treatments-xml/data/03/A1/B8/03A1B80DDE49FFDDFF43B0E4FE07924B.xml

<document ID-DOI="10.11646/phytotaxa.591.1.6" ID-ISSN="1179-3163" ID-Zenodo-Dep="7784280" checkinTime="1680164019933" checkinUser="plazi" docAuthor="Chinh, Vu Tien, Dinh, Dien, Le, Tuan Anh, Doan, Quoc Tuan, Le, Ngoc Tuan, Le, Van Huong, Tran, Nam Thang &amp; Vislobokov, Nikolay A." docDate="2023" docId="03A1B80DDE49FFDDFF43B0E4FE07924B" docLanguage="en" docName="phytotaxa.591.1.6.pdf" docOrigin="Phytotaxa 591 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.4:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="4" docTitle="Aspidistra phongdiensis D. Dien, T. A. Le &amp; Vislobokov 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="67" masterDocId="FF98C075DE48FFDEFFCBB350FFAD9537" masterDocTitle="Aspidistra phongdiensis (Asparagaceae), a new species from Central Vietnam" masterLastPageNumber="70" masterPageNumber="64" pageNumber="65" updateTime="1680164459737" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>Aspidistra phongdiensis (Asparagaceae), a new species from Central Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chinh, Vu Tien</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-5014-8500</mods:nameIdentifier>
<mods:affiliation>Vietnam National Museum of Nature, VAST, 18, Hoang Quoc Viet str., Ha Noi, Vietnam. &amp; tienchinhvu @ gmail. com; https: // orcid. org / 0000 - 0002 - 5014 - 8500</mods:affiliation>
<mods:nameIdentifier type="email">tienchinhvu@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dinh, Dien</mods:namePart>
<mods:affiliation>Phong Dien Nature Reserve, ThuaThien Hue province, Vietnam. &amp; University of Agriculture and Forestry, Hue University, Vietnam.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Le, Tuan Anh</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-5917-3243</mods:nameIdentifier>
<mods:affiliation>Mien Trung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321, Huynh Thuc Khang str., Thua Thien Hue, Vietnam. &amp; tasa 207 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5917 - 3243</mods:affiliation>
<mods:nameIdentifier type="email">tasa207@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Doan, Quoc Tuan</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4664-8972</mods:nameIdentifier>
<mods:affiliation>Faculty of Pharmacy, Hue University of Medicine and Pharmacy, Hue University, Hue City, Vietnam. &amp; doanquoctuan @ hueuni. edu. vn; https: // orcid. org / 0000 - 0003 - 4664 - 8972</mods:affiliation>
<mods:nameIdentifier type="email">doanquoctuan@hueuni.edu.vn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Le, Ngoc Tuan</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7786-5078</mods:nameIdentifier>
<mods:affiliation>Thua Thien Hue Forest Protection Department, 18 Doan Huu Trung St., Hue city, Thua Thien Hue, Vietnam. &amp; tuanngoc _ kbt @ yahoo. com; https: // orcid. org / 0000 - 0001 - 7786 - 5078</mods:affiliation>
<mods:nameIdentifier type="email">tuanngoc_kbt@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Le, Van Huong</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7399-6927</mods:nameIdentifier>
<mods:affiliation>Phong Dien Nature Reserve, ThuaThien Hue province, Vietnam. &amp; huongkbt 751970 @ gmail. com; https: // orcid. org / 0000 - 0001 - 7399 - 6927</mods:affiliation>
<mods:nameIdentifier type="email">huongkbt751970@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tran, Nam Thang</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4404-8009</mods:nameIdentifier>
<mods:affiliation>University of Agriculture and Forestry, Hue University, Vietnam. &amp; tnthang @ hueuni. edu. vn; https: // orcid. org / 0000 - 0003 - 4404 - 8009</mods:affiliation>
<mods:nameIdentifier type="email">tnthang@hueuni.edu.vn</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vislobokov, Nikolay A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-0568-6542</mods:nameIdentifier>
<mods:affiliation>Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia. &amp; n. vislobokov @ gmail. com; https: // orcid. org / 0000 - 0003 - 0568 - 6542</mods:affiliation>
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<mods:title>Phytotaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-30</mods:number>
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<mods:number>591</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>64</mods:start>
<mods:end>70</mods:end>
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<subSubSection box="[136,945,948,975]" pageId="1" pageNumber="65" type="nomenclature">
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<taxonomicName authority="D.Dien, T.A.Le &amp; Vislobokov" authorityName="D. Dien, T. A. Le &amp; Vislobokov" authorityYear="2023" box="[136,758,948,975]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="65" phylum="Tracheophyta" rank="species" species="phongdiensis" status="sp. nov.">
<emphasis bold="true" box="[136,412,948,975]" italics="true" pageId="1" pageNumber="65">Aspidistra phongdiensis</emphasis>
D.Dien, T.A.Le &amp; Vislobokov
</taxonomicName>
, 
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<taxonomicNameLabel box="[770,855,949,974]" pageId="1" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[869,934,948,974]" captionStart="FIGURE 1" captionStartId="2.[136,229,1880,1902]" captionTargetBox="[227,1361,190,1855]" captionTargetId="figure-18@2.[227,1361,190,1855]" captionTargetPageId="2" captionText="FIGURE 1. Aspidistra phongdiensis. A. Habit, scale bar: 10 cm; B. Flower, apical view; C. Sterile bracts of peduncle, scale bar: 1 cm; D. Flower, lateral view, scale bar: 5 cm; E. Perigone, dissected to show stamens and pistil, scale bar: 2 cm; F. Pistil, scale bar: 5 mm; G. Stamens, scale bar: 2 mm; H. Fruits, scale bars: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.7784282" httpUri="https://zenodo.org/record/7784282/files/figure.png" pageId="1" pageNumber="65">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="65" type="diagnosis">
<paragraph blockId="1.[136,1451,988,1190]" pageId="1" pageNumber="65">
Diagnosis: 
<taxonomicName authorityName="D. Dien, T. A. Le &amp; Vislobokov" authorityYear="2023" box="[244,475,988,1009]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="65" phylum="Tracheophyta" rank="species" species="phongdiensis">
<emphasis box="[244,475,988,1009]" italics="true" pageId="1" pageNumber="65">Aspidistra phongdiensis</emphasis>
</taxonomicName>
is morphologically similar to 
<taxonomicName authorityName="Averyanov &amp; Tillich" authorityYear="2014" box="[760,858,988,1009]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="65" phylum="Tracheophyta" rank="species" species="khangii">
<emphasis box="[760,858,988,1009]" italics="true" pageId="1" pageNumber="65">A. khangii</emphasis>
</taxonomicName>
but differs in wider leaf blade (vs. narrowly elliptic), perigone white outside (vs. purple mottled), yellow pollen (vs. white), introrse anthers (vs. latrorse) and widely obconic pistil (vs. deskshaped).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="65" type="materials_examined">
<paragraph blockId="1.[136,1451,988,1190]" pageId="1" pageNumber="65">
<emphasis bold="true" box="[136,194,1096,1117]" pageId="1" pageNumber="65">Type:</emphasis>
— 
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<collectingCountry box="[219,332,1096,1117]" name="Vietnam" pageId="1" pageNumber="65">VIETNAM</collectingCountry>
. 
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: 
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, 
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commune, 
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, 
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, 
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a.s.l.
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, 
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, 
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, 
<collectorName box="[911,1050,1132,1153]" pageId="1" pageNumber="65">Vu Tien Chinh</collectorName>
, 
<collectorName box="[1060,1196,1132,1153]" pageId="1" pageNumber="65">Le Ngoc Tuan</collectorName>
, 
<collectorName box="[1206,1346,1132,1153]" pageId="1" pageNumber="65">Le Van Huong</collectorName>
, 
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, 
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LTA 2337
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(
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!; isotype 
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!)
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.
</paragraph>
<paragraph blockId="1.[136,1452,1236,2092]" pageId="1" pageNumber="65">
Plant perennial, evergreen, herbaceous, rhizomatous, glabrous. Rhizome creeping, epigeous, 
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in diameter, distal part (
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) ascending to vertical with prop roots. Roots grey, 
<quantity box="[993,1113,1272,1299]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="1" pageNumber="65" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
in diameter. Rhizomes with regularly repeating units, each comprising a few distichously arranged phyllomes (cataphylls followed by a foliage leaf). Cataphylls oblong, 
<quantity box="[423,520,1344,1371]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="1" pageNumber="65" unit="cm" value="12.0" valueMax="15.0" valueMin="9.0">9–15 cm</quantity>
long, 
<quantity box="[593,698,1344,1371]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="65" unit="cm" value="1.15" valueMax="1.3" valueMin="1.0">1–1.3 cm</quantity>
wide, rapidly withering. Foliage leaves green, divided into petiole and lamina. Petiole green, adaxially sulcate, 
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long, 
<quantity box="[858,953,1380,1407]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="65" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
in diameter. Lamina green, elliptic, basally cuneate, apically narrowly acuminate, 
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long, 
<quantity box="[748,886,1416,1443]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="1" pageNumber="65" unit="cm" value="9.5" valueMax="10.5" valueMin="8.5">8.5–10.5 cm</quantity>
wide, parallel-veined with median vein prominent on abaxial surface, margin entire. Peduncle (specialized reproductive shoot) arising horizontally and arching down, purple mottled, 
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long, 
<quantity box="[501,594,1488,1515]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="65" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
in diameter, with 5–6 dark green and purple mottled, broadly ovate, 
<quantity box="[1359,1452,1488,1515]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="1" pageNumber="65" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
long, 
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wide distichously arranged sterile bracts. Flower solitary at the top of the peduncle. Perigone broadly obovate, globoid, container-shaped (urceolate with inflexed lobes), down faced, white outside, purple-red inside, fleshy, ca. 
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thick; tube widely campanulate, 
<quantity box="[620,725,1596,1623]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="1" pageNumber="65" unit="cm" value="2.85" valueMax="3.0" valueMin="2.7">2.7–3 cm</quantity>
long, 
<quantity box="[799,904,1596,1623]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" metricValueMax="5.0" metricValueMin="3.6" pageId="1" pageNumber="65" unit="cm" value="4.3" valueMax="5.0" valueMin="3.6">3.6–5 cm</quantity>
in diameter; lobes 6, widely ovate to triangular, incurved forming a star-like perigone opening, 
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in diameter. Stamens 6, inserted at base of perigone tube at the same radii with perigone lobes; anthers subsessile, yellow, ovoid, 
<quantity box="[947,1012,1668,1695]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="65" unit="mm" value="2.0">2 mm</quantity>
long, 
<quantity box="[1083,1218,1668,1695]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="1" pageNumber="65" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
wide, introrse. Pistil uniformly white, widely obconic to club-shaped, 
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long; ovary superior, inconspicuous, 3-locular; style widely obconic, roundish hexagonal in cross section, with 12–13 longitudinal ridges, up to 
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long, 
<quantity box="[1248,1315,1740,1767]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="65" unit="mm" value="4.0">4 mm</quantity>
in diameter at base; stigma disk-shaped, fleshy, 
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in diameter, upper surface tuberose with 3 radial bifurcated grooves, hexagonal, side surface with 20 prominent longitudinal ridges. Fruits dull green to brow, subspherical, 
<quantity box="[1288,1423,1812,1839]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="1" pageNumber="65" unit="mm" value="2.05" valueMax="2.5" valueMin="1.6">1.6–2.5 mm</quantity>
in diameter, with conical protuberances 
<quantity box="[556,670,1848,1875]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="1" pageNumber="65" unit="mm" value="1.75" valueMax="2.5" valueMin="1.0">1–2.5 mm</quantity>
long.
</paragraph>
<paragraph blockId="1.[136,1452,1236,2092]" pageId="1" pageNumber="65">
<materialsCitation collectingDate="2023-03-11" collectionCode="VNMN, SGN, MW" collectorName="Dien Dinh &amp; Quoc Tuan Doan &amp; Vu Tien Chinh &amp; Le Ngoc Tuan &amp; Le Van Huong &amp; Tran Nam Thang &amp; Tuan Anh Le" country="Vietnam" county="Phong Dien Nature Reserve" elevation="350" latitude="16.40409" location="Thua Thien Hue province" longLatPrecision="1" longitude="107.224335" municipality="Phong Xuan" pageId="1" pageNumber="65" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[189,308,1884,1910]" pageId="1" pageNumber="65">
<typeStatus box="[189,302,1884,1910]" pageId="1" pageNumber="65">Paratype</typeStatus>
:
</emphasis>
— 
<collectingCountry box="[337,469,1884,1910]" name="Vietnam" pageId="1" pageNumber="65">VIETNAM</collectingCountry>
. 
<location LSID="urn:lsid:plazi:treatment:03A1B80DDE49FFDDFF43B0E4FE07924B:8ED75FC0DE49FFDFFE2AB40CFCA39241" box="[481,782,1884,1911]" country="Vietnam" county="Phong Dien Nature Reserve" latitude="16.40409" longLatPrecision="1" longitude="107.224335" municipality="Phong Xuan" name="Thua Thien Hue province" pageId="1" pageNumber="65">Thua Thien Hue province</location>
: 
<location LSID="urn:lsid:plazi:treatment:03A1B80DDE49FFDDFF43B0E4FE07924B:8ED75FC0DE49FFDFFCD7B40CFC539241" box="[796,1022,1884,1910]" country="Vietnam" county="Phong Dien Nature Reserve" latitude="16.40409" longLatPrecision="1" longitude="107.224335" municipality="Phong Xuan" name="Phong Dien district" pageId="1" pageNumber="65">Phong Dien district</location>
, 
<collectingMunicipality box="[1036,1179,1884,1911]" pageId="1" pageNumber="65">Phong Xuan</collectingMunicipality>
commune, 
<collectingCounty pageId="1" pageNumber="65">Phong Dien Nature Reserve</collectingCounty>
, 
<geoCoordinate box="[324,492,1921,1948]" degrees="16" direction="north" minutes="24" orientation="latitude" pageId="1" pageNumber="65" precision="1" seconds="14.73" value="16.40409">16°24′14.73″N</geoCoordinate>
, 
<geoCoordinate box="[506,685,1921,1948]" degrees="107" direction="east" minutes="13" orientation="longitude" pageId="1" pageNumber="65" precision="1" seconds="27.59" value="107.224335">107°13′27.59″E</geoCoordinate>
, 
<elevation box="[699,831,1920,1947]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="65" unit="m" value="350.0">
<quantity box="[699,770,1920,1947]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="65" unit="m" value="350.0">350 m</quantity>
a.s.l.
</elevation>
, 
<date box="[844,1017,1920,1946]" pageId="1" pageNumber="65" value="2023-03-11">
<collectingDate box="[844,1017,1920,1946]" pageId="1" pageNumber="65" value="2023-03-11">11 March 2023</collectingDate>
</date>
, 
<emphasis box="[1028,1146,1920,1946]" italics="true" pageId="1" pageNumber="65">
<collectorName box="[1028,1146,1920,1946]" pageId="1" pageNumber="65">Dien Dinh</collectorName>
</emphasis>
, 
<emphasis box="[1160,1353,1920,1946]" italics="true" pageId="1" pageNumber="65">
<collectorName box="[1160,1353,1920,1946]" pageId="1" pageNumber="65">Quoc Tuan Doan</collectorName>
</emphasis>
, 
<emphasis italics="true" pageId="1" pageNumber="65">
<collectorName pageId="1" pageNumber="65">Vu Tien Chinh</collectorName>
</emphasis>
, 
<emphasis box="[219,376,1957,1982]" italics="true" pageId="1" pageNumber="65">
<collectorName box="[219,376,1957,1982]" pageId="1" pageNumber="65">Le Ngoc Tuan</collectorName>
</emphasis>
, 
<emphasis box="[390,550,1957,1982]" italics="true" pageId="1" pageNumber="65">
<collectorName box="[390,550,1957,1982]" pageId="1" pageNumber="65">Le Van Huong</collectorName>
</emphasis>
, 
<emphasis box="[564,756,1956,1982]" italics="true" pageId="1" pageNumber="65">
<collectorName box="[564,756,1956,1982]" pageId="1" pageNumber="65">Tran Nam Thang</collectorName>
</emphasis>
, 
<emphasis box="[769,1010,1956,1982]" italics="true" pageId="1" pageNumber="65">
<collectorName box="[769,912,1956,1982]" pageId="1" pageNumber="65">Tuan Anh Le</collectorName>
LTA s.n.
</emphasis>
(
<collectionCode box="[1024,1111,1956,1982]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="1" pageNumber="65">VNMN</collectionCode>
!, 
<collectionCode box="[1135,1191,1956,1982]" country="0" httpUri="http://grbio.org/cool/fyzf-afsy" name="Southern Institute of Ecology" pageId="1" pageNumber="65" type="Herbarium">SGN</collectionCode>
!, 
<collectionCode box="[1214,1275,1956,1982]" country="Netherlands" name="Museum Wasmann" pageId="1" pageNumber="65">MW</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="67" pageId="1" pageNumber="65" type="etymology">
<paragraph blockId="1.[136,1452,1236,2092]" pageId="1" pageNumber="65">
<emphasis bold="true" box="[189,328,1992,2018]" pageId="1" pageNumber="65">Etymology:</emphasis>
—The specific epithet is derived from its 
<typeStatus box="[813,861,1993,2019]" pageId="1" pageNumber="65">type</typeStatus>
locality, in Phong Dien Nature Reserve, located in Phong Dien district, Thua Thien Hue province, central 
<collectingCountry box="[752,851,2028,2054]" name="Vietnam" pageId="1" pageNumber="65">Vietnam</collectingCountry>
.
</paragraph>
<paragraph blockId="1.[136,1452,1236,2092]" box="[189,645,2064,2092]" pageId="1" pageNumber="65">
<emphasis bold="true" box="[189,405,2064,2090]" pageId="1" pageNumber="65">Vernacular name:</emphasis>
—Tỏi đá phong điễn.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7784282" ID-Zenodo-Dep="7784282" httpUri="https://zenodo.org/record/7784282/files/figure.png" pageId="2" pageNumber="66" startId="2.[136,229,1880,1902]" targetBox="[227,1361,190,1855]" targetPageId="2">
<paragraph blockId="2.[136,1452,1880,1974]" pageId="2" pageNumber="66">
<emphasis bold="true" box="[136,255,1880,1902]" pageId="2" pageNumber="66">FIGURE 1.</emphasis>
<taxonomicName authorityName="D. Dien, T. A. Le &amp; Vislobokov" authorityYear="2023" box="[261,494,1880,1901]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="66" phylum="Tracheophyta" rank="species" species="phongdiensis">
<emphasis box="[261,494,1880,1901]" italics="true" pageId="2" pageNumber="66">Aspidistra phongdiensis</emphasis>
</taxonomicName>
. A. Habit, scale bar: 10 cm; B. Flower, apical view; C. Sterile bracts of peduncle, scale bar: 1 cm; D. Flower, lateral view, scale bar: 5 cm; E. Perigone, dissected to show stamens and pistil, scale bar: 2 cm; F. Pistil, scale bar: 5 mm; G. Stamens, scale bar: 2 mm; H. Fruits, scale bars: 2 cm.
</paragraph>
</caption>
<paragraph blockId="3.[136,1451,158,329]" box="[189,1264,158,184]" pageId="3" pageNumber="67">
<emphasis bold="true" box="[189,323,158,184]" pageId="3" pageNumber="67">Phenology:</emphasis>
—Flowering from March to April, and fruiting from April to May in natural habitat.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="67" type="distribution">
<paragraph blockId="3.[136,1451,158,329]" pageId="3" pageNumber="67">
<emphasis bold="true" box="[189,502,193,220]" pageId="3" pageNumber="67">Distribution and ecology:</emphasis>
— 
<taxonomicName authorityName="D. Dien, T. A. Le &amp; Vislobokov" authorityYear="2023" box="[530,804,194,220]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="phongdiensis">
<emphasis box="[530,804,194,220]" italics="true" pageId="3" pageNumber="67">Aspidistra phongdiensis</emphasis>
</taxonomicName>
is only known from 
<typeStatus box="[1061,1109,194,220]" pageId="3" pageNumber="67">type</typeStatus>
locality where it is common (
<figureCitation box="[144,215,230,256]" captionStart="FIGURE 2" captionStartId="3.[136,229,1281,1303]" captionTargetBox="[177,1423,370,1242]" captionTargetId="figure-100@3.[151,1436,353,1257]" captionTargetPageId="3" captionText="FIGURE 2. The known geographical distribution of Aspidistra phongdiensis in Phong Dien Nature Reserve, located in Phong Dien district, Thua Thien Hue province, central Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7784284" httpUri="https://zenodo.org/record/7784284/files/figure.png" pageId="3" pageNumber="67">Fig. 2</figureCitation>
). The new species grows in primary and secondary evergreen broad-leaved lowland forest on alluvial soils at elevation around 
<quantity box="[328,398,266,293]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="67" unit="m" value="850.0">850 m</quantity>
a.s.l., commonly on shady reclined slopes in assemblage with 
<taxonomicName box="[1087,1189,266,292]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[1087,1149,267,292]" italics="true" pageId="3" pageNumber="67">Piper</emphasis>
sp.
</taxonomicName>
, 
<taxonomicName authority="Hance (1875: 289)" authorityName="Hance" authorityPageNumber="289" authorityYear="1875" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="tetradactylus">
<emphasis box="[1200,1451,266,292]" italics="true" pageId="3" pageNumber="67">Calamus tetradactylus</emphasis>
<bibRefCitation author="Hance, H. F." box="[136,348,302,329]" pageId="3" pageNumber="67" pagination="289 - 290" refId="ref3724" refString="Hance, H. F. (1875) A fourth new Hong Kong Calamus. Journal of Botany, British and Foreign. 13: 289 - 290." type="journal article" year="1875">Hance (1875: 289)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Loureiro (1790: 527)" authorityName="Loureiro" authorityPageNumber="527" authorityYear="1790" box="[360,874,302,329]" class="Magnoliopsida" family="Melastomataceae" genus="Blastus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="cochinchinensis">
<emphasis box="[360,628,302,328]" italics="true" pageId="3" pageNumber="67">Blastus cochinchinensis</emphasis>
<bibRefCitation author="Loureiro, J." box="[635,874,302,329]" pageId="3" pageNumber="67" pagination="357 - 744" refId="ref4243" refString="Loureiro, J. (1790) Flora Cochinchinensis 2. Ulyssipone, pp. 357 - 744." type="book chapter" year="1790">Loureiro (1790: 527)</bibRefCitation>
</taxonomicName>
, etc.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7784284" ID-Zenodo-Dep="7784284" httpUri="https://zenodo.org/record/7784284/files/figure.png" pageId="3" pageNumber="67" startId="3.[136,229,1281,1303]" targetBox="[177,1423,370,1242]" targetPageId="3">
<paragraph blockId="3.[136,1452,1281,1339]" pageId="3" pageNumber="67">
<emphasis bold="true" box="[136,256,1281,1303]" pageId="3" pageNumber="67">FIGURE 2.</emphasis>
The known geographical distribution of 
<taxonomicName authorityName="D. Dien, T. A. Le &amp; Vislobokov" authorityYear="2023" box="[666,901,1281,1302]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="phongdiensis">
<emphasis box="[666,901,1281,1302]" italics="true" pageId="3" pageNumber="67">Aspidistra phongdiensis</emphasis>
</taxonomicName>
in Phong Dien Nature Reserve, located in Phong Dien district, Thua Thien Hue province, central Vietnam.
</paragraph>
</caption>
<paragraph blockId="3.[136,1452,1385,1916]" pageId="3" pageNumber="67">
<emphasis bold="true" box="[189,438,1385,1412]" pageId="3" pageNumber="67">Conservation status:</emphasis>
—Although the new species is located in a protected area, it grows near to the landslide area. Thus, following the IUCN Red List Categories and Criteria (IUCN 2021), it is provisionally assessed as Data Deficient (DD).
</paragraph>
<paragraph blockId="3.[136,1452,1385,1916]" pageId="3" pageNumber="67">
<emphasis bold="true" box="[189,491,1493,1519]" pageId="3" pageNumber="67">Taxonomic relationships:</emphasis>
—The new species is morphologically closest to 
<taxonomicName authorityName="Averyanov &amp; Tillich" authorityYear="2014" box="[1047,1163,1494,1520]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="khangii">
<emphasis box="[1047,1163,1494,1520]" italics="true" pageId="3" pageNumber="67">A. khangii</emphasis>
</taxonomicName>
by shape of perigone but differs in shape of leaf blade (elliptic 
<quantity box="[557,696,1530,1557]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="3" pageNumber="67" unit="cm" value="9.5" valueMax="10.5" valueMin="8.5">8.5–10.5 cm</quantity>
wide vs. narrowly elliptic 
<quantity box="[997,1080,1530,1556]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="3" pageNumber="67" unit="cm" value="4.5" valueMax="7.0" valueMin="2.0">2–7 cm</quantity>
wide), colour of perigone (white outside vs. purple mottled outside), colour of pollen (yellow vs. white), 
<typeStatus box="[931,979,1566,1592]" pageId="3" pageNumber="67">type</typeStatus>
of anther dehiscence (introrse vs. latrorse) and shape of pistil (widely obconic vs. desk-shaped) (
<bibRefCitation author="Averyanov, L. V. &amp; Tillich, H. J." box="[737,1036,1602,1628]" pageId="3" pageNumber="67" pagination="752 - 760" refId="ref2694" refString="Averyanov, L. V. &amp; Tillich, H. J. (2014) Aspidistra albopurpurea, A. khangii, A. lubae and A. stellata spp. nov. (Asparagaceae, Convallariaceae ss) from Indochina. Nordic Journal of Botany 32: 752 - 760. https: // doi. org / 10.1111 / j. 1756 - 1051.2013.00273. x" type="journal article" year="2014">Averyanov &amp; Tillich 2014</bibRefCitation>
, 
<bibRefCitation author="Averyanov, L. V. &amp; Nguyen, K. S. &amp; Son, H. T. &amp; Tillich, H. J. &amp; Maisak, T. V." box="[1045,1313,1602,1628]" pageId="3" pageNumber="67" pagination="02352" refId="ref2873" refString="Averyanov, L. V., Nguyen, K. S., Son, H. T., Tillich, H. J. &amp; Maisak, T. V. (2019 b) New taxa and taxonomic notes in Aspidistra (Convallariaceae s. s.) of Laos and Vietnam. Nordic Journal of Botany 37: e 02352. https: // doi. org / 10.1111 / njb. 01833" type="journal article" year="2019">
Averyanov 
<emphasis box="[1175,1231,1602,1628]" italics="true" pageId="3" pageNumber="67">et al.</emphasis>
2019b
</bibRefCitation>
).
</paragraph>
<paragraph blockId="3.[136,1452,1385,1916]" pageId="3" pageNumber="67">
In flower structure 
<taxonomicName authorityName="D. Dien, T. A. Le &amp; Vislobokov" authorityYear="2023" box="[413,595,1638,1664]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="phongdiensis">
<emphasis box="[413,595,1638,1664]" italics="true" pageId="3" pageNumber="67">A. phongdiensis</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName box="[807,891,1638,1664]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="locii">
<emphasis box="[807,891,1638,1664]" italics="true" pageId="3" pageNumber="67">A. locii</emphasis>
</taxonomicName>
but differs in colour of perigone (white outside, purple-red inside vs. uniformly grass green), presence of six perigone lobes (vs. absence of free lobes), number of stamens (6 vs. 12–21), 
<typeStatus box="[401,449,1710,1736]" pageId="3" pageNumber="67">type</typeStatus>
of anther dehiscence (introrse vs. latrorse) and shape of pistil (widely obconic vs. deskshaped) (
<bibRefCitation author="Bogner, J. &amp; Arnautov, N." box="[238,528,1746,1772]" pageId="3" pageNumber="67" pagination="203 - 208" refId="ref3429" refString="Bogner, J. &amp; Arnautov, N. (2004) Aspidistra locii (Convallariaceae), an unusual new species from Vietnam. Willdenowia 34: 203 - 208. [https: // www. jstor. org / stable / 3997475]" type="journal article" year="2004">Bogner &amp; Arnautov 2004</bibRefCitation>
, 
<bibRefCitation author="Averyanov, L. V. &amp; Nguyen, K. S. &amp; Son, H. T. &amp; Tillich, H. J. &amp; Maisak, T. V." box="[537,805,1746,1772]" pageId="3" pageNumber="67" pagination="02352" refId="ref2873" refString="Averyanov, L. V., Nguyen, K. S., Son, H. T., Tillich, H. J. &amp; Maisak, T. V. (2019 b) New taxa and taxonomic notes in Aspidistra (Convallariaceae s. s.) of Laos and Vietnam. Nordic Journal of Botany 37: e 02352. https: // doi. org / 10.1111 / njb. 01833" type="journal article" year="2019">
Averyanov 
<emphasis box="[667,723,1746,1772]" italics="true" pageId="3" pageNumber="67">et al.</emphasis>
2019b
</bibRefCitation>
).
</paragraph>
<paragraph blockId="3.[136,1452,1385,1916]" pageId="3" pageNumber="67">
In flower coloration the new species is similar to 
<taxonomicName authority="Hu, Cai &amp; Wang" authorityName="Hu, Cai &amp; Wang" box="[739,1171,1782,1808]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="67" phylum="Tracheophyta" rank="species" species="austroyunnanensis">
<emphasis box="[739,978,1782,1808]" italics="true" pageId="3" pageNumber="67">A. austroyunnanensis</emphasis>
Hu, Cai &amp; Wang
</taxonomicName>
in 
<bibRefCitation author="Cai, L. &amp; Peng, S. &amp; Tian, J. &amp; Dao, Z. L. &amp; Wei, N. &amp; Hu, G. W. &amp; Wang, Q. F." box="[1206,1447,1782,1808]" pageId="3" pageNumber="67" pagination="233 - 237" refId="ref3475" refString="Cai, L., Peng, S., Tian, J., Dao, Z. L., Wei, N., Hu, G. W. &amp; Wang, Q. F. (2018) Aspidistra austroyunnanensis (Asparagaceae), a new species from southern Yunnan, China. Phytotaxa 356: 233 - 237. https: // doi. org / 10.11646 / phytotaxa. 356.3.5" type="journal article" year="2018">
Cai 
<emphasis box="[1251,1307,1782,1808]" italics="true" pageId="3" pageNumber="67">et al.</emphasis>
(2018: 233)
</bibRefCitation>
, but differs in perigone lobes orientation (curved inward vs. curved outward), stamens morphology (subsessile anthers vs. with filaments 
<quantity box="[347,442,1854,1880]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="67" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long) and shape of pistil (widely obconic with longitudinal ridges vs. mushroom-shaped smooth) (
<bibRefCitation author="Cai, L. &amp; Peng, S. &amp; Tian, J. &amp; Dao, Z. L. &amp; Wei, N. &amp; Hu, G. W. &amp; Wang, Q. F." box="[243,414,1890,1916]" pageId="3" pageNumber="67" pagination="233 - 237" refId="ref3475" refString="Cai, L., Peng, S., Tian, J., Dao, Z. L., Wei, N., Hu, G. W. &amp; Wang, Q. F. (2018) Aspidistra austroyunnanensis (Asparagaceae), a new species from southern Yunnan, China. Phytotaxa 356: 233 - 237. https: // doi. org / 10.11646 / phytotaxa. 356.3.5" type="journal article" year="2018">
Cai 
<emphasis box="[290,346,1890,1916]" italics="true" pageId="3" pageNumber="67">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>