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<mods:title id="CE0390D919540AA8A9BA9D9AB6643B60">Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)</mods:title>
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<taxonomicName id="658FC8F84B4FFF83E7B8F9AFFD88F983" ID-CoL="84W7P" ID-ENA="1240031" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[151,579,1538,1565]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="euphrosyne">
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<emphasis id="90FB6F694B4FFF83E7B8F9AFFE45F982" bold="true" box="[151,398,1538,1565]" italics="true" pageId="11" pageNumber="12">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B4FFF83E6BAF9AEFD88F983" author="De Man, J. G." box="[405,579,1538,1565]" pageId="11" pageNumber="12" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
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<paragraph id="A230B37B4B4FFF83E7B8F98AFDF2F9DE" blockId="11.[151,579,1538,1601]" box="[151,569,1574,1601]" pageId="11" pageNumber="12">
(
<figureCitation id="3AB4AFFE4B4FFF83E7B0F98AFF26F9DE" box="[159,237,1575,1601]" captionStart="FIGURE 1" captionStartId="4.[151,250,1452,1476]" captionTargetBox="[167,1423,198,1420]" captionTargetId="figure-21@4.[152,1435,181,1427]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus euphrosyne De Man, 1897: syntype, female (cl 46.0 mm) from Java Sea, Indonesia (after De Man 1897, specimen lost) [A–E]; male (cl indet., tl 34.0 mm) from Bangkok, Thailand (after De Man 1898) [F, G]; A—frontal region, dorsal view; B—female minor (left) cheliped, lateral view; C—second pereiopod, lateral view; D—third pereiopod, lateral view; E—sixth pleonite, telson and uropods, dorsal view; F—male minor (right) cheliped, dorsolateral view; G—same, minor chela, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7912294" httpUri="https://zenodo.org/record/7912294/files/figure.png" pageId="11" pageNumber="12">Figs. 1</figureCitation>
, 
<figureCitation id="3AB4AFFE4B4FFF83E7D4F98AFEFDF9DE" box="[251,310,1575,1601]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStart-5="FIGURE 9" captionStart-6="FIGURE 10" captionStartId-0="13.[151,250,1688,1712]" captionStartId-1="14.[151,250,1935,1959]" captionStartId-2="15.[151,250,1807,1831]" captionStartId-3="16.[151,250,1796,1820]" captionStartId-4="19.[151,250,1902,1926]" captionStartId-5="20.[151,250,1875,1899]" captionStartId-6="21.[151,250,1866,1890]" captionTargetBox-0="[151,1435,813,1665]" captionTargetBox-1="[151,1435,379,1912]" captionTargetBox-2="[203,1391,197,1776]" captionTargetBox-3="[151,1424,192,1761]" captionTargetBox-4="[151,1435,181,1877]" captionTargetBox-5="[151,1435,181,1851]" captionTargetBox-6="[159,1429,181,1842]" captionTargetId-0="figure-245@13.[151,1435,813,1665]" captionTargetId-1="figure-95@14.[151,1435,379,1912]" captionTargetId-2="figure-15@15.[183,1404,181,1784]" captionTargetId-3="figure-21@16.[151,1436,181,1772]" captionTargetId-4="figure-15@19.[151,1435,181,1878]" captionTargetId-5="figure-21@20.[151,1435,181,1851]" captionTargetId-6="figure-15@21.[159,1429,181,1842]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="19" captionTargetPageId-5="20" captionTargetPageId-6="21" captionText-0="FIGURE 4. Alpheus euphrosyne De Man, 1897: chelipeds of male (cl indet.) from Sumatra, Indonesia (ZMA Crus.D.202726) showing main features and terminology used in the present study; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (left) cheliped, mesial view; D—same, lateral view. Abbreviations (in alphabetical order): ad—adhesive disks (plaques) on dactylus and palm (small on major chela, highly reduced in minor chela); be—balaeniceps expansion of minor chela dactylus; ddr—dorsal ridge of major chela dactylus (dactylar dorsal ridge); dla—dorsolateral angle of major chela pollex; dlt—dorsolateral tooth of major / minor chela palm; dma—dorsomesial angle of major chela pollex; dmt—distomesial tooth of major / minor chela palm; dp—plunger of major chela dactylus (dactylar plunger); ds—dorsal shoulder of major / minor chela palm; dtg—dorsal transverse groove of major / minor chela palm (also known as dorsal notch); lbd—lateral balaeniceps ridge of minor chela dactylus; lbp—lateral balaeniceps ridge of minor chela pollex; li—linea impressa of major / minor chela palm; lld—lateral longitudinal depression (groove) of major / minor chela palm; ltr—smooth lateral transverse ridge of major chela palm (often poorly developed); mbd—mesial balaeniceps ridge of minor chela dactylus; mbp—mesial balaeniceps ridge of minor chela pollex; mld—mesial longitudinal depression (groove) of major / minor chela palm; msr—mesial subdistal ridge on major chela pollex; mtr—smooth mesial transverse ridge of major chela palm (often poorly developed); pf—fossa (socket) of major chela pollex; vs—ventral shoulder of major / minor chela palm; vtg—ventral transverse groove of major / minor chela palm (also known as ventral notch)." captionText-1="FIGURE 5. Alpheus euphrosyne De Man, 1897: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela in situ, mesial view; D—same, lateral view; E—minor (right) chela in situ, dorsal view. Refer to A for scale." captionText-2="FIGURE 6. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) from Angsila, Thailand (MNHN-IU-2018-5593); A— frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H, I—same, dactylus, mesial (H) and dorsal (I) views; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—second pleopod, appendices masculina and interna, anterior view; M—left uropod, dorsal view; N—right uropod, detail of distolateral angle." captionText-3="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." captionText-4="FIGURE 8. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Satun, Thailand, in lateral (A) and dorsal (B) views (deposition unknown). Photographs courtesy of Rueangrit Promdam." captionText-5="FIGURE 9. Alpheus euphrosyne De Man, 1897: male (cl 16.8 mm) from Pattani, Thailand (ZRC 2014.0672), in dorsal view (A), with details of the frontal area and chelipeds (B). Photographs courtesy of Sukree Hajisamae." captionText-6="FIGURE 10. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Nha Trang Bay, Vietnam (ZMMU) [A]; male (cl indet.) from unknown locality in Taiwan (AST) [B]; male from Dongtou Island, Zhejiang, China (IOCAS-MBM188857) [C]; male (cl indet.) from Siracha, Thailand (ZRC 2000.2137) [D]; male (cl indet.) from Kochi, India (RMNH Crus.D.49747) [E, F]; A, B, C—habitus, dorsal view; D, E—same, lateral view; F—posterior pleon and tail fan, dorsal view. Photographs courtesy of Oleg Savinkin (A), Ming-Shiou Jeng (B), Xinzheng Li (C), Peter K.L. Ng (D) and Juan C.E. Miquel (E, F)." figureDoi-0="http://doi.org/10.5281/zenodo.7912302" figureDoi-1="http://doi.org/10.5281/zenodo.7912304" figureDoi-2="http://doi.org/10.5281/zenodo.7912306" figureDoi-3="http://doi.org/10.5281/zenodo.7912310" figureDoi-4="http://doi.org/10.5281/zenodo.7912312" figureDoi-5="http://doi.org/10.5281/zenodo.7912318" figureDoi-6="http://doi.org/10.5281/zenodo.7912320" httpUri-0="https://zenodo.org/record/7912302/files/figure.png" httpUri-1="https://zenodo.org/record/7912304/files/figure.png" httpUri-2="https://zenodo.org/record/7912306/files/figure.png" httpUri-3="https://zenodo.org/record/7912310/files/figure.png" httpUri-4="https://zenodo.org/record/7912312/files/figure.png" httpUri-5="https://zenodo.org/record/7912318/files/figure.png" httpUri-6="https://zenodo.org/record/7912320/files/figure.png" pageId="11" pageNumber="12">4–10</figureCitation>
, 
<figureCitation id="3AB4AFFE4B4FFF83E66EF98BFEBFF9DE" box="[321,372,1574,1601]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="11" pageNumber="12">51A</figureCitation>
, 
<figureCitation id="3AB4AFFE4B4FFF83E650F98BFE79F9DE" box="[383,434,1574,1601]" captionStart="FIGURE 53" captionStartId="107.[151,250,1859,1883]" captionTargetBox="[239,1349,192,1835]" captionTargetId="figure-15@107.[239,1349,181,1835]" captionTargetPageId="107" captionText="FIGURE 53. Alpheus euphrosyne De Man, 1897 complex in the northern and western Indian Ocean: A—Alpheus cf. mangalis sp. nov. (upper row) and A. euphrosyne (lower row) from an estuary near Kochi, India; B, C—Alpheus cf. eurydactylus De Man, 1920, ovigerious female (cl indet.) from Kochi, India (RMNH.Crus.D.49749), lateral view (B) and pleon and tail fan, dorsal view; D, E—Alpheus sp. euphrosyne complex (?), female photographed in situ in a mangrove at Watamu, Kenya. Photographs courtesy of Mahadevan Harikrishnan (A), Juan C. Miquel (B, C) and Dawn Goebbels (D, E)." figureDoi="http://doi.org/10.5281/zenodo.7912459" httpUri="https://zenodo.org/record/7912459/files/figure.png" pageId="11" pageNumber="12">53A</figureCitation>
, lower row)
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<emphasis id="90FB6F694B4FFF83E7B8F9C6FEAFF91C" box="[151,356,1643,1667]" italics="true" pageId="11" pageNumber="12">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B4FFF83E644F9C1FDE6F91C" author="De Man, J. G." box="[363,557,1643,1667]" pageId="11" pageNumber="12" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man 1897: 745</bibRefCitation>
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, pl. 36, fig. 64; 
<treatmentCitation id="232E956A4B4FFF83E5FEF9C1FC58F91C" author="De Man, J. G." box="[721,915,1643,1667]" page="317" pageId="11" pageNumber="12" year="1898">
<bibRefCitation id="C61ECE8A4B4FFF83E5FEF9C1FC58F91C" author="De Man, J. G." box="[721,915,1643,1667]" pageId="11" pageNumber="12" pagination="309 - 325" refId="ref70903" refString="De Man, J. G. (1898) Note sur quelques especes du genre Alpheus Fabr., appartenant a la section dont l' Alpheus Edwardsii est le representant. Memoires de la Societ Zoologique de France, 11, 309 - 325, pl. 4." type="journal article" year="1898">De Man 1898: 317</bibRefCitation>
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, pl. 4, fig. 2; 
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<bibRefCitation id="C61ECE8A4B4FFF83E332F9C1FB2BF91C" author="De Man, J. G." box="[1053,1248,1643,1667]" pageId="11" pageNumber="12" pagination="133 - 465" refId="ref70992" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga-Expeditie, 39 a 1 (2), 133 - 465; (1915) Supplement - Explanations of plates of Alpheidae, pls. 1 - 23. E. J. Brill, Leiden. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article" year="1911">De Man 1911: 413</bibRefCitation>
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; 
<treatmentCitation id="232E956A4B4FFF83E3C5F9C6FA53F91C" author="Ahmad, N." box="[1258,1432,1643,1667]" page="16" pageId="11" pageNumber="12" year="1957">
<bibRefCitation id="C61ECE8A4B4FFF83E3C5F9C6FA53F91C" author="Ahmad, N." box="[1258,1432,1643,1667]" pageId="11" pageNumber="12" refId="ref68058" refString="Ahmad, N. (1957) Prawn and prawn fishery of East Pakistan. Directorate of Fisheries, Government of East Pakistan, 31 pp." type="book" year="1957">Ahmad 1957: 16</bibRefCitation>
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; 
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(part.?, not fig. 49); 
<treatmentCitation id="232E956A4B4FFF83E59AF927FCBEF93D" author="Thomas, M. M." box="[693,885,1674,1698]" page="667" pageId="11" pageNumber="12" year="1976">
<bibRefCitation id="C61ECE8A4B4FFF83E59AF927FCBEF93D" author="Thomas, M. M." box="[693,885,1674,1698]" pageId="11" pageNumber="12" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976: 667</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="232E956A4B4FFF83E450F927FB83F93D" author="Holthuis, L. B." box="[895,1096,1674,1698]" page="119" pageId="11" pageNumber="12" year="1980">
<bibRefCitation id="C61ECE8A4B4FFF83E450F927FB83F93D" author="Holthuis, L. B." box="[895,1096,1674,1698]" pageId="11" pageNumber="12" refId="ref71792" refString="Holthuis, L. B. (1980) FAO species catalogue. Vol. 1. Shrimps and prawns of the world. An annotated catalogue of species of interest to fisheries. FAO Fisheries Synopsis 125. FAO, Rome, xvii + 271 pp." type="book" year="1980">Holthuis, 1980: 119</bibRefCitation>
</treatmentCitation>
(part.?); (?) 
<treatmentCitation id="232E956A4B4FFF83E3E3F927FABDF93D" author="Becker, K." box="[1228,1398,1674,1698]" page="95" pageId="11" pageNumber="12" year="1996">
<bibRefCitation id="C61ECE8A4B4FFF83E3E3F927FABDF93D" author="Becker, K." box="[1228,1398,1674,1698]" pageId="11" pageNumber="12" pagination="92 - 104" refId="ref69673" refString="Becker, K. (1996) Epibionts on carapaces of some malacostracan crustaceans from the Gulf of Thailand. Journal of Crustacean Biology, 16, 92 - 104. https: // doi. org / 10.2307 / 1548934" type="journal article" year="1996">Becker 1996: 95</bibRefCitation>
</treatmentCitation>
; (?) 
<treatmentCitation id="232E956A4B4FFF83E7E8F905FE77F95F" author="Becker, K. &amp; Hormchong, T. &amp; Wahl, M." box="[199,444,1704,1729]" page="195" pageId="11" pageNumber="12" year="2000">
<bibRefCitation id="C61ECE8A4B4FFF83E7E8F905FE77F95F" author="Becker, K. &amp; Hormchong, T. &amp; Wahl, M." box="[199,444,1704,1729]" pageId="11" pageNumber="12" pagination="193 - 201" refId="ref69714" refString="Becker, K., Hormchong, T. &amp; Wahl, M. (2000) Relevance of crustacean carapace wettability for fouling. Hydrobiologia, 426, 193 - 201. https: // doi. org / 10.1023 / A: 1003918512565" type="journal article" year="2000">
Becker 
<emphasis id="90FB6F694B4FFF83E639F904FE80F95F" box="[278,331,1704,1728]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2000: 195
</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="232E956A4B4FFF83E6E7F905FD51F95E" author="Naiyanetr, P." box="[456,666,1704,1729]" page="173" pageId="11" pageNumber="12" year="2007">
<bibRefCitation id="C61ECE8A4B4FFF83E6E7F905FD51F95E" author="Naiyanetr, P." box="[456,666,1704,1729]" pageId="11" pageNumber="12" refId="ref73355" refString="Naiyanetr, P. (2007) Checklist of crustacean fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">Naiyanetr 2007: 173</bibRefCitation>
</treatmentCitation>
(part.?, referring to 
<bibRefCitation id="C61ECE8A4B4FFF83E442F904FBABF95F" author="Banner, A. H. &amp; Banner, D. M." box="[877,1120,1704,1729]" pageId="11" pageNumber="12" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
); 
<treatmentCitation id="232E956A4B4FFF83E35CF905FF26F940" author="Harikrishnan, M. &amp; Kurup, B. M." page="111" pageId="11" pageNumber="12" year="2008">
<bibRefCitation id="C61ECE8A4B4FFF83E35CF905FF26F940" author="Harikrishnan, M. &amp; Kurup, B. M." pageId="11" pageNumber="12" pagination="1 - 280" refId="ref71572" refString="Harikrishnan, M. &amp; Kurup, B. M. (2008) Potential of snapping shrimps as ornamentals. In: Kurup, B. M., Boopendranth, M. R., Ravindran, K., Sairabanu &amp; Nair, A. G. (Eds.), Ornamental fish, breeding, farming and trade. Department of Fisheries, Government of Kerala, Kerala, pp. 1 - 280." type="book chapter" year="2008">Harikrishnan &amp; Kurup 2008: 111</bibRefCitation>
</treatmentCitation>
, fig. 1 (part., specimen on the left photograph); 
<treatmentCitation id="232E956A4B4FFF83E5FCF965FC77F940" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[723,956,1735,1759]" page="1695" pageId="11" pageNumber="12" year="2019">
<bibRefCitation id="C61ECE8A4B4FFF83E5FCF965FC77F940" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[723,956,1735,1759]" pageId="11" pageNumber="12" pagination="1695 - 1698" refId="ref73799" refString="Paras, N. J., Purushothaman, P., Chinnadurai, S., Renjith R. K., Remesan, M. P., Baiju, M. V. &amp; Edwin, L. (2019) Occurrence of Alpheus euphrosyne de Man, 1897 (Crustacea: Decapoda: Alpheidae) from the south-eastern Arabian Sea, India. Indian Journal of Geo Marine Sciences, 48, 1695 - 1698." type="journal article" year="2019">
Paras 
<emphasis id="90FB6F694B4FFF83E43FF965FCF6F940" box="[784,829,1735,1759]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. 2019: 1695
</bibRefCitation>
</treatmentCitation>
(part., not description), figs. 1–3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B4FFF83E7B8F94BFD4CF961" blockId="11.[151,1437,1643,2035]" box="[151,647,1766,1790]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="829F94554B4FFF83E7B8F94BFD4CF961" box="[151,647,1766,1790]" pageId="11" pageNumber="12">
<emphasis id="90FB6F694B4FFF83E7B8F94BFE42F961" box="[151,393,1766,1790]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="658FC8F84B4FFF83E7B8F94BFEA0F961" ID-CoL="84Z63" baseAuthorityName="De Man" baseAuthorityYear="1897" box="[151,363,1766,1790]" class="Malacostraca" family="Alpheidae" genus="Crangon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="euphrosyne">Crangon euphrosyne</taxonomicName>
.—
</emphasis>
<treatmentCitation id="232E956A4B4FFF83E6A7F94BFDE4F961" author="Suvatti, C." box="[392,559,1766,1790]" page="48" pageId="11" pageNumber="12" year="1937">
<bibRefCitation id="C61ECE8A4B4FFF83E6A7F94BFDE4F961" author="Suvatti, C." box="[392,559,1766,1790]" pageId="11" pageNumber="12" refId="ref74577" refString="Suvatti, C. (1937) A check-list of aquatic fauna of Siam (excluding fishes). Bureau of Fisheries, Bangkok, 116 pp." type="book" year="1937">Suvatti 1937: 48</bibRefCitation>
</treatmentCitation>
(part.?).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B4FFF83E7B8F8A9FBD3F883" blockId="11.[151,1437,1643,2035]" box="[151,1048,1796,1821]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="829F94554B4FFF83E7B8F8A9FBD3F883" box="[151,1048,1796,1821]" pageId="11" pageNumber="12">
<emphasis id="90FB6F694B4FFF83E7B8F8A9FE2FF883" box="[151,484,1796,1820]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="658FC8F84B4FFF83E7B8F8A9FE2BF883" ID-CoL="84W7P" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[151,480,1796,1820]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">Alpheus euphrosyne euphrosyne</taxonomicName>
.
</emphasis>
— 
<bibRefCitation id="C61ECE8A4B4FFF83E6D2F8A9FD43F882" author="Li, X. &amp; Shen, A. &amp; Xu, Z." box="[509,648,1796,1821]" pageId="11" pageNumber="12" refId="ref72773" refString="Li, X., Shen, A. &amp; Xu, Z. (2008) Report on some shrimps from Dongtou Island, Zhejiang, China (Decapoda, Natantia). Crustaceana, 81, 1371 - 1330. https: // doi. org / 10.1163 / 156854008 X 369500" type="book" year="2008">
Li 
<emphasis id="90FB6F694B4FFF83E535F8A8FD8DF883" box="[538,582,1796,1820]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. 2008
</bibRefCitation>
, fig. 2A; 
<treatmentCitation id="232E956A4B4FFF83E5C8F8A9FBDFF883" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." box="[743,1044,1796,1821]" page="195" pageId="11" pageNumber="12" year="2010">
<bibRefCitation id="C61ECE8A4B4FFF83E5C8F8A9FBDFF883" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." box="[743,1044,1796,1821]" pageId="11" pageNumber="12" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B4FFF83E45DF8A8FC6EF883" box="[882,933,1796,1820]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2010: 195
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B4FFF83E7B8F88EFEDEF8E7" blockId="11.[151,1437,1643,2035]" pageId="11" pageNumber="12">
<taxonomicName id="658FC8F84B4FFF83E7B8F88EFEACF8A4" authority="Anker &amp; De Grave 2016" box="[151,359,1827,1851]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="104" phylum="Arthropoda" rank="subSpecies" species="euphrosyne">
<emphasis id="90FB6F694B4FFF83E7B8F88EFEACF8A4" box="[151,359,1827,1851]" italics="true" pageId="11" pageNumber="12">Alpheus euphrosyne</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8BC8D2124B4FFF83E65EF889FE62F8A4" box="[369,425,1828,1851]" pageId="11" pageNumber="12" sensu="lato">sensu</taxonomicNameLabel>
lato.— 
<bibRefCitation id="C61ECE8A4B4FFF83E6D8F88EFCF0F8A4" author="Anker, A. &amp; De Grave, S." box="[503,827,1827,1851]" pageId="11" pageNumber="12" pagination="343 - 454" refId="ref68774" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave 2016: 355</bibRefCitation>
[specimen dredged off the eastern Straits of 
<collectingRegion id="604B7D994B4FFF83E20EF88EFA96F8A4" box="[1313,1373,1827,1851]" country="Malaysia" name="Johor" pageId="11" pageNumber="12">Johor</collectingRegion>
; most Singaporean specimens are 
<taxonomicName id="658FC8F84B4FFF83E6C3F8EFFDA2F8C5" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[492,617,1858,1882]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B4FFF83E6C3F8EFFDA2F8C5" box="[492,617,1858,1882]" italics="true" pageId="11" pageNumber="12">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B4FFF83E55CF8EFFD0CF8C5" bold="true" box="[627,711,1858,1882]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="8BC8D2124B4FFF83E55CF8EFFD0CF8C5" box="[627,711,1858,1882]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="658FC8F84B4FFF83E42FF8EFFC6AF8C5" ID-CoL="85LHH" authority="De Man, 1920" box="[768,929,1858,1882]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B4FFF83E42FF8EFFC6AF8C5" box="[768,929,1858,1882]" italics="true" pageId="11" pageNumber="12">A. eurydactylus</emphasis>
</taxonomicName>
; at least 
<specimenCount id="B48978F24B4FFF83E328F8EFFB5DF8C5" box="[1031,1174,1858,1882]" pageId="11" pageNumber="12" type="generic">one specimen</specimenCount>
is 
<taxonomicName id="658FC8F84B4FFF83E395F8EFFAC6F8C5" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1210,1293,1858,1882]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B4FFF83E395F8EFFAC6F8C5" box="[1210,1293,1858,1882]" italics="true" pageId="11" pageNumber="12">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B4FFF83E238F8EFFAA0F8C5" bold="true" box="[1303,1387,1858,1882]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="8BC8D2124B4FFF83E238F8EFFAA0F8C5" box="[1303,1387,1858,1882]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, see below].
</paragraph>
<paragraph id="A230B37B4B4FFF84E7B8F82DFAD8FE94" blockId="11.[151,1437,1643,2035]" lastBlockId="12.[151,1437,151,451]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<collectingRegion id="604B7D994B4FFF83E7B8F82DFF76F808" box="[151,189,1920,1943]" country="Maldives" name="Uthuru" pageId="11" pageNumber="12">Not</collectingRegion>
<emphasis id="90FB6F694B4FFF83E7ECF8D2FE5DF808" box="[195,406,1919,1943]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="658FC8F84B4FFF83E7ECF8D2FE59F808" ID-CoL="84W7P" authorityName="sensu De Man" authorityYear="1897" box="[195,402,1919,1943]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="euphrosyne">Alpheus euphrosyne</taxonomicName>
.
</emphasis>
— 
<bibRefCitation id="C61ECE8A4B4FFF83E680F8D2FDABF808" author="Johnson, D. S." box="[431,608,1919,1943]" pageId="11" pageNumber="12" pagination="1 - 54" refId="ref72097" refString="Johnson, D. S. (1976) Prawns of the Malacca Straits and Singapore waters. Journal of the Marine Biological Association of India, 18, 1 - 54." type="journal article" year="1976">Johnson 1976: 37</bibRefCitation>
[= 
<taxonomicName id="658FC8F84B4FFF83E5ABF8D2FCC3F808" box="[644,776,1919,1943]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B4FFF83E5ABF8D2FD1DF808" box="[644,726,1919,1943]" italics="true" pageId="11" pageNumber="12">Alpheus</emphasis>
spp.
</taxonomicName>
, most probably 
<taxonomicName id="658FC8F84B4FFF83E486F8D2FB15F808" ID-CoL="85LHH" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[937,1246,1919,1943]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B4FFF83E486F8D2FB8DF808" box="[937,1094,1919,1943]" italics="true" pageId="11" pageNumber="12">A. eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B4FFF83E363F82DFB15F808" author="De Man, J. G." box="[1100,1246,1919,1943]" pageId="11" pageNumber="12" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
or 
<taxonomicName id="658FC8F84B4FFF83E3D0F8D2FAB3F808" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1279,1400,1919,1943]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B4FFF83E3D0F8D2FAB3F808" box="[1279,1400,1919,1943]" italics="true" pageId="11" pageNumber="12">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B4FFF83E251F8D2FF38F829" bold="true" pageId="11" pageNumber="12">
<taxonomicNameLabel id="8BC8D2124B4FFF83E251F8D2FF38F829" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
]; 
<bibRefCitation id="C61ECE8A4B4FFF83E625F833FE1FF829" author="Frith, J. W. &amp; Tantanasiriwong, R. &amp; Bhatia, O." box="[266,468,1950,1974]" pageId="11" pageNumber="12" pagination="1 - 14" refId="ref71449" refString="Frith, J. W., Tantanasiriwong, R. &amp; Bhatia, O. (1976) Zonation of macrofauna on a mangrove shore, Phuket Island, southern Thailand. Phuket Marine Biology Center Research Bulletin, 17, 1 - 14." type="journal article" year="1976">
Frith 
<emphasis id="90FB6F694B4FFF83E66EF832FEBEF829" box="[321,373,1950,1974]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
1976: 18
</bibRefCitation>
[= 
<taxonomicName id="658FC8F84B4FFF83E6D7F833FDB6F82A" box="[504,637,1950,1974]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B4FFF83E6D7F833FD81F829" box="[504,586,1950,1974]" italics="true" pageId="11" pageNumber="12">Alpheus</emphasis>
spp.
