7153DFE2FF8CC" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[199,428,1879,1905]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9201F7153DFE2FF8CC" box="[199,428,1879,1905]" italics="true" pageId="37" pageNumber="38">Scoloplos papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF920084153DFE76F8CC" box="[436,501,1879,1905]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF920084153DFE6EF8CC" bold="true" box="[436,493,1879,1905]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is also similar to
|
||
<taxonomicName id="4C605BB9FFF7FF9203F1153DFB84F8CC" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[705,1031,1879,1905]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="californiensis">
|
||
<emphasis id="B914FC28FFF7FF9203F1153DFCF5F8CC" box="[705,886,1879,1905]" italics="true" pageId="37" pageNumber="38">S. californiensis</emphasis>
|
||
<bibRefCitation id="EFF15DCBFFF7FF92024F153DFB84F8CC" author="Blake, J. A." box="[895,1031,1879,1905]" pageId="37" pageNumber="38" pagination="1 - 61" refId="ref82361" refString="Blake, J. A. (2020) New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea. Zootaxa, 4730 (1), 1 - 61. https: // doi. org / 10.11646 / zootaxa. 4730.1.1" type="journal article" year="2020">Blake, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
from deep water off California and
|
||
<taxonomicName id="4C605BB9FFF7FF9201A71511FE5EF828" authority="Blake, 2020" authorityName="Blake" authorityYear="2020" box="[151,477,1915,1941]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="sparsaciculus">
|
||
<emphasis id="B914FC28FFF7FF9201A71511FECDF828" box="[151,334,1915,1941]" italics="true" pageId="37" pageNumber="38">S. sparsaciculus</emphasis>
|
||
<bibRefCitation id="EFF15DCBFFF7FF9200651511FE5EF828" author="Blake, J. A." box="[341,477,1915,1941]" pageId="37" pageNumber="38" pagination="1 - 61" refId="ref82361" refString="Blake, J. A. (2020) New species and records of deep-water Orbiniidae (Annelida, Polychaeta) from the Eastern Pacific continental slope, abyssal Pacific Ocean, and the South China Sea. Zootaxa, 4730 (1), 1 - 61. https: // doi. org / 10.11646 / zootaxa. 4730.1.1" type="journal article" year="2020">Blake, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
from deep water in the South
|
||
<collectingCountry id="F37760AAFFF7FF9202041511FCF4F828" box="[820,887,1915,1941]" name="China" pageId="37" pageNumber="38">China</collectingCountry>
|
||
Sea in having a reduced number of uncini in the thoracic neuropodia. Both of the two latter species have an extra subpodial lobe, but this is only present in anterior abdominal segments, whereas
|
||
<taxonomicName id="4C605BB9FFF7FF9200C315A9FD02F860" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[499,641,1987,2013]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF7FF9200C315A9FD02F860" box="[499,641,1987,2013]" italics="true" pageId="37" pageNumber="38">S. papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF7FF9203BA15A9FD48F860" box="[650,715,1987,2013]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="B914FC28FFF7FF9203BA15A9FD40F860" bold="true" box="[650,707,1987,2013]" pageId="37" pageNumber="38">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
also has an extra subpodial lobe on posterior thoracic setigers.
|
||
<taxonomicName id="4C605BB9FFFBFF9E01A712FDFEA0FF0C" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[151,291,151,177]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="41" pageNumber="42" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFFBFF9E01A712FDFEA0FF0C" box="[151,291,151,177]" italics="true" pageId="41" pageNumber="42">S. papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFFBFF9E001A12FDFEEAFF0C" box="[298,361,151,177]" pageId="41" pageNumber="42" rank="species">
|
||
<emphasis id="B914FC28FFFBFF9E001A12FDFEE2FF0C" bold="true" box="[298,353,151,177]" pageId="41" pageNumber="42">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has 17–18 thoracic setigers and branchiae from setigers 13–14, whereas
|
||
<taxonomicName id="4C605BB9FFFBFF9E05A912FDFACEFF0C" authorityName="Blake" authorityYear="2020" box="[1177,1357,151,177]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="41" pageNumber="42" phylum="Annelida" rank="species" species="californiensis">
|
||
<emphasis id="B914FC28FFFBFF9E05A912FDFACEFF0C" box="[1177,1357,151,177]" italics="true" pageId="41" pageNumber="42">S. californiensis</emphasis>
|
||
</taxonomicName>
|
||
has 12 thoracic setigers and branchiae from setigers 11–12.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1F70B2FFF4FF9101A715FDFD8AF84A" ID-DOI="http://doi.org/10.5281/zenodo.4544931" ID-Zenodo-Dep="4544931" httpUri="https://zenodo.org/record/4544931/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,1943,1967]" targetBox="[193,1395,186,1911]" targetPageId="38">
|
||
<paragraph id="8BDF203AFFF4FF9101A715FDFD8AF84A" blockId="38.[151,1437,1943,2040]" pageId="38" pageNumber="39">
|
||
<emphasis id="B914FC28FFF4FF9101A715FDFE9BF812" bold="true" box="[151,280,1943,1967]" pageId="38" pageNumber="39">FIGURE 15</emphasis>
|
||
.
