309 lines
37 KiB
XML
309 lines
37 KiB
XML
<document id="85B0F95F992282A8AF0EA42EF6292A5C" ID-CLB-Dataset="46000" ID-DOI="10.11646/zootaxa.3786.3.6" ID-GBIF-Dataset="839c26e6-bb6f-4258-84f3-75174e7fabdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="250549" ID-ZooBank="98B6A0EF-CAFC-4A78-B044-C5B9D6136CFB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460408750808" checkinUser="plazi" docAuthor="Grismer, L. Lee, Wood, Perry L., Anuar, Shahrul, Quah, Evan S. H., Muin, Mohd Abdul, Mohamed, Maketab, Onn, Chan Kin, Sumarli, Alexandra X., Loredo, Ariel I. & Heinz, Heather M." docDate="2014" docId="6E5E8796FFBEFF8DFF41FA8A25A8F8AB" docLanguage="en" docName="zt03786p381.pdf" docOrigin="Zootaxa 3786 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus timur Grismer, Wood, Anuar, Quah, Muin, Mohamed, Onn, Sumarli, Loredo & Heinz, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="377" masterDocId="9267FFEEFFAEFF9FFFD6FFBB2030FFE6" masterDocTitle="The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia" masterLastPageNumber="381" masterPageNumber="359" pageNumber="375" updateTime="1698300232766" updateUser="ExternalLinkService">
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<mods:title id="CC1CB2B32D17889DAEE2506385F329C4">The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia</mods:title>
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<mods:namePart id="774B908E83B6832571A438ED7CF30691">Grismer, L. Lee</mods:namePart>
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<mods:namePart id="2ED39C877B72EC1C4850716181F6C0E7">Wood, Perry L.</mods:namePart>
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<mods:namePart id="5B1A2168EEB72DE87074E266D352C60A">Anuar, Shahrul</mods:namePart>
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<mods:namePart id="74CBBDCDA177A3CBADAD5856EE916337">Quah, Evan S. H.</mods:namePart>
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<mods:namePart id="2571A1AEA4558EC699E9C7EEBC3EE6AA">Muin, Mohd Abdul</mods:namePart>
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<mods:namePart id="BD11FCF994CD237A9A9DD40568AB699A">Mohamed, Maketab</mods:namePart>
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<mods:namePart id="7CC51F1F3A60310BFAFC143969C1C340">Onn, Chan Kin</mods:namePart>
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<mods:namePart id="3653C7A900BEE0FFAB399CA131C324D9">Sumarli, Alexandra X.</mods:namePart>
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<mods:namePart id="C0CD27C7CEEC102524B946840F1EDEA3">Loredo, Ariel I.</mods:namePart>
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<mods:namePart id="C22214E4BB76C071F3513A07B92ECDE5">Heinz, Heather M.</mods:namePart>
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<mods:typeOfResource id="9DE83261B5B76180E3BC2ED7837133C7">text</mods:typeOfResource>
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<mods:title id="4DC0EC371D9023E3638138DD0A0EF3EE">Zootaxa</mods:title>
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<mods:part id="B90CC077494D9E399F6D4FF68D7A05C1">
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<mods:date id="9312FC37CAA7560E62513BCA64223E39">2014</mods:date>
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<mods:detail id="AAD77D16EDC8E1E656C9F44F43D38267" type="volume">
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<mods:number id="D82CA7567B46C067D5AA2A00655BB616">3786</mods:number>
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<mods:number id="62D4FC41FFF55F655D8F6F664808DAAD">3</mods:number>
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<mods:start id="FF99A1C9D4A52CB8364C665C4C0246C4">359</mods:start>
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<mods:end id="9B90B023288CD8ED4EB574BEF3419206">381</mods:end>
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<mods:classification id="5AC169C0C51DE48E0341F3ABB0BC2B45">journal article</mods:classification>
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<mods:identifier id="60A61BC4B08AD151E36EC572C9AC1F8A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="6E5E8796FFBEFF8DFF41FA8A25A8F8AB" ID-DOI="http://doi.org/10.5281/zenodo.6139287" ID-GBIF-Taxon="119422825" ID-Zenodo-Dep="6139287" LSID="urn:lsid:plazi:treatment:6E5E8796FFBEFF8DFF41FA8A25A8F8AB" httpUri="http://treatment.plazi.org/id/6E5E8796FFBEFF8DFF41FA8A25A8F8AB" lastPageId="18" lastPageNumber="377" pageId="16" pageNumber="375">
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<subSubSection id="AEED650BFFBEFF8FFF41FA8A20CFFA76" pageId="16" pageNumber="375" type="nomenclature">
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<paragraph id="E6483680FFBEFF8FFF41FA8A21DCFAAD" blockId="16.[151,525,1329,1424]" box="[151,492,1329,1355]" pageId="16" pageNumber="375">
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<heading id="BD0081ECFFBEFF8FFF41FA8A21DCFAAD" bold="true" box="[151,492,1329,1355]" fontSize="11" level="1" pageId="16" pageNumber="375" reason="1">
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<emphasis id="D483EA92FFBEFF8FFF41FA8A21DCFAAD" bold="true" box="[151,492,1329,1355]" pageId="16" pageNumber="375">