</taxonomicName>
, most probably 
<taxonomicName id="658FC8F84B4FFF83E430F833FB9DF829" ID-CoL="85LHH" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[799,1110,1950,1974]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B4FFF83E430F833FC76F829" box="[799,957,1950,1974]" italics="true" pageId="11" pageNumber="12">A. eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B4FFF83E4EBF833FB9DF829" author="De Man, J. G." box="[964,1110,1950,1974]" pageId="11" pageNumber="12" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
or 
<taxonomicName id="658FC8F84B4FFF83E357F833FB0CF829" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1144,1223,1950,1974]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B4FFF83E357F833FB0CF829" box="[1144,1223,1950,1974]" italics="true" pageId="11" pageNumber="12">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B4FFF83E3E1F833FAD5F829" bold="true" box="[1230,1310,1950,1974]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="8BC8D2124B4FFF83E3E1F833FAD5F829" box="[1230,1310,1950,1974]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
]; 
<bibRefCitation id="C61ECE8A4B4FFF83E21AF833FEE9F84B" author="Yeo, D. C. J. &amp; Ng, P. K. L." pageId="11" pageNumber="12" pagination="37 - 63" refId="ref75202" refString="Yeo, D. C. J. &amp; Ng, P. K. L. (1996) A new species of freshwater snapping shrimp, Alpheus cyanoteles (Decapoda: Caridea: Alpheidae) from Peninsular Malaysia and a redescription of Alpheus paludicola Kemp, 1915. Raffles Bulletin of Zoology, 44, 37 - 63." type="journal article" year="1996">Yeo &amp; Ng 1996: 58</bibRefCitation>
[= 
<taxonomicName id="658FC8F84B4FFF83E667F811FE05F84B" box="[328,462,1980,2004]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B4FFF83E667F811FE51F84B" box="[328,410,1980,2004]" italics="true" pageId="11" pageNumber="12">Alpheus</emphasis>
spp.
</taxonomicName>
, including 
<taxonomicName id="658FC8F84B4FFF83E56EF810FD2BF84B" ID-CoL="85LHH" authorityName="De Man" authorityYear="1920" box="[577,736,1980,2004]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B4FFF83E56EF810FD2BF84B" box="[577,736,1980,2004]" italics="true" pageId="11" pageNumber="12">A. eurydactylus</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B4FFF83E43BF810FC44F84B" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[788,911,1980,2004]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B4FFF83E43BF810FC44F84B" box="[788,911,1980,2004]" italics="true" pageId="11" pageNumber="12">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B4FFF83E4B8F810FC23F84A" bold="true" box="[919,1000,1981,2005]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="8BC8D2124B4FFF83E4B8F810FC23F84A" box="[919,1000,1981,2005]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, fig. 8d = 
<taxonomicName id="658FC8F84B4FFF83E377F810FB18F84B" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1112,1235,1980,2004]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B4FFF83E377F810FB18F84B" box="[1112,1235,1980,2004]" italics="true" pageId="11" pageNumber="12">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B4FFF83E3F5F810FAE7F84A" bold="true" box="[1242,1324,1981,2005]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="8BC8D2124B4FFF83E3F5F810FAE7F84A" box="[1242,1324,1981,2005]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
]; 
<bibRefCitation id="C61ECE8A4B4FFF83E26BF810FE5FF86C" author="Bruce, A. J. &amp; Coombes, K. E." pageId="11" pageNumber="12" pagination="301 - 337" refId="ref69806" refString="Bruce, A. J. &amp; Coombes, K. E. (1997) An annotated check-list of the caridean shrimps (Crustacea: Decapoda) of Darwin Harbour, with descriptions of three new species of Periclimenes [Palaemonidae: Pontoniinae]. In: Hanley, J. R., Caswell, G., Megirian, D. &amp; Larson, H. K. (Eds.), Proceedings of the sixth international marine biological workshop. The marine fauna and flora of Darwin Harbour, Northern Territory, Australia. Museums and Art Galleries of the Northern Territory and the Australian Marine Sciences Association, Darwin, pp. 301 - 337." type="book chapter" year="1997">Bruce &amp; Coombes 1997: 321</bibRefCitation>
[= 
<taxonomicName id="658FC8F84B4FFF83E697F876FDF7F86C" box="[440,572,2011,2035]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B4FFF83E697F876FDC1F86C" box="[440,522,2011,2035]" italics="true" pageId="11" pageNumber="12">Alpheus</emphasis>
spp.
</taxonomicName>
, most probably 
<taxonomicName id="658FC8F84B4FFF83E5F0F876FBDDF86C" ID-CoL="85LHH" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[735,1046,2011,2035]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B4FFF83E5F0F876FCB7F86C" box="[735,892,2011,2035]" italics="true" pageId="11" pageNumber="12">A. eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B4FFF83E4ABF871FBDDF86C" author="De Man, J. G." box="[900,1046,2011,2035]" pageId="11" pageNumber="12" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
or 
<taxonomicName id="658FC8F84B4FFF83E317F876FB79F86C" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1080,1202,2011,2035]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B4FFF83E317F876FB79F86C" box="[1080,1202,2011,2035]" italics="true" pageId="11" pageNumber="12">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B4FFF83E397F876FAC2F86C" bold="true" box="[1208,1289,2011,2035]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="8BC8D2124B4FFF83E397F876FAC2F86C" box="[1208,1289,2011,2035]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
]; 
<bibRefCitation id="C61ECE8A4B4FFF84E230F876FF26FF30" author="Edgar, G. J." lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" refId="ref71209" refString="Edgar, G. J. (1997) Australian marine life. The plants and animals of temperate waters. Reed, Victoria, 544 pp. [revised edition published in 2001]" type="book" year="1997">Edgar 1997: 194</bibRefCitation>
(colour photo only) [= 
<taxonomicName id="658FC8F84B48FF84E6CAFF3AFD4AFF31" box="[485,641,151,175]" class="Malacostraca" family="Alpheidae" genus="Synalpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B48FF84E6CAFF3AFD93FF30" box="[485,600,151,175]" italics="true" pageId="12" pageNumber="13">Synalpheus</emphasis>
sp.
</taxonomicName>
]; 
<bibRefCitation id="C61ECE8A4B48FF84E5BAFF3AFC8EFF30" author="Shokita, S." box="[661,837,151,175]" pageId="12" pageNumber="13" pagination="76 - 110" refId="ref74339" refString="Shokita, S. (2004) Chapter 5. The role of aquatic organisms in mangrove ecosystems. In: Vannucci, M. (Ed.), Mangrove management and conservation: present and future. United Nations University Press, Tokyo, pp. 76 - 110." type="book chapter" year="2004">Shokita 2004: 82</bibRefCitation>
[= 
<taxonomicName id="658FC8F84B48FF84E442FF3AFC38FF31" box="[877,1011,151,175]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B48FF84E442FF3AFC74FF30" box="[877,959,151,175]" italics="true" pageId="12" pageNumber="13">Alpheus</emphasis>
spp.
</taxonomicName>
, most probably 
<taxonomicName id="658FC8F84B48FF84E3B5FF3AFF31FF52" ID-CoL="85LHH" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B48FF84E3B5FF3AFAF2FF30" box="[1178,1337,151,175]" italics="true" pageId="12" pageNumber="13">A. eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B48FF84E26DFF3AFF31FF52" author="De Man, J. G." pageId="12" pageNumber="13" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
or 
<taxonomicName id="658FC8F84B48FF84E635FF1BFEA2FF52" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[282,361,181,205]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B48FF84E635FF1BFEA2FF52" box="[282,361,181,205]" italics="true" pageId="12" pageNumber="13">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B48FF84E641FF18FE75FF52" bold="true" box="[366,446,181,205]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="8BC8D2124B48FF84E641FF18FE75FF52" box="[366,446,181,205]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
]; 
<bibRefCitation id="C61ECE8A4B48FF84E6FCFF18FDB2FF52" author="Wada, K." box="[467,633,181,205]" pageId="12" pageNumber="13" pagination="183 - 188" refId="ref74798" refString="Wada, K. (2013) Benthic animals in mangrove swamp: a review. Global Environmental Research, 17 / 2013, 183 - 188." type="journal article" year="2013">Wada 2013: 184</bibRefCitation>
, fig. 5 [= 
<taxonomicName id="658FC8F84B48FF84E5F5FF18FC96FF52" box="[730,861,181,205]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B48FF84E5F5FF18FCE7FF52" box="[730,812,181,205]" italics="true" pageId="12" pageNumber="13">Alpheus</emphasis>
spp.
</taxonomicName>
, most probably 
<taxonomicName id="658FC8F84B48FF84E4D2FF1BFB80FF52" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1021,1099,181,205]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B48FF84E4D2FF1BFB80FF52" box="[1021,1099,181,205]" italics="true" pageId="12" pageNumber="13">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B48FF84E37EFF18FB6AFF52" bold="true" box="[1105,1185,181,205]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="8BC8D2124B48FF84E37EFF18FB6AFF52" box="[1105,1185,181,205]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
]; 
<bibRefCitation id="C61ECE8A4B48FF84E399FF1BFA56FF52" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[1206,1437,181,205]" pageId="12" pageNumber="13" pagination="1695 - 1698" refId="ref73799" refString="Paras, N. J., Purushothaman, P., Chinnadurai, S., Renjith R. K., Remesan, M. P., Baiju, M. V. &amp; Edwin, L. (2019) Occurrence of Alpheus euphrosyne de Man, 1897 (Crustacea: Decapoda: Alpheidae) from the south-eastern Arabian Sea, India. Indian Journal of Geo Marine Sciences, 48, 1695 - 1698." type="journal article" year="2019">
Paras 
<emphasis id="90FB6F694B48FF84E3DCFF1BFAD4FF52" box="[1267,1311,181,205]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. 2019: 1695
</bibRefCitation>
(part., text, not figs. 1–3) [= 
<taxonomicName id="658FC8F84B48FF84E6CBFF79FBE4FF73" ID-CoL="84W7P" authority="sensu Banner &amp; Banner (1982)" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[484,1071,212,236]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">
<emphasis id="90FB6F694B48FF84E6CBFF79FCE0FF73" box="[484,811,212,236]" italics="true" pageId="12" pageNumber="13">A. euphrosyne euphrosyne sensu</emphasis>
<bibRefCitation id="C61ECE8A4B48FF84E41EFF79FBE4FF73" author="Banner, D. M. &amp; Banner, A. H." box="[817,1071,212,236]" pageId="12" pageNumber="13" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982)</bibRefCitation>
</taxonomicName>
, see below]; 
<bibRefCitation id="C61ECE8A4B48FF84E39EFF79FAA4FF73" author="Sha, Z. &amp; Wang, Y. &amp; Cui, D." box="[1201,1391,212,236]" pageId="12" pageNumber="13" refId="ref74259" refString="Sha, Z., Wang, Y. &amp; Cui, D. (2019) The Alpheidae from China seas. Crustacea: Decapoda: Caridea. Springer, Singapore, 323 pp. https: // doi. org / 10.1007 / 978 - 981 - 15 - 0648 - 2 _ 2" type="book" year="2019">
Sha 
<emphasis id="90FB6F694B48FF84E3F2FF78FADBFF73" box="[1245,1296,212,236]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2019: 87
</bibRefCitation>
, fig. 2.48 [= 
<taxonomicName id="658FC8F84B48FF84E638FF5EFE50FE95" box="[279,411,243,267]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B48FF84E638FF5EFEA2FE94" box="[279,361,243,267]" italics="true" pageId="12" pageNumber="13">Alpheus</emphasis>
spp.
</taxonomicName>
, possibly 
<taxonomicName id="658FC8F84B48FF84E52FFF5EFCF3FE94" ID-CoL="85LHH" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" box="[512,824,243,267]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B48FF84E52FFF5EFD56FE94" box="[512,669,243,267]" italics="true" pageId="12" pageNumber="13">A. eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B48FF84E58BFF5EFCF3FE94" author="De Man, J. G." box="[676,824,243,267]" pageId="12" pageNumber="13" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B48FF84E46DFF5EFC7FFE95" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[834,948,243,266]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nomurai" status="sp. nov.">
<emphasis id="90FB6F694B48FF84E46DFF5EFC7FFE95" box="[834,948,243,266]" italics="true" pageId="12" pageNumber="13">A. nomurai</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B48FF84E495FF5EFBC0FE94" bold="true" box="[954,1035,243,267]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="8BC8D2124B48FF84E495FF5EFBC0FE94" box="[954,1035,243,267]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
or other taxa, see below].
</paragraph>
<paragraph id="A230B37B4B48FF84E7B8FEBFFC6FFEF9" blockId="12.[151,1437,151,451]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="829F94554B48FF84E7B8FEBFFC6FFEF9" pageId="12" pageNumber="13">
<collectingRegion id="604B7D994B48FF84E7B8FEBFFF76FEB6" box="[151,189,274,297]" country="Maldives" name="Uthuru" pageId="12" pageNumber="13">Not</collectingRegion>
<emphasis id="90FB6F694B48FF84E7EAFEBCFDE7FEB6" box="[197,556,273,297]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="658FC8F84B48FF84E7EAFEBCFDC6FEB6" ID-CoL="84W7P" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[197,525,273,297]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">Alpheus euphrosyne euphrosyne</taxonomicName>
.—
</emphasis>
(?) 
<treatmentCitation id="232E956A4B48FF84E560FEBFFCA1FEB6" author="Banner, D. M. &amp; Banner, A. H." box="[591,874,273,297]" page="23" pageId="12" pageNumber="13" year="1981">
<bibRefCitation id="C61ECE8A4B48FF84E560FEBFFCA1FEB6" author="Banner, D. M. &amp; Banner, A. H." box="[591,874,273,297]" pageId="12" pageNumber="13" pagination="1 - 99" refId="ref69349" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981: 23</bibRefCitation>
</treatmentCitation>
, 92; 
<treatmentCitation id="232E956A4B48FF84E4B0FEBFFB0DFEB6" author="Banner, D. M. &amp; Banner, A. H." box="[927,1222,273,297]" page="232" pageId="12" pageNumber="13" year="1982">
<bibRefCitation id="C61ECE8A4B48FF84E4B0FEBFFB0DFEB6" author="Banner, D. M. &amp; Banner, A. H." box="[927,1222,273,297]" pageId="12" pageNumber="13" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982: 232</bibRefCitation>
</treatmentCitation>
, fig. 73, tab. 5 (part.?); (?) 
<treatmentCitation id="232E956A4B48FF84E624FE9DFDE9FED7" author="Banner, A. H. &amp; Banner, D. M." box="[267,546,304,328]" page="30" pageId="12" pageNumber="13" year="1983">
<bibRefCitation id="C61ECE8A4B48FF84E624FE9DFDE9FED7" author="Banner, A. H. &amp; Banner, D. M." box="[267,546,304,328]" pageId="12" pageNumber="13" pagination="1 - 164" refId="ref69267" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983: 30</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="232E956A4B48FF84E502FE9DFC8BFED7" author="Banner, D. M. &amp; Banner, A. H." box="[557,832,304,328]" page="16" pageId="12" pageNumber="13" year="1985">
<bibRefCitation id="C61ECE8A4B48FF84E502FE9DFC8BFED7" author="Banner, D. M. &amp; Banner, A. H." box="[557,832,304,328]" pageId="12" pageNumber="13" pagination="1 - 79" refId="ref69455" refString="Banner, D. M. &amp; Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's &quot; The Decapoda of the Siboga Expedition, Part II. Family Alpheidae &quot; (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner &amp; Banner 1985: 16</bibRefCitation>
</treatmentCitation>
(part.); 
<treatmentCitation id="232E956A4B48FF84E4BCFE9DFBF8FED7" author="Chace, F. A. Jr." box="[915,1075,304,328]" page="27" pageId="12" pageNumber="13" year="1988">
<bibRefCitation id="C61ECE8A4B48FF84E4BCFE9DFBF8FED7" author="Chace, F. A. Jr." box="[915,1075,304,328]" pageId="12" pageNumber="13" pagination="1 - 99" refId="ref69928" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988: 27</bibRefCitation>
</treatmentCitation>
(part.) [= 
<taxonomicName id="658FC8F84B48FF84E3B3FE9DFAEAFED8" box="[1180,1313,304,328]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="90FB6F694B48FF84E3B3FE9DFB25FED7" box="[1180,1262,304,328]" italics="true" pageId="12" pageNumber="13">Alpheus</emphasis>
spp.
</taxonomicName>
, possibly 
<taxonomicName id="658FC8F84B48FF84E2A9FE9DFE28FEF8" ID-CoL="85LHH" authority="De Man, 1920" authorityName="De Man" authorityYear="1920" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B48FF84E2A9FE9DFE83FEF8" italics="true" pageId="12" pageNumber="13">A. eurydactylus</emphasis>
<bibRefCitation id="C61ECE8A4B48FF84E660FEE2FE28FEF8" author="De Man, J. G." box="[335,483,335,359]" pageId="12" pageNumber="13" pagination="103 - 109" refId="ref71067" refString="De Man, J. G. (1920) Diagnoses of some new species of Penaeidae and Alpheidae with remarks on two known species of the genus Penaeopsis A. M. - Edw. from the Indian Archipelago. Zoologische Mededelingen, Leiden, 5, 103 - 109." type="journal article" year="1920">De Man, 1920</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="658FC8F84B48FF84E6C2FEE2FDF7FEF8" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[493,572,335,359]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B48FF84E6C2FEE2FDF7FEF8" box="[493,572,335,359]" italics="true" pageId="12" pageNumber="13">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B48FF84E56DFEE2FD58FEF8" bold="true" box="[578,659,335,359]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="8BC8D2124B48FF84E56DFEE2FD58FEF8" box="[578,659,335,359]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="658FC8F84B48FF84E5EAFEE2FCF5FEF8" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[709,830,335,359]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B48FF84E5EAFEE2FCF5FEF8" box="[709,830,335,359]" italics="true" pageId="12" pageNumber="13">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B48FF84E46BFEE2FC5FFEF8" bold="true" box="[836,916,335,359]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="8BC8D2124B48FF84E46BFEE2FC5FFEF8" box="[836,916,335,359]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B48FF84E7B8FEC3FB60FE1A" blockId="12.[151,1437,151,451]" box="[151,1195,365,389]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="829F94554B48FF84E7B8FEC3FB60FE1A" box="[151,1195,365,389]" pageId="12" pageNumber="13">
<collectingRegion id="604B7D994B48FF84E7B8FEC3FF76FE1A" box="[151,189,366,389]" country="Maldives" name="Uthuru" pageId="12" pageNumber="13">Not</collectingRegion>
<emphasis id="90FB6F694B48FF84E7EBFEC0FE2CFE1A" box="[196,487,365,389]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="658FC8F84B48FF84E7EBFEC0FE02FE1A" authority="Banner &amp; Banner 1982" box="[196,457,365,389]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="langi">Alpheus euphrosyne langi</taxonomicName>
.—
</emphasis>
<treatmentCitation id="232E956A4B48FF84E6C8FEC3FCCDFE1A" author="Banner, D. M. &amp; Banner, A. H." box="[487,774,365,389]" page="238" pageId="12" pageNumber="13" year="1982">
<bibRefCitation id="C61ECE8A4B48FF84E6C8FEC3FCCDFE1A" author="Banner, D. M. &amp; Banner, A. H." box="[487,774,365,389]" pageId="12" pageNumber="13" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982: 238</bibRefCitation>
</treatmentCitation>
[= 
<taxonomicName id="658FC8F84B48FF84E406FEC3FB54FE1A" ID-CoL="85LN7" authority="de Rochebrune, 1883" authorityName="de Rochebrune" authorityYear="1883" box="[809,1183,365,389]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pontederiae">
<emphasis id="90FB6F694B48FF84E406FEC3FC75FE1A" box="[809,958,365,389]" italics="true" pageId="12" pageNumber="13">A. pontederiae</emphasis>
de 
<bibRefCitation id="C61ECE8A4B48FF84E4CCFEC0FB54FE1A" author="Rochebrune, A. T. de" box="[995,1183,365,389]" pageId="12" pageNumber="13" pagination="167 - 182" refId="ref74104" refString="Rochebrune, A. T. de (1883) Diagnoses d'arthropodes nouveaux propres la Senegambie. Bulletin de la Societe Philomathique de Paris, Serie 7, 7, 167 - 182." type="journal article" year="1883">Rochebrune, 1883</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B48FF84E7B8FE21FAF7FE3B" blockId="12.[151,1437,151,451]" box="[151,1340,396,420]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="829F94554B48FF84E7B8FE21FAF7FE3B" box="[151,1340,396,420]" pageId="12" pageNumber="13">
<treatmentCitation id="232E956A4B48FF84E7B8FE21FDCAFE3B" author="Kazmi, M. A." box="[151,513,396,420]" page="170" pageId="12" pageNumber="13" year="1974">
<taxonomicName id="658FC8F84B48FF84E7B8FE21FDCAFE3B" ID-CoL="85LQ9" authority="Kazmi 1974: 170" authorityName="Kazmi" authorityPageNumber="170" authorityYear="1974" box="[151,513,396,420]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tirmiziae">
<emphasis id="90FB6F694B48FF84E7B8FE21FE82FE3B" box="[151,329,396,420]" italics="true" pageId="12" pageNumber="13">Alpheus tirmiziae</emphasis>
<bibRefCitation id="C61ECE8A4B48FF84E67FFE21FDCAFE3B" author="Kazmi, M. A." box="[336,513,396,420]" pageId="12" pageNumber="13" pagination="170 - 174" refId="ref72166" refString="Kazmi, M. A. (1974) A new species of the genus Alpheus from the Bay of Bengal (Decapoda, Caridea). Crustaceana, 27, 170 - 174. https: // doi. org / 10.1163 / 156854074 X 00389" type="journal article" year="1974">Kazmi 1974: 170</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1–4; 
<treatmentCitation id="232E956A4B48FF84E55DFE21FC58FE3B" author="Banner, D. M. &amp; Banner, A. H." box="[626,915,396,420]" page="233" pageId="12" pageNumber="13" year="1978">
<bibRefCitation id="C61ECE8A4B48FF84E55DFE21FC58FE3B" author="Banner, D. M. &amp; Banner, A. H." box="[626,915,396,420]" pageId="12" pageNumber="13" pagination="215 - 257" refId="ref69308" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner 1978: 233</bibRefCitation>
</treatmentCitation>
, fig. 2i–m; 
<treatmentCitation id="232E956A4B48FF84E327FE21FB1EFE3B" author="Sha, Z. &amp; Wang, Y. &amp; Cui, D." box="[1032,1237,396,420]" page="122" pageId="12" pageNumber="13" year="2019">
<bibRefCitation id="C61ECE8A4B48FF84E327FE21FB1EFE3B" author="Sha, Z. &amp; Wang, Y. &amp; Cui, D." box="[1032,1237,396,420]" pageId="12" pageNumber="13" refId="ref74259" refString="Sha, Z., Wang, Y. &amp; Cui, D. (2019) The Alpheidae from China seas. Crustacea: Decapoda: Caridea. Springer, Singapore, 323 pp. https: // doi. org / 10.1007 / 978 - 981 - 15 - 0648 - 2 _ 2" type="book" year="2019">
Sha 
<emphasis id="90FB6F694B48FF84E31BFE20FBAAFE3B" box="[1076,1121,396,420]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. 2019: 122
</bibRefCitation>
</treatmentCitation>
, fig. 2.67.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A230B37B4B48FF84E7B8FE06FD2DFE5D" blockId="12.[151,1437,151,451]" box="[151,742,426,451]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="829F94554B48FF84E7B8FE06FD2DFE5D" box="[151,742,426,451]" pageId="12" pageNumber="13">
<taxonomicName id="658FC8F84B48FF84E7B8FE06FE89FE5D" authorityName="Kazmi" authorityYear="1974" box="[151,322,427,451]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tirmizae">
<emphasis id="90FB6F694B48FF84E7B8FE06FE89FE5D" box="[151,322,427,451]" italics="true" pageId="12" pageNumber="13">Alpheus tirmizae</emphasis>
</taxonomicName>
(
<emphasis id="90FB6F694B48FF84E67EFE06FE56FE5C" box="[337,413,427,451]" italics="true" pageId="12" pageNumber="13">lap. cal</emphasis>
.).— 
<treatmentCitation id="232E956A4B48FF84E6E9FE06FD29FE5D" author="Kazmi, M. A. &amp; Kazmi, Q. B." box="[454,738,426,451]" page="154" pageId="12" pageNumber="13" year="1979">
<bibRefCitation id="C61ECE8A4B48FF84E6E9FE06FD29FE5D" author="Kazmi, M. A. &amp; Kazmi, Q. B." box="[454,738,426,451]" pageId="12" pageNumber="13" pagination="151 - 157" refId="ref72213" refString="Kazmi, M. A. &amp; Kazmi, Q. B. (1979) A check-list of marine caridean prawns of Pakistan. Biologia, 25, 151 - 157." type="journal article" year="1979">Kazmi &amp; Kazmi 1979: 154</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B48FF85E7B8FE5CFDD9FE82" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="A230B37B4B48FF84E7B8FE5CFDF5FDB0" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<emphasis id="90FB6F694B48FF84E7B8FE5CFE8BFD94" bold="true" box="[151,320,497,523]" pageId="12" pageNumber="13">Type material</emphasis>
. 
<materialsCitation id="12E7B9264B48FF84E67EFE5CFDF5FDB0" ID-GBIF-Occurrence="4104078436" collectionCode="RMNH" collectorName="Crus. D. &amp; P. Buitendijk" country="Indonesia" location="Java" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Jawa Timur" typeStatus="neotype">
<typeStatus id="7D340DD94B48FF84E67EFE5CFE7FFD94" box="[337,436,497,523]" pageId="12" pageNumber="13" type="neotype">Neotype</typeStatus>
: male (cl 
<quantity id="65771E9E4B48FF84E507FE5CFD5AFD94" box="[552,657,497,523]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="12" pageNumber="13" unit="mm" value="16.6">16.6 mm</quantity>
, tl 
<quantity id="65771E9E4B48FF84E597FE5CFCEAFD94" box="[696,801,497,524]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" pageId="12" pageNumber="13" unit="mm" value="48.5">48.5 mm</quantity>
, chl 26.0 mm), 
<collectionCode id="C49E2BBE4B48FF84E4F4FE5CFBE4FD94" box="[987,1071,497,523]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13" type="Museum">RMNH</collectionCode>
<collectorName id="0F7AD6AD4B48FF84E317FE5CFB46FD94" box="[1080,1165,497,523]" pageId="12" pageNumber="13">Crus.D.</collectorName>
49669, 
<collectingCountry id="DA98F3EB4B48FF84E3C9FE5CFA9DFD94" box="[1254,1366,497,523]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
, 
<collectingRegion id="604B7D994B48FF84E24CFE5CFA5CFD94" box="[1379,1431,497,523]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="13">Java</collectingRegion>
, Surabaya, leg. 
<collectorName id="0F7AD6AD4B48FF84E611FDB8FE1AFDB0" box="[318,465,533,559]" pageId="12" pageNumber="13">P. Buitendijk</collectorName>
, 02.1929.
</materialsCitation>
</paragraph>
<paragraph id="A230B37B4B48FF84E7E8FD94FB06FD7C" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<emphasis id="90FB6F694B48FF84E7E8FD94FDE7FDCC" bold="true" box="[199,556,569,595]" pageId="12" pageNumber="13">Additional material examined</emphasis>
. 
<materialsCitation id="12E7B9264B48FF84E515FD94FBC8FDE8" ID-GBIF-Occurrence="4104078340" collectionCode="ZMA" collectorName="Jacquin" country="Indonesia" location="Postillon Islands" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Selatan">
<emphasis id="90FB6F694B48FF84E515FD94FD63FDCC" box="[570,680,569,595]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="DA98F3EB4B48FF84E515FD94FD63FDCC" box="[570,680,569,595]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B48FF84E598FD94FCD3FDCC" box="[695,792,569,595]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl indet., missing both chelipeds), 
<collectionCode id="C49E2BBE4B48FF84E388FD94FB2FFDCC" box="[1191,1252,569,595]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="12" pageNumber="13">ZMA</collectionCode>
<collectorName id="0F7AD6AD4B48FF84E3C5FD94FAF5FDCC" box="[1258,1342,569,595]" pageId="12" pageNumber="13">Crus.D.</collectorName>
202727, 
<collectorName id="0F7AD6AD4B48FF84E7B8FDF0FE97FDE8" box="[151,348,605,631]" pageId="12" pageNumber="13">Siboga-Expeditie</collectorName>
, 
<location id="A750E5A04B48FF84E645FDF0FDEDFDE8" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E645FDF0FDEDFDE8" box="[362,550,605,631]" country="Indonesia" name="Postillon Islands" pageId="12" pageNumber="13" stateProvince="Sulawesi Selatan">Postillon Islands</location>
(
<collectingRegion id="604B7D994B48FF84E518FDF0FD35FDE8" box="[567,766,605,631]" country="Indonesia" name="Sulawesi Selatan" pageId="12" pageNumber="13">Sulawesi Selatan</collectingRegion>
), leg. 
<collectorName id="0F7AD6AD4B48FF84E469FDF0FC55FDE8" box="[838,926,605,631]" pageId="12" pageNumber="13">Jacquin</collectorName>
, no date
</materialsCitation>
; 
<materialsCitation id="12E7B9264B48FF84E33EFDF0FEE6FD20" ID-GBIF-Occurrence="4104078514" collectionCode="ZMA" collectorName="J. D. F. Hardenberg" country="Indonesia" determinerName="H. Balss" location="Sumatra" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara">
<specimenCount id="B48978F24B48FF84E33EFDF0FB95FDE8" box="[1041,1118,605,631]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl indet.), 1 ov. female (cl indet., missing both chelipeds), 
<collectionCode id="C49E2BBE4B48FF84E6D3FD2CFDF2FD04" box="[508,569,641,667]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="12" pageNumber="13">ZMA</collectionCode>
<collectorName id="0F7AD6AD4B48FF84E56FFD2CFD5FFD04" box="[576,660,641,667]" pageId="12" pageNumber="13">Crus.D.</collectorName>
202726, 
<collectingRegion id="604B7D994B48FF84E5D6FD2CFC92FD04" box="[761,857,641,667]" country="Indonesia" name="Sumatera Utara" pageId="12" pageNumber="13">Sumatra</collectingRegion>
, 
<collectorName id="0F7AD6AD4B48FF84E44AFD2CFBEFFD04" box="[869,1060,641,667]" pageId="12" pageNumber="13">Bagansi Api-Api</collectorName>
, leg. 
<collectorName id="0F7AD6AD4B48FF84E34FFD2CFAFBFD04" box="[1120,1328,641,667]" pageId="12" pageNumber="13">J.D.F. Hardenberg</collectorName>
, 01.1929, det. 
<determinerName id="57EC5FBF4B48FF84E7E6FD08FEE6FD20" box="[201,301,677,703]" pageId="12" pageNumber="13">H. Balss</determinerName>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B48FF84E615FD08FB02FD7C" ID-GBIF-Occurrence="4104078534" collectionCode="ZMA" collectorName="Kleiweg &amp; de Zwaan" country="Indonesia" location="Kleiweg de Zwaan" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B48FF84E615FD08FE57FD20" box="[314,412,677,703]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E6E4FD08FDF8FD20" box="[459,563,677,704]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="12" pageNumber="13" unit="mm" value="15.5">15.5 mm</quantity>
, missing both chelipeds), 1 ov. female (cl 
<quantity id="65771E9E4B48FF84E337FD08FB4BFD20" box="[1048,1152,677,703]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="12" pageNumber="13" unit="mm" value="16.2">16.2 mm</quantity>
, missing major cheliped, identification somewhat tentative), 
<collectionCode id="C49E2BBE4B48FF84E50BFD64FDAAFD7C" box="[548,609,713,739]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="12" pageNumber="13">ZMA</collectionCode>
<collectorName id="0F7AD6AD4B48FF84E548FD64FD77FD7C" box="[615,700,713,739]" pageId="12" pageNumber="13">Crus.D.</collectorName>
202793, 
<collectorName id="0F7AD6AD4B48FF84E40EFD64FC9CFD7C" box="[801,855,713,739]" pageId="12" pageNumber="13">Nias</collectorName>
, leg. 
<collectorName id="0F7AD6AD4B48FF84E4BDFD64FBA3FD7C" box="[914,1128,713,739]" pageId="12" pageNumber="13">
<location id="A750E5A04B48FF84E4BDFD64FBA3FD7C" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E4BDFD64FBA3FD7C" box="[914,1128,713,739]" country="Indonesia" name="Kleiweg de Zwaan" pageId="12" pageNumber="13">Kleiweg de Zwaan</location>
</collectorName>
, no date
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B48FF84E7E8FD40FBA1FC08" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<materialsCitation id="12E7B9264B48FF84E7E8FD40FE43FCB4" ID-GBIF-Occurrence="4104078447" collectingDate="1998-12-11" collectionCode="ZRC" collectorName="Y. Cai &amp; N. K. Ng" country="China" county="Nanao County" location="Houzhai" pageId="12" pageNumber="13" specimenCode="ZRC 2014.0677" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong">
<emphasis id="90FB6F694B48FF84E7E8FD40FEC0FC98" box="[199,267,749,775]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="DA98F3EB4B48FF84E7E8FD40FEC0FC98" box="[199,267,749,775]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B48FF84E634FD40FEADFC98" box="[283,358,749,775]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E6BDFD40FE31FC98" box="[402,506,749,775]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="12" pageNumber="13" unit="mm" value="14.3">14.3 mm</quantity>
), 
<specimenCode id="F2291B004B48FF84E522FD40FD0EFC98" box="[525,709,749,775]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E522FD40FD88FC98" box="[525,579,749,775]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2014.0677
</specimenCode>
, 
<collectingRegion id="604B7D994B48FF84E5FFFD40FC9DFC98" box="[720,854,749,775]" country="China" name="Guangdong" pageId="12" pageNumber="13">Guangdong</collectingRegion>
, 
<collectingCounty id="4B51CBF74B48FF84E44EFD40FBCEFC98" box="[865,1029,749,775]" pageId="12" pageNumber="13">Nanao County</collectingCounty>
, 
<location id="A750E5A04B48FF84E33FFD40FBB9FC98" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E33FFD40FBB9FC98" box="[1040,1138,749,775]" country="China" county="Nanao County" name="Houzhai" pageId="12" pageNumber="13" stateProvince="Guangdong">Houzhai</location>
, fish market, leg. 
<collectorName id="0F7AD6AD4B48FF84E215FD40FAB4FC98" box="[1338,1407,749,775]" pageId="12" pageNumber="13">Y. Cai</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B48FF84E7B8FCBCFF30FCB4" box="[151,251,785,811]" pageId="12" pageNumber="13">N.K. Ng</collectorName>
, 
<date id="D63195BB4B48FF84E628FCBCFE43FCB4" box="[263,392,785,811]" pageId="12" pageNumber="13" value="1998-12-11">
<collectingDate id="C6756C534B48FF84E628FCBCFE43FCB4" box="[263,392,785,811]" pageId="12" pageNumber="13" value="1998-12-11">12.11.1998</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B48FF84E6BBFCBCFB2FFCB4" ID-GBIF-Occurrence="4104078400" box="[404,1252,785,811]" collectionCode="USNM" collectorName="Light" country="China" pageId="12" pageNumber="13" specimenCode="USNM 62106" specimenCount="1" specimenCount-male="1">
<specimenCount id="B48978F24B48FF84E6BBFCBCFE2AFCB4" box="[404,481,785,811]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 23.0 mm), 
<specimenCode id="F2291B004B48FF84E5A3FCBCFCE4FCB4" box="[652,815,785,811]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13" type="Museum">
<collectionCode id="C49E2BBE4B48FF84E5A3FCBCFD16FCB4" box="[652,733,785,811]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="12" pageNumber="13">USNM</collectionCode>
62106
</specimenCode>
, locality not specified, don. S.F. 
<collectorName id="0F7AD6AD4B48FF84E38BFCBCFB2FFCB4" box="[1188,1252,785,811]" pageId="12" pageNumber="13">Light</collectorName>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B48FF84E3DEFCBCFF1CFCEC" ID-GBIF-Occurrence="4104078421" collectionCode="USNM" collectorName="Shantou &amp; S. F. Light" country="China" location="Fuzhou" pageId="12" pageNumber="13" specimenCode="USNM 62107" specimenCount="1" specimenCount-male="1" stateProvince="Fujian">
<specimenCount id="B48978F24B48FF84E3DEFCBCFAF6FCB4" box="[1265,1341,785,811]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E244FCBCFF0CFCD0" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="12" pageNumber="13" unit="mm" value="20.5">20.5 mm</quantity>
), 
<specimenCode id="F2291B004B48FF84E7F2FC98FE4BFCD0" box="[221,384,821,847]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13" type="Museum">
<collectionCode id="C49E2BBE4B48FF84E7F2FC98FEE5FCD0" box="[221,302,821,847]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="12" pageNumber="13">USNM</collectionCode>
62107
</specimenCode>
, 
<collectingRegion id="604B7D994B48FF84E6A1FC98FE12FCD0" box="[398,473,821,847]" country="China" name="Fujian" pageId="12" pageNumber="13">Fujian</collectingRegion>
, “Santu north of Foochow” (possibly 
<collectorName id="0F7AD6AD4B48FF84E4BFFC98FC24FCD0" box="[912,1007,821,847]" pageId="12" pageNumber="13">Shantou</collectorName>
, south of 
<location id="A750E5A04B48FF84E34BFC98FB77FCD0" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E34BFC98FB77FCD0" box="[1124,1212,821,847]" country="China" name="Fuzhou" pageId="12" pageNumber="13" stateProvince="Fujian">Fuzhou</location>
), no date, don. 
<collectorName id="0F7AD6AD4B48FF84E25EFC98FF1CFCEC" pageId="12" pageNumber="13">S.F. Light</collectorName>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B48FF84E7CDFCF4FED2FC08" ID-GBIF-Occurrence="4104078518" collectingDate="1923-04-16" collectionCode="USNM" collectorName="S. F. Light" country="China" location="Amoy" municipality="Xiamen" pageId="12" pageNumber="13" specimenCode="USNM 62109" specimenCount="1" specimenCount-female="1" stateProvince="Fujian">
<specimenCount id="B48978F24B48FF84E7CDFCF4FE88FCEC" box="[226,323,857,883]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E641FCF4FE1EFCEC" box="[366,469,857,883]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="12" pageNumber="13" unit="mm" value="13.7">13.7 mm</quantity>
), 
<specimenCode id="F2291B004B48FF84E6C7FCF4FD43FCEC" box="[488,648,857,883]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13" type="Museum">
<collectionCode id="C49E2BBE4B48FF84E6C7FCF4FDF2FCEC" box="[488,569,857,883]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="12" pageNumber="13">USNM</collectionCode>
62109
</specimenCode>
, 
<collectingRegion id="604B7D994B48FF84E5BCFCF4FD15FCEC" box="[659,734,857,883]" country="China" name="Fujian" pageId="12" pageNumber="13">Fujian</collectingRegion>
, 
<collectingMunicipality id="425429014B48FF84E5C7FCF4FC8FFCEC" box="[744,836,857,883]" pageId="12" pageNumber="13">Xiamen</collectingMunicipality>
, 
<location id="A750E5A04B48FF84E463FCF4FC58FCEC" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E463FCF4FC58FCEC" box="[844,915,857,883]" country="China" municipality="Xiamen" name="Amoy" pageId="12" pageNumber="13" stateProvince="Fujian">Amoy</location>
, outer harbour, 
<collectingMethod id="7BCECB6C4B48FF84E36DFCF4FB53FCEC" box="[1090,1176,857,883]" pageId="12" pageNumber="13">net trap</collectingMethod>
, leg. &amp; don. 
<collectorName id="0F7AD6AD4B48FF84E208FCF4FA52FCEC" box="[1319,1433,857,883]" pageId="12" pageNumber="13">S.F. Light</collectorName>
, 
<date id="D63195BB4B48FF84E7B8FCD0FED2FC08" box="[151,281,893,919]" pageId="12" pageNumber="13" value="1923-04-16">
<collectingDate id="C6756C534B48FF84E7B8FCD0FED2FC08" box="[151,281,893,919]" pageId="12" pageNumber="13" value="1923-04-16">16.04.1923</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B48FF84E60BFCD0FD5EFC08" ID-GBIF-Occurrence="4104078388" box="[292,661,893,920]" collectionCode="MCZ" country="China" pageId="12" pageNumber="13" specimenCode="MCZ 1743" specimenCount="1" specimenCount-male="1">
<specimenCount id="B48978F24B48FF84E60BFCD0FEA4FC08" box="[292,367,893,919]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E6B4FCD0FDC8FC08" box="[411,515,893,920]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="12" pageNumber="13" unit="mm" value="17.5">17.5 mm</quantity>
), 
<specimenCode id="F2291B004B48FF84E539FCD0FD5EFC08" box="[534,661,893,919]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="13" type="Museum">
<collectionCode id="C49E2BBE4B48FF84E539FCD0FD98FC08" box="[534,595,893,919]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" pageId="12" pageNumber="13">MCZ</collectionCode>
1743
</specimenCode>
</materialsCitation>
, 
<collectingCountry id="DA98F3EB4B48FF84E58FFCD0FCEDFC08" box="[672,806,893,919]" name="Hong Kong" pageId="12" pageNumber="13">Hong Kong</collectingCountry>
, leg. 
<collectorName id="0F7AD6AD4B48FF84E44EFCD0FC37FC08" box="[865,1020,893,919]" pageId="12" pageNumber="13">Capt. Putnam</collectorName>
, 03.1861.
</paragraph>
<paragraph id="A230B37B4B48FF84E7E8FC0FFA9DFC40" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<materialsCitation id="12E7B9264B48FF84E7E8FC0FFF03FC40" ID-GBIF-Occurrence="4104078481" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; K. Lim" country="Taiwan" location="Tungkang" municipality="County" pageId="12" pageNumber="13" specimenCode="ZRC 1999.0723" specimenCount="1" specimenCount-female="1" stateProvince="Kaohsiung">
<emphasis id="90FB6F694B48FF84E7E8FC0FFED3FC24" box="[199,280,930,955]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="DA98F3EB4B48FF84E7E8FC0FFED3FC24" box="[199,280,930,955]" name="Taiwan" pageId="12" pageNumber="13">Taiwan</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B48FF84E607FC0CFE42FC24" box="[296,393,929,955]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 10.0 mm), 
<specimenCode id="F2291B004B48FF84E51FFC0CFD2CFC24" box="[560,743,929,955]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E51FFC0CFDADFC24" box="[560,614,929,955]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
1999.0723
</specimenCode>
, 
<collectingRegion id="604B7D994B48FF84E5DCFC0CFCA7FC24" box="[755,876,929,955]" country="Taiwan" name="Kaohsiung" pageId="12" pageNumber="13">Kaohsiung</collectingRegion>
<collectingMunicipality id="425429014B48FF84E45CFC0CFC0CFC24" box="[883,967,929,955]" pageId="12" pageNumber="13">County</collectingMunicipality>
, 
<location id="A750E5A04B48FF84E4FEFC0CFB8FFC24" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E4FEFC0CFB8FFC24" box="[977,1092,929,955]" country="Taiwan" municipality="County" name="Tungkang" pageId="12" pageNumber="13" stateProvince="Kaohsiung">Tungkang</location>
, fish port, leg. 
<collectorName id="0F7AD6AD4B48FF84E3C2FC0CFA96FC24" box="[1261,1373,929,955]" pageId="12" pageNumber="13">P.K.L. Ng</collectorName>
&amp; 
<collectorName id="0F7AD6AD4B48FF84E2AEFC0CFF03FC40" pageId="12" pageNumber="13">K. Lim</collectorName>
</materialsCitation>
, 05.1999; 
<materialsCitation id="12E7B9264B48FF84E611FC68FBEFFC40" ID-GBIF-Occurrence="4104078529" box="[318,1060,965,992]" collectionCode="USNM" collectorName="M. Maki" country="Taiwan" location="Yensui River" pageId="12" pageNumber="13" specimenCode="USNM 59044" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B48FF84E611FC68FE54FC40" box="[318,415,965,991]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E6E5FC68FDF9FC40" box="[458,562,965,991]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="12" pageNumber="13" unit="mm" value="13.8">13.8 mm</quantity>
), 
<specimenCode id="F2291B004B48FF84E56AFC68FD2DFC7F" box="[581,742,965,992]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13" type="Museum">
<collectionCode id="C49E2BBE4B48FF84E56AFC68FD5DFC40" box="[581,662,965,991]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="12" pageNumber="13">USNM</collectionCode>
59044
</specimenCode>
, 
<location id="A750E5A04B48FF84E5DFFC68FC4FFC40" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E5DFFC68FC4FFC40" box="[752,900,965,991]" country="Taiwan" name="Yensui River" pageId="12" pageNumber="13">Yensui River</location>
, leg. 
<collectorName id="0F7AD6AD4B48FF84E490FC68FBEFFC40" box="[959,1060,965,991]" pageId="12" pageNumber="13">M. Maki</collectorName>
</materialsCitation>
, don. M. Oshima, 01.1923.
</paragraph>
<paragraph id="A230B37B4B48FF84E7E8FC47FED6FBB7" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<materialsCitation id="12E7B9264B48FF84E7E8FC47FED2FBB7" ID-GBIF-Occurrence="4104078557" collectingDate="1995-08-21" collectorName="H. Duy" country="Vietnam" elevation="25" location="Vietnam" pageId="12" pageNumber="13" specimenCount="4" specimenCount-male="4">
<emphasis id="90FB6F694B48FF84E7E8FC47FEE9FB9C" box="[199,290,1002,1027]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="DA98F3EB4B48FF84E7E8FC47FEE9FB9C" box="[199,290,1002,1027]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B48FF84E61AFC44FE44FB9C" box="[309,399,1001,1027]" pageId="12" pageNumber="13" type="male">4 males</specimenCount>
(cl 12.5–18.0 mm), MNHN-IU-2018-5730, Ha Tien, Sau, depth: 
<quantity id="65771E9E4B48FF84E3BCFC44FB37FB9C" box="[1171,1276,1001,1027]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="12" pageNumber="13" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">
<elevation id="29A254484B48FF84E3BCFC44FB37FB9C" box="[1171,1276,1001,1027]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="12" pageNumber="13" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">20–30 m</elevation>
</quantity>
, leg. 
<collectorName id="0F7AD6AD4B48FF84E26FFC44FA53FB9C" box="[1344,1432,1001,1027]" pageId="12" pageNumber="13">H. Duy</collectorName>
, 
<date id="D63195BB4B48FF84E7B8FBA0FED2FBB7" box="[151,281,1037,1064]" pageId="12" pageNumber="13" value="1995-08-21">
<collectingDate id="C6756C534B48FF84E7B8FBA0FED2FBB7" box="[151,281,1037,1064]" pageId="12" pageNumber="13" value="1995-08-21">21.08.1995</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B48FF84E7E8FB9CFE62FBF0" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<materialsCitation id="12E7B9264B48FF84E7E8FB9CFE6EFBF0" ID-GBIF-Occurrence="4104078332" collectingDate="2006-03-04" collectingDateMax="2006-03-29" collectingDateMin="2006-03-04" collectionCode="T, R" country="Cambodia" location="Prek Romeas" pageId="12" pageNumber="13" specimenCode="ZRC 2013.0458" specimenCount="1" specimenCount-male="1" stateProvince="Kampot">
<emphasis id="90FB6F694B48FF84E7E8FB9CFEF7FBD4" box="[199,316,1073,1099]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="DA98F3EB4B48FF84E7E8FB9CFEF7FBD4" box="[199,316,1073,1099]" name="Cambodia" pageId="12" pageNumber="13">Cambodia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B48FF84E663FB9CFE5CFBD4" box="[332,407,1073,1099]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E6EAFB9CFDE5FBD4" box="[453,558,1073,1099]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="12" pageNumber="13" unit="mm" value="18.9">18.9 mm</quantity>
), 
<specimenCode id="F2291B004B48FF84E56DFB9CFD31FBD3" box="[578,762,1073,1100]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E56DFB9CFDB3FBD4" box="[578,632,1073,1099]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2013.0458
</specimenCode>
, 
<collectingRegion id="604B7D994B48FF84E429FB9CFCABFBD4" box="[774,864,1073,1099]" country="Cambodia" name="Kampot" pageId="12" pageNumber="13">Kampot</collectingRegion>
and 
<location id="A750E5A04B48FF84E4B6FB9CFBE5FBD4" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E4B6FB9CFBE5FBD4" box="[921,1070,1073,1099]" country="Cambodia" name="Prek Romeas" pageId="12" pageNumber="13" stateProvince="Kampot">Prek Romeas</location>
fish landings, leg. 
<collectionCode id="C49E2BBE4B48FF84E228FB9CFAD1FBD4" box="[1287,1306,1073,1099]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="13">T</collectionCode>
. 
<collectionCode id="C49E2BBE4B48FF84E232FB9CFAF9FBD4" box="[1309,1330,1073,1099]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
. Roberts, 
<date id="D63195BB4B48FF84E7B8FBF8FEDCFBF0" box="[151,279,1109,1135]" pageId="12" pageNumber="13" value="2006-03-29">
<collectingDate id="C6756C534B48FF84E7B8FBF8FEDCFBF0" box="[151,279,1109,1135]" pageId="12" pageNumber="13" value="2006-03-29">29.03.2006</collectingDate>
</date>
– 
<date id="D63195BB4B48FF84E60AFBF8FE6EFBF0" box="[293,421,1109,1135]" pageId="12" pageNumber="13" value="2006-03-04">
<collectingDate id="C6756C534B48FF84E60AFBF8FE6EFBF0" box="[293,421,1109,1135]" pageId="12" pageNumber="13" value="2006-03-04">03.04.2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B48FF84E7E8FBD4FBD1F884" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<materialsCitation id="12E7B9264B48FF84E7E8FBD4FDAAFB28" ID-GBIF-Occurrence="4104078334" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-male="3" stateProvince="Songkhla">
Thailand: 
<specimenCount id="B48978F24B48FF84E614FBD4FE03FB0C" box="[315,456,1145,1172]" pageId="12" pageNumber="13" type="generic">5 specimens</specimenCount>
, including 
<specimenCount id="B48978F24B48FF84E56BFBD4FD51FB0C" box="[580,666,1145,1171]" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
(cl 16.0–16.9 mm), 
<specimenCode id="F2291B004B48FF84E4AEFBD4FBF3FB0B" box="[897,1080,1145,1172]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E4AEFBD4FC7CFB0C" box="[897,951,1145,1171]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2000.2150
</specimenCode>
, Gulf of Thailand, off Khanom, trawl, fish port, leg. P.K.L. Ng, 12.1984;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E546FB30FEB7FB44" ID-GBIF-Occurrence="4104078452" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B48978F24B48FF84E546FB30FD7EFB28" box="[617,693,1181,1207]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 19.3 mm), MNHN-IU-2018-5752, same collection data as for previous specimens;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E6ABFB6CFDC0FA9F" ID-GBIF-Occurrence="4104078331" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B48978F24B48FF84E6ABFB6CFE1BFB44" box="[388,464,1217,1243]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 14.0 mm), 
<specimenCode id="F2291B004B48FF84E555FB6CFCF9FB44" box="[634,818,1217,1243]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E555FB6CFD7BFB44" box="[634,688,1217,1243]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2003.0619
</specimenCode>
, Songkhla Lake (Thale Sap), Sakom crab landing, leg. P.K.L. Ng et al., 15– 
<date id="D63195BB4B48FF84E6ABFB48FDCCFA9F" box="[388,519,1253,1280]" pageId="12" pageNumber="13" value="2003-10-17">
<collectingDate id="C6756C534B48FF84E6ABFB48FDCCFA9F" box="[388,519,1253,1280]" pageId="12" pageNumber="13" value="2003-10-17">17.10.2003</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E53BFB48FEB6FABC" ID-GBIF-Occurrence="4104078394" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B48978F24B48FF84E53BFB48FDAAFB60" box="[532,609,1253,1279]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl indet.), 
<specimenCode id="F2291B004B48FF84E5C6FB48FC69FB60" box="[745,930,1253,1279]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E5C6FB48FCD4FB60" box="[745,799,1253,1279]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2000.2137
</specimenCode>
, Gulf of Thailand, Siracha, fish market, leg. P.K.L. Ng, 02.2000;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E6A9FAA4FC23FAD8" ID-GBIF-Occurrence="4104078313" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B48978F24B48FF84E6A9FAA4FE14FABC" box="[390,479,1289,1315]" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
(cl 20.2–21.7 mm), 1 ov. female (cl 21.9 mm), 
<specimenCode id="F2291B004B48FF84E322FAA4FB0CFABB" box="[1037,1223,1289,1316]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E322FAA4FB88FABC" box="[1037,1091,1289,1315]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2000.0957
</specimenCode>
, Gulf of Thailand, Chonburi, Angsila and Siracha, fish ports, leg. P.K.L. Ng et al., 
<date id="D63195BB4B48FF84E44DFA80FC2FFAD8" box="[866,996,1325,1351]" pageId="12" pageNumber="13" value="2000-02-20">
<collectingDate id="C6756C534B48FF84E44DFA80FC2FFAD8" box="[866,996,1325,1351]" pageId="12" pageNumber="13" value="2000-02-20">20.02.2000</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E4C0FA80FEAEFA10" ID-GBIF-Occurrence="4104078371" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-female="2" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B48978F24B48FF84E4C0FA80FB8EFAD8" box="[1007,1093,1325,1351]" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
(cl 15.0–16.9 mm), 
<collectionCode id="C49E2BBE4B48FF84E204FA80FA53FAD8" box="[1323,1432,1325,1351]" collectionName="United Kingdom, Oxford, University Museum of Natural History" pageId="12" pageNumber="13">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B48FF84E7B8FAFCFF75FAF4" box="[151,190,1361,1387]" country="Austria" name="Zoological Collection, University of Vienna" pageId="12" pageNumber="13" type="University or college">ZC</collectionCode>
. 2019.06.42 [ex-ZRC 2000.0956], Gulf of Thailand, Chonburi, Angsila and Siracha, fish ports, leg. P.K.L. Ng et al., 
<date id="D63195BB4B48FF84E7F0FAD8FEAAFA10" box="[223,353,1397,1423]" pageId="12" pageNumber="13" value="2000-02-20">
<collectingDate id="C6756C534B48FF84E7F0FAD8FEAAFA10" box="[223,353,1397,1423]" pageId="12" pageNumber="13" value="2000-02-20">20.02.2000</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E642FAD8FDFDFA2B" ID-GBIF-Occurrence="4104078558" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-female="2" specimenCount-male="3" stateProvince="Songkhla">
&gt; 
<specimenCount id="B48978F24B48FF84E653FAD8FDDEFA10" box="[380,533,1397,1423]" pageId="12" pageNumber="13" type="generic">20 specimens</specimenCount>
of both sexes (cl indet.), 
<specimenCode id="F2291B004B48FF84E418FAD8FC24FA10" box="[823,1007,1397,1423]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E418FAD8FCA6FA10" box="[823,877,1397,1423]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
1998.1066
</specimenCode>
, Gulf of Thailand, Chonburi, Angsila, fish port, leg. P.K.L. Ng, 
<date id="D63195BB4B48FF84E69FFA34FDF9FA2B" box="[432,562,1433,1460]" pageId="12" pageNumber="13" value="1998-09-25">
<collectingDate id="C6756C534B48FF84E69FFA34FDF9FA2B" box="[432,562,1433,1460]" pageId="12" pageNumber="13" value="1998-09-25">25.09.1998</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E513FA34FC8FFA48" ID-GBIF-Occurrence="4104078440" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-female="2" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B48978F24B48FF84E513FA34FD4CFA2C" box="[572,647,1433,1459]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 16.4 mm), 
<specimenCount id="B48978F24B48FF84E402FA34FC46FA2C" box="[813,909,1433,1459]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 18.0 mm), MNHN-IU-2018-5593 (ex-ZRC 1998.1066), same collection data as for previous specimens;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E463FA10FDF2F980" ID-GBIF-Occurrence="4104078496" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-female="2" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B48978F24B48FF84E463FA10FC6CFA48" box="[844,935,1469,1495]" pageId="12" pageNumber="13" type="male">2 males</specimenCount>
, 
<specimenCount id="B48978F24B48FF84E49CFA10FBDDFA48" box="[947,1046,1469,1495]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B48FF84E3B3FA10FB3BFA48" box="[1180,1264,1469,1495]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13" type="Museum">RMNH</collectionCode>
Crus.D.42009, Gulf of Thailand, Chonburi, ca. 
<locationDeviation id="B382AA594B48FF84E523FA4CFCD4FA64" box="[524,799,1505,1531]" location="Bangkok" pageId="12" pageNumber="13">100 km SE of Bangkok</locationDeviation>
, between Si Racha and Naklua, fishermen, 
<date id="D63195BB4B48FF84E238FA4CFA53FA63" box="[1303,1432,1505,1532]" pageId="12" pageNumber="13" value="1985-03-14">
<collectingDate id="C6756C534B48FF84E238FA4CFA53FA63" box="[1303,1432,1505,1532]" pageId="12" pageNumber="13" value="1985-03-14">14.03.1985</collectingDate>
</date>
, leg. A.C.J. Burgers &amp; L.B. Holthuis;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E56DF9A8FBF1F9DC" ID-GBIF-Occurrence="4104078560" collectingDate="2003-10-17" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" county="Khanom" location="Gulf" municipality="Lake" pageId="12" pageNumber="13" specimenCode="ZRC 2000.2150" specimenCount="8" specimenCount-female="2" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B48978F24B48FF84E56DF9A8FD59F980" box="[578,658,1541,1567]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
, 
<specimenCount id="B48978F24B48FF84E5B0F9A8FCC9F980" box="[671,770,1541,1567]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B48FF84E4A6F9A8FC16F980" box="[905,989,1541,1567]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13" type="Museum">RMNH</collectionCode>
Crus.D.42426, Gulf of Thailand, Surat Thani, off river mouth of Don Sak, depth: 4 m, 
<date id="D63195BB4B48FF84E598F984FCF3F9DC" box="[695,824,1577,1603]" pageId="12" pageNumber="13" value="1992-10-20">
<collectingDate id="C6756C534B48FF84E598F984FCF3F9DC" box="[695,824,1577,1603]" pageId="12" pageNumber="13" value="1992-10-20">20.10.1992</collectingDate>
</date>
, leg. S. Chaitiamvong;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E36DF984FE8CF914" ID-GBIF-Occurrence="4104078315" collectingDate="1992-10-20" collectionCode="ZRC" collectorName="Don Sak &amp; S. Chaitiamvong" country="Thailand" elevation="4" location="Gulf" pageId="12" pageNumber="13" specimenCode="ZRC 1999.2006" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Surat Thani">
<specimenCount id="B48978F24B48FF84E36DF984FB45F9DC" box="[1090,1166,1577,1603]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 21.9 mm), 
<specimenCount id="B48978F24B48FF84E215F984FA57F9DC" box="[1338,1436,1577,1603]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 13.2 mm), MNHN-IU-2018-5751 (ex 
<specimenCode id="F2291B004B48FF84E541F9E0FCE3F9F8" box="[622,808,1613,1639]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E541F9E0FD6FF9F8" box="[622,676,1613,1639]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
1999.2006
</specimenCode>
), Gulf of Thailand, Siracha, fish port, leg. P.K.L. Ng et al., 
<date id="D63195BB4B48FF84E7EDF9DCFE88F914" box="[194,323,1649,1675]" pageId="12" pageNumber="13" value="1999-10-11">
<collectingDate id="C6756C534B48FF84E7EDF9DCFE88F914" box="[194,323,1649,1675]" pageId="12" pageNumber="13" value="1999-10-11">10.11.1999</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E661F9DCFDDCF930" ID-GBIF-Occurrence="4104078491" collectingDate="1992-10-20" collectionCode="ZRC" collectorName="Don Sak &amp; S. Chaitiamvong" country="Thailand" elevation="4" location="Gulf" pageId="12" pageNumber="13" specimenCode="ZRC 1999.2006" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Surat Thani">
<specimenCount id="B48978F24B48FF84E661F9DCFE52F914" box="[334,409,1649,1675]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 16.1 mm), 
<specimenCount id="B48978F24B48FF84E56FF9DCFD6AF914" box="[576,673,1649,1675]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 14.5 mm), 
<specimenCode id="F2291B004B48FF84E468F9DCFC34F913" box="[839,1023,1649,1676]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E468F9DCFCB6F914" box="[839,893,1649,1675]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
1999.2005
</specimenCode>
, Gulf of Thailand, Angsila, fish port, leg. P.K.L. Ng et al., 
<date id="D63195BB4B48FF84E6BDF938FDD8F930" box="[402,531,1685,1711]" pageId="12" pageNumber="13" value="1999-10-11">
<collectingDate id="C6756C534B48FF84E6BDF938FDD8F930" box="[402,531,1685,1711]" pageId="12" pageNumber="13" value="1999-10-11">10.11.1999</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E50EF938FD93F94C" ID-GBIF-Occurrence="4104078461" collectingDate="1992-10-20" collectingDateMax="2001-02-17" collectingDateMin="1926-06-14" collectionCode="ZRC" collectorName="Don Sak &amp; S. Chaitiamvong" country="Thailand" elevation="4" location="Gulf" municipality="Lem Sing" pageId="12" pageNumber="13" specimenCode="ZRC 1999.2006" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Surat Thani">
<specimenCount id="B48978F24B48FF84E50EF938FDB2F930" box="[545,633,1685,1711]" pageId="12" pageNumber="13" type="male">2 males</specimenCount>
(cl 15.5 mm, 17.5 mm), 
<specimenCount id="B48978F24B48FF84E48DF938FBCDF930" box="[930,1030,1685,1711]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 16.0 mm), 
<specimenCode id="F2291B004B48FF84E398F938FA90F92F" box="[1207,1371,1685,1712]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="13" type="Museum">
<collectionCode id="C49E2BBE4B48FF84E398F938FAC3F930" box="[1207,1288,1685,1711]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="12" pageNumber="13">USNM</collectionCode>
95528
</specimenCode>
, Gulf of Thailand, off Lem Sing, 
<date id="D63195BB4B48FF84E6FCF914FD9EF94C" box="[467,597,1721,1747]" pageId="12" pageNumber="13" value="1926-06-14">
<collectingDate id="C6756C534B48FF84E6FCF914FD9EF94C" box="[467,597,1721,1747]" pageId="12" pageNumber="13" value="1926-06-14">14.06.1926</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E54EF914FDB4F968" ID-GBIF-Occurrence="4104078494" collectingDate="1926-06-14" collectingDateMax="2001-02-17" collectingDateMin="1926-06-14" collectionCode="OUMNH, ZC" collectorName="P. K. L." country="Thailand" location="Andaman Sea" municipality="Lem Sing" pageId="12" pageNumber="13" specimenCode="ZRC 2008.0600" specimenCount="2" specimenCount-male="2" stateProvince="Phuket">
<specimenCount id="B48978F24B48FF84E54EF914FD65F94C" box="[609,686,1721,1747]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 21.5 mm), 
<specimenCode id="F2291B004B48FF84E473F914FBDDF94C" box="[860,1046,1721,1747]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E473F914FC59F94C" box="[860,914,1721,1747]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2008.0600
</specimenCode>
, Andaman Sea off Phuket, Phuket fish port, leg. P.K.L. Ng et al., 
<date id="D63195BB4B48FF84E6D5F970FDB7F968" box="[506,636,1757,1783]" pageId="12" pageNumber="13" value="2001-02-17">
<collectingDate id="C6756C534B48FF84E6D5F970FDB7F968" box="[506,636,1757,1783]" pageId="12" pageNumber="13" value="2001-02-17">17.02.2001</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="12E7B9264B48FF84E5A7F970FBD1F884" ID-GBIF-Occurrence="4104078431" collectingDate="1926-06-14" collectingDateMax="2001-02-17" collectingDateMin="1926-06-14" collectionCode="OUMNH, ZC" collectorName="S. Chiatiamuong" country="Thailand" location="Andaman Sea" municipality="Lem Sing" pageId="12" pageNumber="13" specimenCode="ZRC 2008.0600" specimenCount="2" specimenCount-male="2" stateProvince="Phuket">
<specimenCount id="B48978F24B48FF84E5A7F970FD1FF968" box="[648,724,1757,1783]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 20.2 mm), 
<collectionCode id="C49E2BBE4B48FF84E4AFF970FC3BF968" box="[896,1008,1757,1783]" collectionName="United Kingdom, Oxford, University Museum of Natural History" pageId="12" pageNumber="13">OUMNH</collectionCode>
. 
<collectionCode id="C49E2BBE4B48FF84E4DCF970FBD3F968" box="[1011,1048,1757,1783]" country="Austria" name="Zoological Collection, University of Vienna" pageId="12" pageNumber="13" type="University or college">ZC</collectionCode>
. 2019.06.43 [ex-ZRC 1998.1152], Andaman Sea off Phuket, Phuket fish port, leg. S. Chiatiamuong et al., 12.1998.
</materialsCitation>
</paragraph>
<paragraph id="A230B37B4B48FF84E7E8F888FEE4F818" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<materialsCitation id="12E7B9264B48FF84E7E8F888FE2AF8FC" ID-GBIF-Occurrence="4104078345" collectingDate="1961-02-06" collectionCode="RMNH, R" collectorName="E. R. Alfred" country="Malaysia" location="Penang" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Pulau Pinang">
<emphasis id="90FB6F694B48FF84E7E8F888FEE4F8A0" box="[199,303,1829,1855]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="DA98F3EB4B48FF84E7E8F888FEE4F8A0" box="[199,303,1829,1855]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B48FF84E66BF888FE5EF8A0" box="[324,405,1829,1855]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl indet., missing major cheliped), 
<collectionCode id="C49E2BBE4B48FF84E460F888FC68F8A0" box="[847,931,1829,1855]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="13" type="Museum">RMNH</collectionCode>
<collectorName id="0F7AD6AD4B48FF84E49EF888FBCEF8A0" box="[945,1029,1829,1855]" pageId="12" pageNumber="13">Crus.D.</collectorName>
49740, 
<collectingRegion id="604B7D994B48FF84E34DF888FB7CF8A0" box="[1122,1207,1829,1855]" country="Malaysia" name="Pulau Pinang" pageId="12" pageNumber="13">Penang</collectingRegion>
, 
<collectorName id="0F7AD6AD4B48FF84E3E7F888FA53F8A0" box="[1224,1432,1829,1855]" pageId="12" pageNumber="13">Kuala Jalan Baru</collectorName>
, 
<date id="D63195BB4B48FF84E7B8F8E4FED3F8FC" box="[151,280,1865,1891]" pageId="12" pageNumber="13" value="1961-02-06">
<collectingDate id="C6756C534B48FF84E7B8F8E4FED3F8FC" box="[151,280,1865,1891]" pageId="12" pageNumber="13" value="1961-02-06">02.06.1961</collectingDate>
</date>
, leg. E. 
<collectionCode id="C49E2BBE4B48FF84E65DF8E4FE43F8FC" box="[370,392,1865,1891]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
. 
<collectorName id="0F7AD6AD4B48FF84E6BBF8E4FE2AF8FC" box="[404,481,1865,1891]" pageId="12" pageNumber="13">Alfred</collectorName>
</materialsCitation>
; 
<materialsCitation id="12E7B9264B48FF84E6C0F8E4FEE0F818" ID-GBIF-Occurrence="4104078350" collectionCode="V" collectorName="C. Chong" country="Malaysia" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1">
<specimenCount id="B48978F24B48FF84E6C0F8E4FD9FF8FC" box="[495,596,1865,1891]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E5A9F8E4FD3AF8FC" box="[646,753,1865,1892]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="12" pageNumber="13" unit="mm" value="21.5">21.5 mm</quantity>
), MNHN-IU-2018-5594, Lumut, no further data, don. 
<collectionCode id="C49E2BBE4B48FF84E244F8E4FA4AF8FC" box="[1387,1409,1865,1891]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="13" type="Museum">V</collectionCode>
. 
<collectorName id="0F7AD6AD4B48FF84E2AAF8E4FF2FF818" pageId="12" pageNumber="13">C. Chong</collectorName>
, 1999
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B48FF84E7E8F83CFCE7F86B" blockId="12.[151,1437,497,2036]" pageId="12" pageNumber="13">
<materialsCitation id="12E7B9264B48FF84E7E8F83CFB93F834" ID-GBIF-Occurrence="4104078540" box="[199,1112,1937,1963]" collectionCode="ZRC" collectorName="Siglap" country="Singapore" location="Bedok area" pageId="12" pageNumber="13" specimenCode="ZRC 2008.0632" specimenCount="1" specimenCount-male="1">
<emphasis id="90FB6F694B48FF84E7E8F83CFEF2F834" box="[199,313,1937,1963]" italics="true" pageId="12" pageNumber="13">
<collectingCountry id="DA98F3EB4B48FF84E7E8F83CFEF2F834" box="[199,313,1937,1963]" name="Singapore" pageId="12" pageNumber="13">Singapore</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B48FF84E666F83CFE5DF834" box="[329,406,1937,1963]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E6EBF83CFDE5F834" box="[452,558,1937,1963]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="12" pageNumber="13" unit="mm" value="23.6">23.6 mm</quantity>
), 
<specimenCode id="F2291B004B48FF84E56DF83CFD30F834" box="[578,763,1937,1963]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E56DF83CFDB3F834" box="[578,632,1937,1963]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2008.0632
</specimenCode>
, 
<collectorName id="0F7AD6AD4B48FF84E428F83CFC84F834" box="[775,847,1937,1963]" pageId="12" pageNumber="13">Siglap</collectorName>
(
<location id="A750E5A04B48FF84E471F83CFC2EF834" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E471F83CFC2EF834" box="[862,997,1937,1963]" country="Singapore" name="Bedok area" pageId="12" pageNumber="13">Bedok area</location>
), 06.1933
</materialsCitation>
; 
<materialsCitation id="12E7B9264B48FF84E34BF83CFCE3F86B" ID-GBIF-Occurrence="4104078370" collectingDate="2014-03-25" collectionCode="ZRC" collectorName="S. C. Lim; A. Anker &amp; C. K. Chim" country="Singapore" elevation="12" location="Pulau Tekong" pageId="12" pageNumber="13" specimenCode="ZRC 2014.0671" specimenCount="1" specimenCount-male="1">
<specimenCount id="B48978F24B48FF84E34BF83CFB7BF834" box="[1124,1200,1937,1963]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(cl 
<quantity id="65771E9E4B48FF84E3F0F83CFAF3F834" box="[1247,1336,1937,1963]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="12" pageNumber="13" unit="mm" value="6.7">6.7 mm</quantity>
, missing both chelipeds), 
<specimenCode id="F2291B004B48FF84E67FF818FDC3F850" box="[336,520,1973,1999]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="13">
<collectionCode id="C49E2BBE4B48FF84E67FF818FE4DF850" box="[336,390,1973,1999]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="12" pageNumber="13">ZRC</collectionCode>
2014.0671
</specimenCode>
, south of 
<location id="A750E5A04B48FF84E55BF818FCDAF850" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B48FF84E55BF818FCDAF850" box="[628,785,1973,1999]" country="Singapore" name="Pulau Tekong" pageId="12" pageNumber="13">Pulau Tekong</location>
, MPA grid 0422, depth: 
<quantity id="65771E9E4B48FF84E30FF818FB52F850" box="[1056,1177,1973,1999]" metricMagnitude="1" metricUnit="m" metricValue="1.1949999999999998" metricValueMax="1.2" metricValueMin="1.19" pageId="12" pageNumber="13" unit="m" value="11.95" valueMax="12.0" valueMin="11.9">
<elevation id="29A254484B48FF84E30FF818FB52F850" box="[1056,1177,1973,1999]" metricMagnitude="1" metricUnit="m" metricValue="1.1949999999999998" metricValueMax="1.2" metricValueMin="1.19" pageId="12" pageNumber="13" unit="m" value="11.95" valueMax="12.0" valueMin="11.9">11.9–12 m</elevation>
</quantity>
, mud, clay, rectangular dredge, leg. 
<collectorName id="0F7AD6AD4B48FF84E60DF874FE41F86C" box="[290,394,2009,2035]" pageId="12" pageNumber="13">S.C. Lim</collectorName>
, A. Anker &amp; 
<collectorName id="0F7AD6AD4B48FF84E531F874FD50F86C" box="[542,667,2009,2035]" pageId="12" pageNumber="13">C.K. Chim</collectorName>
, 
<date id="D63195BB4B48FF84E588F874FCE3F86B" box="[679,808,2009,2036]" pageId="12" pageNumber="13" value="2014-03-25">
<collectingDate id="C6756C534B48FF84E588F874FCE3F86B" box="[679,808,2009,2036]" pageId="12" pageNumber="13" value="2014-03-25">25.03.2014</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B49FF85E7E8FF3AFC35FF4A" blockId="13.[151,1437,151,285]" pageId="13" pageNumber="14">
<materialsCitation id="12E7B9264B49FF85E7E8FF3AFAC3FF2E" ID-GBIF-Occurrence="4104078420" box="[199,1288,151,177]" collectingDate="1979-03-28" collectionCode="RMNH" collectorName="Crus. D." country="India" location="India" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<emphasis id="90FB6F694B49FF85E7E8FF3AFEC9FF2E" box="[199,258,151,177]" italics="true" pageId="13" pageNumber="14">
<collectingCountry id="DA98F3EB4B49FF85E7E8FF3AFEC9FF2E" box="[199,258,151,177]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
</emphasis>
: 
<specimenCount id="B48978F24B49FF85E63BFF3AFEAAFF2E" box="[276,353,151,177]" pageId="13" pageNumber="14" type="male">1 male</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B49FF85E6C4FF3AFDF4FF2E" box="[491,575,151,177]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="14" type="Museum">RMNH</collectionCode>
Crus.D.49747, Kochi (Cochin), 
<date id="D63195BB4B49FF85E496FF3AFBF2FF2E" box="[953,1081,151,177]" pageId="13" pageNumber="14" value="1979-03-28">
<collectingDate id="C6756C534B49FF85E496FF3AFBF2FF2E" box="[953,1081,151,177]" pageId="13" pageNumber="14" value="1979-03-28">28.03.1979</collectingDate>
</date>
, leg. 
<collectorName id="0F7AD6AD4B49FF85E355FF3AFACFFF2E" box="[1146,1284,151,177]" pageId="13" pageNumber="14">J.C. Miquel</collectorName>
;
</materialsCitation>
<materialsCitation id="12E7B9264B49FF85E23EFF3AFC31FF4A" ID-GBIF-Occurrence="4104078430" collectingDate="1979-03-28" collectionCode="RMNH" collectorName="Crus. D." country="India" location="India" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="B48978F24B49FF85E23EFF3AFABEFF2E" box="[1297,1397,151,177]" pageId="13" pageNumber="14" type="female">1 female</specimenCount>
(cl indet.), 
<collectionCode id="C49E2BBE4B49FF85E7C1FF16FE89FF4A" box="[238,322,187,213]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="14" type="Museum">RMNH</collectionCode>
Crus.D.49748, same collection data as for previous specimen
</materialsCitation>
.
</paragraph>
<paragraph id="A230B37B4B49FF85E7E8FF72FDD9FE82" blockId="13.[151,1437,151,285]" pageId="13" pageNumber="14">
<materialsCitation id="12E7B9264B49FF85E7E8FF72FD18FF66" ID-GBIF-Occurrence="4104078473" box="[199,723,222,249]" collectionCode="USNM" country="Pakistan" location="Pakistan" pageId="13" pageNumber="14" specimenCode="USNM 63515" specimenCount="1" specimenCount-female="1">
<emphasis id="90FB6F694B49FF85E7E8FF72FEE2FF66" box="[199,297,223,249]" italics="true" pageId="13" pageNumber="14">
<collectingCountry id="DA98F3EB4B49FF85E7E8FF72FEE2FF66" box="[199,297,223,249]" name="Pakistan" pageId="13" pageNumber="14">Pakistan</collectingCountry>
</emphasis>
. 
<specimenCount id="B48978F24B49FF85E619FF72FE5EFF66" box="[310,405,223,249]" pageId="13" pageNumber="14" type="female">1 female</specimenCount>
(cl 
<quantity id="65771E9E4B49FF85E692FF72FDE8FF66" box="[445,547,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="13" pageNumber="14" unit="mm" value="11.4">11.4 mm</quantity>
), 
<specimenCode id="F2291B004B49FF85E51BFF72FD18FF66" box="[564,723,222,249]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="14" type="Museum">
<collectionCode id="C49E2BBE4B49FF85E51BFF72FD4EFF66" box="[564,645,223,249]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="13" pageNumber="14">USNM</collectionCode>
63515
</specimenCode>
</materialsCitation>
, 
<materialsCitation id="12E7B9264B49FF85E5F3FF72FDC7FE82" ID-GBIF-Occurrence="4104078391" collectorName="H. Coutiere" country="Pakistan" location="Kurrachee Bay" pageId="13" pageNumber="14" specimenCount="1">
<collectingCountry id="DA98F3EB4B49FF85E5F3FF72FCF5FF66" box="[732,830,223,249]" name="Pakistan" pageId="13" pageNumber="14">Pakistan</collectingCountry>
, 
<location id="A750E5A04B49FF85E467FF72FC24FF66" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B49FF85E467FF72FC24FF66" box="[840,1007,223,249]" country="Pakistan" name="Kurrachee Bay" pageId="13" pageNumber="14">Kurrachee Bay</location>
(= Karachi), no further data [identified by 
<collectorName id="0F7AD6AD4B49FF85E795FEAEFEF7FE82" box="[186,316,259,285]" pageId="13" pageNumber="14">H. Coutière</collectorName>
as 
<taxonomicName id="658FC8F84B49FF85E64DFEAEFDCAFE82" authorityName="sensu De Man" authorityYear="1897" box="[354,513,259,285]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B49FF85E64DFEAEFDCAFE82" box="[354,513,259,285]" italics="true" pageId="13" pageNumber="14">A. euphrosyne</emphasis>
</taxonomicName>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B49FF9AE7E8FEC3FDEAFCA6" lastPageId="18" lastPageNumber="19" pageId="13" pageNumber="14" type="description">
<paragraph id="A230B37B4B49FF85E7E8FEC3FDF3FD86" blockId="13.[151,1437,366,789]" pageId="13" pageNumber="14">
<emphasis id="90FB6F694B49FF85E7E8FEC3FEA6FE17" bold="true" box="[199,365,366,392]" pageId="13" pageNumber="14">Redescription</emphasis>
. See 
<figureCitation id="3AB4AFFE4B49FF85E682FEC2FDD1FE16" box="[429,538,367,393]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="13.[151,250,1688,1712]" captionStartId-1="14.[151,250,1935,1959]" captionStartId-2="15.[151,250,1807,1831]" captionStartId-3="16.[151,250,1796,1820]" captionTargetBox-0="[151,1435,813,1665]" captionTargetBox-1="[151,1435,379,1912]" captionTargetBox-2="[203,1391,197,1776]" captionTargetBox-3="[151,1424,192,1761]" captionTargetId-0="figure-245@13.[151,1435,813,1665]" captionTargetId-1="figure-95@14.[151,1435,379,1912]" captionTargetId-2="figure-15@15.[183,1404,181,1784]" captionTargetId-3="figure-21@16.[151,1436,181,1772]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionText-0="FIGURE 4. Alpheus euphrosyne De Man, 1897: chelipeds of male (cl indet.) from Sumatra, Indonesia (ZMA Crus.D.202726) showing main features and terminology used in the present study; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (left) cheliped, mesial view; D—same, lateral view. Abbreviations (in alphabetical order): ad—adhesive disks (plaques) on dactylus and palm (small on major chela, highly reduced in minor chela); be—balaeniceps expansion of minor chela dactylus; ddr—dorsal ridge of major chela dactylus (dactylar dorsal ridge); dla—dorsolateral angle of major chela pollex; dlt—dorsolateral tooth of major / minor chela palm; dma—dorsomesial angle of major chela pollex; dmt—distomesial tooth of major / minor chela palm; dp—plunger of major chela dactylus (dactylar plunger); ds—dorsal shoulder of major / minor chela palm; dtg—dorsal transverse groove of major / minor chela palm (also known as dorsal notch); lbd—lateral balaeniceps ridge of minor chela dactylus; lbp—lateral balaeniceps ridge of minor chela pollex; li—linea impressa of major / minor chela palm; lld—lateral longitudinal depression (groove) of major / minor chela palm; ltr—smooth lateral transverse ridge of major chela palm (often poorly developed); mbd—mesial balaeniceps ridge of minor chela dactylus; mbp—mesial balaeniceps ridge of minor chela pollex; mld—mesial longitudinal depression (groove) of major / minor chela palm; msr—mesial subdistal ridge on major chela pollex; mtr—smooth mesial transverse ridge of major chela palm (often poorly developed); pf—fossa (socket) of major chela pollex; vs—ventral shoulder of major / minor chela palm; vtg—ventral transverse groove of major / minor chela palm (also known as ventral notch)." captionText-1="FIGURE 5. Alpheus euphrosyne De Man, 1897: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela in situ, mesial view; D—same, lateral view; E—minor (right) chela in situ, dorsal view. Refer to A for scale." captionText-2="FIGURE 6. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) from Angsila, Thailand (MNHN-IU-2018-5593); A— frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H, I—same, dactylus, mesial (H) and dorsal (I) views; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—second pleopod, appendices masculina and interna, anterior view; M—left uropod, dorsal view; N—right uropod, detail of distolateral angle." captionText-3="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.7912302" figureDoi-1="http://doi.org/10.5281/zenodo.7912304" figureDoi-2="http://doi.org/10.5281/zenodo.7912306" figureDoi-3="http://doi.org/10.5281/zenodo.7912310" httpUri-0="https://zenodo.org/record/7912302/files/figure.png" httpUri-1="https://zenodo.org/record/7912304/files/figure.png" httpUri-2="https://zenodo.org/record/7912306/files/figure.png" httpUri-3="https://zenodo.org/record/7912310/files/figure.png" pageId="13" pageNumber="14">Figs. 4–7</figureCitation>
. Large-sized species of 
<taxonomicName id="658FC8F84B49FF85E405FEC2FC4FFE16" authorityName="Fabricius" authorityYear="1798" box="[810,900,367,393]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="90FB6F694B49FF85E405FEC2FC4FFE16" box="[810,900,367,393]" italics="true" pageId="13" pageNumber="14">Alpheus</emphasis>
</taxonomicName>
(maximal cl 
<quantity id="65771E9E4B49FF85E333FEC2FB48FE16" box="[1052,1155,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="13" pageNumber="14" unit="mm" value="23.6">23.6 mm</quantity>
, tl ~72.0 mm). Carapace glabrous, finely pitted, especially around cardiac region, without pubescence, with slight grooves. Rostrum short, triangular, distally acute, reaching at most to mid-length of first article of antennular peduncle; rostral carina absent or present as low broad elevation; rostro-orbital furrows indistinct or extremely shallow. Pterygostomial angle broadly rounded; cardiac notch deep.
</paragraph>
<paragraph id="A230B37B4B49FF85E7E8FD8EFB63FC8A" blockId="13.[151,1437,366,789]" pageId="13" pageNumber="14">Pleon smooth, finely and scarcely pitted, without pubescence. Telson very broad, ovate-rectangular, slightly tapering towards posterior margin, about 1.3–1.4 times as long as maximal width near proximal margin; lateral margins with shallow concavity on proximal half; dorsal surface finely pitted, with longitudinal median depression densely furnished with setae along its margins, and with two pairs of short spiniform setae inserted in deep pits at considerable distance from lateral margins, first pair near telson mid-length, second pair between 0.7 and 0.8 of telson length, and with numerous erect simple setae; posterior margin broadly rounded, with two pairs of small spiniform setae at posterolateral angles, lateral shorter than mesial (often broken or missing).</paragraph>
<caption id="F6F0E3F34B49FF86E7B8F935FDC8FEA0" ID-DOI="http://doi.org/10.5281/zenodo.7912302" ID-Zenodo-Dep="7912302" httpUri="https://zenodo.org/record/7912302/files/figure.png" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" startId="13.[151,250,1688,1712]" targetBox="[151,1435,813,1665]" targetPageId="13" targetType="figure">
<paragraph id="A230B37B4B49FF85E7B8F935FA57F86B" blockId="13.[151,1437,1688,2036]" pageId="13" pageNumber="14">
<emphasis id="90FB6F694B49FF85E7B8F935FEC6F92F" bold="true" box="[151,269,1688,1712]" pageId="13" pageNumber="14">FIGURE 4</emphasis>
. 
<taxonomicName id="658FC8F84B49FF85E635F935FD48F92F" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[282,643,1688,1712]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B49FF85E635F935FE2DF92F" box="[282,486,1688,1712]" italics="true" pageId="13" pageNumber="14">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B49FF85E6C2F934FD48F92F" author="De Man, J. G." box="[493,643,1688,1712]" pageId="13" pageNumber="14" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: chelipeds of male (cl indet.) from Sumatra, Indonesia (ZMA Crus.D.202726) showing main features and terminology used in the present study; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (left) cheliped, mesial view; D—same, lateral view. Abbreviations (in alphabetical order): ad—adhesive disks (plaques) on dactylus and palm (small on major chela, highly reduced in minor chela); be—balaeniceps expansion of minor chela dactylus; ddr—dorsal ridge of major chela dactylus (dactylar dorsal ridge); dla—dorsolateral angle of major chela pollex; dlt—dorsolateral tooth of major / minor chela palm; dma—dorsomesial angle of major chela pollex; dmt—distomesial tooth of major / minor chela palm; dp—plunger of major chela dactylus (dactylar plunger); ds—dorsal shoulder of major / minor chela palm; dtg—dorsal transverse groove of major / minor chela palm (also known as dorsal notch); lbd—lateral balaeniceps ridge of minor chela dactylus; lbp—lateral balaeniceps ridge of minor chela pollex; li—linea impressa of major / minor chela palm; lld—lateral longitudinal depression (groove) of major / minor chela palm; ltr—smooth lateral transverse ridge of major chela
</paragraph>
<paragraph id="A230B37B4B4AFF86E7B8FF3AFDC8FEA0" blockId="14.[151,1436,151,319]" pageId="14" pageNumber="15">palm (often poorly developed); mbd—mesial balaeniceps ridge of minor chela dactylus; mbp—mesial balaeniceps ridge of minor chela pollex; mld—mesial longitudinal depression (groove) of major / minor chela palm; msr—mesial subdistal ridge on major chela pollex; mtr—smooth mesial transverse ridge of major chela palm (often poorly developed); pf—fossa (socket) of major chela pollex; vs—ventral shoulder of major / minor chela palm; vtg—ventral transverse groove of major / minor chela palm (also known as ventral notch).</paragraph>
</caption>
<caption id="F6F0E3F34B4AFF86E7B8F822FC97F870" ID-DOI="http://doi.org/10.5281/zenodo.7912304" ID-Zenodo-Dep="7912304" httpUri="https://zenodo.org/record/7912304/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1935,1959]" targetBox="[151,1435,379,1912]" targetPageId="14" targetType="figure">
<paragraph id="A230B37B4B4AFF86E7B8F822FC97F870" blockId="14.[151,1437,1935,2032]" pageId="14" pageNumber="15">
<emphasis id="90FB6F694B4AFF86E7B8F822FEC4F837" bold="true" box="[151,271,1935,1960]" pageId="14" pageNumber="15">FIGURE 5</emphasis>
. 
<taxonomicName id="658FC8F84B4AFF86E632F822FD40F837" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[285,651,1935,1960]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B4AFF86E632F822FE20F838" box="[285,491,1935,1959]" italics="true" pageId="14" pageNumber="15">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B4AFF86E6DDF83DFD40F837" author="De Man, J. G." box="[498,651,1936,1960]" pageId="14" pageNumber="15" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela 
<emphasis id="90FB6F694B4AFF86E30CF819FBAAF854" box="[1059,1121,1972,1995]" italics="true" pageId="14" pageNumber="15">in situ</emphasis>
, mesial view; D—same, lateral view; E—minor (right) chela 
<emphasis id="90FB6F694B4AFF86E6EBF875FDC8F870" box="[452,515,2008,2031]" italics="true" pageId="14" pageNumber="15">in situ</emphasis>
, dorsal view. Refer to A for scale.
</paragraph>
</caption>
<caption id="F6F0E3F34B4BFF87E7B8F8A2FD97F843" ID-DOI="http://doi.org/10.5281/zenodo.7912306" ID-Zenodo-Dep="7912306" httpUri="https://zenodo.org/record/7912306/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1807,1831]" targetBox="[203,1391,197,1776]" targetPageId="15" targetType="figure">
<paragraph id="A230B37B4B4BFF87E7B8F8A2FD97F843" blockId="15.[151,1437,1807,2012]" pageId="15" pageNumber="16">
<emphasis id="90FB6F694B4BFF87E7B8F8A2FEDBF8B7" bold="true" box="[151,272,1807,1832]" pageId="15" pageNumber="16">FIGURE 6</emphasis>
. 
<taxonomicName id="658FC8F84B4BFF87E60FF8BDFD58F8B7" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[288,659,1808,1832]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B4BFF87E60FF8BDFE24F8B7" box="[288,495,1808,1832]" italics="true" pageId="15" pageNumber="16">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B4BFF87E6D7F8BDFD58F8B7" author="De Man, J. G." box="[504,659,1808,1832]" pageId="15" pageNumber="16" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: male (cl 16.4 mm) from Angsila, Thailand (MNHN-IU-2018-5593); A— frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H, I—same, dactylus, mesial (H) and dorsal (I) views; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—second pleopod, appendices masculina and interna, anterior view; M—left uropod, dorsal view; N—right uropod, detail of distolateral angle.
</paragraph>
</caption>
<caption id="F6F0E3F34B54FF98E7B8F8A9FEDFF833" ID-DOI="http://doi.org/10.5281/zenodo.7912310" ID-Zenodo-Dep="7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1796,1820]" targetBox="[151,1424,192,1761]" targetPageId="16" targetType="figure">
<paragraph id="A230B37B4B54FF98E7B8F8A9FEDFF833" blockId="16.[151,1437,1796,1964]" pageId="16" pageNumber="17">
<emphasis id="90FB6F694B54FF98E7B8F8A9FEC7F883" bold="true" box="[151,268,1796,1820]" pageId="16" pageNumber="17">FIGURE 7</emphasis>
. 
<taxonomicName id="658FC8F84B54FF98E637F8A9FDB6F883" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[280,637,1796,1820]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B54FF98E637F8A9FE28F883" box="[280,483,1796,1820]" italics="true" pageId="16" pageNumber="17">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B54FF98E6C6F8A8FDB6F883" author="De Man, J. G." box="[489,637,1796,1820]" pageId="16" pageNumber="17" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view.
</paragraph>
</caption>
<paragraph id="A230B37B4B55FF99E7E8FF3AFCECFEFA" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">Antennular peduncle with stylocerite broad, markedly convex laterally, with acute tip, latter not overreaching distal margin of first article; ventromesial carina with small, anteriorly directed, sharp tooth; second article about 2.2–2.5 times as long as wide. Antennal peduncle with distoventral margin of basicerite unarmed or with minute, blunt or subacute tooth; carpocerite not reaching beyond scaphocerite; scaphocerite with lateral margin almost straight; blade very broad, separated from broad distolateral tooth by deep cleft; anterior margin of blade broadly rounded and reaching somewhat beyond distolateral tooth.</paragraph>
<paragraph id="A230B37B4B55FF99E7E8FEC2FE27FD86" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">Third maxilliped with antepenultimate article somewhat broadened; penultimate article elongate, about three times as long as wide, with long setae on ventrolateral margin, some surpassing ultimate article; ultimate article with dense transverse rows of short serrulate setae on mesial surface and much longer setae on dorsal and lateral surfaces, especially near or on apex; coxal lateral plate bluntly protruding dorsally; exopod fleshy, broad, reaching end of antepenultimate article.</paragraph>
<paragraph id="A230B37B4B55FF99E7E8FD8EFD68FB3E" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
Major cheliped of 
<taxonomicName id="658FC8F84B55FF99E6B6FD8EFDEDFDA2" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[409,550,547,573]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="90FB6F694B55FF99E6B6FD8EFDEDFDA2" box="[409,550,547,573]" italics="true" pageId="17" pageNumber="18">A. edwardsii</emphasis>
</taxonomicName>
- 
<typeStatus id="7D340DD94B55FF99E501FD8EFDA8FDA2" box="[558,611,547,573]" pageId="17" pageNumber="18">type</typeStatus>
. Merus stout, about twice as long as wide; ventromesial margin unarmed distally. Carpus very short, cup-shaped. Chela large, massive, with fingers about 0.6–0.7 length of palm; surface partly heavily granulated. Mesial face of palm with distinct mesial transverse ridge in proximal half, extending from proximal edge of subtriangular mesial longitudinal groove to ventral surface of palm, although not reaching it; large areas of mesial surface, including mesial longitudinal groove and part of mesial transverse ridge, densely covered with granules. Lateral face of palm mostly smooth, with relatively small field of granules present on low ridge between lateral longitudinal groove and dorsal edge of ventral shoulder; lateral longitudinal groove deep, subrectangular; proximal half with low transverse ridge. Dorsal shoulder overhanging transverse groove, i.e., distinctly protruding above groove in form of blunt tooth. Ventral shoulder pronounced, slightly protruding in lateral view, broadly rounded; surface covered with granules. Pollex with mesial face distinctly granulated over most of its surface; both mesial subdistal ridge and adjacent proximal protuberance prominent, extending subdistal angle of pollex; area ventral to mesial subdistal ridge conspicuously grooved; distomesial angle distinctly superior to 90°, blunt; lateral face distinctly granulated, except close to ventral edge; distolateral angle about 90°, broadly rounded. Dactylus with prominent dorsal ridge, latter distinctly twisted mesially and protruding proximally; mesial surface strongly granulated; lateral surface moderately granulated, mostly proximally so, with deep depression above plunger; dactylar plunger relatively short, with anterior margin gently sloping and merging into (i.e., almost in continuation with) ventral margin of dactylus; distal surface of plunger obliquely flattened, its mesial surface with deep transverse channel. Adhesive discs small.
</paragraph>
<paragraph id="A230B37B4B55FF99E7E8FB06FDECF9EA" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<collectingRegion id="604B7D994B55FF99E7E8FB06FECAFB5A" box="[199,257,1195,1221]" country="Maldives" name="Maale" pageId="17" pageNumber="18">Male</collectingRegion>
minor cheliped with chela strongly balaeniceps. Merus noticeably slenderer than that of major cheliped, about three times as long as wide; ventral margin with distomesial angle unarmed. Chela moderately swollen, slender in lateral view, with fingers about as long as palm. Palm subcylindrical, slightly compressed, feebly swollen; mesial face smooth proximally, distinctly granulated distally, with low bump-like elevation subdistally; mesial longitudinal groove well marked, deep; lateral face of palm with small field of granules distally, below lateral longitudinal groove; lateral longitudinal groove well marked, deep, with lower margin slightly elevated; dorsal shoulder pronounced, rounded, in form of blunt tooth slightly overhanging deep, broad dorsal transverse groove; ventral transverse groove moderately deep, bordered by distinct ventral shoulder on lateral surface. Pollex with mesial surfaces granulated proximally and with row of balaeniceps setae on low crest extending from base to mid-length of pollex; lateral surface mostly smooth, with similar row of balaeniceps setae as on mesial surface. Dactylus with well-developed proximal balaeniceps expansion, about 1.7–1.8 as long as maximal width; lateral and mesial dactylar ridges each with dense rows of balaeniceps setae. Cutting edges of both fingers shallowly excavated on each side of sharp, blade-like ridge.
</paragraph>
<paragraph id="A230B37B4B55FF99E7E8F9D2FF37F89A" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">Female minor cheliped simple, not balaeniceps. Chela slender, not particularly swollen, with fingers 1.5–2.0 times as long as palm. Palm granulated, more on mesial than on lateral side, without mesial or lateral longitudinal grooves. Fingers simple, with sharp cutting edges, slightly gaping when closed; fingertips strongly curved and crossing.</paragraph>
<paragraph id="A230B37B4B55FF9AE7E8F8A2FD02FE82" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,150,1977]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">Second pereiopod with ratio of carpal subarticles approximately equal to 4: 3: 1: 1: 1.5. Third pereiopod moderately robust; ischium armed with short, stout spiniform seta on ventrolateral surface near proximal margin, sometimes unarmed; merus about 4.5 times as long as maximal width, unarmed; propodus with two to four more or less stout spiniform setae on ventral margin and one pair of spiniform setae on distoventral margin adjacent to dactylus; dactylus about 0.3 length of propodus, spatulate, conspicuously expanded, slightly wider than propodus in dorsal view, with dense row of setae and tufts of setae along median ridge. Fourth pereiopod generally similar to third pereiopod, slightly slenderer; merus about 5.0 times as long as maximal width; dactylus about 0.3 length of propodus. Fifth pereiopod much slenderer than third and fourth pereiopods; ischium unarmed; propodus with four slender spiniform setae and well-developed setal brush; dactylus spatulate, less expanded compared to that of third or fourth pereiopod, about 0.4 length of propodus.</paragraph>
<paragraph id="A230B37B4B56FF9AE7E8FE8AFDACFE32" blockId="18.[151,1437,150,1977]" pageId="18" pageNumber="19">
<collectingRegion id="604B7D994B56FF9AE7E8FE8AFECAFEDE" box="[199,257,295,321]" country="Maldives" name="Maale" pageId="18" pageNumber="19">Male</collectingRegion>
second pleopod with appendix masculina 0.6–0.8 length of appendix interna. Uropod with each protopodal lobe ending in sharp tooth distally; exopod and endopod broad, ovate; diaeresis of exopod complete, straight for most part, with small subacute or blunt tooth near small distolateral spiniform seta and adjacent distolateral tooth; lateral margin of exopod broadly convex.
</paragraph>
<paragraph id="A230B37B4B56FF9AE7E8FE1AFBF2FE4E" blockId="18.[151,1437,150,1977]" box="[199,1081,439,465]" pageId="18" pageNumber="19">
Eggs numerous (&gt; 
<quantity id="65771E9E4B56FF9AE6BCFE1AFE17FE4E" box="[403,476,439,465]" metricMagnitude="0" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="19" unit="in" value="200.0">200 in</quantity>
large females), small; egg diameter less than 
<quantity id="65771E9E4B56FF9AE4F4FE1AFBFFFE4E" box="[987,1076,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="mm" value="0.6">0.6 mm</quantity>
.
</paragraph>
<paragraph id="A230B37B4B56FF9AE7E8FE77FDEAFCA6" blockId="18.[151,1437,150,1977]" pageId="18" pageNumber="19">
<emphasis id="90FB6F694B56FF9AE7E8FE77FEB5FE6A" bold="true" box="[199,382,474,501]" pageId="18" pageNumber="19">Colour pattern</emphasis>
. Body pale beige to greyish yellow; carapace with dark grey-blue anterolateral margin and two diffuse, dark brown, transverse bands, one across mid-length and one near posterior margin; each pleonite with dark brown to grey-brown band running transversely near posterior margin; antennular flagella bright blue (lateral) or reddish purple (mesial); antennal flagella bright blue; mesial face of major chela olive-green with large pale yellow or pale brown-orange areas, and two large, more or less conspicuous, violet-blue patches on palm; fingers distally pale orange-pink; dactylar plunger with whitish blue area; minor chela pale olive-green, darker and with purplish tinge on fingers; walking legs pink to reddish; pleopods reddish; telson pale greenish yellow, darker posteriorly; uropods pale yellow proximally, dark blue distally (
<figureCitation id="3AB4AFFE4B56FF9AE45EFD7AFC26FD6E" box="[881,1005,727,753]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="19.[151,250,1902,1926]" captionStartId-1="20.[151,250,1875,1899]" captionStartId-2="21.[151,250,1866,1890]" captionTargetBox-0="[151,1435,181,1877]" captionTargetBox-1="[151,1435,181,1851]" captionTargetBox-2="[159,1429,181,1842]" captionTargetId-0="figure-15@19.[151,1435,181,1878]" captionTargetId-1="figure-21@20.[151,1435,181,1851]" captionTargetId-2="figure-15@21.[159,1429,181,1842]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 8. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Satun, Thailand, in lateral (A) and dorsal (B) views (deposition unknown). Photographs courtesy of Rueangrit Promdam." captionText-1="FIGURE 9. Alpheus euphrosyne De Man, 1897: male (cl 16.8 mm) from Pattani, Thailand (ZRC 2014.0672), in dorsal view (A), with details of the frontal area and chelipeds (B). Photographs courtesy of Sukree Hajisamae." captionText-2="FIGURE 10. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Nha Trang Bay, Vietnam (ZMMU) [A]; male (cl indet.) from unknown locality in Taiwan (AST) [B]; male from Dongtou Island, Zhejiang, China (IOCAS-MBM188857) [C]; male (cl indet.) from Siracha, Thailand (ZRC 2000.2137) [D]; male (cl indet.) from Kochi, India (RMNH Crus.D.49747) [E, F]; A, B, C—habitus, dorsal view; D, E—same, lateral view; F—posterior pleon and tail fan, dorsal view. Photographs courtesy of Oleg Savinkin (A), Ming-Shiou Jeng (B), Xinzheng Li (C), Peter K.L. Ng (D) and Juan C.E. Miquel (E, F)." figureDoi-0="http://doi.org/10.5281/zenodo.7912312" figureDoi-1="http://doi.org/10.5281/zenodo.7912318" figureDoi-2="http://doi.org/10.5281/zenodo.7912320" httpUri-0="https://zenodo.org/record/7912312/files/figure.png" httpUri-1="https://zenodo.org/record/7912318/files/figure.png" httpUri-2="https://zenodo.org/record/7912320/files/figure.png" pageId="18" pageNumber="19">Figs. 8–10</figureCitation>
; see also black-and-white photograph in 
<bibRefCitation id="C61ECE8A4B56FF9AE79BFD56FE84FC8A" author="Li, X. &amp; Shen, A. &amp; Xu, Z." box="[180,335,763,789]" pageId="18" pageNumber="19" refId="ref72773" refString="Li, X., Shen, A. &amp; Xu, Z. (2008) Report on some shrimps from Dongtou Island, Zhejiang, China (Decapoda, Natantia). Crustaceana, 81, 1371 - 1330. https: // doi. org / 10.1163 / 156854008 X 369500" type="book" year="2008">
Li 
<emphasis id="90FB6F694B56FF9AE7FBFD56FEC7FC8A" box="[212,268,763,789]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2008
</bibRefCitation>
: fig. 2A, and colour photographs in 
<bibRefCitation id="C61ECE8A4B56FF9AE5CBFD56FBEEFC8A" author="Harikrishnan, M. &amp; Kurup, B. M." box="[740,1061,763,789]" pageId="18" pageNumber="19" pagination="1 - 280" refId="ref71572" refString="Harikrishnan, M. &amp; Kurup, B. M. (2008) Potential of snapping shrimps as ornamentals. In: Kurup, B. M., Boopendranth, M. R., Ravindran, K., Sairabanu &amp; Nair, A. G. (Eds.), Ornamental fish, breeding, farming and trade. Department of Fisheries, Government of Kerala, Kerala, pp. 1 - 280." type="book chapter" year="2008">Harikrishnan &amp; Kurup 2008</bibRefCitation>
: fig. 1 and 
<bibRefCitation id="C61ECE8A4B56FF9AE38EFD56FAABFC8A" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[1185,1376,763,789]" pageId="18" pageNumber="19" pagination="1695 - 1698" refId="ref73799" refString="Paras, N. J., Purushothaman, P., Chinnadurai, S., Renjith R. K., Remesan, M. P., Baiju, M. V. &amp; Edwin, L. (2019) Occurrence of Alpheus euphrosyne de Man, 1897 (Crustacea: Decapoda: Alpheidae) from the south-eastern Arabian Sea, India. Indian Journal of Geo Marine Sciences, 48, 1695 - 1698." type="journal article" year="2019">
Paras 
<emphasis id="90FB6F694B56FF9AE3CAFD56FAD5FC8A" box="[1253,1310,763,789]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2019
</bibRefCitation>
: figs. 1–3, specimen with faded colours).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B56FF9AE7E8FCEEFB95FCC2" box="[199,1118,834,861]" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="A230B37B4B56FF9AE7E8FCEEFB95FCC2" blockId="18.[151,1437,150,1977]" box="[199,1118,834,861]" pageId="18" pageNumber="19">
<emphasis id="90FB6F694B56FF9AE7E8FCEEFEABFCC3" bold="true" box="[199,352,834,861]" pageId="18" pageNumber="19">Type locality</emphasis>
. 
<materialsCitation id="12E7B9264B56FF9AE641FCEEFB95FCC2" ID-GBIF-Occurrence="4104078524" box="[366,1118,835,861]" country="Indonesia" location="Surabaya" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Jawa Timur" typeStatus="neotype">
<collectingRegion id="604B7D994B56FF9AE641FCEEFE6BFCC2" box="[366,416,835,861]" country="Indonesia" name="Jawa Timur" pageId="18" pageNumber="19">Java</collectingRegion>
Sea (
<bibRefCitation id="C61ECE8A4B56FF9AE6F1FCEEFDB6FCC2" author="De Man, J. G." box="[478,637,835,861]" pageId="18" pageNumber="19" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man 1897</bibRefCitation>
). 
<typeStatus id="7D340DD94B56FF9AE5BEFCEEFD3BFCC2" box="[657,752,835,861]" pageId="18" pageNumber="19" type="neotype">Neotype</typeStatus>
from 
<location id="A750E5A04B56FF9AE41BFCEEFC54FCC2" LSID="urn:lsid:plazi:treatment:2A26026D4B4FFF9FE7B8F9AFFC84FF66:A750E5A04B56FF9AE41BFCEEFC54FCC2" box="[820,927,835,861]" country="Indonesia" name="Surabaya" pageId="18" pageNumber="19" stateProvince="Jawa Timur">Surabaya</location>
, Java, 
<collectingCountry id="DA98F3EB4B56FF9AE4C5FCEEFB90FCC2" box="[1002,1115,835,861]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B56FF9AE7E8FCCBFDBAFBC6" pageId="18" pageNumber="19" type="distribution">
<paragraph id="A230B37B4B56FF9AE7E8FCCBFDBAFBC6" blockId="18.[151,1437,150,1977]" pageId="18" pageNumber="19">
<emphasis id="90FB6F694B56FF9AE7E8FCCBFE92FC1F" bold="true" box="[199,345,870,896]" pageId="18" pageNumber="19">Distribution</emphasis>
. Tropical Indo-West Pacific from the northern Arabian Sea to South 
<collectingCountry id="DA98F3EB4B56FF9AE3ABFCCAFB0CFC1E" box="[1156,1223,871,897]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
Sea and Java Sea (
<figureCitation id="3AB4AFFE4B56FF9AE7B0FC26FEC3FC3A" box="[159,264,906,933]" captionStart="FIGURE 51" captionStartId="105.[151,250,1943,1967]" captionTargetBox="[155,1433,181,1920]" captionTargetId="figure-15@105.[155,1433,181,1920]" captionTargetPageId="105" captionText="FIGURE 51. Distributional maps for species of Alpheus treated in this study: A—A. eurphrosyne De Man, 1897; B—A. eurydactylus De Man, 1920; C—A. richardsoni Yaldwyn, 1971; D—A. microrhynchus De Man, 1897; E—A. cyanoteles Yeo &amp; Ng, 1996; F—A. nomurai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7912455" httpUri="https://zenodo.org/record/7912455/files/figure.png" pageId="18" pageNumber="19">Fig. 51A</figureCitation>
), including 
<collectingCountry id="DA98F3EB4B56FF9AE6A0FC26FE25FC3A" box="[399,494,907,933]" name="Pakistan" pageId="18" pageNumber="19">Pakistan</collectingCountry>
(Karachi), 
<collectingCountry id="DA98F3EB4B56FF9AE55FFC26FD61FC3A" box="[624,682,907,933]" name="India" pageId="18" pageNumber="19">India</collectingCountry>
(
<collectingCountry id="DA98F3EB4B56FF9AE595FC26FCC8FC3A" box="[698,771,907,933]" name="India" pageId="18" pageNumber="19">Kochi</collectingCountry>
, Vembanad Lake, Gulf of Mannar, Palk Bay), 
<collectingCountry id="DA98F3EB4B56FF9AE23CFC26FA52FC3A" box="[1299,1433,907,933]" name="Bangladesh" pageId="18" pageNumber="19">Bangladesh</collectingCountry>
, 
<collectingCountry id="DA98F3EB4B56FF9AE7B8FC02FF30FC56" box="[151,251,943,969]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
(Andaman Sea: 
<collectingRegion id="604B7D994B56FF9AE69AFC02FDCDFC56" box="[437,518,943,969]" country="Thailand" name="Phuket" pageId="18" pageNumber="19">Phuket</collectingRegion>
; Gulf of 
<collectingCountry id="DA98F3EB4B56FF9AE547FC02FD1BFC56" box="[616,720,943,969]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
: 
<collectingRegion id="604B7D994B56FF9AE5F5FC02FCE0FC56" box="[730,811,943,969]" country="Thailand" name="Pattani" pageId="18" pageNumber="19">Pattani</collectingRegion>
, Angsila, Khanom, 
<collectingRegion id="604B7D994B56FF9AE329FC02FB40FC56" box="[1030,1163,943,969]" country="Thailand" name="Surat Thani" pageId="18" pageNumber="19">Surat Thani</collectingRegion>
, 
<collectingRegion id="604B7D994B56FF9AE3B9FC02FB12FC56" box="[1174,1241,943,969]" country="Thailand" name="Satun" pageId="18" pageNumber="19">Satun</collectingRegion>
, 
<collectingRegion id="604B7D994B56FF9AE3CBFC02FA85FC56" box="[1252,1358,943,969]" country="Thailand" name="Songkhla" pageId="18" pageNumber="19">Songkhla</collectingRegion>
Lake), 
<collectingCountry id="DA98F3EB4B56FF9AE7B8FC7EFEC1FC72" box="[151,266,979,1005]" name="Cambodia" pageId="18" pageNumber="19">Cambodia</collectingCountry>
(
<collectingRegion id="604B7D994B56FF9AE631FC7EFEB5FC72" box="[286,382,979,1005]" country="Cambodia" name="Kampot" pageId="18" pageNumber="19">Kampot</collectingRegion>
), 
<collectingCountry id="DA98F3EB4B56FF9AE6B9FC7EFE3EFC72" box="[406,501,979,1005]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
(Nhatrang Bay), 
<collectingCountry id="DA98F3EB4B56FF9AE5E7FC7EFCE4FC72" box="[712,815,979,1005]" name="Malaysia" pageId="18" pageNumber="19">Malaysia</collectingCountry>
(Andaman Sea: 
<collectingRegion id="604B7D994B56FF9AE4D5FC7EFB9AFC72" box="[1018,1105,979,1005]" country="Malaysia" name="Pulau Pinang" pageId="18" pageNumber="19">Penang</collectingRegion>
), 
<collectingCountry id="DA98F3EB4B56FF9AE346FC7EFB1DFC72" box="[1129,1238,979,1005]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
(
<collectingRegion id="604B7D994B56FF9AE3C5FC7EFAEBFC72" box="[1258,1312,979,1005]" country="Indonesia" name="Jawa Timur" pageId="18" pageNumber="19">Java</collectingRegion>
, 
<collectingRegion id="604B7D994B56FF9AE21FFC7EFA53FC72" box="[1328,1432,979,1005]" country="Indonesia" name="Sulawesi Utara" pageId="18" pageNumber="19">Sulawesi</collectingRegion>
, 
<collectingRegion id="604B7D994B56FF9AE7B8FC5AFF3CFB8E" box="[151,247,1015,1041]" country="Indonesia" name="Sumatera Utara" pageId="18" pageNumber="19">Sumatra</collectingRegion>
, Nias?), 
<collectingCountry id="DA98F3EB4B56FF9AE64CFC5AFE1EFB8E" box="[355,469,1015,1041]" name="Singapore" pageId="18" pageNumber="19">Singapore</collectingCountry>
(Bedok, Pulau Tekong), southern 
<collectingCountry id="DA98F3EB4B56FF9AE443FC5AFC64FB8E" box="[876,943,1015,1041]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
(
<collectingRegion id="604B7D994B56FF9AE4EDFC5AFB82FB8E" box="[962,1097,1015,1041]" country="China" name="Guangdong" pageId="18" pageNumber="19">Guangdong</collectingRegion>
, 
<collectingRegion id="604B7D994B56FF9AE376FC5AFB0BFB8E" box="[1113,1216,1015,1041]" country="China" name="Zhejiang" pageId="18" pageNumber="19">Zhejiang</collectingRegion>
, 
<collectingCountry id="DA98F3EB4B56FF9AE3FFFC5AFA97FB8E" box="[1232,1372,1015,1041]" name="Hong Kong" pageId="18" pageNumber="19">Hong Kong</collectingCountry>
), and 
<collectingCountry id="DA98F3EB4B56FF9AE7B8FBB6FF22FBAA" box="[151,233,1051,1077]" name="Taiwan" pageId="18" pageNumber="19">Taiwan</collectingCountry>
(Yensui, 
<collectingRegion id="604B7D994B56FF9AE679FBB6FE1AFBAA" box="[342,465,1051,1077]" country="Taiwan" name="Kaohsiung" pageId="18" pageNumber="19">Kaohsiung</collectingRegion>
) (
<bibRefCitation id="C61ECE8A4B56FF9AE6C7FBB6FD43FBAA" author="De Man, J. G." box="[488,648,1051,1077]" pageId="18" pageNumber="19" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man 1897</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B56FF9AE5BAFBB6FCE9FBAA" author="Kazmi, M. A." box="[661,802,1051,1077]" pageId="18" pageNumber="19" pagination="170 - 174" refId="ref72166" refString="Kazmi, M. A. (1974) A new species of the genus Alpheus from the Bay of Bengal (Decapoda, Caridea). Crustaceana, 27, 170 - 174. https: // doi. org / 10.1163 / 156854074 X 00389" type="journal article" year="1974">Kazmi 1974</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B56FF9AE400FBB6FBF2FBAA" author="Banner, D. M. &amp; Banner, A. H." box="[815,1081,1051,1077]" pageId="18" pageNumber="19" pagination="215 - 257" refId="ref69308" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner 1978</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B56FF9AE36AFBB6FB2AFBAA" author="Li, X. &amp; Shen, A. &amp; Xu, Z." box="[1093,1249,1051,1077]" pageId="18" pageNumber="19" refId="ref72773" refString="Li, X., Shen, A. &amp; Xu, Z. (2008) Report on some shrimps from Dongtou Island, Zhejiang, China (Decapoda, Natantia). Crustaceana, 81, 1371 - 1330. https: // doi. org / 10.1163 / 156854008 X 369500" type="book" year="2008">
Li 
<emphasis id="90FB6F694B56FF9AE349FBB6FB53FBAA" box="[1126,1176,1051,1077]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. 2008
</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B56FF9AE3C1FBB6FF3CFBC6" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." pageId="18" pageNumber="19" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B56FF9AE2A7FBB6FF7FFBC6" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2010
</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B56FF9AE62DFB92FE74FBC6" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[258,447,1087,1113]" pageId="18" pageNumber="19" pagination="1695 - 1698" refId="ref73799" refString="Paras, N. J., Purushothaman, P., Chinnadurai, S., Renjith R. K., Remesan, M. P., Baiju, M. V. &amp; Edwin, L. (2019) Occurrence of Alpheus euphrosyne de Man, 1897 (Crustacea: Decapoda: Alpheidae) from the south-eastern Arabian Sea, India. Indian Journal of Geo Marine Sciences, 48, 1695 - 1698." type="journal article" year="2019">
Paras 
<emphasis id="90FB6F694B56FF9AE66AFB92FEBCFBC6" box="[325,375,1087,1113]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. 2019
</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B56FF9AE7E8FBCFFAC8FBE2" box="[199,1283,1122,1149]" pageId="18" pageNumber="19" type="vernacular_names">
<paragraph id="A230B37B4B56FF9AE7E8FBCFFAC8FBE2" blockId="18.[151,1437,150,1977]" box="[199,1283,1122,1149]" pageId="18" pageNumber="19">
<emphasis id="90FB6F694B56FF9AE7E8FBCFFE35FBE2" bold="true" box="[199,510,1122,1149]" pageId="18" pageNumber="19">Common names proposed</emphasis>
. Euphrosyne snapping shrimp; De Man’s estuarine snapping shrimp.
</paragraph>
</subSubSection>
<subSubSection id="EA95E0F04B56FF9AE7E8FB2AFC19F826" pageId="18" pageNumber="19" type="biology_ecology">
<paragraph id="A230B37B4B56FF9AE7E8FB2AFE34FA5E" blockId="18.[151,1437,150,1977]" pageId="18" pageNumber="19">
<emphasis id="90FB6F694B56FF9AE7E8FB2AFE7EFB3F" bold="true" box="[199,437,1158,1185]" pageId="18" pageNumber="19">Ecology and biology</emphasis>
. 
<taxonomicName id="658FC8F84B56FF9AE6EEFB2AFD6AFB3E" authorityName="sensu De Man" authorityYear="1897" box="[449,673,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B56FF9AE6EEFB2AFD6AFB3E" box="[449,673,1159,1185]" italics="true" pageId="18" pageNumber="19">Alpheus euphrosyne</emphasis>
</taxonomicName>
appears to be mainly a marine / estuarine snapping shrimp associated with subtidal sandy, sandy-muddy and muddy bottoms.Therefore, it can be considered as a polyhaline and mesohaline species, typically encountered within or near large estuarine areas. However, 
<taxonomicName id="658FC8F84B56FF9AE4D8FB62FB5DFB76" authorityName="sensu De Man" authorityYear="1897" box="[1015,1174,1231,1257]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B56FF9AE4D8FB62FB5DFB76" box="[1015,1174,1231,1257]" italics="true" pageId="18" pageNumber="19">A. euphrosyne</emphasis>
</taxonomicName>
is capable of tolerating very low salinities and even freshwater conditions, at least in laboratory experiments (
<bibRefCitation id="C61ECE8A4B56FF9AE378FB5EFA5CFA92" author="Harikrishnan, M. &amp; Kurup, B. M." box="[1111,1431,1267,1293]" pageId="18" pageNumber="19" pagination="1 - 280" refId="ref71572" refString="Harikrishnan, M. &amp; Kurup, B. M. (2008) Potential of snapping shrimps as ornamentals. In: Kurup, B. M., Boopendranth, M. R., Ravindran, K., Sairabanu &amp; Nair, A. G. (Eds.), Ornamental fish, breeding, farming and trade. Department of Fisheries, Government of Kerala, Kerala, pp. 1 - 280." type="book chapter" year="2008">Harikrishnan &amp; Kurup 2008</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B56FF9AE7B8FABAFE67FAAE" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." box="[151,428,1303,1329]" pageId="18" pageNumber="19" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B56FF9AE61FFABAFEA2FAAE" box="[304,361,1303,1329]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2010
</bibRefCitation>
). The exact depth range of 
<taxonomicName id="658FC8F84B56FF9AE5F3FABAFCB0FAAE" authorityName="sensu De Man" authorityYear="1897" box="[732,891,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B56FF9AE5F3FABAFCB0FAAE" box="[732,891,1303,1329]" italics="true" pageId="18" pageNumber="19">A. euphrosyne</emphasis>
</taxonomicName>
is unknown as most specimens examined herein came from crab/prawn/fish landings or fish markets. Others were dredged from a depth of 
<quantity id="65771E9E4B56FF9AE3ACFA97FB26FACA" box="[1155,1261,1338,1365]" metricMagnitude="1" metricUnit="m" metricValue="1.775" metricValueMax="2.7" metricValueMin="0.85" pageId="18" pageNumber="19" unit="m" value="17.75" valueMax="27.0" valueMin="8.5">8.5–27 m</quantity>
(
<bibRefCitation id="C61ECE8A4B56FF9AE3D5FA96FA5CFACA" author="Thomas, M. M." box="[1274,1431,1339,1365]" pageId="18" pageNumber="19" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B56FF9AE7B8FAF2FE69FAE6" author="Banner, D. M. &amp; Banner, A. H." box="[151,418,1375,1401]" pageId="18" pageNumber="19" pagination="215 - 257" refId="ref69308" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner 1978</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B56FF9AE680FAF2FDBBFAE6" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[431,624,1375,1401]" pageId="18" pageNumber="19" pagination="1695 - 1698" refId="ref73799" refString="Paras, N. J., Purushothaman, P., Chinnadurai, S., Renjith R. K., Remesan, M. P., Baiju, M. V. &amp; Edwin, L. (2019) Occurrence of Alpheus euphrosyne de Man, 1897 (Crustacea: Decapoda: Alpheidae) from the south-eastern Arabian Sea, India. Indian Journal of Geo Marine Sciences, 48, 1695 - 1698." type="journal article" year="2019">
Paras 
<emphasis id="90FB6F694B56FF9AE6DBFAF2FDEDFAE6" box="[500,550,1375,1401]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. 2019
</bibRefCitation>
; present study) or collected with indigenous stake nets (
<bibRefCitation id="C61ECE8A4B56FF9AE3C3FAF2FF32FA02" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." pageId="18" pageNumber="19" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B56FF9AE2A7FAF2FF66FA02" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. 2010
</bibRefCitation>
). However, the identity of 
<taxonomicName id="658FC8F84B56FF9AE508FA2EFD0CFA02" authorityName="sensu De Man" authorityYear="1897" box="[551,711,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B56FF9AE508FA2EFD0CFA02" box="[551,711,1411,1437]" italics="true" pageId="18" pageNumber="19">A. euphrosyne</emphasis>
</taxonomicName>
in 
<bibRefCitation id="C61ECE8A4B56FF9AE5C2FA2EFBF4FA02" author="Harikrishnan, M. &amp; Kurup, B. M." box="[749,1087,1411,1437]" pageId="18" pageNumber="19" pagination="1 - 280" refId="ref71572" refString="Harikrishnan, M. &amp; Kurup, B. M. (2008) Potential of snapping shrimps as ornamentals. In: Kurup, B. M., Boopendranth, M. R., Ravindran, K., Sairabanu &amp; Nair, A. G. (Eds.), Ornamental fish, breeding, farming and trade. Department of Fisheries, Government of Kerala, Kerala, pp. 1 - 280." type="book chapter" year="2008">Harikrishnan &amp; Kurup (2008)</bibRefCitation>
and 
<bibRefCitation id="C61ECE8A4B56FF9AE358FA2EFA57FA02" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." box="[1143,1436,1411,1437]" pageId="18" pageNumber="19" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B56FF9AE23EFA2EFA81FA02" box="[1297,1354,1411,1437]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2010)
</bibRefCitation>
needs confirmation (see below).
</paragraph>
<paragraph id="A230B37B4B56FF9AE7E8FA66FD63F922" blockId="18.[151,1437,150,1977]" pageId="18" pageNumber="19">
In the lower 
<collectingCountry id="DA98F3EB4B56FF9AE67EFA66FE5EFA7A" box="[337,405,1483,1509]" name="India" pageId="18" pageNumber="19">Kochi</collectingCountry>
(
<collectingCountry id="DA98F3EB4B56FF9AE68DFA66FE3DFA7A" box="[418,502,1483,1509]" name="India" pageId="18" pageNumber="19">Cochin</collectingCountry>
) estuary in southern 
<collectingCountry id="DA98F3EB4B56FF9AE5F4FA66FCDCFA7A" box="[731,791,1483,1509]" name="India" pageId="18" pageNumber="19">India</collectingCountry>
, 
<bibRefCitation id="C61ECE8A4B56FF9AE40EFA66FBF4FA7A" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." box="[801,1087,1483,1509]" pageId="18" pageNumber="19" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B56FF9AE497FA66FC24FA7A" box="[952,1007,1483,1509]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2010)
</bibRefCitation>
observed a high preponderance of 
<taxonomicName id="658FC8F84B56FF9AE796FA42FE90F996" authorityName="sensu De Man" authorityYear="1897" box="[185,347,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B56FF9AE796FA42FE90F996" box="[185,347,1519,1545]" italics="true" pageId="18" pageNumber="19">A. euphrosyne</emphasis>
</taxonomicName>
in post- and pre-monsoon months, i.e., in salinity conditions varying from polyhaline in pre-monsoon months to mesohaline (5–18 
<superScript id="55FA1E334B56FF9AE57FF9BFFD93F9BF" attach="right" box="[592,600,1554,1568]" fontSize="6" pageId="18" pageNumber="19">0</superScript>
/ 
<subScript id="3E0BB13E4B56FF9AE54FF9B1FDBBF9B5" attach="none" box="[608,624,1564,1578]" fontSize="6" pageId="18" pageNumber="19">00</subScript>
) in post-monsoon months, suggesting an inward migration of primarily marine species to highly fertile estuarine areas for feeding and breeding. These authors also studied some aspects of the reproductive biology of 
<taxonomicName id="658FC8F84B56FF9AE6E3F9F6FDA1F9EA" authorityName="sensu De Man" authorityYear="1897" box="[460,618,1627,1653]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B56FF9AE6E3F9F6FDA1F9EA" box="[460,618,1627,1653]" italics="true" pageId="18" pageNumber="19">A. euphrosyne</emphasis>
</taxonomicName>
, examining 1309 females with cl 
<quantity id="65771E9E4B56FF9AE4CEF9F7FBB3F9EA" box="[993,1144,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.15" metricValueMin="0.85" pageId="18" pageNumber="19" unit="mm" value="15.0" valueMax="21.5" valueMin="8.5">8.5–21.5 mm</quantity>
. The large number of eggs (141–1553 eggs per brood, mean fecundity 480 272) and their small size (diameter range: 
<quantity id="65771E9E4B56FF9AE3BBF9D3FAF2F906" box="[1172,1337,1662,1689]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="5.3" metricValueMin="1.2" pageId="18" pageNumber="19" unit="mm" value="0.325" valueMax="0.53" valueMin="0.12">0.12–0.53 mm</quantity>
) suggest an extended larval development in this species.
</paragraph>
<paragraph id="A230B37B4B56FF9AE7E8F96AFC19F826" blockId="18.[151,1437,150,1977]" pageId="18" pageNumber="19">
<taxonomicName id="658FC8F84B56FF9AE7E8F96AFE55F97E" authorityName="De Man" authorityYear="1897" box="[199,414,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euphosyne">
<emphasis id="90FB6F694B56FF9AE7E8F96AFE55F97E" box="[199,414,1735,1761]" italics="true" pageId="18" pageNumber="19">Alpheus euphosyne</emphasis>
</taxonomicName>
is one of the largest snapping shrimps known to date, with the carapace length (cl) reaching 
<quantity id="65771E9E4B56FF9AE7B8F946FF37F89A" box="[151,252,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="18" pageNumber="19" unit="mm" value="22.6">22.6 mm</quantity>
and total body length (tl) up to 
<quantity id="65771E9E4B56FF9AE548F946FD7CF89A" box="[615,695,1771,1797]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="18" pageNumber="19" unit="mm" value="71.0">71 mm</quantity>
(
<bibRefCitation id="C61ECE8A4B56FF9AE5E9F946FCADF89A" author="Thomas, M. M." box="[710,870,1771,1797]" pageId="18" pageNumber="19" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976</bibRefCitation>
). The largest specimen in the present material is a female from 
<collectingCountry id="DA98F3EB4B56FF9AE609F8A2FE41F8B6" box="[294,394,1807,1833]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
(ZRC 2000.0957) with cl 
<quantity id="65771E9E4B56FF9AE59FF8A2FD35F8B6" box="[688,766,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="19" unit="mm" value="22.0">22 mm</quantity>
and tl 
<quantity id="65771E9E4B56FF9AE466F8A2FC52F8B6" box="[841,921,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="18" pageNumber="19" unit="mm" value="66.0">66 mm</quantity>
, with several other adults exceeding cl 
<quantity id="65771E9E4B56FF9AE261F8A2FA56F8B6" box="[1358,1437,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="mm" value="20.0">20 mm</quantity>
and tl 
<quantity id="65771E9E4B56FF9AE7CFF89EFEF8F8D2" box="[224,307,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="mm" value="60.0">60 mm</quantity>
; in the largest specimens (
<figureCitation id="3AB4AFFE4B56FF9AE54FF89EFD6EF8D2" box="[608,677,1843,1869]" captionStart="FIGURE 8" captionStartId="19.[151,250,1902,1926]" captionTargetBox="[151,1435,181,1877]" captionTargetId="figure-15@19.[151,1435,181,1878]" captionTargetPageId="19" captionText="FIGURE 8. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Satun, Thailand, in lateral (A) and dorsal (B) views (deposition unknown). Photographs courtesy of Rueangrit Promdam." figureDoi="http://doi.org/10.5281/zenodo.7912312" httpUri="https://zenodo.org/record/7912312/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
), the major chela length (chl) overreaches 
<quantity id="65771E9E4B56FF9AE3A1F89FFB2AF8D2" box="[1166,1249,1842,1869]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="19" unit="mm" value="35.0">35 mm</quantity>
. Due to its large size, 
<taxonomicName id="658FC8F84B56FF9AE7FEF8FAFEBAF8EE" authorityName="sensu De Man" authorityYear="1897" box="[209,369,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B56FF9AE7FEF8FAFEBAF8EE" box="[209,369,1879,1905]" italics="true" pageId="18" pageNumber="19">A. euphrosyne</emphasis>
</taxonomicName>
is occasionally sold on fish / crab landings and seafood markets, mainly in 
<collectingCountry id="DA98F3EB4B56FF9AE3E6F8FAFAE6F8EE" box="[1225,1325,1879,1905]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
(Angsila, Siracha, 
<collectingRegion id="604B7D994B56FF9AE7D6F8D6FEA8F80A" box="[249,355,1915,1941]" country="Thailand" name="Songkhla" pageId="18" pageNumber="19">Songkhla</collectingRegion>
Lake, 
<collectingRegion id="604B7D994B56FF9AE69FF8D6FDCAF80A" box="[432,513,1915,1941]" country="Thailand" name="Phuket" pageId="18" pageNumber="19">Phuket</collectingRegion>
; see also 
<figureCitation id="3AB4AFFE4B56FF9AE543F8D6FD19F80A" box="[620,722,1915,1941]" captionStart="FIGURE 10" captionStartId="21.[151,250,1866,1890]" captionTargetBox="[159,1429,181,1842]" captionTargetId="figure-15@21.[159,1429,181,1842]" captionTargetPageId="21" captionText="FIGURE 10. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Nha Trang Bay, Vietnam (ZMMU) [A]; male (cl indet.) from unknown locality in Taiwan (AST) [B]; male from Dongtou Island, Zhejiang, China (IOCAS-MBM188857) [C]; male (cl indet.) from Siracha, Thailand (ZRC 2000.2137) [D]; male (cl indet.) from Kochi, India (RMNH Crus.D.49747) [E, F]; A, B, C—habitus, dorsal view; D, E—same, lateral view; F—posterior pleon and tail fan, dorsal view. Photographs courtesy of Oleg Savinkin (A), Ming-Shiou Jeng (B), Xinzheng Li (C), Peter K.L. Ng (D) and Juan C.E. Miquel (E, F)." figureDoi="http://doi.org/10.5281/zenodo.7912320" httpUri="https://zenodo.org/record/7912320/files/figure.png" pageId="18" pageNumber="19">Fig. 10D</figureCitation>
), but also in 
<collectingCountry id="DA98F3EB4B56FF9AE44CF8D6FC6DF80A" box="[867,934,1915,1941]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
(Houzhai), 
<collectingCountry id="DA98F3EB4B56FF9AE303F8D6FB40F80A" box="[1068,1163,1915,1941]" name="Vietnam" pageId="18" pageNumber="19">Vietnam</collectingCountry>
(Nha Trang), 
<collectingCountry id="DA98F3EB4B56FF9AE206F8D6FA57F80A" box="[1321,1436,1915,1941]" name="Cambodia" pageId="18" pageNumber="19">Cambodia</collectingCountry>
(
<collectingRegion id="604B7D994B56FF9AE7B0F832FF36F826" box="[159,253,1951,1977]" country="Cambodia" name="Kampot" pageId="18" pageNumber="19">Kampot</collectingRegion>
) and 
<collectingCountry id="DA98F3EB4B56FF9AE615F832FEBFF826" box="[314,372,1951,1977]" name="India" pageId="18" pageNumber="19">India</collectingCountry>
(Pamban, Mandapam) (
<bibRefCitation id="C61ECE8A4B56FF9AE5ADF832FCEBF826" author="Thomas, M. M." box="[642,800,1951,1977]" pageId="18" pageNumber="19" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<caption id="F6F0E3F34B57FF9BE7B8F8C3FC9FF835" ID-DOI="http://doi.org/10.5281/zenodo.7912312" ID-Zenodo-Dep="7912312" httpUri="https://zenodo.org/record/7912312/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1902,1926]" targetBox="[151,1435,181,1877]" targetPageId="19" targetType="figure">
<paragraph id="A230B37B4B57FF9BE7B8F8C3FC9FF835" blockId="19.[151,1442,1902,1963]" pageId="19" pageNumber="20">
<emphasis id="90FB6F694B57FF9BE7B8F8C3FEDBF819" bold="true" box="[151,272,1902,1926]" pageId="19" pageNumber="20">FIGURE 8</emphasis>
. 
<taxonomicName id="658FC8F84B57FF9BE630F8C3FD5BF819" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[287,656,1902,1926]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B57FF9BE630F8C3FE26F819" box="[287,493,1902,1926]" italics="true" pageId="19" pageNumber="20">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B57FF9BE6D9F8C2FD5BF819" author="De Man, J. G." box="[502,656,1902,1926]" pageId="19" pageNumber="20" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: male (cl indet.) from Satun, Thailand, in lateral (A) and dorsal (B) views (deposition unknown). Photographs courtesy of Rueangrit Promdam.
</paragraph>
</caption>
<caption id="F6F0E3F34B50FF9CE7B8F8FEFBB2F810" ID-DOI="http://doi.org/10.5281/zenodo.7912318" ID-Zenodo-Dep="7912318" httpUri="https://zenodo.org/record/7912318/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1875,1899]" targetBox="[151,1435,181,1851]" targetPageId="20" targetType="figure">
<paragraph id="A230B37B4B50FF9CE7B8F8FEFBB2F810" blockId="20.[151,1436,1875,1935]" pageId="20" pageNumber="21">
<emphasis id="90FB6F694B50FF9CE7B8F8FEFEC5F8F4" bold="true" box="[151,270,1875,1899]" pageId="20" pageNumber="21">FIGURE 9</emphasis>
. 
<taxonomicName id="658FC8F84B50FF9CE633F8FEFD43F8F4" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[284,648,1875,1899]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B50FF9CE633F8FEFE22F8F4" box="[284,489,1875,1899]" italics="true" pageId="20" pageNumber="21">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B50FF9CE6DFF8FEFD43F8F4" author="De Man, J. G." box="[496,648,1875,1899]" pageId="20" pageNumber="21" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: male (cl 16.8 mm) from Pattani, Thailand (ZRC 2014.0672), in dorsal view (A), with details of the frontal area and chelipeds (B). Photographs courtesy of Sukree Hajisamae.
</paragraph>
</caption>
<caption id="F6F0E3F34B51FF9DE7B8F8E7FB69F86D" ID-DOI="http://doi.org/10.5281/zenodo.7912320" ID-Zenodo-Dep="7912320" httpUri="https://zenodo.org/record/7912320/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1866,1890]" targetBox="[159,1429,181,1842]" targetPageId="21" targetType="figure">
<paragraph id="A230B37B4B51FF9DE7B8F8E7FB69F86D" blockId="21.[151,1437,1866,2035]" pageId="21" pageNumber="22">
<emphasis id="90FB6F694B51FF9DE7B8F8E7FED2F8FD" bold="true" box="[151,281,1866,1890]" pageId="21" pageNumber="22">FIGURE 10</emphasis>
. 
<taxonomicName id="658FC8F84B51FF9DE609F8E7FD47F8FD" authority="De Man, 1897" authorityName="sensu De Man" authorityYear="1897" box="[294,652,1866,1890]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B51FF9DE609F8E7FE3AF8FD" box="[294,497,1866,1890]" italics="true" pageId="21" pageNumber="22">Alpheus euphrosyne</emphasis>
<bibRefCitation id="C61ECE8A4B51FF9DE6D8F8E6FD47F8FD" author="De Man, J. G." box="[503,652,1866,1890]" pageId="21" pageNumber="22" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man, 1897</bibRefCitation>
</taxonomicName>
: male (cl indet.) from Nha Trang Bay, Vietnam (ZMMU) [A]; male (cl indet.) from unknown locality in Taiwan (AST) [B]; male from Dongtou Island, Zhejiang, China (IOCAS-MBM188857) [C]; male (cl indet.) from Siracha, Thailand (ZRC 2000.2137) [D]; male (cl indet.) from Kochi, India (RMNH Crus.D.49747) [E, F]; A, B, C—habitus, dorsal view; D, E—same, lateral view; F—posterior pleon and tail fan, dorsal view. Photographs courtesy of Oleg Savinkin (A), Ming-Shiou Jeng (B), Xinzheng Li (C), Peter K.L. Ng (D) and Juan C.E. Miquel (E, F).
</paragraph>
</caption>
<subSubSection id="EA95E0F04B52FF9FE7E8FF3BFC84FF66" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="discussion">
<paragraph id="A230B37B4B52FF9EE7E8FF3BFB70FD6E" blockId="22.[151,1437,150,2013]" pageId="22" pageNumber="23">
<emphasis id="90FB6F694B52FF9EE7E8FF3BFE71FF2E" bold="true" box="[199,442,150,177]" pageId="22" pageNumber="23">Taxonomic remarks</emphasis>
. 
<taxonomicName id="658FC8F84B52FF9EE6E4FF3AFD7BFF2E" authorityName="sensu De Man" authorityYear="1897" box="[459,688,151,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE6E4FF3AFD7BFF2E" box="[459,688,151,177]" italics="true" pageId="22" pageNumber="23">Alpheus euphrosyne</emphasis>
</taxonomicName>
can be most easily recognised by several features on the major cheliped (
<figureCitation id="3AB4AFFE4B52FF9EE626FF16FE5FFF4A" box="[265,404,186,213]" captionStart="FIGURE 5" captionStartId="14.[151,250,1935,1959]" captionTargetBox="[151,1435,379,1912]" captionTargetId="figure-95@14.[151,1435,379,1912]" captionTargetPageId="14" captionText="FIGURE 5. Alpheus euphrosyne De Man, 1897: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela in situ, mesial view; D—same, lateral view; E—minor (right) chela in situ, dorsal view. Refer to A for scale." figureDoi="http://doi.org/10.5281/zenodo.7912304" httpUri="https://zenodo.org/record/7912304/files/figure.png" pageId="22" pageNumber="23">Figs. 5C, D</figureCitation>
, 
<figureCitation id="3AB4AFFE4B52FF9EE68DFF16FE23FF4A" box="[418,488,187,213]" captionStart="FIGURE 7" captionStartId="16.[151,250,1796,1820]" captionTargetBox="[151,1424,192,1761]" captionTargetId="figure-21@16.[151,1436,181,1772]" captionTargetPageId="16" captionText="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="22" pageNumber="23">7A–D</figureCitation>
) and telson (
<figureCitation id="3AB4AFFE4B52FF9EE5ADFF16FD10FF4A" box="[642,731,187,213]" captionStart="FIGURE 6" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[203,1391,197,1776]" captionTargetId="figure-15@15.[183,1404,181,1784]" captionTargetPageId="15" captionText="FIGURE 6. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) from Angsila, Thailand (MNHN-IU-2018-5593); A— frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H, I—same, dactylus, mesial (H) and dorsal (I) views; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—second pleopod, appendices masculina and interna, anterior view; M—left uropod, dorsal view; N—right uropod, detail of distolateral angle." figureDoi="http://doi.org/10.5281/zenodo.7912306" httpUri="https://zenodo.org/record/7912306/files/figure.png" pageId="22" pageNumber="23">Fig. 6B</figureCitation>
) and, when alive or fresh, by its unmistakable and therefore diagnostic colour pattern (
<figureCitation id="3AB4AFFE4B52FF9EE6E6FF72FD83FF66" box="[457,584,223,249]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="19.[151,250,1902,1926]" captionStartId-1="20.[151,250,1875,1899]" captionStartId-2="21.[151,250,1866,1890]" captionTargetBox-0="[151,1435,181,1877]" captionTargetBox-1="[151,1435,181,1851]" captionTargetBox-2="[159,1429,181,1842]" captionTargetId-0="figure-15@19.[151,1435,181,1878]" captionTargetId-1="figure-21@20.[151,1435,181,1851]" captionTargetId-2="figure-15@21.[159,1429,181,1842]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 8. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Satun, Thailand, in lateral (A) and dorsal (B) views (deposition unknown). Photographs courtesy of Rueangrit Promdam." captionText-1="FIGURE 9. Alpheus euphrosyne De Man, 1897: male (cl 16.8 mm) from Pattani, Thailand (ZRC 2014.0672), in dorsal view (A), with details of the frontal area and chelipeds (B). Photographs courtesy of Sukree Hajisamae." captionText-2="FIGURE 10. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Nha Trang Bay, Vietnam (ZMMU) [A]; male (cl indet.) from unknown locality in Taiwan (AST) [B]; male from Dongtou Island, Zhejiang, China (IOCAS-MBM188857) [C]; male (cl indet.) from Siracha, Thailand (ZRC 2000.2137) [D]; male (cl indet.) from Kochi, India (RMNH Crus.D.49747) [E, F]; A, B, C—habitus, dorsal view; D, E—same, lateral view; F—posterior pleon and tail fan, dorsal view. Photographs courtesy of Oleg Savinkin (A), Ming-Shiou Jeng (B), Xinzheng Li (C), Peter K.L. Ng (D) and Juan C.E. Miquel (E, F)." figureDoi-0="http://doi.org/10.5281/zenodo.7912312" figureDoi-1="http://doi.org/10.5281/zenodo.7912318" figureDoi-2="http://doi.org/10.5281/zenodo.7912320" httpUri-0="https://zenodo.org/record/7912312/files/figure.png" httpUri-1="https://zenodo.org/record/7912318/files/figure.png" httpUri-2="https://zenodo.org/record/7912320/files/figure.png" pageId="22" pageNumber="23">Figs. 8–10</figureCitation>
). The most important distinguishing features of the major cheliped of 
<taxonomicName id="658FC8F84B52FF9EE2ABFF72FEDCFE82" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE2ABFF72FEDCFE82" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
are the absence of a sharp tooth on the mesioventral margin of the merus; the presence of a prominent sharp ridge preceded by a conspicuous bump-like elevation on the mesial face of the pollex; the dorsal shoulder forming a blunt tooth overhanging the adjacent transverse groove; the mesial palmar face with large areas covered by numerous, very conspicuous granules; and the dactylus with a very short, distally truncate plunger and prominent, mesially protruding, proximal ridge (
<figureCitation id="3AB4AFFE4B52FF9EE514FE3EFD09FE32" box="[571,706,402,429]" captionStart="FIGURE 5" captionStartId="14.[151,250,1935,1959]" captionTargetBox="[151,1435,379,1912]" captionTargetId="figure-95@14.[151,1435,379,1912]" captionTargetPageId="14" captionText="FIGURE 5. Alpheus euphrosyne De Man, 1897: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela in situ, mesial view; D—same, lateral view; E—minor (right) chela in situ, dorsal view. Refer to A for scale." figureDoi="http://doi.org/10.5281/zenodo.7912304" httpUri="https://zenodo.org/record/7912304/files/figure.png" pageId="22" pageNumber="23">Figs. 5C, D</figureCitation>
, 
<figureCitation id="3AB4AFFE4B52FF9EE5E0FE3EFCDCFE32" box="[719,791,403,429]" captionStart="FIGURE 7" captionStartId="16.[151,250,1796,1820]" captionTargetBox="[151,1424,192,1761]" captionTargetId="figure-21@16.[151,1436,181,1772]" captionTargetPageId="16" captionText="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="22" pageNumber="23">7A–D</figureCitation>
). The male minor cheliped is fully balaeniceps, i.e., with a strongly expanded dactylus and well-developed rows of balaeniceps setae, and with a pronounced sculpture on the palm, consisting of a deep dorsal groove and shallower ventral transversal groove (the former usually being slightly overhung by dorsal shoulder), and well-marked longitudinal grooves on both sides (
<figureCitation id="3AB4AFFE4B52FF9EE3B6FE52FB32FD86" box="[1177,1273,510,537]" captionStart="FIGURE 5" captionStartId="14.[151,250,1935,1959]" captionTargetBox="[151,1435,379,1912]" captionTargetId="figure-95@14.[151,1435,379,1912]" captionTargetPageId="14" captionText="FIGURE 5. Alpheus euphrosyne De Man, 1897: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela in situ, mesial view; D—same, lateral view; E—minor (right) chela in situ, dorsal view. Refer to A for scale." figureDoi="http://doi.org/10.5281/zenodo.7912304" httpUri="https://zenodo.org/record/7912304/files/figure.png" pageId="22" pageNumber="23">Figs. 5E</figureCitation>
, 
<figureCitation id="3AB4AFFE4B52FF9EE22BFE52FA80FD86" box="[1284,1355,511,537]" captionStart="FIGURE 7" captionStartId="16.[151,250,1796,1820]" captionTargetBox="[151,1424,192,1761]" captionTargetId="figure-21@16.[151,1436,181,1772]" captionTargetPageId="16" captionText="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="22" pageNumber="23">7E–G</figureCitation>
). Other important features of 
<taxonomicName id="658FC8F84B52FF9EE6BFFD8EFDFAFDA2" authorityName="sensu De Man" authorityYear="1897" box="[400,561,547,573]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE6BFFD8EFDFAFDA2" box="[400,561,547,573]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
are the absence of a rostral carina; the extremely shallow, barely noticeable rostro-orbital furrows; the very broad scaphocerite; the antennal basicerite unarmed or armed with a minute subacute tooth; the ischium of the third pereiopod typically armed with a small spiniform seta (not illustrated by 
<bibRefCitation id="C61ECE8A4B52FF9EE210FDC6FF19FD36" author="De Man, J. G." pageId="22" pageNumber="23" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man 1897</bibRefCitation>
, see 
<figureCitation id="3AB4AFFE4B52FF9EE625FD22FEA9FD36" box="[266,354,655,681]" captionStart="FIGURE 1" captionStartId="4.[151,250,1452,1476]" captionTargetBox="[167,1423,198,1420]" captionTargetId="figure-21@4.[152,1435,181,1427]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus euphrosyne De Man, 1897: syntype, female (cl 46.0 mm) from Java Sea, Indonesia (after De Man 1897, specimen lost) [A–E]; male (cl indet., tl 34.0 mm) from Bangkok, Thailand (after De Man 1898) [F, G]; A—frontal region, dorsal view; B—female minor (left) cheliped, lateral view; C—second pereiopod, lateral view; D—third pereiopod, lateral view; E—sixth pleonite, telson and uropods, dorsal view; F—male minor (right) cheliped, dorsolateral view; G—same, minor chela, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7912294" httpUri="https://zenodo.org/record/7912294/files/figure.png" pageId="22" pageNumber="23">Fig. 1D</figureCitation>
); the third and fourth pereiopods with dactyli expanded, spatulate; and the dorsal surface of the telson with a longitudinal median depression, with its lateral edges densely furnished with setae (
<figureCitation id="3AB4AFFE4B52FF9EE3F6FD1EFF69FD6E" captionStart="FIGURE 6" captionStartId="15.[151,250,1807,1831]" captionTargetBox="[203,1391,197,1776]" captionTargetId="figure-15@15.[183,1404,181,1784]" captionTargetPageId="15" captionText="FIGURE 6. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) from Angsila, Thailand (MNHN-IU-2018-5593); A— frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H, I—same, dactylus, mesial (H) and dorsal (I) views; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—second pleopod, appendices masculina and interna, anterior view; M—left uropod, dorsal view; N—right uropod, detail of distolateral angle." figureDoi="http://doi.org/10.5281/zenodo.7912306" httpUri="https://zenodo.org/record/7912306/files/figure.png" pageId="22" pageNumber="23">Fig. 6A, B, D, G, I</figureCitation>
). For separation of 
<taxonomicName id="658FC8F84B52FF9EE651FD7AFDD6FD6E" authorityName="sensu De Man" authorityYear="1897" box="[382,541,727,753]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE651FD7AFDD6FD6E" box="[382,541,727,753]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
from closely related species see below and refer to 
<tableCitation id="EF0D86C04B52FF9EE34EFD7AFB7EFD6E" box="[1121,1205,727,753]" captionStart="TABLE 1" captionStartId="93.[145,233,465,491]" captionTargetPageId="93" captionText="TABLE 1. Comparison of main taxonomically useful characters (morphology, colour pattern, larval development, ecology) in the Alpheus euphrosyne De Man, 1897—A. micro-" httpUri="http://table.plazi.org/id/F6F0E3F34B19FFD5E7BEFE7CF8F8FD90" pageId="22" pageNumber="23" tableUuid="F6F0E3F34B19FFD5E7BEFE7CF8F8FD90">Table 1</tableCitation>
.
</paragraph>
<paragraph id="A230B37B4B52FF9EE7E8FD56FE97FB76" blockId="22.[151,1437,150,2013]" pageId="22" pageNumber="23">
All material reported as 
<taxonomicName id="658FC8F84B52FF9EE6F3FD56FDAAFC8B" authorityName="Kazmi" authorityYear="1974" box="[476,609,763,789]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirmiziae">
<emphasis id="90FB6F694B52FF9EE6F3FD56FDAAFC8B" box="[476,609,763,789]" italics="true" pageId="22" pageNumber="23">A. tirmiziae</emphasis>
</taxonomicName>
(
<bibRefCitation id="C61ECE8A4B52FF9EE55FFD56FCCBFC8A" author="Kazmi, M. A." box="[624,768,763,789]" pageId="22" pageNumber="23" pagination="170 - 174" refId="ref72166" refString="Kazmi, M. A. (1974) A new species of the genus Alpheus from the Bay of Bengal (Decapoda, Caridea). Crustaceana, 27, 170 - 174. https: // doi. org / 10.1163 / 156854074 X 00389" type="journal article" year="1974">Kazmi 1974</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B52FF9EE422FD56FBC0FC8A" author="Kazmi, M. A. &amp; Kazmi, Q. B." box="[781,1035,763,789]" pageId="22" pageNumber="23" pagination="151 - 157" refId="ref72213" refString="Kazmi, M. A. &amp; Kazmi, Q. B. (1979) A check-list of marine caridean prawns of Pakistan. Biologia, 25, 151 - 157." type="journal article" year="1979">Kazmi &amp; Kazmi 1979</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B52FF9EE337FD56FAEFFC8A" author="Banner, D. M. &amp; Banner, A. H." box="[1048,1316,763,789]" pageId="22" pageNumber="23" pagination="215 - 257" refId="ref69308" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner 1978</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B52FF9EE21EFD56FF1AFCA6" author="Sha, Z. &amp; Wang, Y. &amp; Cui, D." pageId="22" pageNumber="23" refId="ref74259" refString="Sha, Z., Wang, Y. &amp; Cui, D. (2019) The Alpheidae from China seas. Crustacea: Decapoda: Caridea. Springer, Singapore, 323 pp. https: // doi. org / 10.1007 / 978 - 981 - 15 - 0648 - 2 _ 2" type="book" year="2019">
Sha 
<emphasis id="90FB6F694B52FF9EE24CFD56FA5EFC8A" box="[1379,1429,763,789]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. 2019
</bibRefCitation>
) is herein referred to 
<taxonomicName id="658FC8F84B52FF9EE6E2FCB2FDA5FCA6" authorityName="sensu De Man" authorityYear="1897" box="[461,622,799,825]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE6E2FCB2FDA5FCA6" box="[461,622,799,825]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
. The female 
<typeStatus id="7D340DD94B52FF9EE428FCB2FCA2FCA6" box="[775,873,799,825]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="658FC8F84B52FF9EE4BDFCB2FBDCFCA7" authorityName="Kazmi" authorityYear="1974" box="[914,1047,799,825]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirmiziae">
<emphasis id="90FB6F694B52FF9EE4BDFCB2FBDCFCA7" box="[914,1047,799,825]" italics="true" pageId="22" pageNumber="23">A. tirmiziae</emphasis>
</taxonomicName>
dredged in the Bay of Bengal off 
<collectingCountry id="DA98F3EB4B52FF9EE7B8FCEEFED1FCC2" box="[151,282,835,861]" name="Bangladesh" pageId="22" pageNumber="23">Bangladesh</collectingCountry>
has a somewhat aberrant, most probably damaged rostro-orbital area (
<bibRefCitation id="C61ECE8A4B52FF9EE316FCEEFA8DFCC2" author="Banner, D. M. &amp; Banner, A. H." box="[1081,1350,835,861]" pageId="22" pageNumber="23" pagination="215 - 257" refId="ref69308" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner 1978</bibRefCitation>
), but in most other characters agrees very well with female specimens of 
<taxonomicName id="658FC8F84B52FF9EE4A7FCCAFBE2FC1E" authorityName="sensu De Man" authorityYear="1897" box="[904,1065,871,897]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE4A7FCCAFBE2FC1E" box="[904,1065,871,897]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
from 
<collectingCountry id="DA98F3EB4B52FF9EE35EFCCAFB1EFC1E" box="[1137,1237,871,897]" name="Thailand" pageId="22" pageNumber="23">Thailand</collectingCountry>
and 
<collectingCountry id="DA98F3EB4B52FF9EE220FCCAFAAEFC1E" box="[1295,1381,871,897]" name="Taiwan" pageId="22" pageNumber="23">Taiwan</collectingCountry>
. The main difference between 
<taxonomicName id="658FC8F84B52FF9EE696FC26FDF5FC3B" authorityName="Kazmi" authorityYear="1974" box="[441,574,907,933]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirmiziae">
<emphasis id="90FB6F694B52FF9EE696FC26FDF5FC3B" box="[441,574,907,933]" italics="true" pageId="22" pageNumber="23">A. tirmiziae</emphasis>
</taxonomicName>
and 
<taxonomicName id="658FC8F84B52FF9EE557FC26FCD2FC3A" authorityName="sensu De Man" authorityYear="1897" box="[632,793,907,933]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE557FC26FCD2FC3A" box="[632,793,907,933]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
lies in the relative length of the minor cheliped fingers, which are about twice as long as the palm in the 
<typeStatus id="7D340DD94B52FF9EE59FFC02FCD9FC56" box="[688,786,943,969]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="658FC8F84B52FF9EE419FC02FC73FC57" authorityName="Kazmi" authorityYear="1974" box="[822,952,943,969]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirmiziae">
<emphasis id="90FB6F694B52FF9EE419FC02FC73FC57" box="[822,952,943,969]" italics="true" pageId="22" pageNumber="23">A. tirmiziae</emphasis>
</taxonomicName>
(
<bibRefCitation id="C61ECE8A4B52FF9EE4EAFC02FB9FFC56" author="Kazmi, M. A." box="[965,1108,943,969]" pageId="22" pageNumber="23" pagination="170 - 174" refId="ref72166" refString="Kazmi, M. A. (1974) A new species of the genus Alpheus from the Bay of Bengal (Decapoda, Caridea). Crustaceana, 27, 170 - 174. https: // doi. org / 10.1163 / 156854074 X 00389" type="journal article" year="1974">Kazmi 1974</bibRefCitation>
: fig. 4B) 
<emphasis id="90FB6F694B52FF9EE394FC02FB12FC57" box="[1211,1241,943,968]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
at most 1.6 times as long as the palm in De Man’s type specimen from Java and 
<specimenCount id="B48978F24B52FF9EE479FC7EFC27FC72" box="[854,1004,979,1005]" pageId="22" pageNumber="23" type="female">three females</specimenCount>
from 
<collectingCountry id="DA98F3EB4B52FF9EE31FFC7EFB5FFC72" box="[1072,1172,979,1005]" name="Thailand" pageId="22" pageNumber="23">Thailand</collectingCountry>
(
<figureCitation id="3AB4AFFE4B52FF9EE38CFC7EFACEFC72" box="[1187,1285,979,1005]" captionStart="FIGURE 1" captionStartId="4.[151,250,1452,1476]" captionTargetBox="[167,1423,198,1420]" captionTargetId="figure-21@4.[152,1435,181,1427]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus euphrosyne De Man, 1897: syntype, female (cl 46.0 mm) from Java Sea, Indonesia (after De Man 1897, specimen lost) [A–E]; male (cl indet., tl 34.0 mm) from Bangkok, Thailand (after De Man 1898) [F, G]; A—frontal region, dorsal view; B—female minor (left) cheliped, lateral view; C—second pereiopod, lateral view; D—third pereiopod, lateral view; E—sixth pleonite, telson and uropods, dorsal view; F—male minor (right) cheliped, dorsolateral view; G—same, minor chela, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7912294" httpUri="https://zenodo.org/record/7912294/files/figure.png" pageId="22" pageNumber="23">Figs. 1B</figureCitation>
, 
<figureCitation id="3AB4AFFE4B52FF9EE23DFC7EFAFCFC72" box="[1298,1335,979,1005]" captionStart="FIGURE 7" captionStartId="16.[151,250,1796,1820]" captionTargetBox="[151,1424,192,1761]" captionTargetId="figure-21@16.[151,1436,181,1772]" captionTargetPageId="16" captionText="FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7912310" httpUri="https://zenodo.org/record/7912310/files/figure.png" pageId="22" pageNumber="23">7H</figureCitation>
). 
<bibRefCitation id="C61ECE8A4B52FF9EE264FC7EFE93FB8E" author="Banner, D. M. &amp; Banner, A. H." pageId="22" pageNumber="23" pagination="215 - 257" refId="ref69308" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner (1978)</bibRefCitation>
noted some other differences between the 
<typeStatus id="7D340DD94B52FF9EE412FC5AFC54FB8E" box="[829,927,1015,1041]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="658FC8F84B52FF9EE4E9FC5AFB81FB8F" authorityName="Kazmi" authorityYear="1974" box="[966,1098,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirmiziae">
<emphasis id="90FB6F694B52FF9EE4E9FC5AFB81FB8F" box="[966,1098,1015,1041]" italics="true" pageId="22" pageNumber="23">A. tirmiziae</emphasis>
</taxonomicName>
and their material from 
<collectingCountry id="DA98F3EB4B52FF9EE271FC5AFF12FBAA" name="Hong Kong" pageId="22" pageNumber="23">Hong Kong</collectingCountry>
. However, the most important difference that these authors found is directly related to sexual dimorphism of the minor cheliped, which in 
<taxonomicName id="658FC8F84B52FF9EE52AFB92FD6EFBC6" authorityName="sensu De Man" authorityYear="1897" box="[517,677,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE52AFB92FD6EFBC6" box="[517,677,1087,1113]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
and all related species is strongly balaeniceps in males and simple in females. The minute pits present on the carapace and pleon of the 
<typeStatus id="7D340DD94B52FF9EE489FBCEFBC3FBE2" box="[934,1032,1123,1149]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="658FC8F84B52FF9EE31FFBCEFB7FFBE3" authorityName="Kazmi" authorityYear="1974" box="[1072,1204,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirmiziae">
<emphasis id="90FB6F694B52FF9EE31FFBCEFB7FFBE3" box="[1072,1204,1123,1149]" italics="true" pageId="22" pageNumber="23">A. tirmiziae</emphasis>
</taxonomicName>
were overlooked by 
<bibRefCitation id="C61ECE8A4B52FF9EE7B8FB2AFE64FB3E" author="Banner, D. M. &amp; Banner, A. H." box="[151,431,1159,1185]" pageId="22" pageNumber="23" pagination="215 - 257" refId="ref69308" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">Banner &amp; Banner (1978)</bibRefCitation>
; these pits are clearly present in the 
<typeStatus id="7D340DD94B52FF9EE46EFB2AFC51FB3E" box="[833,922,1159,1185]" pageId="22" pageNumber="23" type="neotype">neotype</typeStatus>
of 
<taxonomicName id="658FC8F84B52FF9EE491FB2AFB96FB3E" authorityName="sensu De Man" authorityYear="1897" box="[958,1117,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE491FB2AFB96FB3E" box="[958,1117,1159,1185]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
and several other specimens from 
<collectingCountry id="DA98F3EB4B52FF9EE7FCFB06FEFCFB5A" box="[211,311,1195,1221]" name="Thailand" pageId="22" pageNumber="23">Thailand</collectingCountry>
and 
<collectingCountry id="DA98F3EB4B52FF9EE644FB06FE2BFB5A" box="[363,480,1195,1221]" name="Singapore" pageId="22" pageNumber="23">Singapore</collectingCountry>
. The above presented evidence justifies the placement of 
<taxonomicName id="658FC8F84B52FF9EE373FB06FB15FB5B" authorityName="Kazmi" authorityYear="1974" box="[1116,1246,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirmiziae">
<emphasis id="90FB6F694B52FF9EE373FB06FB15FB5B" box="[1116,1246,1195,1221]" italics="true" pageId="22" pageNumber="23">A. tirmiziae</emphasis>
</taxonomicName>
in the synonymy of 
<taxonomicName id="658FC8F84B52FF9EE799FB62FE9EFB76" authorityName="sensu De Man" authorityYear="1897" box="[182,341,1231,1257]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE799FB62FE9EFB76" box="[182,341,1231,1257]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A230B37B4B52FF9EE7E8FB5EFC83F89A" blockId="22.[151,1437,150,2013]" pageId="22" pageNumber="23">
As already mentioned, many previous records of 
<taxonomicName id="658FC8F84B52FF9EE5C9FB5EFC4FFA92" authorityName="sensu De Man" authorityYear="1897" box="[742,900,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE5C9FB5EFC4FFA92" box="[742,900,1267,1293]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
do not correspond to 
<taxonomicName id="658FC8F84B52FF9EE35DFB5EFADBFA92" authorityName="sensu De Man" authorityYear="1897" box="[1138,1296,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE35DFB5EFADBFA92" box="[1138,1296,1267,1293]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
as redefined in the present study. For instance, material from 
<collectingCountry id="DA98F3EB4B52FF9EE5EEFABAFCEEFAAE" box="[705,805,1303,1329]" name="Thailand" pageId="22" pageNumber="23">Thailand</collectingCountry>
reported as 
<taxonomicName id="658FC8F84B52FF9EE49CFABAFB9FFAAE" authorityName="sensu De Man" authorityYear="1897" box="[947,1108,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE49CFABAFB9FFAAE" box="[947,1108,1303,1329]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
by 
<bibRefCitation id="C61ECE8A4B52FF9EE3AEFABAFA57FAAE" author="Banner, A. H. &amp; Banner, D. M." box="[1153,1436,1303,1329]" pageId="22" pageNumber="23" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966)</bibRefCitation>
is most likely 
<taxonomicName id="658FC8F84B52FF9EE611FA96FE24FACA" authorityName="De Man" authorityYear="1920" box="[318,495,1339,1365]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B52FF9EE611FA96FE24FACA" box="[318,495,1339,1365]" italics="true" pageId="22" pageNumber="23">A. eurydactylus</emphasis>
</taxonomicName>
or 
<taxonomicName id="658FC8F84B52FF9EE535FA96FDBFFACA" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[538,628,1339,1365]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="takla" status="sp. nov.">
<emphasis id="90FB6F694B52FF9EE535FA96FDBFFACA" box="[538,628,1339,1365]" italics="true" pageId="22" pageNumber="23">A. takla</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B52FF9EE551FA96FD11FACA" bold="true" box="[638,730,1339,1365]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="8BC8D2124B52FF9EE551FA96FD11FACA" box="[638,730,1339,1365]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see below). Some Australian specimens reported by 
<bibRefCitation id="C61ECE8A4B52FF9EE264FA96FEABFAE6" author="Banner, D. M. &amp; Banner, A. H." pageId="22" pageNumber="23" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982)</bibRefCitation>
as 
<taxonomicName id="658FC8F84B52FF9EE6A0FAF2FD75FAE6" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[399,702,1375,1401]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">
<emphasis id="90FB6F694B52FF9EE6A0FAF2FD75FAE6" box="[399,702,1375,1401]" italics="true" pageId="22" pageNumber="23">A. euphrosyne euphrosyne</emphasis>
</taxonomicName>
were reexamined and assigned to one of the below described new species. Most records of 
<taxonomicName id="658FC8F84B52FF9EE6DDFA2EFD5FFA02" authorityName="sensu De Man" authorityYear="1897" box="[498,660,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE6DDFA2EFD5FFA02" box="[498,660,1411,1437]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
(sometimes as 
<taxonomicName id="658FC8F84B52FF9EE464FA2EFBBCFA02" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[843,1143,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">
<emphasis id="90FB6F694B52FF9EE464FA2EFBBCFA02" box="[843,1143,1411,1437]" italics="true" pageId="22" pageNumber="23">A. euphrosyne euphrosyne</emphasis>
</taxonomicName>
) from 
<collectingCountry id="DA98F3EB4B52FF9EE3E5FA2EFACFFA02" box="[1226,1284,1411,1437]" name="India" pageId="22" pageNumber="23">India</collectingCountry>
appear to be correct, especially those accompanied by a description of the colour pattern or colour photographs (
<bibRefCitation id="C61ECE8A4B52FF9EE3D6FA0AFA5CFA5E" author="Thomas, M. M." box="[1273,1431,1447,1473]" pageId="22" pageNumber="23" pagination="666 - 669" refId="ref74643" refString="Thomas, M. M. (1976) New records of four alpheid shrimp from the Indian waters. Journal of the Marine Biological Association of India, 18, 666 - 669." type="journal article" year="1976">Thomas 1976</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B52FF9EE7B8FA66FE1CFA7A" author="Harikrishnan, M. &amp; Kurup, B. M." box="[151,471,1483,1509]" pageId="22" pageNumber="23" pagination="1 - 280" refId="ref71572" refString="Harikrishnan, M. &amp; Kurup, B. M. (2008) Potential of snapping shrimps as ornamentals. In: Kurup, B. M., Boopendranth, M. R., Ravindran, K., Sairabanu &amp; Nair, A. G. (Eds.), Ornamental fish, breeding, farming and trade. Department of Fisheries, Government of Kerala, Kerala, pp. 1 - 280." type="book chapter" year="2008">Harikrishnan &amp; Kurup 2008</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B52FF9EE6CDFA66FD3DFA7A" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." box="[482,758,1483,1509]" pageId="22" pageNumber="23" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B52FF9EE554FA66FD66FA7A" box="[635,685,1483,1509]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. 2010
</bibRefCitation>
; 
<bibRefCitation id="C61ECE8A4B52FF9EE42EFA66FC0BFA7A" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[769,960,1483,1509]" pageId="22" pageNumber="23" pagination="1695 - 1698" refId="ref73799" refString="Paras, N. J., Purushothaman, P., Chinnadurai, S., Renjith R. K., Remesan, M. P., Baiju, M. V. &amp; Edwin, L. (2019) Occurrence of Alpheus euphrosyne de Man, 1897 (Crustacea: Decapoda: Alpheidae) from the south-eastern Arabian Sea, India. Indian Journal of Geo Marine Sciences, 48, 1695 - 1698." type="journal article" year="2019">
Paras 
<emphasis id="90FB6F694B52FF9EE46AFA66FCB6FA7A" box="[837,893,1483,1509]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2019
</bibRefCitation>
). The “new record” of 
<taxonomicName id="658FC8F84B52FF9EE3EFFA66FA94FA7A" authorityName="sensu De Man" authorityYear="1897" box="[1216,1375,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE3EFFA66FA94FA7A" box="[1216,1375,1483,1509]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
from 
<collectingCountry id="DA98F3EB4B52FF9EE7B8FA42FF10F996" box="[151,219,1519,1545]" name="India" pageId="22" pageNumber="23">Kochi</collectingCountry>
by 
<bibRefCitation id="C61ECE8A4B52FF9EE629FA42FE1EF996" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[262,469,1519,1545]" pageId="22" pageNumber="23" pagination="1695 - 1698" refId="ref73799" refString="Paras, N. J., Purushothaman, P., Chinnadurai, S., Renjith R. K., Remesan, M. P., Baiju, M. V. &amp; Edwin, L. (2019) Occurrence of Alpheus euphrosyne de Man, 1897 (Crustacea: Decapoda: Alpheidae) from the south-eastern Arabian Sea, India. Indian Journal of Geo Marine Sciences, 48, 1695 - 1698." type="journal article" year="2019">
Paras 
<emphasis id="90FB6F694B52FF9EE665FA42FEB7F996" box="[330,380,1519,1545]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. (2019)
</bibRefCitation>
was not new as the species was previously reported from this area by 
<bibRefCitation id="C61ECE8A4B52FF9EE3C2FA42FEFBF9B2" author="Harikrishnan, M. &amp; Kurup, B. M." pageId="22" pageNumber="23" pagination="1 - 280" refId="ref71572" refString="Harikrishnan, M. &amp; Kurup, B. M. (2008) Potential of snapping shrimps as ornamentals. In: Kurup, B. M., Boopendranth, M. R., Ravindran, K., Sairabanu &amp; Nair, A. G. (Eds.), Ornamental fish, breeding, farming and trade. Department of Fisheries, Government of Kerala, Kerala, pp. 1 - 280." type="book chapter" year="2008">Harikrishnan &amp; Kurup (2008)</bibRefCitation>
and 
<bibRefCitation id="C61ECE8A4B52FF9EE64AF9BEFD42F9B2" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." box="[357,649,1555,1581]" pageId="22" pageNumber="23" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B52FF9EE6D2F9BEFDE5F9B2" box="[509,558,1555,1581]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. (2010)
</bibRefCitation>
. In addition, the description of the 
<collectingCountry id="DA98F3EB4B52FF9EE323F9BEFB9BF9B2" box="[1036,1104,1555,1581]" name="India" pageId="22" pageNumber="23">Kochi</collectingCountry>
material was largely adapted, in a simplified form, from 
<bibRefCitation id="C61ECE8A4B52FF9EE6E6F99AFD2EF9CE" author="Banner, D. M. &amp; Banner, A. H." box="[457,741,1591,1617]" pageId="22" pageNumber="23" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner (1982)</bibRefCitation>
and therefore does not refer to 
<taxonomicName id="658FC8F84B52FF9EE37FF99AFF2EF9EA" authority="sensu De Man (1897)" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE37FF99AFAFCF9CF" box="[1104,1335,1591,1617]" italics="true" pageId="22" pageNumber="23">A. euphrosyne sensu</emphasis>
<bibRefCitation id="C61ECE8A4B52FF9EE26FF99AFF2EF9EA" author="De Man, J. G." pageId="22" pageNumber="23" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man (1897)</bibRefCitation>
</taxonomicName>
. However, the photographs provided by 
<bibRefCitation id="C61ECE8A4B52FF9EE5EFF9F6FC5CF9EA" author="Paras, N. J. &amp; Purushothaman, P. &amp; Chinnadurai, S. &amp; Renjith R. K. &amp; Remesan, M. P. &amp; Baiju, M. V. &amp; Edwin, L." box="[704,919,1627,1653]" pageId="22" pageNumber="23" pagination="1695 - 1698" refId="ref73799" refString="Paras, N. J., Purushothaman, P., Chinnadurai, S., Renjith R. K., Remesan, M. P., Baiju, M. V. &amp; Edwin, L. (2019) Occurrence of Alpheus euphrosyne de Man, 1897 (Crustacea: Decapoda: Alpheidae) from the south-eastern Arabian Sea, India. Indian Journal of Geo Marine Sciences, 48, 1695 - 1698." type="journal article" year="2019">
Paras 
<emphasis id="90FB6F694B52FF9EE428F9F6FCF0F9EA" box="[775,827,1627,1653]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. (2019)
</bibRefCitation>
indeed show male and female specimens of 
<taxonomicName id="658FC8F84B52FF9EE7B8F9D2FEF2F906" authorityName="sensu De Man" authorityYear="1897" box="[151,313,1663,1689]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE7B8F9D2FEF2F906" box="[151,313,1663,1689]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
, as redefined here. Nevertheless, since the presence of other species from the 
<emphasis id="90FB6F694B52FF9EE3E9F9D2FEF6F922" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="658FC8F84B52FF9EE3E9F9D2FAA3F906" authorityName="sensu De Man" authorityYear="1897" box="[1222,1384,1663,1689]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">A. euphrosyne</taxonomicName>
— 
<taxonomicName id="658FC8F84B52FF9EE2ABF9D2FEF6F922" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="microrhynchus">A. microrhynchus</taxonomicName>
</emphasis>
complex in 
<collectingCountry id="DA98F3EB4B52FF9EE6FCF90EFDC4F922" box="[467,527,1699,1725]" name="India" pageId="22" pageNumber="23">India</collectingCountry>
, possibly 
<taxonomicName id="658FC8F84B52FF9EE5AAF90EFCFCF922" authorityName="De Man" authorityYear="1920" box="[645,823,1699,1725]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B52FF9EE5AAF90EFCFCF922" box="[645,823,1699,1725]" italics="true" pageId="22" pageNumber="23">A. eurydactylus</emphasis>
</taxonomicName>
and/or 
<taxonomicName id="658FC8F84B52FF9EE4BCF90EFBD6F922" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[915,1053,1699,1725]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mangalis" status="sp. nov.">
<emphasis id="90FB6F694B52FF9EE4BCF90EFBD6F922" box="[915,1053,1699,1725]" italics="true" pageId="22" pageNumber="23">A. mangalis</emphasis>
</taxonomicName>
<emphasis id="90FB6F694B52FF9EE308F90EFB48F922" bold="true" box="[1063,1155,1699,1725]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="8BC8D2124B52FF9EE308F90EFB48F922" box="[1063,1155,1699,1725]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is extremely likely (see below and 
<figureCitation id="3AB4AFFE4B52FF9EE635F96AFEBAF97E" box="[282,369,1734,1761]" captionStart="FIGURE 53" captionStartId="107.[151,250,1859,1883]" captionTargetBox="[239,1349,192,1835]" captionTargetId="figure-15@107.[239,1349,181,1835]" captionTargetPageId="107" captionText="FIGURE 53. Alpheus euphrosyne De Man, 1897 complex in the northern and western Indian Ocean: A—Alpheus cf. mangalis sp. nov. (upper row) and A. euphrosyne (lower row) from an estuary near Kochi, India; B, C—Alpheus cf. eurydactylus De Man, 1920, ovigerious female (cl indet.) from Kochi, India (RMNH.Crus.D.49749), lateral view (B) and pleon and tail fan, dorsal view; D, E—Alpheus sp. euphrosyne complex (?), female photographed in situ in a mangrove at Watamu, Kenya. Photographs courtesy of Mahadevan Harikrishnan (A), Juan C. Miquel (B, C) and Dawn Goebbels (D, E)." figureDoi="http://doi.org/10.5281/zenodo.7912459" httpUri="https://zenodo.org/record/7912459/files/figure.png" pageId="22" pageNumber="23">Fig. 53</figureCitation>
), all biological and ecological data on 
<taxonomicName id="658FC8F84B52FF9EE411F96AFC2AF97E" authorityName="sensu De Man" authorityYear="1897" box="[830,993,1735,1761]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE411F96AFC2AF97E" box="[830,993,1735,1761]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
in 
<bibRefCitation id="C61ECE8A4B52FF9EE322F96AFAA2F97E" author="Harikrishnan, M. &amp; Kurup, B. M." box="[1037,1385,1735,1761]" pageId="22" pageNumber="23" pagination="1 - 280" refId="ref71572" refString="Harikrishnan, M. &amp; Kurup, B. M. (2008) Potential of snapping shrimps as ornamentals. In: Kurup, B. M., Boopendranth, M. R., Ravindran, K., Sairabanu &amp; Nair, A. G. (Eds.), Ornamental fish, breeding, farming and trade. Department of Fisheries, Government of Kerala, Kerala, pp. 1 - 280." type="book chapter" year="2008">Harikrishnan &amp; Kurup (2008)</bibRefCitation>
and 
<bibRefCitation id="C61ECE8A4B52FF9EE7B8F946FE71F89A" author="Harikrishnan, M. &amp; Unnikrishnan, U. &amp; Smija Maju, M. &amp; Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B." box="[151,442,1771,1797]" pageId="22" pageNumber="23" pagination="195 - 202" refId="ref71652" refString="Harikrishnan, M., Unnikrishnan, U., Smija Maju, M., Reena Greeshma, A. R. &amp; Madhusoodana Kurup, B. (2010) Size at sexual maturity, egg number and reproductive output of the snapping shrimp Alpheus euphrosyne euphrosyne De Man, 1897. Invertebrate Reproduction &amp; Development, 54, 195 - 202. https: // doi. org / 10.1080 / 07924259.2010.9652333" type="journal article" year="2010">
Harikrishnan 
<emphasis id="90FB6F694B52FF9EE61FF946FEA2F89A" box="[304,361,1771,1797]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
(2010)
</bibRefCitation>
must be treated with some caution.
</paragraph>
<paragraph id="A230B37B4B52FF9FE7E8F8A2FC84FF66" blockId="22.[151,1437,150,2013]" lastBlockId="23.[151,1437,151,249]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
The identity of material from the Red Sea, Persian Gulf and East Africa reported as 
<taxonomicName id="658FC8F84B52FF9EE359F8A2FA57F8B6" authorityName="sensu Banner &amp; Banner" authorityYear="1982" box="[1142,1436,1807,1833]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="euphrosyne" subSpecies="euphrosyne">
<emphasis id="90FB6F694B52FF9EE359F8A2FA57F8B6" box="[1142,1436,1807,1833]" italics="true" pageId="22" pageNumber="23">A. euphrosyne euphrosyne</emphasis>
</taxonomicName>
by 
<bibRefCitation id="C61ECE8A4B52FF9EE794F89EFE06F8D2" author="Banner, D. M. &amp; Banner, A. H." box="[187,461,1843,1869]" pageId="22" pageNumber="23" pagination="1 - 99" refId="ref69349" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner (1981</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B52FF9EE6F6F89EFDDFF8D2" author="Banner, A. H. &amp; Banner, D. M." box="[473,532,1843,1869]" pageId="22" pageNumber="23" pagination="1 - 164" refId="ref69267" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">1983</bibRefCitation>
), without illustrations or morphological details, remains unknown. Based on the collection data for the Kenyan specimen in 
<bibRefCitation id="C61ECE8A4B52FF9EE5BEF8FAFC64F8EE" author="Banner, A. H. &amp; Banner, D. M." box="[657,943,1879,1905]" pageId="22" pageNumber="23" pagination="1 - 164" refId="ref69267" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner (1983)</bibRefCitation>
and the concept of 
<taxonomicName id="658FC8F84B52FF9EE3B7F8FAFAF1F8EE" authorityName="sensu De Man" authorityYear="1897" box="[1176,1338,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE3B7F8FAFAF1F8EE" box="[1176,1338,1879,1905]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
adopted by 
<bibRefCitation id="C61ECE8A4B52FF9EE793F8D6FE1AF80A" author="Banner, A. H. &amp; Banner, D. M." box="[188,465,1915,1941]" pageId="22" pageNumber="23" pagination="1 - 168" refId="ref69176" refString="Banner, A. H. &amp; Banner, D. M. (1966) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner (1966</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B52FF9EE6F2F8D6FDD3F80A" author="Banner, D. M. &amp; Banner, A. H." box="[477,536,1915,1941]" pageId="22" pageNumber="23" pagination="215 - 257" refId="ref69308" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14, 215 - 257." type="journal article" year="1978">1978</bibRefCitation>
, 
<bibRefCitation id="C61ECE8A4B52FF9EE50AF8D6FDAAF80A" author="Banner, D. M. &amp; Banner, A. H." box="[549,609,1915,1941]" pageId="22" pageNumber="23" pagination="1 - 357" refId="ref69391" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
), the material from the western Indian Ocean is not 
<taxonomicName id="658FC8F84B52FF9EE39AF8D6FE86F826" authority="sensu De Man (1897)" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE39AF8D6FA57F80B" box="[1205,1436,1915,1941]" italics="true" pageId="22" pageNumber="23">A. euphrosyne sensu</emphasis>
<bibRefCitation id="C61ECE8A4B52FF9EE7B8F832FE86F826" author="De Man, J. G." box="[151,333,1951,1977]" pageId="22" pageNumber="23" pagination="725 - 790" refId="ref70820" refString="De Man, J. G. (1897) Bericht u ¨ ber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Ku ¨ sten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Fu ¨ nfter Theil. Z oologische Jahrbu ¨ cher. Abtheilung fu ¨ r Systematik, Geographie und Biologie der Thiere, 9, 725 - 790, pls. 34 - 37 (in volume 10)." type="journal article" year="1897">De Man (1897)</bibRefCitation>
</taxonomicName>
. The same is true for the material from 
<collectingCountry id="DA98F3EB4B52FF9EE409F832FCA2F826" box="[806,873,1951,1977]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
reported as 
<taxonomicName id="658FC8F84B52FF9EE4D2F832FB54F826" authorityName="sensu De Man" authorityYear="1897" box="[1021,1183,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B52FF9EE4D2F832FB54F826" box="[1021,1183,1951,1977]" italics="true" pageId="22" pageNumber="23">A. euphrosyne</emphasis>
</taxonomicName>
by 
<bibRefCitation id="C61ECE8A4B52FF9EE3FFF832FA53F826" author="Sha, Z. &amp; Wang, Y. &amp; Cui, D." box="[1232,1432,1951,1977]" pageId="22" pageNumber="23" refId="ref74259" refString="Sha, Z., Wang, Y. &amp; Cui, D. (2019) The Alpheidae from China seas. Crustacea: Decapoda: Caridea. Springer, Singapore, 323 pp. https: // doi. org / 10.1007 / 978 - 981 - 15 - 0648 - 2 _ 2" type="book" year="2019">
Sha 
<emphasis id="90FB6F694B52FF9EE22BF832FA8BF826" box="[1284,1344,1951,1977]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
(2019)
</bibRefCitation>
; their material ranged from 
<collectingRegion id="604B7D994B52FF9EE6E2F86EFDD5F842" box="[461,542,1987,2013]" country="China" name="Hainan" pageId="22" pageNumber="23">Hainan</collectingRegion>
to Qingdao and included Xisha Islands in the South-China Sea. However, the superficial description and crude illustrations accompanying the Chinese records are unusable for identification of (possibly more than one) species involved. The possibility that some of the Chinese material reported as 
<taxonomicName id="658FC8F84B53FF9FE2ABFF16FEDCFF66" authorityName="sensu De Man" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="90FB6F694B53FF9FE2ABFF16FEDCFF66" italics="true" pageId="23" pageNumber="24">A. euphrosyne</emphasis>
</taxonomicName>
may represent 
<taxonomicName id="658FC8F84B53FF9FE6ECFF72FDBAFF66" authorityName="De Man" authorityYear="1920" box="[451,625,223,249]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="eurydactylus">
<emphasis id="90FB6F694B53FF9FE6ECFF72FDBAFF66" box="[451,625,223,249]" italics="true" pageId="23" pageNumber="24">A. eurydactylus</emphasis>
</taxonomicName>
is discussed below.
</paragraph>
</subSubSection>
</treatment>
</document>