|
||
<taxonomicName id="4C605BB9FFF4FF91001415F2FE72F80D" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[292,497,1944,1968]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="38" pageNumber="39" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF4FF91001415F2FE72F80D" box="[292,497,1944,1968]" italics="true" pageId="38" pageNumber="39">Scoloplos papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF4FF9100C615F2FDADF812" box="[502,558,1944,1968]" pageId="38" pageNumber="39" rank="species">
|
||
<emphasis id="B914FC28FFF4FF9100C615F2FDABF80D" bold="true" box="[502,552,1944,1968]" pageId="38" pageNumber="39">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Paratype (USNM 1621019): A, anterior end, dorsal view; B, anterior end, ventral view; C, pre-setiger region, ventral view; D, thoracic setiger 7, anterior view; E, thoracic setiger 15, posterior view; F–G, thoracic neuropodial uncini; H, furcate setae.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F70B2FFF5FF9001A7152CFBA0F81B" ID-DOI="http://doi.org/10.5281/zenodo.4544933" ID-Zenodo-Dep="4544933" httpUri="https://zenodo.org/record/4544933/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1862,1886]" targetBox="[161,1423,185,1824]" targetPageId="39">
|
||
<paragraph id="8BDF203AFFF5FF9001A7152CFBA0F81B" blockId="39.[151,1436,1862,1959]" pageId="39" pageNumber="40">
|
||
<emphasis id="B914FC28FFF5FF9001A7152CFE9BF8E3" bold="true" box="[151,280,1862,1886]" pageId="39" pageNumber="40">FIGURE 16</emphasis>
|
||
.
|
||
<taxonomicName id="4C605BB9FFF5FF900014152CFE72F8E3" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[292,497,1862,1886]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFF5FF900014152CFE72F8E3" box="[292,497,1862,1886]" italics="true" pageId="39" pageNumber="40">Scoloplos papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFF5FF9000C6152CFDADF8E3" box="[502,558,1862,1886]" pageId="39" pageNumber="40" rank="species">
|
||
<emphasis id="B914FC28FFF5FF9000C6152CFDABF8E3" bold="true" box="[502,552,1862,1886]" pageId="39" pageNumber="40">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A, anterior abdominal setiger, posterior view with inset showing separate branchia with bifid tip; B, middle abdominal setiger, posterior view; C, far posterior abdominal setiger, posterior view; D, posterior end with pygidium, dorsal view. A–C, paratype (USNM 1621018); D, holotype (USNM 1621017).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F70B2FFFAFF9F01A71534FB56F85F" ID-DOI="http://doi.org/10.5281/zenodo.4544935" ID-Zenodo-Dep="4544935" httpUri="https://zenodo.org/record/4544935/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1886,1910]" targetBox="[217,1372,189,1855]" targetPageId="40">
|
||
<paragraph id="8BDF203AFFFAFF9F01A71534FB56F85F" blockId="40.[151,1437,1886,2018]" pageId="40" pageNumber="41">
|
||
<emphasis id="B914FC28FFFAFF9F01A71534FE9BF8CB" bold="true" box="[151,280,1886,1910]" pageId="40" pageNumber="41">FIGURE 17</emphasis>
|
||
.
