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<mods:title id="93BAC192A1EF98561E48B6F9EFBB04FF">Illustrated overview and identification key to Cameroonian Ceratocanthinae beetles (Coleoptera: Scarabaeoidea: Hybosoridae) with description of four new species</mods:title>
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<mods:namePart id="989692974CB3464A1879F852000384D3">Gill, Bruce D.</mods:namePart>
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<mods:namePart id="8D8D5486FEC4BE2003D8043A4075F218">Grebennikov, Vasily V.</mods:namePart>
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<treatment id="03A1C115FFD3FFA4FF3FFA77D99EF84C" ID-DOI="http://doi.org/10.5281/zenodo.6183909" ID-GBIF-Taxon="119409503" ID-Zenodo-Dep="6183909" LSID="urn:lsid:plazi:treatment:03A1C115FFD3FFA4FF3FFA77D99EF84C" httpUri="http://treatment.plazi.org/id/03A1C115FFD3FFA4FF3FFA77D99EF84C" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="C3122388FFD3FFA7FF3FFA77DCE6FA07" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BB77003FFD3FFA7FF3FFA77DEADFA61" blockId="17.[151,690,1433,1493]" box="[151,690,1433,1460]" pageId="17" pageNumber="18">
<heading id="D0FFC76FFFD3FFA7FF3FFA77DEADFA61" bold="true" box="[151,690,1433,1460]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="4C080B80FFD3FFA7FF3FFA77DEADFA61" ID-CoL="3Z9JX" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" box="[151,690,1433,1460]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="demirei">
<emphasis id="B97CAC11FFD3FFA7FF3FFA77DEADFA61" bold="true" box="[151,690,1433,1460]" pageId="17" pageNumber="18">
<emphasis id="B97CAC11FFD3FFA7FF3FFA77DE1EFA66" bold="true" box="[151,513,1434,1460]" italics="true" pageId="17" pageNumber="18">Melanophilharmostes demirei</emphasis>
<bibRefCitation id="EF990DF2FFD3FFA7FDA0FA74DEADFA61" author="Paulian" box="[520,690,1433,1459]" pageId="17" pageNumber="18" refString="Paulian, R. (1977) Revision des Ceratocanthidae (Coleoptera, Scarabaeidae). I. - Les formes africaines. Revue de Zoologie Africaine, 91, 253 - 316." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB77003FFD3FFA7FF3FFA50DCE6FA07" blockId="17.[151,690,1433,1493]" box="[151,249,1469,1493]" pageId="17" pageNumber="18">
(
<figureCitation id="13336C86FFD3FFA7FF37FA50DCEFFA07" box="[159,240,1469,1493]" captionStart="FIGURE 15" captionStartId="18.[151,250,1036,1059]" captionTargetBox="[181,1410,219,966]" captionTargetId="figure@18.[151,1436,193,1015]" captionTargetPageId="18" captionText="FIGURE 15. Melanophilharmostes demirei Paulian, habitus. A—distended specimen, dorsal, B D—rolled up specimen, ventral (B), dorsal (C), lateral (D)." httpUri="https://zenodo.org/record/277656/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3122388FFD3FFA7FF3FF9E9DF71F992" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BB77003FFD3FFA7FF3FF9E9DF71F992" blockId="17.[151,1436,1540,1600]" pageId="17" pageNumber="18">
<emphasis id="B97CAC11FFD3FFA7FF3FF9E9DD19F9CF" bold="true" box="[151,262,1540,1565]" pageId="17" pageNumber="18">Material.</emphasis>
<specimenCount id="9D0EBB8AFFD3FFA7FEBCF9E8DDA8F9CF" box="[276,439,1541,1565]" pageId="17" pageNumber="18" type="generic">33 specimens</specimenCount>
,
<collectingCountry id="F31F3093FFD3FFA7FE62F9E8DE20F9CF" box="[458,575,1541,1565]" name="Cameroon" pageId="17" pageNumber="18">Cameroon</collectingCountry>
S.-West Prov., Mt. Kupé at Nyasoso,
<geoCoordinate id="EE3C16C4FFD3FFA7FBB7F9E8D8AFF9CF" box="[1055,1200,1541,1565]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="4.8192167">N04°49.153</geoCoordinate>
<geoCoordinate id="EE3C16C4FFD3FFA7FB17F9E8D941F9CF" box="[1215,1374,1541,1565]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="9.70755">E009°42.453</geoCoordinate>
,
<date id="FFB656C3FFD3FFA7FADAF9E8DD13F992" pageId="17" pageNumber="18" value="2006-05-19" valueMax="2006-05-21" valueMin="2006-05-19">19 21.V.2006</date>
, 1550m, V. Grebennikov leg. (
<collectionCode id="ED19E8C6FFD3FFA7FDCBF9C5DEACF992" box="[611,691,1576,1600]" pageId="17" pageNumber="18">ABCB</collectionCode>
,
<collectionCode id="ED19E8C6FFD3FFA7FD17F9C5DF0FF992" box="[703,784,1576,1600]" pageId="17" pageNumber="18">BDGC</collectionCode>
,
<collectionCode id="ED19E8C6FFD3FFA7FCB2F9C5DF7DF992" box="[794,866,1576,1600]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="17" pageNumber="18">CNCI</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF77208BFFD0FFA4FF3FFBE1DDD0FB90" httpUri="https://zenodo.org/record/277656/files/figure.png" pageId="18" pageNumber="19" targetBox="[181,1410,219,966]" targetPageId="18">
<paragraph id="8BB77003FFD0FFA4FF3FFBE1DDD0FB90" blockId="18.[151,1436,1036,1090]" pageId="18" pageNumber="19">
<emphasis id="B97CAC11FFD0FFA4FF3FFBE1DD3FFBF0" bold="true" box="[151,288,1036,1059]" pageId="18" pageNumber="19">FIGURE 15.</emphasis>
<taxonomicName id="4C080B80FFD0FFA4FE8EFBE0DEB3FBF1" authority="Paulian" authorityName="Paulian" box="[294,684,1037,1059]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="demirei">
<emphasis id="B97CAC11FFD0FFA4FE8EFBE0DE48FBF1" box="[294,599,1037,1059]" italics="true" pageId="18" pageNumber="19">Melanophilharmostes demirei</emphasis>
Paulian
</taxonomicName>
, habitus. A—distended specimen, dorsal, BD—rolled up specimen, ventral (B), dorsal (C), lateral (D).
</paragraph>
</caption>
<subSubSection id="C3122388FFD0FFA4FF6DFB83D99EF84C" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BB77003FFD0FFA4FF6DFB83D99EF84C" blockId="18.[151,1437,1134,1951]" pageId="18" pageNumber="19">
<emphasis id="B97CAC11FFD0FFA4FF6DFB83DD26FB54" bold="true" box="[197,313,1134,1158]" pageId="18" pageNumber="19">Remarks.</emphasis>
This flightless species is apparently endemic to
<collectingCountry id="F31F3093FFD0FFA4FCFCFB83DFD3FB54" box="[852,972,1134,1158]" name="Cameroon" pageId="18" pageNumber="19">Cameroon</collectingCountry>
. Representatives of the Mt. Kupé population differ slightly from the
<typeStatus id="54B3CEA1FFD0FFA4FE7EFB7FDE24FB78" box="[470,571,1170,1194]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
, which originated from Mt. Febé (some
<quantity id="4CF0DDE6FFD0FFA4FBAFFB7FD842FB78" box="[1031,1117,1170,1194]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="km" value="300.0">300 km</quantity>
eastwards). The differences are mainly in the punctures of the pronotum and elytra, which are larger and sparser in the Mt. Kupé population. While the aedeagus does not show appreciable differences, the lack of additional material from the
<typeStatus id="54B3CEA1FFD0FFA4FAA7FB36D920FB21" box="[1295,1343,1243,1267]" pageId="18" pageNumber="19">type</typeStatus>
locality makes it impossible to draw any conclusion about any possible specific separation of these two populations. The current placement of this taxon within the genus
<taxonomicName id="4C080B80FFD0FFA4FD74FACFDFCFFAE8" box="[732,976,1314,1338]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FD74FACFDFCFFAE8" box="[732,976,1314,1338]" italics="true" pageId="18" pageNumber="19">Melanophilharmostes</emphasis>
</taxonomicName>
is uncertain.
<taxonomicName id="4C080B80FFD0FFA4FBDCFACFD977FAE8" box="[1140,1384,1314,1338]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FBDCFACFD977FAE8" box="[1140,1384,1314,1338]" italics="true" pageId="18" pageNumber="19">Melanophilharmostes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C080B80FFD0FFA4FF3FFAABDDB9FA8C" box="[151,422,1350,1374]" class="Insecta" family="Hybosoridae" genus="Pseudopterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FF3FFAABDDB9FA8C" box="[151,422,1350,1374]" italics="true" pageId="18" pageNumber="19">Pseudopterorthochaetes</emphasis>
</taxonomicName>
are similar to each other and there is some confusion in their generic definitions.
<bibRefCitation id="EF990DF2FFD0FFA4FAE0FAABDCFCFA50" author="Paulian" pageId="18" pageNumber="19" refString="Paulian, R. (1977) Revision des Ceratocanthidae (Coleoptera, Scarabaeidae). I. - Les formes africaines. Revue de Zoologie Africaine, 91, 253 - 316." type="journal article" year="1977">Paulian (1977)</bibRefCitation>
, in the key to the African genera of
<taxonomicName id="4C080B80FFD0FFA4FDD5FA87DF29FA50" box="[637,822,1386,1410]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Ceratocanthinae">Ceratocanthinae</taxonomicName>
, distinguished the two genera on the basis of the presence/absence of a lateral carina delimiting the pseudoepipleuron and the development of apical teeth of the protibiae. However these characters were somewhat misinterpreted and/or overemphasized and were not followed consistently as, for example, in
<taxonomicName id="4C080B80FFD0FFA4FDA0FA3ADE9BFA3C" box="[520,644,1494,1518]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="demirei">
<emphasis id="B97CAC11FFD0FFA4FDA0FA3ADE9BFA3C" box="[520,644,1494,1518]" italics="true" pageId="18" pageNumber="19">M. demirei</emphasis>
</taxonomicName>
. This species clearly shares several key characters with
<taxonomicName id="4C080B80FFD0FFA4FABCFA3BDD37F9C0" class="Insecta" family="Hybosoridae" genus="Pseudopterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FABCFA3BDD37F9C0" italics="true" pageId="18" pageNumber="19">Pseudopterorthochaetes</emphasis>
</taxonomicName>
and strongly differs from the majority of the species currently assigned to
<taxonomicName id="4C080B80FFD0FFA4FB09FA17D98AF9C0" box="[1185,1429,1530,1554]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FB09FA17D98AF9C0" box="[1185,1429,1530,1554]" italics="true" pageId="18" pageNumber="19">Melanophilharmostes</emphasis>
</taxonomicName>
, namely from the group of species comprising the
<typeStatus id="54B3CEA1FFD0FFA4FD63F9F2DEE4F9E5" box="[715,763,1567,1591]" pageId="18" pageNumber="19">type</typeStatus>
species of
<taxonomicName id="4C080B80FFD0FFA4FCD4F9F3D86FF9E4" box="[892,1136,1566,1590]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FCD4F9F3D86FF9E4" box="[892,1136,1566,1590]" italics="true" pageId="18" pageNumber="19">Melanophilharmostes</emphasis>
</taxonomicName>
:
<taxonomicName id="4C080B80FFD0FFA4FBD7F9F2D988F9E4" authority="Paulian, 1968" authorityName="Paulian" authorityYear="1968" box="[1151,1431,1566,1590]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="zicsii">
<emphasis id="B97CAC11FFD0FFA4FBD7F9F2D8FFF9E4" box="[1151,1248,1567,1590]" italics="true" pageId="18" pageNumber="19">M. zicsii</emphasis>
(Paulian, 1968)
</taxonomicName>
, which includes at least
<taxonomicName id="4C080B80FFD0FFA4FE33F9AEDEFFF988" authority="Paulian, 1974" authorityName="Paulian" authorityYear="1974" box="[411,736,1602,1626]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B97CAC11FFD0FFA4FE33F9AEDE36F988" box="[411,553,1603,1626]" italics="true" pageId="18" pageNumber="19">M. carinatus</emphasis>
(Paulian, 1974)
</taxonomicName>
,
<taxonomicName id="4C080B80FFD0FFA4FD44F9AED806F988" authority="Paulian, 1974" authorityName="Paulian" authorityYear="1974" box="[748,1049,1602,1626]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ghanae">
<emphasis id="B97CAC11FFD0FFA4FD44F9AEDF7BF988" box="[748,868,1602,1626]" italics="true" pageId="18" pageNumber="19">M. ghanae</emphasis>
(Paulian, 1974)
</taxonomicName>
,
<taxonomicName id="4C080B80FFD0FFA4FB8CF9AED96DF988" authority="Petrovitz, 1968" authorityName="Petrovitz" authorityYear="1968" box="[1060,1394,1602,1626]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="palustris">
<emphasis id="B97CAC11FFD0FFA4FB8CF9AED8B4F988" box="[1060,1195,1602,1626]" italics="true" pageId="18" pageNumber="19">M. palustris</emphasis>
(Petrovitz, 1968)
</taxonomicName>
,
<taxonomicName id="4C080B80FFD0FFA4FAD5F9AEDD85F9AC" authority="Paulian, 1937" authorityName="Paulian" authorityYear="1937" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="posthi">
<emphasis id="B97CAC11FFD0FFA4FAD5F9AEDCC3F9AC" italics="true" pageId="18" pageNumber="19">M. posthi</emphasis>
(Paulian, 1937)
</taxonomicName>
,
<taxonomicName id="4C080B80FFD0FFA4FE03F98ADF38F9AC" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" box="[427,807,1638,1662]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pseudoposthi">
<emphasis id="B97CAC11FFD0FFA4FE03F98ADE76F9AC" box="[427,617,1638,1662]" italics="true" pageId="18" pageNumber="19">M. pseudoposthi</emphasis>
(
<bibRefCitation id="EF990DF2FFD0FFA4FDD4F98BDF3FF9AC" author="Paulian" box="[636,800,1638,1662]" pageId="18" pageNumber="19" refString="Paulian, R. (1977) Revision des Ceratocanthidae (Coleoptera, Scarabaeidae). I. - Les formes africaines. Revue de Zoologie Africaine, 91, 253 - 316." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C080B80FFD0FFA4FC90F98AD886F9AC" authority="Paulian, 1946" authorityName="Paulian" authorityYear="1946" box="[824,1177,1638,1662]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="B97CAC11FFD0FFA4FC90F98ADFC5F9AC" box="[824,986,1639,1662]" italics="true" pageId="18" pageNumber="19">M. puncticeps</emphasis>
(Paulian, 1946)
</taxonomicName>
,
<taxonomicName id="4C080B80FFD0FFA4FB01F98ADCC5F970" authority="Paulian, 1968" authorityName="Paulian" authorityYear="1968" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vincenti">
<emphasis id="B97CAC11FFD0FFA4FB01F98AD933F9AC" box="[1193,1324,1639,1662]" italics="true" pageId="18" pageNumber="19">M. vincenti</emphasis>
(Paulian, 1968)
</taxonomicName>
,
<taxonomicName id="4C080B80FFD0FFA4FF41F966DD54F970" box="[233,331,1675,1698]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="zicsii">
<emphasis id="B97CAC11FFD0FFA4FF41F966DD54F970" box="[233,331,1675,1698]" italics="true" pageId="18" pageNumber="19">M. zicsii</emphasis>
</taxonomicName>
, and probably also
<taxonomicName id="4C080B80FFD0FFA4FD9AF966DF8DF970" authority="Paulian, 1974" authorityName="Paulian" authorityYear="1974" box="[562,914,1674,1698]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bicarinatus">
<emphasis id="B97CAC11FFD0FFA4FD9AF966DEC5F970" box="[562,730,1674,1698]" italics="true" pageId="18" pageNumber="19">M. bicarinatus</emphasis>
(Paulian, 1974)
</taxonomicName>
and
<taxonomicName id="4C080B80FFD0FFA4FC66F966D908F970" authority="Paulian, 1968" authorityName="Paulian" authorityYear="1968" box="[974,1303,1674,1698]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ocellatus">
<emphasis id="B97CAC11FFD0FFA4FC66F966D844F970" box="[974,1115,1674,1698]" italics="true" pageId="18" pageNumber="19">M. ocellatus</emphasis>
(Paulian, 1968)
</taxonomicName>
. A detailed morphological analysis suggests that other characters could be more reliable in defining the genera than the ones used by Paulian, especially the sexual dimorphism of the mesotibiae and protibiae. The mesotibiae of
<taxonomicName id="4C080B80FFD0FFA4FAB9F93FDD15F8DC" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FAB9F93FDD15F8DC" italics="true" pageId="18" pageNumber="19">Melanophilharmostes</emphasis>
</taxonomicName>
are subrectangular and the male mesotibiae have the inner apical spur bent inwards at a right angle, while in
<taxonomicName id="4C080B80FFD0FFA4FF52F8F7DE16F8E0" box="[250,521,1818,1842]" class="Insecta" family="Hybosoridae" genus="Pseudopterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FF52F8F7DE16F8E0" box="[250,521,1818,1842]" italics="true" pageId="18" pageNumber="19">Pseudopterorthochaetes</emphasis>
</taxonomicName>
species the male mesotibiae are narrowed slightly apically and lack the inwardly bent inner apical spur. As a result of the adoption of these new characters there are some species (including
<taxonomicName id="4C080B80FFD0FFA4FAD5F8D2DCF4F8A8" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="demirei">
<emphasis id="B97CAC11FFD0FFA4FAD5F8D2DCF4F8A8" italics="true" pageId="18" pageNumber="19">M. demirei</emphasis>
</taxonomicName>
) assigned to
<taxonomicName id="4C080B80FFD0FFA4FE24F88FDE9FF8A8" box="[396,640,1890,1914]" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FE24F88FDE9FF8A8" box="[396,640,1890,1914]" italics="true" pageId="18" pageNumber="19">Melanophilharmostes</emphasis>
</taxonomicName>
, which actually should be transferred to
<taxonomicName id="4C080B80FFD0FFA4FBCEF88FD96AF8A8" box="[1126,1397,1890,1914]" class="Insecta" family="Hybosoridae" genus="Pseudopterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B97CAC11FFD0FFA4FBCEF88FD96AF8A8" box="[1126,1397,1890,1914]" italics="true" pageId="18" pageNumber="19">Pseudopterorthochaetes</emphasis>
</taxonomicName>
. A complete redefinition of these two genera and the resulting new combinations will be the topic of a future paper.
</paragraph>
</subSubSection>
</treatment>
</document>