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||
<taxonomicName id="21F74D03FFBEFF8FFF41FA8A21B7FAAD" box="[151,391,1329,1355]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="375" phylum="Chordata" rank="species" species="timur" status="sp. nov.">
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<emphasis id="D483EA92FFBEFF8FFF41FA8A21B7FAAD" bold="true" box="[151,391,1329,1355]" italics="true" pageId="16" pageNumber="375">Cyrtodactylus timur</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CFB057E9FFBEFF8FFE58FA8A21DCFAAD" box="[398,492,1329,1355]" pageId="16" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E6483680FFBEFF8FFF41FAEF20CFFA76" blockId="16.[151,525,1329,1424]" pageId="16" pageNumber="375">
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||
Banjaran
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||
<taxonomicName id="21F74D03FFBEFF8FFED5FAEF2178FA8B" box="[259,328,1364,1389]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="379" phylum="Chordata" rank="species" species="timur" status="sp. nov.">Timur</taxonomicName>
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Bent-toed Gecko
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<figureCitation id="7ECC2A05FFBEFF8FFF41FACC20D4FA76" box="[151,228,1399,1424]" captionStart="FIGURE 5" captionStartId="17.[151,250,1519,1541]" captionTargetBox="[151,1435,645,1497]" captionTargetId="figure@17.[151,1436,645,1497]" captionTargetPageId="17" captionText="FIGURE 5. Upper left: adult male holotype (LSUHC 11207) of Cyrtotdactylus timur sp. nov. from Punca Air, Gunung Tebu, Terengganu, Peninsular Malaysia. Upper right: female paratype LSUHC 11185. Middle left: uncataloged hatching LSUDPC 8249. Lower left: adult male paratype LSUHC 11183. Lower right: juvenile female paratype LSUHC 11184." httpUri="https://zenodo.org/record/250554/files/figure.png" pageId="16" pageNumber="375">Figs. 5</figureCitation>
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,
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<figureCitation id="7ECC2A05FFBEFF8FFF27FACC20CFFA76" box="[241,255,1399,1424]" captionStart="FIGURE 6" captionStartId="19.[151,250,1728,1750]" captionTargetBox="[162,1436,193,1700]" captionTargetId="figure@19.[162,1447,193,1701]" captionTargetPageId="19" captionText="FIGURE 6. Microhabitat of Cyrtodactylus timur sp. nov. at the type locality of Punca Air, Gunung Tebu, Terengganu, Peninsular Malaysia." httpUri="https://zenodo.org/record/250555/files/figure.png" pageId="16" pageNumber="375">6</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="AEED650BFFBEFF8FFF41FA0423E6F9A2" pageId="16" pageNumber="375" type="materials_examined">
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<paragraph id="E6483680FFBEFF8FFF41FA0424F6FA1B" blockId="16.[151,1436,1471,2037]" pageId="16" pageNumber="375">
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<emphasis id="D483EA92FFBEFF8FFF41FA04213BFA3E" bold="true" box="[151,267,1471,1496]" pageId="16" pageNumber="375">
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<typeStatus id="394C8822FFBEFF8FFF41FA042136FA3E" box="[151,262,1471,1496]" pageId="16" pageNumber="375" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult male
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<collectionCode id="80E6AE45FFBEFF8FFE49FA7B21CBFA3E" box="[415,507,1472,1496]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="16" pageNumber="375">LSUHC</collectionCode>
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11207 collected on
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<date id="92491040FFBEFF8FFD3CFA7B2343FA3E" box="[746,883,1471,1496]" pageId="16" pageNumber="375" value="2013-07-01">1 July 2013</date>
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||
by Shahrul Anuar at 2350 hrs from Punca Air, Gunung Tebu, Terengganu, Peninsular
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<collectingCountry id="9EE07610FFBEFF8FFD9AFA5F2283FA1B" box="[588,691,1508,1533]" name="Malaysia" pageId="16" pageNumber="375">Malaysia</collectingCountry>
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(5°
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<geoCoordinate id="83C35047FFBEFF8FFD35FA5F230DFA1A" box="[739,829,1508,1533]" direction="north" orientation="latitude" pageId="16" pageNumber="375" precision="555" value="36.11">36.11 N</geoCoordinate>
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102°
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<geoCoordinate id="83C35047FFBEFF8FFC56FA5F23EAFA1A" box="[896,986,1508,1533]" direction="east" orientation="longitude" pageId="16" pageNumber="375" precision="555" value="36.19">36.19 E</geoCoordinate>
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; at
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<quantity id="210F9B65FFBEFF8FFBD7FA5F2478FA1A" box="[1025,1096,1508,1533]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="16" pageNumber="375" unit="m" value="650.0">650 m</quantity>
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elevation).