|
||
<taxonomicName id="4C605BB9FFFAFF9F00141534FE72F8CB" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[292,497,1886,1910]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="40" pageNumber="41" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFFAFF9F00141534FE72F8CB" box="[292,497,1886,1910]" italics="true" pageId="40" pageNumber="41">Scoloplos papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFFAFF9F00C71534FDACF8C8" box="[503,559,1886,1910]" pageId="40" pageNumber="41" rank="species">
|
||
<emphasis id="B914FC28FFFAFF9F00C71534FDAAF8CB" bold="true" box="[503,553,1886,1910]" pageId="40" pageNumber="41">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A, anterior end of holotype, dorsolateral view; B, anterior end of paratype ventrolateral view; C, furcate notoseta and camerated capillaries, abdominal setiger; D, anterior abdominal segments dorsal view with pigmented interramal swellings denoted with arrows; E, middle abdominal segments, lateral view; F, posterior end and pygidium. A, D–F, holotype, (USNM 1621017); B–C, G paratype, (USNM 1621018). A–F stained with Shirlastain A.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37A73B1FFFBFF9E01F712B4FD98FE48" pageId="41" pageNumber="42" type="biology_ecology">
|
||
<paragraph id="8BDF203AFFFBFF9E01F712B4FD98FE48" blockId="41.[151,1437,151,609]" pageId="41" pageNumber="42">
|
||
<emphasis id="B914FC28FFFBFF9E01F712B4FEA2FF45" bold="true" box="[199,289,222,248]" pageId="41" pageNumber="42">Biology</emphasis>
|
||
. The upper continental slope off
|
||
<collectingRegion id="49A4EED8FFFBFF9E03A412B5FD4DFF44" box="[660,718,223,249]" country="South Africa" name="Western Cape" pageId="41" pageNumber="42">Cape</collectingRegion>
|
||
Hatteras,
|
||
<collectingRegion id="49A4EED8FFFBFF9E020F12B5FC69FF44" box="[831,1002,223,249]" country="United States of America" name="North Carolina" pageId="41" pageNumber="42">North Carolina</collectingRegion>
|
||
, where
|
||
<taxonomicName id="4C605BB9FFFBFF9E057112B5FAA7FF44" authority="Blake, 2021" authorityName="Blake" authorityYear="2021" box="[1089,1316,223,249]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="41" pageNumber="42" phylum="Annelida" rank="species" species="papillatus" status="sp. nov.">
|
||
<emphasis id="B914FC28FFFBFF9E057112B5FAA7FF44" box="[1089,1316,223,249]" italics="true" pageId="41" pageNumber="42">Scoloplos papillatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2274153FFFBFF9E041B12B5FAE9FF44" box="[1323,1386,223,249]" pageId="41" pageNumber="42" rank="species">
|
||
<emphasis id="B914FC28FFFBFF9E041B12B5FAE1FF44" bold="true" box="[1323,1378,223,249]" pageId="41" pageNumber="42">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
was discovered is unusual in having high infaunal densities of 24,055 to 61,244 (mean
|
||
<emphasis id="B914FC28FFFBFF9E05601369FBE0FEA1" box="[1104,1123,259,284]" italics="true" pageId="41" pageNumber="42">=</emphasis>
|
||
46,255) individuals per m
|
||
<superScript id="7C158D72FFFBFF9E04A41368FA1FFEAD" attach="left" box="[1428,1436,258,272]" fontSize="6" pageId="41" pageNumber="42">2</superScript>
|
||
(
|
||
<bibRefCitation id="EFF15DCBFFFBFF9E01AF134DFE17FEFC" author="Blake, J. A. & Hilbig, B." box="[159,404,295,321]" pageId="41" pageNumber="42" pagination="875 - 899" refId="ref82852" refString="Blake, J. A. & Hilbig, B. (1994) Dense infaunal assemblages on the continental slope off Cape Hatteras, North Carolina. Deep- Sea Research II, 41, 875 - 899. https: // doi. org / 10.1016 / 0967 - 0645 (94) 90052 - 3" type="journal article" year="1994">Blake & Hilbig 1994</bibRefCitation>
|
||
). The fauna is dominated by polychaetes such as
|
||
<taxonomicName id="4C605BB9FFFBFF9E02F7134DFAD8FEFC" authority="Rathke, 1843" authorityName="Rathke" authorityYear="1843" box="[967,1371,295,321]" class="Polychaeta" family="Scalibregmatidae" genus="Scalibregma" kingdom="Animalia" pageId="41" pageNumber="42" phylum="Annelida" rank="species" species="inflatum">
|
||
<emphasis id="B914FC28FFFBFF9E02F7134DFB3AFEFC" box="[967,1209,295,321]" italics="true" pageId="41" pageNumber="42">Scalibregma inflatum</emphasis>
|
||
Rathke, 1843
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C605BB9FFFBFF9E0457134DFD09FED8" authority="Webster & Benedict, 1887" authorityName="Webster & Benedict" authorityYear="1887" class="Polychaeta" family="Cossuridae" genus="Cossura" kingdom="Animalia" pageId="41" pageNumber="42" phylum="Annelida" rank="species" species="longocirrata">
|
||
<emphasis id="B914FC28FFFBFF9E0457134DFEDEFED8" italics="true" pageId="41" pageNumber="42">Cossura longocirrata</emphasis>
|
||
Webster & Benedict, 1887
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C605BB9FFFBFF9E03F31321FB6AFED8" authority="Webster & Benedict, 1887" authorityName="Webster & Benedict" authorityYear="1887" box="[707,1257,331,357]" class="Polychaeta" family="Paraonidae" genus="Aricidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Spionida" pageId="41" pageNumber="42" phylum="Annelida" rank="species" species="quadrilobata">
|
||
<emphasis id="B914FC28FFFBFF9E03F31321FC38FED8" box="[707,955,331,357]" italics="true" pageId="41" pageNumber="42">Aricidea quadrilobata</emphasis>
|
||
Webster & Benedict, 1887
|
||
</taxonomicName>
|
||
, species that are more typical of shelf depths. Rhoads & Hecker (1994) found that organic matter in the
|
||
<collectingRegion id="49A4EED8FFFBFF9E056D1305FB14FE34" box="[1117,1175,367,393]" country="South Africa" name="Western Cape" pageId="41" pageNumber="42">Cape</collectingRegion>
|
||
Hatteras sediments is a mixture of both marine and terrestrially derived carbon that is more typical of continental shelf sediments, suggesting that the high percentage of refractory organic matter favors the survival of preadapted shelf species over those from adjacent slope environments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A73B1FFFBFF9E01F71395FD32FD80" pageId="41" pageNumber="42" type="etymology">
|
||
<paragraph id="8BDF203AFFFBFF9E01F71395FD32FD80" blockId="41.[151,1437,151,609]" pageId="41" pageNumber="42">
|
||
<emphasis id="B914FC28FFFBFF9E01F71395FECBFDA4" bold="true" box="[199,328,511,537]" pageId="41" pageNumber="42">Etymology</emphasis>
|
||
. Latin, for
|
||
<emphasis id="B914FC28FFFBFF9E00F41395FD90FDA4" box="[452,531,511,537]" italics="true" pageId="41" pageNumber="42">papilla</emphasis>
|
||
, a nipple or bud, referring to the subpodial papillae that occur on upper and lower margins of the subpodial flanges of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A73B1FFFBFF9E01F7102CFB12FDDC" box="[199,1169,582,609]" pageId="41" pageNumber="42" type="distribution">
|
||
<paragraph id="8BDF203AFFFBFF9E01F7102CFB12FDDC" blockId="41.[151,1437,151,609]" box="[199,1169,582,609]" pageId="41" pageNumber="42">
|
||
<emphasis id="B914FC28FFFBFF9E01F7102CFEDAFDDD" bold="true" box="[199,345,582,608]" pageId="41" pageNumber="42">Distribution</emphasis>
|
||
. Off
|
||
<collectingRegion id="49A4EED8FFFBFF9E00A5102DFE4CFDDC" box="[405,463,583,609]" country="South Africa" name="Western Cape" pageId="41" pageNumber="42">Cape</collectingRegion>
|
||
Hatteras,
|
||
<collectingRegion id="49A4EED8FFFBFF9E0371102DFD6FFDDC" box="[577,748,583,609]" country="United States of America" name="North Carolina" pageId="41" pageNumber="42">North Carolina</collectingRegion>
|
||
, upper continental slope,
|
||
<quantity id="4C988DDFFFFBFF9E053B102DFB0EFDDC" box="[1035,1165,583,609]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" metricValueMax="8.12" metricValueMin="6.2" pageId="41" pageNumber="42" unit="m" value="716.0" valueMax="812.0" valueMin="620.0">
|
||
620–
|
||
<specimenCount id="9D66EBB3FFFBFF9E0573102DFB0EFDDC" box="[1091,1165,583,609]" pageId="41" pageNumber="42" type="generic">812 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |