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<document id="41EE4A6AE6F6E61277AE3B8AE0B9F794" ID-CLB-Dataset="5556" ID-DOI="10.11646/zootaxa.4991.1.4" ID-GBIF-Dataset="8676cee2-bcd9-4c05-8420-af7951a7c387" ID-ISSN="1175-5326" ID-Zenodo-Dep="5027542" ID-ZooBank="6C803B9C-9C70-4C3D-AEFD-C085072789CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624550476717" checkinUser="plazi" docAuthor="Heller, Klaus-Gerhard, Puskás, Gellért, Szövényi, Gergely &amp; Chobanov, Dragan P." docDate="2021" docId="E041878C7133FFEE06B4FC44166DF895" docLanguage="en" docName="zootaxa.4991.1.4.pdf" docOrigin="Zootaxa 4991 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Uromenus innocentii" docType="treatment" docVersion="9" lastPageNumber="104" masterDocId="1C78FFF47138FFE50623FFCF1717FFD9" masterDocTitle="Songs in the genus Uromenus (Orthoptera: Tettigoniidae: Ephippigerini): A review with new information about some species" masterLastPageNumber="115" masterPageNumber="93" pageNumber="104" updateTime="1699020347932" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C0E06F3EBBA269D9DEF18607364CC6B6">Songs in the genus Uromenus (Orthoptera: Tettigoniidae: Ephippigerini): A review with new information about some species</mods:title>
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<mods:namePart id="859DB7645E2E8AADC9A9508B7183208B">Heller, Klaus-Gerhard</mods:namePart>
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<mods:affiliation id="FE4BC78136E7BF8239D96F7525AB6B45">Nagy Fuvaros u. 6, 1084 Budapest, Hungary; saksup @ gmail. com; https: // orcid. org / 0000 - 0001 - 8008 - 749 X</mods:affiliation>
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<mods:namePart id="B77178AF959E94B577F2216C2341703A">Szövényi, Gergely</mods:namePart>
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<mods:affiliation id="1BCFBBA9BE5C2F801D85AB1F24C9F8BE">Department of Systematic Zoology and Ecology, ELTE, Eötvös Loránd University, Budapest, 1117, Hungary; gergely. szovenyi @ ttk. elte. hu; https: // orcid. org / 0000 - 0001 - 9632 - 4066</mods:affiliation>
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<mods:namePart id="8AD5DCBDD256C84CE176AEF312B0D165">Chobanov, Dragan P.</mods:namePart>
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<mods:affiliation id="3E982DEC28E1D3322F6D8575334ACE45">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Boulevard, 1000 Sofia, Bulgaria. dchobanov @ gmail. com; https: // orcid. org / 0000 - 0002 - 1642 - 0363</mods:affiliation>
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<taxonomicName id="AFE84D197133FFEE06B4FC44168FFC7D" ID-CoL="7DVMN" baseAuthorityName="E.Bonnet &amp; Finot" baseAuthorityYear="1885" box="[151,408,906,933]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="104" phylum="Arthropoda" rank="species" species="innocentii">
<emphasis id="5A9CEA887133FFEE06B4FC44168FFC7D" bold="true" box="[151,408,906,933]" italics="true" pageId="11" pageNumber="104">Uromenus innocentii</emphasis>
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(Finot &amp; Bonnet in
<bibRefCitation id="0C794B6B7133FFEE04B7FC45148CFC7D" author="Bonnet, E. &amp; Finot, A." box="[660,923,906,932]" pageId="11" pageNumber="104" pagination="193 - 232" refId="ref9037" refString="Bonnet, E. &amp; Finot, A. (1885) Les Orthopteres de la regence de Tunis. Revue des Sciences Naturelles, Montpellier, Series 3, 4, 193 - 232 + 333 - 465." type="journal article" year="1885">Bonnet &amp; Finot 1885</bibRefCitation>
)
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<materialsCitation id="D8803CC77133FFEE06B4FC1D13FBFC35" ID-GBIF-Occurrence="3320086301" box="[151,1260,978,1005]" collectingDate="2013-06-03" collectorName="Chobanov." country="Morocco" elevation="730" latitude="34.46791" location="Mezguitem" longLatPrecision="1" longitude="-3.68883" pageId="11" pageNumber="104" specimenCount="1">
<collectingCountry id="10FF760A7133FFEE06B4FC1D163FFC35" box="[151,296,978,1004]" name="Morocco" pageId="11" pageNumber="104">MOROCCO</collectingCountry>
, S of
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,
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,
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,
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a.s.l.
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,
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<collectingDate id="0C12E9B27133FFEE059EFC1D1320FC35" box="[957,1079,978,1004]" pageId="11" pageNumber="104" value="2013-06-03">03 vi 2013</collectingDate>
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, leg.
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<paragraph id="6857369A7133FFEE06E4FC3916CAFB81" blockId="11.[151,1437,978,1869]" pageId="11" pageNumber="104">
The nightly calling song consists of echemes with 28±2 (n=15) syllables (
<figureCitation id="F0D32A1F7133FFEE05D7FC39132FFBC9" box="[1012,1080,1014,1040]" captionStart="FIGURE 2" captionStartId="4.[151,250,1977,2001]" captionTargetBox="[163,1428,327,1941]" captionTargetId="figure-107@4.[151,1436,309,1954]" captionTargetPageId="4" captionText="FIGURE 2. Oscillograms of male calling songs of the Uromenus species studied here, 10 second sections." figureDoi="http://doi.org/10.5281/zenodo.5027550" httpUri="https://zenodo.org/record/5027550/files/figure.png" pageId="11" pageNumber="104">Fig. 2</figureCitation>
; syllable period 129±4 ms; SRR 7.7 Hz). In these syllables, lasting 84±3 ms, probably only during the closing movement of the tegmina impulses (19±2) are produced (
<figureCitation id="F0D32A1F7133FFEE07A8FBF116D8FB81" box="[395,463,1086,1112]" captionStart="FIGURE 3" captionStartId="5.[151,250,1924,1948]" captionTargetBox="[157,1424,193,1889]" captionTargetId="figure-19@5.[151,1436,181,1901]" captionTargetPageId="5" captionText="FIGURE 3. Oscillograms of male calling songs of Uromenus species studied here, 2 second sections." figureDoi="http://doi.org/10.5281/zenodo.5027558" httpUri="https://zenodo.org/record/5027558/files/figure.png" pageId="11" pageNumber="104">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="6857369A7133FFEE06E4FBAD13B1FBA4" blockId="11.[151,1437,978,1869]" box="[199,1190,1122,1149]" pageId="11" pageNumber="104">
The frequency spectrum of the male song has a broad maximum around 20 kHz (
<figureCitation id="F0D32A1F7133FFEE0277FBAD138FFBA4" box="[1108,1176,1122,1149]" captionStart="FIGURE 5" captionStartId="7.[151,260,1447,1473]" captionTargetBox="[161,1427,187,1422]" captionTargetId="figure-19@7.[151,1436,179,1423]" captionTargetPageId="7" captionText="FIGURE 5. Power spectra of the calling songs of the Uromenus species studied here." figureDoi="http://doi.org/10.5281/zenodo.5027564" httpUri="https://zenodo.org/record/5027564/files/figure.png" pageId="11" pageNumber="104">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="6857369A7133FFEE06E4FB4916FFFAA1" blockId="11.[151,1437,978,1869]" pageId="11" pageNumber="104">
The tegmina (
<figureCitation id="F0D32A1F7133FFEE0741FB4916A2FB78" box="[354,437,1158,1185]" captionStart="FIGURE 6" captionStartId="9.[151,260,1205,1231]" captionTargetBox="[161,1432,186,1177]" captionTargetId="figure-19@9.[151,1436,179,1181]" captionTargetPageId="9" captionText="FIGURE 6. Male tegmina in Uromenus. A—U. elegans (Crete), B—U. dyrrhachiacus, C—U. finoti, D—U. galvagnii, E—U. robustus, F—U. tobboganensis and G—U. innocentii." figureDoi="http://doi.org/10.5281/zenodo.5027566" httpUri="https://zenodo.org/record/5027566/files/figure.png" pageId="11" pageNumber="104">Fig. 6F</figureCitation>
) differ in shape slightly from the other studied species. In the posterior region of the right tegmen there is a network of cells only slightly smaller than the small roundish mirror. The same pattern can be seen in the left tegmen. The male stridulatory file carries 56 teeth (
<figureCitation id="F0D32A1F7133FFEE051DFB011495FB30" box="[830,898,1230,1257]" captionStart="FIGURE 7" captionStartId="9.[151,260,1961,1987]" captionTargetBox="[154,1433,1292,1934]" captionTargetId="figure-59@9.[151,1436,1289,1937]" captionTargetPageId="9" captionText="FIGURE 7. Male stridulatory files in Uromenus. A—U. elegans (Crete), B—U. dyrrhachiacus, C—U. finoti, D—U. galvagnii, E—U. robustus, F—U. tobboganensis, G—U. melillae, H—U. innocentii, I—U. choumarae." figureDoi="http://doi.org/10.5281/zenodo.5027572" httpUri="https://zenodo.org/record/5027572/files/figure.png" pageId="11" pageNumber="104">Fig. 7</figureCitation>
;
<tableCitation id="256A03217133FFEE05AEFB0114C4FB31" box="[909,979,1230,1256]" captionStart="TABLE 2" captionStartId="16.[151,239,151,175]" captionTargetPageId="16" captionText="TABLE 2. Measurements of stridulatory files of the Uromenus species studied here" httpUri="http://table.plazi.org/id/3C9766127128FFF506B4FF581303FF76" pageId="11" pageNumber="104" tableUuid="3C9766127128FFF506B4FF581303FF76">Tab. 2</tableCitation>
) a distinctly lower number than the other species of the genus (except
<taxonomicName id="AFE84D197133FFEE07F1FB3C1520FAD5" box="[466,567,1267,1292]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="104" phylum="Arthropoda" rank="species" species="annae">
<emphasis id="5A9CEA887133FFEE07F1FB3C1520FAD5" box="[466,567,1267,1292]" italics="true" pageId="11" pageNumber="104">U. annae</emphasis>
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). In the middle of the file the inter-tooth intervals are about
<quantity id="AF109B7F7133FFEE02E8FB3D1201FAD4" box="[1227,1302,1266,1293]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="11" pageNumber="104" unit="mm" value="83.0">83 µm</quantity>
. The female stridulatory file (
<figureCitation id="F0D32A1F7133FFEE0770FAD916B1FAE9" box="[339,422,1302,1328]" captionStart="FIGURE 9" captionStartId="13.[151,260,1850,1876]" captionTargetBox="[158,1425,354,1810]" captionTargetId="figure-98@13.[151,1436,342,1825]" captionTargetPageId="13" captionText="FIGURE 9. Female tegmina in Uromenus. A—U. elegans (Crete), B—U. dyrrhachiacus, C—U. rugosicollis, D—U. agarenus, E—U. finoti, F—U. tobboganensis and G—U. innocentii. Note the long stridulatory vein on the upper side of the right tegmen crossing the mirror-like translucent tegmen membrane." figureDoi="http://doi.org/10.5281/zenodo.5027578" httpUri="https://zenodo.org/record/5027578/files/figure.png" pageId="11" pageNumber="104">Fig. 9F</figureCitation>
) is less strongly developed than in the other species. In the lateral part the teeth are smaller than in the central part which itself is quite short. In the inner part there are more net-like veins mostly covered with pegs than in the other species.
</paragraph>
<paragraph id="6857369A7133FFEE06E4FA4D147AF965" blockId="11.[151,1437,978,1869]" pageId="11" pageNumber="104">
<emphasis id="5A9CEA887133FFEE06E4FA4D1676FA45" bold="true" box="[199,353,1410,1436]" pageId="11" pageNumber="104">Morphology.</emphasis>
The species was described under
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<emphasis id="5A9CEA887133FFEE04C3FA4D1479FA45" box="[736,878,1410,1436]" italics="true" pageId="11" pageNumber="104">Steropleurus</emphasis>
</taxonomicName>
and considered by
<bibRefCitation id="0C794B6B7133FFEE0269FA4D13F6FA44" author="Nadig, A." box="[1098,1249,1410,1437]" pageId="11" pageNumber="104" pagination="109 - 183" refId="ref10461" refString="Nadig, A. (1995) Die marokkanischen Arten der Gattung Steropleurus Bolivar, 1878 (Orthoptera Ephippigeridae). Atti dell'Accademia Roveretana degli Agiati, Atti anni accademici, 244 [1994] 4 B, 109 - 183." type="journal article" year="1995">Nadig (1995)</bibRefCitation>
forming its own group. The species has a very typical dorsal pattern of dark green, ochre to whitish, and darker (brownish-violet) spots and stripes (
<figureCitation id="F0D32A1F7133FFEE0743FA0516A0FA3D" box="[352,439,1482,1508]" captionStart="FIGURE 1" captionStartId="2.[151,250,1871,1895]" captionTargetBox="[151,1436,561,1848]" captionTargetId="figure-106@2.[151,1436,561,1848]" captionTargetPageId="2" captionText="FIGURE 1. General appearance and habitats of Uromenus species. A—U. elegans (Italy, Lazio), BD—U. elegans (Crete; photos B and C taken by Ionuţ Iorgu), E, F—U. dyrrhachiacus, G—U. agarenus, HJ—U. finoti, K, L—U. robustus, M, N—U. tobboganensis, O—U. maroccanus, P, Q—U. melillae, R—U. innocentii, S, T—U. choumarae." figureDoi="http://doi.org/10.5281/zenodo.5027546" httpUri="https://zenodo.org/record/5027546/files/figure.png" pageId="11" pageNumber="104">Fig. 1R</figureCitation>
). The occiput is black (not uncommon in
<taxonomicName id="AFE84D197133FFEE05A8FA05133CFA3C" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[907,1067,1482,1509]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="104" phylum="Arthropoda" rank="tribe" tribe="Ephippigerini">Ephippigerini</taxonomicName>
). Male cerci have a typical shape with a long, sinuately outcurved apical part (after the internal tooth) (
<figureCitation id="F0D32A1F7133FFEE05BCFA211314F9D1" box="[927,1027,1518,1544]" captionStart="FIGURE 10" captionStartId="14.[151,260,1927,1953]" captionTargetBox="[173,1414,378,1902]" captionTargetId="figure-109@14.[173,1414,378,1903]" captionTargetPageId="14" captionText="FIGURE 10. Cerci of the Moroccan species of Uromenus studied here and of U. dyrrhachiacus. A—U. finoti, B—U. galvagnii, C—U. robustus, D—U. tobboganensis, E—U. melillae, F—U. innocentii, G—U. choumarae, H—U. dyrrhachiacus" figureDoi="http://doi.org/10.5281/zenodo.5027580" httpUri="https://zenodo.org/record/5027580/files/figure.png" pageId="11" pageNumber="104">Fig. 10F</figureCitation>
<figureCitation id="F0D32A1F7133FFEE0220FA21133BF9D1" box="[1027,1068,1518,1544]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1871,1895]" captionStartId-1="4.[151,250,1977,2001]" captionTargetBox-0="[151,1436,561,1848]" captionTargetBox-1="[163,1428,327,1941]" captionTargetId-0="figure-106@2.[151,1436,561,1848]" captionTargetId-1="figure-107@4.[151,1436,309,1954]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. General appearance and habitats of Uromenus species. A—U. elegans (Italy, Lazio), BD—U. elegans (Crete; photos B and C taken by Ionuţ Iorgu), E, F—U. dyrrhachiacus, G—U. agarenus, HJ—U. finoti, K, L—U. robustus, M, N—U. tobboganensis, O—U. maroccanus, P, Q—U. melillae, R—U. innocentii, S, T—U. choumarae." captionText-1="FIGURE 2. Oscillograms of male calling songs of the Uromenus species studied here, 10 second sections." figureDoi-0="http://doi.org/10.5281/zenodo.5027546" figureDoi-1="http://doi.org/10.5281/zenodo.5027550" httpUri-0="https://zenodo.org/record/5027546/files/figure.png" httpUri-1="https://zenodo.org/record/5027550/files/figure.png" pageId="11" pageNumber="104">12</figureCitation>
). We follow
<bibRefCitation id="0C794B6B7133FFEE0298FA20125DF9D1" author="Barat, J." box="[1211,1354,1518,1545]" pageId="11" pageNumber="104" pagination="331 - 334" refId="ref8887" refString="Barat, J. (2017) On the presence of Uromenus innocentii (Finot &amp; Bonnet, 1885) in the western sector of the Sahara desert (Orthoptera, Tettigoniidae, Ephippigerini). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 60, 331 - 334." type="journal article" year="2017">Barat (2017)</bibRefCitation>
and do not recognise subspecies in this widespread species. Our specimens differ in tooth number distinctly from Nadigs Central Moroccan specimens (56 vs 8090 teeth;
<bibRefCitation id="0C794B6B7133FFEE0498F9F91429F989" author="Nadig, A." box="[699,830,1590,1616]" pageId="11" pageNumber="104" pagination="919 - 1016" refId="ref10416" refString="Nadig, A. (1994) Revision der Gattung Uromenus Bolivar, 1878 (Orthoptera: Ephippigeridae). Revue Suisse de Zoologie, 101, 919 - 1016. https: // doi. org / 10.5962 / bhl. part. 79936" type="journal article" year="1994">Nadig 1994</bibRefCitation>
) but the type localities from both described subspecies are situated East of
<collectingCountry id="10FF760A7133FFEE0759F99416F4F9AC" box="[378,483,1627,1653]" name="Morocco" pageId="11" pageNumber="104">Morocco</collectingCountry>
. The
<typeStatus id="B75388387133FFEE0401F994156EF9AC" box="[546,633,1627,1653]" pageId="11" pageNumber="104" type="syntype">syntype</typeStatus>
of
<emphasis id="5A9CEA887133FFEE0482F994145BF9AD" box="[673,844,1627,1652]" italics="true" pageId="11" pageNumber="104">
U. i.
<taxonomicName id="AFE84D197133FFEE04FFF994145BF9AD" baseAuthorityName="E.Bonnet &amp; Finot" baseAuthorityYear="1885" box="[732,844,1627,1652]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="104" phylum="Arthropoda" rank="species" species="innocentii">innocentii</taxonomicName>
</emphasis>
in Paris bears a label (
<bibRefCitation id="0C794B6B7133FFEE0271F9951220F9AD" author="Cigliano, M. &amp; Braun, H. &amp; Eades, D. &amp; Otte, D." box="[1106,1335,1626,1653]" pageId="11" pageNumber="104" refId="ref9522" refString="Cigliano, M., Braun, H., Eades, D. &amp; Otte, D. (2021) Orthoptera Species File. Version 5.0 / 5.0. Available from: http: // Orthoptera. SpeciesFile. org / (accessed 30 June 2021)" type="url" year="2021">
Cigliano
<emphasis id="5A9CEA887133FFEE029EF99413F8F9AD" box="[1213,1263,1626,1652]" italics="true" pageId="11" pageNumber="104">et al</emphasis>
. 2021
</bibRefCitation>
) reading „
<collectingCountry id="10FF760A7133FFEE0682F9B117EEF941" box="[161,249,1662,1688]" name="Tunisia" pageId="11" pageNumber="104">Tunisie</collectingCountry>
, Djebel Berda“, now in
<collectingCountry id="10FF760A7133FFEE0425F9B1154DF941" box="[518,602,1662,1688]" name="Algeria" pageId="11" pageNumber="104">Algeria</collectingCountry>
in the Aures Mts., while
<emphasis id="5A9CEA887133FFEE0555F9B01317F941" box="[886,1024,1662,1688]" italics="true" pageId="11" pageNumber="104">U. i. lobatus</emphasis>
<bibRefCitation id="0C794B6B7133FFEE022BF9B113A6F941" author="Saussure, H. de" box="[1032,1201,1662,1688]" pageId="11" pageNumber="104" pagination="183 - 250" refId="ref10909" refString="Saussure, H. de (1898) Analecta entomologica I. Orthopterologica. Revue Suisse de Zoologie, 5, 183 - 250 + 787 - 809. [1897] https: // doi. org / 10.5962 / bhl. part. 117929" type="journal article" year="1898">Saussure, 1898</bibRefCitation>
was collected in Aïn Séfra,
<collectingCountry id="10FF760A7133FFEE06FCF96D1621F965" box="[223,310,1698,1724]" name="Algeria" pageId="11" pageNumber="104">Algeria</collectingCountry>
, in the Saharan Atlas near to the Moroccan border.
</paragraph>
</subSubSection>
<subSubSection id="20F265117133FFEE06E4F909166DF895" pageId="11" pageNumber="104" type="biology_ecology">
<paragraph id="6857369A7133FFEE06E4F909166DF895" blockId="11.[151,1437,978,1869]" pageId="11" pageNumber="104">
<emphasis id="5A9CEA887133FFEE06E4F909163BF939" bold="true" box="[199,300,1734,1760]" pageId="11" pageNumber="104">Ecology.</emphasis>
The species occurs widely within the Mediterranean climate zone reaching even far south the Sahara desert (Hot desert climate) (
<bibRefCitation id="0C794B6B7133FFEE07F7F9251540F8DD" author="Barat, J." box="[468,599,1770,1796]" pageId="11" pageNumber="104" pagination="331 - 334" refId="ref8887" refString="Barat, J. (2017) On the presence of Uromenus innocentii (Finot &amp; Bonnet, 1885) in the western sector of the Sahara desert (Orthoptera, Tettigoniidae, Ephippigerini). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 60, 331 - 334." type="journal article" year="2017">Barat 2017</bibRefCitation>
). Our observation is from a dry ruderal habitat within the Köppen climate
<typeStatus id="B75388387133FFEE06B4F8C017D0F8F0" box="[151,199,1807,1833]" pageId="11" pageNumber="104">type</typeStatus>
BSk (Cold semi-arid) towards BWk (Cold desert) in the climatic shadow of the westwards rising Rif mountains (
<bibRefCitation id="0C794B6B7133FFEE06BDF8FD1679F895" author="Verner, D. &amp; Treguer, D. &amp; Redwood, J. &amp; Christensen, J. &amp; McDonnell, R. &amp; Elbert, C. &amp; Konishi, Y. &amp; Belghazi, S." box="[158,366,1842,1869]" pageId="11" pageNumber="104" refId="ref11113" refString="Verner, D., Treguer, D., Redwood, J., Christensen, J., McDonnell, R., Elbert, C., Konishi, Y. &amp; Belghazi, S. (2018) Climate Variability, Drought, and Drought Management in Morocco's Agricultural Sector. World Bank Group, Washington, D. C. Available from: http: // documents. worldbank. org / curated / en / 353801538414553978 / Climate-Variability-Drought-and-Drought- Management-in-Moroccos-Agricultural-Sector (accessed 8 June 2021) https: // doi. org / 10.1596 / 30603" type="url" year="2018">
Verner
<emphasis id="5A9CEA887133FFEE06D1F8FC1634F895" box="[242,291,1842,1868]" italics="true" pageId="11" pageNumber="104">et al</emphasis>
. 2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>