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</paragraph>
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<paragraph id="E6483680FFBEFF8FFF11F9BC23E6F9A2" blockId="16.[151,1436,1471,2037]" pageId="16" pageNumber="375">
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<emphasis id="D483EA92FFBEFF8FFF11F9BC2176F9C6" bold="true" box="[199,326,1543,1568]" pageId="16" pageNumber="375">
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<typeStatus id="394C8822FFBEFF8FFF11F9BC2171F9C6" box="[199,321,1543,1568]" pageId="16" pageNumber="375" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="80E6AE45FFBEFF8FFE86F9B3219CF9C6" box="[336,428,1544,1568]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="16" pageNumber="375">LSUHC</collectionCode>
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11183–85 have the same collection data as the
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<typeStatus id="394C8822FFBEFF8FFC0BF9BC2473F9C6" box="[989,1091,1543,1568]" pageId="16" pageNumber="375" type="holotype">holotype</typeStatus>
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.
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<collectionCode id="80E6AE45FFBEFF8FFB87F9B3249DF9C6" box="[1105,1197,1544,1568]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="16" pageNumber="375">LSUHC</collectionCode>
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11288 has the same collection data as the
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<typeStatus id="394C8822FFBEFF8FFE5FF99721DBF9A3" box="[393,491,1580,1605]" pageId="16" pageNumber="375" type="holotype">holotype</typeStatus>
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except that it was collected on
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<date id="92491040FFBEFF8FFC9AF99723E2F9A2" box="[844,978,1580,1605]" pageId="16" pageNumber="375" value="2013-07-02">2 July 2013</date>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEED650BFFBEFF8FFF11F9F424C9F8A6" pageId="16" pageNumber="375" type="diagnosis">
|
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<paragraph id="E6483680FFBEFF8FFF11F9F424C9F8A6" blockId="16.[151,1436,1471,2037]" pageId="16" pageNumber="375">
|
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<emphasis id="D483EA92FFBEFF8FFF11F9F42171F98E" bold="true" box="[199,321,1615,1640]" pageId="16" pageNumber="375">Diagnosis.</emphasis>
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<taxonomicName id="21F74D03FFBEFF8FFE9BF9F42202F98E" box="[333,562,1615,1640]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="375" phylum="Chordata" rank="species" species="timur" status="sp. nov.">
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<emphasis id="D483EA92FFBEFF8FFE9BF9F42202F98E" box="[333,562,1615,1640]" italics="true" pageId="16" pageNumber="375">Cyrtodactylus timur</emphasis>
|
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</taxonomicName>
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||
<taxonomicNameLabel id="CFB057E9FFBEFF8FFDE9F9F422AEF98E" box="[575,670,1615,1640]" pageId="16" pageNumber="375" rank="species">
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||
<emphasis id="D483EA92FFBEFF8FFDE9F9F422AEF98E" bold="true" box="[575,670,1615,1640]" pageId="16" pageNumber="375">sp. nov.</emphasis>
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||
</taxonomicNameLabel>
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can be differentiated from all other species of
|
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<taxonomicName id="21F74D03FFBEFF8FFB01F9F42544F98E" box="[1239,1396,1615,1640]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="375" phylum="Chordata" rank="genus">
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<emphasis id="D483EA92FFBEFF8FFB01F9F42544F98E" box="[1239,1396,1615,1640]" italics="true" pageId="16" pageNumber="375">Cyrtodactylus</emphasis>
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</taxonomicName>
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by having 10–12 supralabials; 8–10 infralabials; weak body tuberculation; no tubercles on ventral surface of forelimbs, gular region, or in ventrolateral body folds; 38–43 paravertebral tubercles; 21–24 longitudinal tubercle rows; 31–40 ventral scales; 21–25 subdigital lamellae on fourth toe; 21 or 22 femoroprecloacal pores; deep precloacal groove; four dark dorsal body bands; body band/interspace ratio 1.00–1.25; no scattered white tubercles on dorsum; 8–10 dark caudal bands on original tail; white caudal bands nearly immaculate; and a maximum SVL of
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<quantity id="210F9B65FFBEFF8FFF63F89C211AF8D9" box="[181,298,1831,1856]" metricMagnitude="-1" metricUnit="m" metricValue="1.205" pageId="16" pageNumber="375" unit="mm" value="120.5">120.5 mm</quantity>
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. These characters are scored across all species of the
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<taxonomicName id="21F74D03FFBEFF8FFCA9F8932423F8A6" box="[895,1043,1831,1856]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="375" phylum="Chordata" rank="species" species="pulchellus">
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<emphasis id="D483EA92FFBEFF8FFCA9F8932423F8A6" box="[895,1043,1831,1856]" italics="true" pageId="16" pageNumber="375">C. pulchellus</emphasis>
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</taxonomicName>
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complex in Table 5.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="AEED650BFFBEFF8DFF11F8F720C0FB23" lastPageId="18" lastPageNumber="377" pageId="16" pageNumber="375" type="description">
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<paragraph id="E6483680FFBEFF8EFF11F8F72310FD86" blockId="16.[151,1436,1471,2037]" lastBlockId="17.[151,1436,151,608]" lastPageId="17" lastPageNumber="376" pageId="16" pageNumber="375">
|
||
<emphasis id="D483EA92FFBEFF8FFF11F8F721D9F883" bold="true" box="[199,489,1868,1893]" pageId="16" pageNumber="375">
|
||
Description of
|
||
<typeStatus id="394C8822FFBEFF8FFEAAF8F721D5F883" box="[380,485,1868,1893]" pageId="16" pageNumber="375" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male SVL
|
||
<quantity id="210F9B65FFBEFF8FFD6DF8F72301F882" box="[699,817,1868,1892]" metricMagnitude="-1" metricUnit="m" metricValue="1.177" pageId="16" pageNumber="375" unit="mm" value="117.7">117.7 mm</quantity>
|
||
; head large, moderate in length (HL/SVL 0.28), wide (HW/HL 0.73), somewhat flattened (HD/HL 0.41), distinct from neck, triangular in dorsal profile; lores concave anteriorly, inflated posteriorly; frontal and prefrontal regions deeply concave; canthus rostralis sharply rounded anteriorly; snout elongate (ES/HL 0.43), rounded in dorsal profile, laterally constricted; eye large (ED/HL 0.23); ear opening subelliptical, moderate in size (EL/HL 0.05), vertically oriented; eye to ear distance greater than diameter of eye; rostral rectangular, divided dorsally by a small azygous scale; an inverted Y-shaped furrow, bordered posteriorly by left and right supranasals, and one medial postrostral (=internasal), bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large anterior supranasal and smaller posterior supranasal, posteriorly by one postnasal, ventrally by first supralabial; 12(R), 10(L) rectangular supralabials extending to just beyond upturn of labial margin, tapering abruptly below midpoint of eye; first supralabial largest; 8(R,L) infralabials tapering in size posteriorly; scales of rostrum and lores flat, larger than granular scales on top of head and occiput, those on posterior portion of canthus rostralis slightly larger; scales on posterior portion of occiput intermixed with small tubercles; no tubercles on top of head between eyes; large, boney frontal ridges bordering orbit confluent with boney, transverse, parietal ridge; dorsal superciliaries elongate, smooth, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by left and right, rectangular postmentals which contact medially for 40% of their length; single row of slightly enlarged, elongate sublabials extending posteriorly to third infralabials; small, granular to flat, gular scales grading posteriorly into larger, flat, smooth, imbricate, pectoral and ventral scales.
|
||
</paragraph>
|
||
<caption id="B2886608FFBFFF8EFF41FA5424D2F9A4" httpUri="https://zenodo.org/record/250554/files/figure.png" pageId="17" pageNumber="376" targetBox="[151,1435,645,1497]" targetPageId="17">
|
||
<paragraph id="E6483680FFBFFF8EFF41FA5424D2F9A4" blockId="17.[151,1436,1519,1602]" pageId="17" pageNumber="376">
|
||
<emphasis id="D483EA92FFBFFF8EFF41FA542123F9E3" bold="true" box="[151,275,1519,1541]" pageId="17" pageNumber="376">FIGURE 5.</emphasis>
|
||
Upper left: adult male holotype (LSUHC 11207) of
|
||
<taxonomicName id="21F74D03FFBFFF8EFCFBFA542431F9E3" box="[813,1025,1519,1541]" class="Reptilia" family="Gekkonidae" genus="Cyrtotdactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="376" phylum="Chordata" rank="species" species="timur" status="sp. nov.">
|
||
<emphasis id="D483EA92FFBFFF8EFCFBFA542431F9E3" box="[813,1025,1519,1541]" italics="true" pageId="17" pageNumber="376">Cyrtotdactylus timur</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CFB057E9FFBFFF8EFBD1FA542468F9E3" box="[1031,1112,1519,1541]" pageId="17" pageNumber="376" rank="species">
|
||
<emphasis id="D483EA92FFBFFF8EFBD1FA542468F9E3" bold="true" box="[1031,1112,1519,1541]" pageId="17" pageNumber="376">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from Punca Air, Gunung Tebu, Terengganu, Peninsular Malaysia. Upper right: female paratype LSUHC 11185. Middle left: uncataloged hatching LSUDPC 8249. Lower left: adult male paratype LSUHC 11183. Lower right: juvenile female paratype LSUHC 11184.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E6483680FFBFFF8EFF11F9D420C1F886" blockId="17.[151,1436,1647,2032]" pageId="17" pageNumber="376">Body relatively short (AG/SVL 0.46) with well-defined, non-tuberculate, ventrolateral folds; dorsal scales small, granular, interspersed with larger, unicainate, regularly arranged, low tubercles; smaller intervening tubercles absent; tubercles extend from occiput to caudal constriction and onto base of tail where they fade out; caudal tubercles absent from posterior nine-tenths of tail; tubercles on occiput and nape small, those on body larger; approximately 21 longitudinal rows of tubercles at midbody; 38 paravertebral tubercles; 31 flat imbricate ventral scales between ventrolateral body folds, ventral scales larger than dorsal scales; precloacal scales large, smooth.</paragraph>
|
||
<paragraph id="E6483680FFBFFF8DFF11F8D02201FDDA" blockId="17.[151,1436,1647,2032]" lastBlockId="18.[151,1436,151,1869]" lastPageId="18" lastPageNumber="377" pageId="17" pageNumber="376">Forelimbs moderate, relatively short (FL/SVL 0.18); scales on dorsal surfaces of forelimbs juxtaposed, intermixed with larger tubercles; scales of ventral surface of forearm flat, subimbricate, lacking tubercles; palmar scales weakly rounded; digits well-developed, inflected at basal, interphalangeal joints; subdigital lamellae rectangular proximal to joint inflection, only slightly expanded distal to inflection; digits more narrow distal to joints; claws well-developed, sheathed by a dorsal and ventral scale; hind limbs more robust than forelimbs, moderate in length (TBL/SVL 0.21), larger tubercles on dorsal surface of thigh separated by smaller granular scales, tubercles on dorsal surfaces of foreleg same size as those on thigh; ventral scales of thigh flat, smooth, imbricate, larger than dorsal granular scales; ventral, tibial scales flat, smooth, imbricate; single row of greatly enlarged, flat, rectangular, imbricate, femoral scales extend from knee to knee through precloacal region where they are continuous with enlarged, precloacal scales; 21 contiguous, pore-bearing femoro-precloacal scales forming an inverted T bearing a deep, precloacal groove in which 12 pore-bearing scales are found (six on left and right); postfemoral scales immediately posterior to enlarged scale row smaller, forming an abrupt union on posteroventral margin of thigh; plantar scales low, slightly raised; digits well-developed, inflected at basal, interphalangeal joints; subdigital lamellae proximal to joint inflection rectangular, only slightly expanded distal to inflection; digits more narrow distal to joints; claws well-developed, sheathed by a dorsal and ventral scale; 20(R) 21(L) subdigital lamellae on 4th toe.</paragraph>
|
||
<paragraph id="E6483680FFBCFF8DFF11FDFC21D1FD16" blockId="18.[151,1436,151,1869]" pageId="18" pageNumber="377">
|
||
Tail original,
|
||
<quantity id="210F9B65FFBCFF8DFE8BFDF321FFFDB9" box="[349,463,584,608]" metricMagnitude="-1" metricUnit="m" metricValue="1.433" pageId="18" pageNumber="377" unit="mm" value="143.3">143.3 mm</quantity>
|
||
in length,
|
||
<quantity id="210F9B65FFBCFF8DFD90FDF32298FDB9" box="[582,680,584,608]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="18" pageNumber="377" unit="mm" value="11.2">11.2 mm</quantity>
|
||
in width at base, tapering to a point; dorsal scales of tail flat, nearly square; tail weakly segmented with 8–10 transverse scale rows per segment; subcaudal region bearing a median row of large transverse scales; very shallow dorsal and lateral caudal furrows extend entire length of tail; base of tail bearing hemipenial swellings; three small, postcloacal tubercles on hemipenial swellings; postcloacal scales smooth, flat, large, imbricate.
|
||
</paragraph>
|
||
<paragraph id="E6483680FFBCFF8DFF11FD4724C7FC2E" blockId="18.[151,1436,151,1869]" pageId="18" pageNumber="377">
|
||
<emphasis id="D483EA92FFBCFF8DFF11FD472233FCF3" bold="true" box="[199,515,764,789]" pageId="18" pageNumber="377">
|
||
Coloration in life (
|
||
<figureCitation id="7ECC2A05FFBCFF8DFE75FD4721C5FCF3" box="[419,501,764,789]" captionStart="FIGURE 5" captionStartId="17.[151,250,1519,1541]" captionTargetBox="[151,1435,645,1497]" captionTargetId="figure@17.[151,1436,645,1497]" captionTargetPageId="17" captionText="FIGURE 5. Upper left: adult male holotype (LSUHC 11207) of Cyrtotdactylus timur sp. nov. from Punca Air, Gunung Tebu, Terengganu, Peninsular Malaysia. Upper right: female paratype LSUHC 11185. Middle left: uncataloged hatching LSUDPC 8249. Lower left: adult male paratype LSUHC 11183. Lower right: juvenile female paratype LSUHC 11184." httpUri="https://zenodo.org/record/250554/files/figure.png" pageId="18" pageNumber="377">Figs. 5</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Dorsal ground color of head, body, and limbs tan, immaculate; wide, dark-brown nuchal loop edged anteriorly and posteriorly by thin, faint yellow line; four wide, brown, body bands between nuchal loop and hind limb insertions weakly edged anteriorly and posteriorly by thin, faint, yellowish lines; band/ interspace ratio 1.00; no markings on posterior margin of thigh; eight white caudal bands infused with brown pigment, one-half to one-third width of dark-brown caudal bands; ventral surface of head, body, and limbs beige, immaculate except for black stipples in each scale; dark-brown and white caudal bands encircle tail.
|
||
</paragraph>
|
||
<paragraph id="E6483680FFBCFF8DFF11FC6F20C0FB23" blockId="18.[151,1436,151,1869]" pageId="18" pageNumber="377">
|
||
<emphasis id="D483EA92FFBCFF8DFF11FC6F21F1FC0B" bold="true" box="[199,449,980,1005]" pageId="18" pageNumber="377">
|
||
Variation (
|
||
<figureCitation id="7ECC2A05FFBCFF8DFE90FC6F21A6FC0B" box="[326,406,980,1005]" captionStart="FIGURE 5" captionStartId="17.[151,250,1519,1541]" captionTargetBox="[151,1435,645,1497]" captionTargetId="figure@17.[151,1436,645,1497]" captionTargetPageId="17" captionText="FIGURE 5. Upper left: adult male holotype (LSUHC 11207) of Cyrtotdactylus timur sp. nov. from Punca Air, Gunung Tebu, Terengganu, Peninsular Malaysia. Upper right: female paratype LSUHC 11185. Middle left: uncataloged hatching LSUDPC 8249. Lower left: adult male paratype LSUHC 11183. Lower right: juvenile female paratype LSUHC 11184." httpUri="https://zenodo.org/record/250554/files/figure.png" pageId="18" pageNumber="377">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="7ECC2A05FFBCFF8DFE75FC6F2182FC0B" box="[419,434,980,1005]" captionStart="FIGURE 6" captionStartId="19.[151,250,1728,1750]" captionTargetBox="[162,1436,193,1700]" captionTargetId="figure@19.[162,1447,193,1701]" captionTargetPageId="19" captionText="FIGURE 6. Microhabitat of Cyrtodactylus timur sp. nov. at the type locality of Punca Air, Gunung Tebu, Terengganu, Peninsular Malaysia." httpUri="https://zenodo.org/record/250555/files/figure.png" pageId="18" pageNumber="377">6</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<typeStatus id="394C8822FFBCFF8DFE1EFC6E2205FC0B" box="[456,565,981,1005]" pageId="18" pageNumber="377" type="paratype">Paratypes</typeStatus>
|
||
LSUHC 11183–85 resemble the
|
||
<typeStatus id="394C8822FFBCFF8DFC7EFC6F243AFC0B" box="[936,1034,980,1005]" pageId="18" pageNumber="377" type="holotype">holotype</typeStatus>
|
||
in patterning but are slightly darker in overall coloration. LSUHC 11184 is a juvenile with a more contrasting color pattern and lighter ground color. LSUHC 11288 is considerably darker than the remainder of the
|
||
<typeStatus id="394C8822FFBCFF8DFCAEFBA62398FBD3" box="[888,936,1053,1077]" pageId="18" pageNumber="377">type</typeStatus>
|
||
series and the thin whitish lines edging the anterior and posterior margins of the body bands are more discontinuous, appearing more as a series of transverse spots. LSUHC 11183 has a completely regenerated, unicolor brown tail. A small juvenile (LSUHC 10886, SVL
|
||
<quantity id="210F9B65FFBCFF8DFF41FB3C20CCFB46" box="[151,252,1159,1184]" metricMagnitude="-2" metricUnit="m" metricValue="6.029999999999999" pageId="18" pageNumber="377" unit="mm" value="60.3">60.3 mm</quantity>
|
||
) not part of the
|
||
<typeStatus id="394C8822FFBCFF8DFE65FB3321D3FB46" box="[435,483,1160,1184]" pageId="18" pageNumber="377">type</typeStatus>
|
||
series has a very bold, contrasting banding pattern. Meristic differences are listed in Table 6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEED650BFFBCFF8DFF11FB7424B7FAEB" pageId="18" pageNumber="377" type="distribution">
|
||
<paragraph id="E6483680FFBCFF8DFF11FB7424B7FAEB" blockId="18.[151,1436,151,1869]" pageId="18" pageNumber="377">
|
||
<emphasis id="D483EA92FFBCFF8DFF11FB742150FB0E" bold="true" box="[199,352,1231,1256]" pageId="18" pageNumber="377">Distribution.</emphasis>
|
||
This species may be endemic to Gunung Tebu, Terengganu, Peninsular
|
||
<collectingCountry id="9EE07610FFBCFF8DFB46FB7424CAFB0E" box="[1168,1274,1231,1256]" name="Malaysia" pageId="18" pageNumber="377">Malaysia</collectingCountry>
|
||
. It is expected that its distribution is more extensive both above and below the
|
||
<typeStatus id="394C8822FFBCFF8DFCB5FB4E23A3FAEB" box="[867,915,1269,1293]" pageId="18" pageNumber="377">type</typeStatus>
|
||
locality at Punca Air.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEED650BFFBCFF8DFF11FAAC25A8F8AB" pageId="18" pageNumber="377" type="etymology">
|
||
<paragraph id="E6483680FFBCFF8DFF11FAAC241DFA9E" blockId="18.[151,1436,151,1869]" pageId="18" pageNumber="377">
|
||
<emphasis id="D483EA92FFBCFF8DFF11FAAC217EFAD6" bold="true" box="[199,334,1303,1328]" pageId="18" pageNumber="377">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<emphasis id="D483EA92FFBCFF8DFD9EFAA322A4FAC9" box="[584,660,1304,1327]" italics="true" pageId="18" pageNumber="377">
|
||
<taxonomicName id="21F74D03FFBCFF8DFD9EFAA322BCFAC9" box="[584,652,1304,1327]" class="Reptilia" family="Gekkonidae" genus="Cyrtotdactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="377" phylum="Chordata" rank="species" species="timur">timur</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
is an invariable noun in apposition in reference to this species being found in the Banjaran
|
||
<taxonomicName id="21F74D03FFBCFF8DFE58FA8721E3FAB3" box="[398,467,1340,1365]" class="Reptilia" family="Gekkonidae" genus="Cyrtotdactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="379" phylum="Chordata" rank="species" species="timur" status="sp. nov.">Timur</taxonomicName>
|
||
of northeastern Peninsular
|
||
<collectingCountry id="9EE07610FFBCFF8DFCD3FA87235FFAB3" box="[773,879,1340,1365]" name="Malaysia" pageId="18" pageNumber="377">Malaysia</collectingCountry>
|
||
. “
|
||
<taxonomicName id="21F74D03FFBCFF8DFC55FA8723FDFAB3" box="[899,973,1340,1365]" class="Reptilia" family="Gekkonidae" genus="Cyrtotdactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="379" phylum="Chordata" rank="species" species="timur" status="sp. nov.">Timur</taxonomicName>
|
||
” means east in the Malay language and is in reference to the mountain range being on the east coast of Peninsular
|
||
<collectingCountry id="9EE07610FFBCFF8DFC69FAE42419FA9E" box="[959,1065,1375,1400]" name="Malaysia" pageId="18" pageNumber="377">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E6483680FFBCFF8DFF11FA3F2167F9CA" blockId="18.[151,1436,151,1869]" pageId="18" pageNumber="377">
|
||
<emphasis id="D483EA92FFBCFF8DFF11FA3F21BCFA7B" bold="true" box="[199,396,1412,1437]" pageId="18" pageNumber="377">Natural History.</emphasis>
|
||
All adults were collected at night in hill dipterocarp forest in the vicinity of a rocky stream (
|
||
<figureCitation id="7ECC2A05FFBCFF8DFF49FA1C20D5FA26" box="[159,229,1447,1472]" captionStart="FIGURE 6" captionStartId="19.[151,250,1728,1750]" captionTargetBox="[162,1436,193,1700]" captionTargetId="figure@19.[162,1447,193,1701]" captionTargetPageId="19" captionText="FIGURE 6. Microhabitat of Cyrtodactylus timur sp. nov. at the type locality of Punca Air, Gunung Tebu, Terengganu, Peninsular Malaysia." httpUri="https://zenodo.org/record/250555/files/figure.png" pageId="18" pageNumber="377">Fig. 6</figureCitation>
|
||
) between
|
||
<quantity id="210F9B65FFBCFF8DFE89FA1C219AFA26" box="[351,426,1447,1472]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="18" pageNumber="377" unit="m" value="640.0">640 m</quantity>
|
||
and
|
||
<quantity id="210F9B65FFBCFF8DFE30FA132200FA26" box="[486,560,1448,1472]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="377" unit="m" value="700.0">700 m</quantity>
|
||
in elevation. Lizards were most common on the granite boulders along the stream but others were seen on the bases of trees and one individual was observed on an earthen bank. A small juvenile (LSUHC 10886, SVL
|
||
<quantity id="210F9B65FFBCFF8DFE21FA54226CF9EE" box="[503,604,1519,1544]" metricMagnitude="-2" metricUnit="m" metricValue="6.029999999999999" pageId="18" pageNumber="377" unit="mm" value="60.3">60.3 mm</quantity>
|
||
) was found on a twig at the base of a large tree deep in the forest and not along the stream.
|
||
</paragraph>
|
||
<paragraph id="E6483680FFBCFF8DFF11F98C25A8F8AB" blockId="18.[151,1436,151,1869]" pageId="18" pageNumber="377">
|
||
<emphasis id="D483EA92FFBCFF8DFF11F98C223AF9B6" bold="true" box="[199,522,1591,1616]" pageId="18" pageNumber="377">Comparisons (Tables 5, 6).</emphasis>
|
||
<taxonomicName id="21F74D03FFBCFF8DFDC5F98C22C6F9B6" box="[531,758,1591,1616]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="377" phylum="Chordata" rank="species" species="timur" status="sp. nov.">
|
||
<emphasis id="D483EA92FFBCFF8DFDC5F98C22C6F9B6" box="[531,758,1591,1616]" italics="true" pageId="18" pageNumber="377">Cyrtodactylus timur</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CFB057E9FFBCFF8DFD29F98C236BF9B6" box="[767,859,1591,1616]" pageId="18" pageNumber="377" rank="species">
|
||
<emphasis id="D483EA92FFBCFF8DFD29F98C236BF9B6" bold="true" box="[767,859,1591,1616]" pageId="18" pageNumber="377">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is differentiated from all other members of the
|
||
<taxonomicName id="21F74D03FFBCFF8DFA54F983213AF993" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="377" phylum="Chordata" rank="species" species="pulchellus">
|
||
<emphasis id="D483EA92FFBCFF8DFA54F983213AF993" italics="true" pageId="18" pageNumber="377">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
complex by having a combination of moderately sized body tubercles and no tubercles in gular region, ventrolateral body fold, or on ventral surfaces of forelimbs; 38–43 paravertebral tubercles; 21–24 longitudinal rows of body tubercles; 31–40 ventral scales; 20–25 subdigital lamellae on the fourth toe; 21 or 22 femoroprecloacal pores; four dorsal body bands between nuchal loop and hind limb insertions; body band/interspace ratio 1.00–1.25; no scattered pattern of white dorsal tubercles; 8–10 dark caudal bands on original tail; white caudal bands infused with brown pigment in adults; and a maximum SVL of
|
||
<quantity id="210F9B65FFBCFF8DFCDDF8B423B1F8CE" box="[779,897,1807,1832]" metricMagnitude="-1" metricUnit="m" metricValue="1.205" pageId="18" pageNumber="377" unit="mm" value="120.5">120.5 mm</quantity>
|
||
. Within the
|
||
<taxonomicName id="21F74D03FFBCFF8DFBDCF8AB24AEF8CE" box="[1034,1182,1807,1832]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="377" phylum="Chordata" rank="species" species="pulchellus">
|
||
<emphasis id="D483EA92FFBCFF8DFBDCF8AB24AEF8CE" box="[1034,1182,1807,1832]" italics="true" pageId="18" pageNumber="377">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
complex,
|
||
<taxonomicName id="21F74D03FFBCFF8DFAC3F8AB2543F8CE" box="[1301,1395,1808,1832]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="377" phylum="Chordata" rank="species" species="timur" status="sp. nov.">
|
||
<emphasis id="D483EA92FFBCFF8DFAC3F8AB2543F8CE" box="[1301,1395,1808,1832]" italics="true" pageId="18" pageNumber="377">C. timur</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CFB057E9FFBCFF8DFAACF8B420F8F8AB" pageId="18" pageNumber="377" rank="species">
|
||
<emphasis id="D483EA92FFBCFF8DFAACF8B420F8F8AB" bold="true" pageId="18" pageNumber="377">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is the sister species to
|
||
<taxonomicName id="21F74D03FFBCFF8DFE1EF88E22B8F8AB" box="[456,648,1844,1869]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="377" phylum="Chordata" rank="species" species="jelawangensis">
|
||
<emphasis id="D483EA92FFBCFF8DFE1EF88E22B8F8AB" box="[456,648,1844,1869]" italics="true" pageId="18" pageNumber="377">C. jelawangensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="7ECC2A05FFBCFF8DFD41F88F22ECF8AA" box="[663,732,1844,1869]" captionStart="FIGURE 2" captionStartId="7.[151,250,1956,1978]" captionTargetBox="[195,1317,215,1930]" captionTargetId="figure@7.[178,1409,195,1934]" captionTargetPageId="7" captionText="FIGURE 2. Likelihood topology (– ln L 15911.137535) of the Cyrtodactylus pulchellus complex with Maximum Likelihood bootstrap support values (ML) and Bayesian Posterior probabilities (BPP) respectively." httpUri="https://zenodo.org/record/250551/files/figure.png" pageId="18" pageNumber="377">Fig. 2</figureCitation>
|
||
). See comparisons between these two species above (Table 6).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |