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<document id="285A93E9CE1EFBA929B915FAC4BFADC2" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A13475324FDACFBCCFC41FD43" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus rivinus Moret 2020, sp. nov." docType="treatment" docUuid="95AF0791-2EC1-42FD-862D-02F7BCFB1419" docUuidSource="ZooBank" docVersion="8" lastPageNumber="18" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="16" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="0B5D9115A84445410D69B6D90B1A9263" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F2930257241AC73BD997BFD0AF07F7EC">
<mods:title id="A70D9D276831897E4C1D017C68D1229D">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="0917C39FFE670154146BA9E5D8B300C5" type="personal">
<mods:role id="8E9E57B4888C518DD5A480D10B776BA1">
<mods:roleTerm id="84AADE0F5D8410F8545B0C87F1FE19A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B1F836B644FEFC6104BCC769095FA648">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="F006F8197116C94158FA78FAAE98A7F9" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="EF0BC72FA68BE6D58328B7403E10A8A0">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="A0B4DDBF3D4A5B54D8F58F9A2C63F9E8" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="B4A24E0144821363AAED586EBB2ACA5A" type="personal">
<mods:role id="290A79249E8BEA172C26B75AC9CA94CA">
<mods:roleTerm id="30DE9C775A3C6817FA8BF9B7876AFFBC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="986E4F3EE890B8ABF329F7A978A886C8">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="606D49650EB09BD24127DC0458B3D415" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="7EBCD1AE9DB82CAB015E55BDDAC58472">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="9371BFF60ADC2BC667A7E9BE363F2CB7" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E7F016BEFBBD1D2E0E60563FB4E1DB59">text</mods:typeOfResource>
<mods:relatedItem id="3D1FD517BC05F23DFBC03B0D2FCDFD75" type="host">
<mods:titleInfo id="C4F7C8825ABB278EF3F08459855D82AF">
<mods:title id="230BD48BDEC3379EE20467F996F957A5">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="76BB5C169B8C02F17D551F314B73D2EA">
<mods:date id="413036A22FE956C1D3D0E97567987CA6">2020</mods:date>
<mods:detail id="7060E20E22780B9D0D2F4EBCCBFDAD6E" type="pubDate">
<mods:number id="F7A76F91603C4D778E430A47CC2FDCE6">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="25BB00FC1B317F029EBE8101E0D71D35" type="volume">
<mods:number id="57138971A801D264D71B28D8EB7B2C74">646</mods:number>
</mods:detail>
<mods:extent id="BAF2EB2503322199E79FD0B49FD8B983" unit="page">
<mods:start id="A0C079E09760C915BC8A4AE76C16F60E">1</mods:start>
<mods:end id="2A5B604EB76747779EF7964166505394">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4F7D644A00932D5EB50044215B1F55BC">journal article</mods:classification>
<mods:identifier id="D5860CBD822A78B32F1D8430162CB859" type="CLB-Dataset">22092</mods:identifier>
<mods:identifier id="3E1C39CE18F4CEFB83E8F09A6306C3E1" type="DOI">10.5852/ejt.2020.646</mods:identifier>
<mods:identifier id="B685D64FE6952A8256AAEFF411A38F2F" type="GBIF-Dataset">c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b</mods:identifier>
<mods:identifier id="C0B567D12DD67672085C442C7F6419E0" type="Zenodo-Dep">3829682</mods:identifier>
<mods:identifier id="A65EE8B8263046227E0B281FC2799EA2" type="ZooBank">4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B</mods:identifier>
</mods:mods>
<treatment id="03B7726A13475324FDACFBCCFC41FD43" ID-DOI="http://doi.org/10.5281/zenodo.3848397" ID-GBIF-Taxon="164160412" ID-Zenodo-Dep="3848397" LSID="urn:lsid:zoobank.org:act:95AF0791-2EC1-42FD-862D-02F7BCFB1419" httpUri="http://treatment.plazi.org/id/03B7726A13475324FDACFBCCFC41FD43" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="C30490F71347533AFDACFBCCFB45FBF6" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BA1C37C1347533AFDACFBCCFC51FBD2" blockId="15.[368,1219,1078,1175]" box="[605,983,1078,1105]" pageId="15" pageNumber="16">
<heading id="D0E974101347533AFDACFBCCFC51FBD2" box="[605,983,1078,1105]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<taxonomicName id="4C1EB8FF1347533AFDACFBCCFCF3FBD2" ID-CoL="8MCD9" authority="Moret" authorityName="Moret" authorityYear="2020" box="[605,885,1078,1105]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="rivinus" status="sp. nov.">
<emphasis id="B96A1F6E1347533AFDACFBCCFCA0FBD2" bold="true" box="[605,806,1078,1105]" italics="true" pageId="15" pageNumber="16">Dyscolus rivinus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151347533AFC8DFBCDFC51FBD2" box="[892,983,1079,1105]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C1347533AFE81FBA0FB45FBF6" blockId="15.[368,1219,1078,1175]" box="[368,1219,1114,1141]" pageId="15" pageNumber="16">
<uri id="FF8FCF7E1347533AFE81FBA0FB45FBF6" box="[368,1219,1114,1141]" pageId="15" pageNumber="16">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A91347533AFD7EFBA0FB45FBF6" box="[655,1219,1114,1141]" pageId="15" pageNumber="16">95AF0791-2EC1-42FD-862D-02F7BCFB1419</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71347533AFD28FB87FCDDFB14" box="[729,859,1149,1175]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BA1C37C1347533AFD28FB87FCDDFB14" blockId="15.[368,1219,1078,1175]" box="[729,859,1149,1175]" pageId="15" pageNumber="16">
<figureCitation id="1325DFF91347533AFD28FB87FCDDFB14" box="[729,859,1149,1175]" captionStart="Figs 7–12. 7–8" captionStartId="16.[189,241,1890,1916]" captionTargetBox="[279,1304,274,1828]" captionTargetId="figure@16.[264,1324,262,1850]" captionTargetPageId="16" captionText="Figs 7–12. 7–8. Dyscolus aquator Moret sp. nov. 7. Habitus of the male holotype (QCAZ). 8. Aedeagus, median lobe in lateral view. 9–10. D. incommunis Moret sp. nov. 9. Aedeagus, median lobe in lateral view. 10. Habitus of the male holotype (QCAZ). 11–12. D. rivinus Moret sp. nov. 11. Apex of the elytra. 12. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829688" httpUri="https://zenodo.org/record/3829688/files/figure.png" pageId="15" pageNumber="16">Figs 11–12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71347533AFF4CFB45FDD0FA84" box="[189,598,1215,1287]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BA1C37C1347533AFF4CFB45FEC2FB5A" blockId="15.[189,324,1215,1241]" box="[189,324,1215,1241]" pageId="15" pageNumber="16">
<heading id="D0E974101347533AFF4CFB45FEC2FB5A" bold="true" box="[189,324,1215,1241]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B96A1F6E1347533AFF4CFB45FEC2FB5A" bold="true" box="[189,324,1215,1241]" pageId="15" pageNumber="16">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1347533AFF4CFB17FDD0FA84" blockId="15.[189,598,1261,1287]" box="[189,598,1261,1287]" pageId="15" pageNumber="16">Latin adjective meaning ‘riparian’.</paragraph>
</subSubSection>
<subSubSection id="C30490F713475324FF4CFAD5FC41FD43" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BA1C37C1347533AFF4CFAD5FEEAFACB" blockId="15.[189,364,1326,1353]" box="[189,364,1326,1353]" pageId="15" pageNumber="16">
<heading id="D0E974101347533AFF4CFAD5FEEAFACB" bold="true" box="[189,364,1326,1353]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B96A1F6E1347533AFF4CFAD5FEEAFACB" bold="true" box="[189,364,1326,1353]" pageId="15" pageNumber="16">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1347533AFF1BFAA7FEDAFAF4" blockId="15.[189,1282,1373,1435]" box="[234,348,1373,1399]" pageId="15" pageNumber="16">
<emphasis id="B96A1F6E1347533AFF1BFAA7FEDAFAF4" bold="true" box="[234,348,1373,1399]" pageId="15" pageNumber="16">
<typeStatus id="54A57DDE1347533AFF1BFAA7FEDAFAF4" box="[234,348,1373,1399]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C1347533AFF4CFA7AFA84FA19" blockId="15.[189,1282,1373,1435]" box="[189,1282,1406,1435]" pageId="15" pageNumber="16">
<materialsCitation id="3B76C9211347533AFF4CFA7AFA84FA19" ID-GBIF-Occurrence="2620303339" box="[189,1282,1406,1435]" collectingDate="1998-07-07" collectionCode="QCAZ" collectorName="I. Tapia" country="Ecuador" elevation="2000" location="Otonga" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Cotopaxi Province" typeStatus="holotype">
<collectingCountry id="F30983EC1347533AFF4CFA7AFECDFA19" box="[189,331,1408,1434]" name="Ecuador" pageId="15" pageNumber="16">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F51347533AFE95FA84FEFDFA19" box="[356,379,1406,1434]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E1347533AFE78FA7AFDE1FA19" box="[393,615,1408,1434]" country="Ecuador" name="Cotopaxi" pageId="15" pageNumber="16">Cotopaxi Province</collectingRegion>
,
<location id="8EC195A71347533AFD82FA7AFD4BFA19" LSID="urn:lsid:plazi:treatment:03B7726A13475324FDACFBCCFC41FD43:8EC195A71347533AFD82FA7AFD4BFA19" box="[627,717,1408,1434]" country="Ecuador" name="Otonga" pageId="15" pageNumber="16" stateProvince="Cotopaxi Province">Otonga</location>
;
<elevation id="0033244F1347533AFD28FA7AFCF4FA19" box="[729,882,1408,1435]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="m" value="2000.0">
<quantity id="4CE66E991347533AFD28FA7AFCB4FA18" box="[729,818,1408,1435]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1347533AFC8EFA7BFB82FA19" box="[895,1028,1408,1435]" pageId="15" pageNumber="16" value="1998-07-07">
<collectingDate id="EFE41C541347533AFC8EFA7BFB82FA19" box="[895,1028,1408,1435]" pageId="15" pageNumber="16" value="1998-07-07">7 Jul. 1998</collectingDate>
</date>
;
<collectorName id="26EBA6AA1347533AFBE1FA7BFBEFFA19" box="[1040,1129,1408,1435]" pageId="15" pageNumber="16">I. Tapia</collectorName>
leg.;
<collectionCode id="ED0F5BB91347533AFB5AFA7AFB79FA19" box="[1195,1279,1408,1434]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="15" pageNumber="16" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C1347533AFF1BFA38FE43FA5F" blockId="15.[189,1398,1472,1571]" box="[234,453,1472,1500]" pageId="15" pageNumber="16">
<emphasis id="B96A1F6E1347533AFF1BFA38FEE1FA5F" bold="true" box="[234,359,1474,1500]" pageId="15" pageNumber="16">
<typeStatus id="54A57DDE1347533AFF1BFA38FEE1FA5F" box="[234,359,1474,1500]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
(3 ♂♂)
</paragraph>
<paragraph id="8BA1C37C1347533AFF35FA1FFC51F9A1" blockId="15.[189,1398,1472,1571]" pageId="15" pageNumber="16">
<materialsCitation id="3B76C9211347533AFF35FA1FFB29FA7C" ID-GBIF-Occurrence="2620303305" box="[196,1199,1507,1535]" collectingDate="1998-07-07" collectionCode="QCAZ" collectorName="I. Tapia" country="Ecuador" elevation="2000" location="Cotopaxi Province" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Cotopaxi Province" typeStatus="paratype">
<collectingCountry id="F30983EC1347533AFF35FA1FFED4FA7C" box="[196,338,1509,1535]" name="Ecuador" pageId="15" pageNumber="16">ECUADOR</collectingCountry>
–
<emphasis id="B96A1F6E1347533AFE9EFA1FFDD0FA7C" bold="true" box="[367,598,1509,1535]" pageId="15" pageNumber="16">
<collectingRegion id="49DA0D9E1347533AFE9EFA1FFDD0FA7C" box="[367,598,1509,1535]" country="Ecuador" name="Cotopaxi" pageId="15" pageNumber="16">Cotopaxi Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1808F51347533AFD9EFA1FFD1DFA7C" box="[623,667,1507,1535]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB91347533AFBAEFA1FFB29FA7C" box="[1119,1199,1509,1535]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="15" pageNumber="16" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211347533AFB36FA1FFC52F9A1" ID-GBIF-Occurrence="2620303383" collectedFrom="berge de ruisseau" collectingDate="2001-07-03" collectingDateMax="2001-07-05" collectingDateMin="2001-07-03" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" elevation="1950" location="Otonga" pageId="15" pageNumber="16" specimenCount="2" specimenCount-male="2" stateProvince="Cotopaxi" typeStatus="paratype">
<specimenCount id="9D1808F51347533AFB36FA1FFA8CFA7C" box="[1223,1290,1507,1535]" pageId="15" pageNumber="16" type="male">2 ♂♂</specimenCount>
;
<location id="8EC195A71347533AFAE9FA1FFAF4FA7C" LSID="urn:lsid:plazi:treatment:03B7726A13475324FDACFBCCFC41FD43:8EC195A71347533AFAE9FA1FFAF4FA7C" box="[1304,1394,1509,1535]" country="Ecuador" name="Otonga" pageId="15" pageNumber="16" stateProvince="Cotopaxi">Otonga</location>
;
<elevation id="0033244F1347533AFF4CF9F2FED0F9A1" box="[189,342,1544,1571]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="15" pageNumber="16" unit="m" value="1950.0">
<quantity id="4CE66E991347533AFF4CF9F2FE90F9A0" box="[189,278,1544,1571]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="15" pageNumber="16" unit="m" value="1950.0">1950 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1347533AFE92F9F2FD83F9A1" box="[355,517,1544,1571]" pageId="15" pageNumber="16" value="2001-07-03" valueMax="2001-07-05" valueMin="2001-07-03">
<collectingDate id="EFE41C541347533AFE92F9F2FD83F9A1" box="[355,517,1544,1571]" pageId="15" pageNumber="16" value="2001-07-03" valueMax="2001-07-05" valueMin="2001-07-03">3–5 Jul. 2001</collectingDate>
</date>
;
<collectorName id="26EBA6AA1347533AFDE3F9F3FDF3F9A0" box="[530,629,1545,1571]" pageId="15" pageNumber="16">P. Moret</collectorName>
leg.;
<collectedFrom id="380280811347533AFD46F9F2FC00F9A1" box="[695,902,1544,1570]" pageId="15" pageNumber="16">berge de ruisseau</collectedFrom>
;
<collectionCode id="ED0F5BB91347533AFC63F9F2FC52F9A1" box="[914,980,1544,1570]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="15" pageNumber="16">CPM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1C37C1347533AFF4CF9B3FE42F9E0" blockId="15.[189,452,1609,1635]" box="[189,452,1609,1635]" pageId="15" pageNumber="16">
<heading id="D0E974101347533AFF4CF9B3FE42F9E0" bold="true" box="[189,452,1609,1635]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B96A1F6E1347533AFF4CF9B3FE42F9E0" bold="true" box="[189,452,1609,1635]" pageId="15" pageNumber="16">Diagnostic escription</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1347533AFF4CF982FCBDF8E6" blockId="15.[189,1399,1656,1894]" pageId="15" pageNumber="16">
Wingless. Body length: 10.0–
<quantity id="4CE66E991347533AFDEDF982FD01F910" box="[540,647,1656,1683]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="15" pageNumber="16" unit="mm" value="11.7">11.7 mm</quantity>
. Very similar externally to
<taxonomicName id="4C1EB8FF1347533AFC39F983FB4DF910" authority="Moret" authorityName="Moret" authorityYear="2020" box="[968,1227,1657,1683]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="incommunis" status="sp. nov.">
<emphasis id="B96A1F6E1347533AFC39F983FBFDF911" box="[968,1147,1657,1682]" italics="true" pageId="15" pageNumber="16">D. incommunis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151347533AFB22F983FAB7F910" box="[1235,1329,1657,1683]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, from which it differs in the following features. Base of the pronotum narrower, lateral margins less expanded near base, hind angles more obtuse and more rounded. Apex of elytra ended with a small but protruding and acute tooth, preceded externally by a short sinuation (
<figureCitation id="1325DFF91347533AFC62F918FC68F97F" box="[915,1006,1762,1788]" captionStart="Figs 7–12. 7–8" captionStartId="16.[189,241,1890,1916]" captionTargetBox="[279,1304,274,1828]" captionTargetId="figure@16.[264,1324,262,1850]" captionTargetPageId="16" captionText="Figs 7–12. 7–8. Dyscolus aquator Moret sp. nov. 7. Habitus of the male holotype (QCAZ). 8. Aedeagus, median lobe in lateral view. 9–10. D. incommunis Moret sp. nov. 9. Aedeagus, median lobe in lateral view. 10. Habitus of the male holotype (QCAZ). 11–12. D. rivinus Moret sp. nov. 11. Apex of the elytra. 12. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829688" httpUri="https://zenodo.org/record/3829688/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
). Fourth metatarsomere without subapical setae. Male genitalia (
<figureCitation id="1325DFF91347533AFDCCF8FFFD12F89C" box="[573,660,1797,1823]" captionStart="Figs 7–12. 7–8" captionStartId="16.[189,241,1890,1916]" captionTargetBox="[279,1304,274,1828]" captionTargetId="figure@16.[264,1324,262,1850]" captionTargetPageId="16" captionText="Figs 7–12. 7–8. Dyscolus aquator Moret sp. nov. 7. Habitus of the male holotype (QCAZ). 8. Aedeagus, median lobe in lateral view. 9–10. D. incommunis Moret sp. nov. 9. Aedeagus, median lobe in lateral view. 10. Habitus of the male holotype (QCAZ). 11–12. D. rivinus Moret sp. nov. 11. Apex of the elytra. 12. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829688" httpUri="https://zenodo.org/record/3829688/files/figure.png" pageId="15" pageNumber="16">Fig. 12</figureCitation>
) with apex of the median lobe shorter, less acute, not sinuate in lateral view, endophallus unarmed, without a dense squamose area near apex, at most with a single feebly sclerotized squama. Female genitalia unknown.
</paragraph>
<caption id="DF6193F413585325FF4CF898FD29F865" ID-DOI="http://doi.org/10.5281/zenodo.3829688" ID-Zenodo-Dep="3829688" httpUri="https://zenodo.org/record/3829688/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,241,1890,1916]" targetBox="[279,1304,274,1828]" targetPageId="16">
<paragraph id="8BA1C37C13585325FF4CF898FD29F865" blockId="16.[189,1399,1890,2023]" pageId="16" pageNumber="17">
<emphasis id="B96A1F6E13585325FF4CF898FEEAF8FE" bold="true" box="[189,364,1890,1917]" pageId="16" pageNumber="17">Figs 7–12. 7–8</emphasis>
.
<taxonomicName id="4C1EB8FF13585325FE8BF899FD11F8FE" authority="Moret" authorityName="Moret" authorityYear="2020" box="[378,663,1891,1917]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aquator" status="sp. nov.">
<emphasis id="B96A1F6E13585325FE8BF899FDCEF8FE" box="[378,584,1891,1917]" italics="true" pageId="16" pageNumber="17">Dyscolus aquator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513585325FD6CF899FD70F8FE" box="[669,758,1891,1917]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96A1F6E13585325FD0DF899FC8DF8FE" bold="true" box="[764,779,1891,1917]" pageId="16" pageNumber="17">7</emphasis>
. Habitus of the male holotype (QCAZ).
<emphasis id="B96A1F6E13585325FB10F899FB76F8FE" bold="true" box="[1249,1264,1891,1917]" pageId="16" pageNumber="17">8</emphasis>
. Aedeagus, median lobe in lateral view.
<emphasis id="B96A1F6E13585325FDE3F87CFDCBF823" bold="true" box="[530,589,1926,1952]" pageId="16" pageNumber="17">9–10</emphasis>
.
<taxonomicName id="4C1EB8FF13585325FDACF87DFCE4F823" authority="Moret" authorityName="Moret" authorityYear="2020" box="[605,866,1926,1952]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="incommunis" status="sp. nov.">
<emphasis id="B96A1F6E13585325FDACF87DFC97F823" box="[605,785,1927,1952]" italics="true" pageId="16" pageNumber="17">D. incommunis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513585325FC9AF87CFC41F823" box="[875,967,1926,1952]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96A1F6E13585325FC21F87CFC59F823" bold="true" box="[976,991,1926,1952]" pageId="16" pageNumber="17">9</emphasis>
. Aedeagus, median lobe in lateral view.
<emphasis id="B96A1F6E13585325FEF3F853FE99F840" bold="true" box="[258,287,1961,1987]" pageId="16" pageNumber="17">10</emphasis>
. Habitus of the male holotype (QCAZ).
<emphasis id="B96A1F6E13585325FD07F853FCB8F840" bold="true" box="[758,830,1961,1987]" pageId="16" pageNumber="17">11–12</emphasis>
.
<taxonomicName id="4C1EB8FF13585325FCBDF850FB89F847" authority="Moret" authorityName="Moret" authorityYear="2020" box="[844,1039,1962,1988]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rivinus" status="sp. nov.">
<emphasis id="B96A1F6E13585325FCBDF850FC46F840" box="[844,960,1962,1987]" italics="true" pageId="16" pageNumber="17">D. rivinus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513585325FBE4F850FBE9F847" box="[1045,1135,1962,1988]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96A1F6E13585325FB84F853FB09F840" bold="true" box="[1141,1167,1961,1987]" pageId="16" pageNumber="17">11</emphasis>
. Apex of the elytra.
<emphasis id="B96A1F6E13585325FF4CF836FF5CF865" bold="true" box="[189,218,1996,2022]" pageId="16" pageNumber="17">12</emphasis>
. Aedeagus, median lobe in lateral view.
</paragraph>
</caption>
<paragraph id="8BA1C37C13595324FF4CFEF1FEE5FEA6" blockId="17.[189,355,267,293]" box="[189,355,267,293]" pageId="17" pageNumber="18">
<heading id="D0E9741013595324FF4CFEF1FEE5FEA6" bold="true" box="[189,355,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B96A1F6E13595324FF4CFEF1FEE5FEA6" bold="true" box="[189,355,267,293]" pageId="17" pageNumber="18">Comparisons</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13595324FF4CFEC3FD9BFE19" blockId="17.[189,1399,313,410]" pageId="17" pageNumber="18">
Closely related to
<taxonomicName id="4C1EB8FF13595324FE6EFEC0FD2EFED7" authority="Moret" authorityName="Moret" authorityYear="2020" box="[415,680,314,340]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="incommunis" status="sp. nov.">
<emphasis id="B96A1F6E13595324FE6EFEC0FDD3FED0" box="[415,597,314,339]" italics="true" pageId="17" pageNumber="18">D. incommunis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513595324FD42FEC0FC92FED7" box="[691,788,314,340]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can be distinguished by a few but constant characters, especially the rounded hind angles of the pronotum, the apical tooth of the elytra and the unarmed endophallus.
</paragraph>
<paragraph id="8BA1C37C13595324FF4CFE3BFE9BFE58" blockId="17.[189,285,449,475]" box="[189,285,449,475]" pageId="17" pageNumber="18">
<heading id="D0E9741013595324FF4CFE3BFE9BFE58" bold="true" box="[189,285,449,475]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B96A1F6E13595324FF4CFE3BFE9BFE58" bold="true" box="[189,285,449,475]" pageId="17" pageNumber="18">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13595324FF4CFE0AFA94FDAD" blockId="17.[189,1398,496,558]" pageId="17" pageNumber="18">
Riparian in the montane forest on the Western slope of the
<taxonomicName id="4C1EB8FF13595324FC73FE0AFC56FD89" authorityName="Stal" authorityYear="1866" box="[898,976,496,522]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="54" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
, at around
<quantity id="4CE66E9913595324FBA7FE0AFB7CFD89" box="[1110,1274,496,522]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="17" pageNumber="18" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">1900–2000 m</quantity>
a.s.l. This hygrophile specialisation differs from the more mesic habitat of
<taxonomicName id="4C1EB8FF13595324FC41FDEEFB36FDAD" authority="Moret" authorityName="Moret" authorityYear="2020" box="[944,1200,532,558]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="incommunis" status="sp. nov.">
<emphasis id="B96A1F6E13595324FC41FDEEFBE7FDAE" box="[944,1121,532,557]" italics="true" pageId="17" pageNumber="18">D. incommunis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513595324FB49FDEEFA94FDAD" box="[1208,1298,532,558]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BA1C37C13595324FF4CFDAEFE6DFDED" blockId="17.[189,491,596,622]" box="[189,491,596,622]" pageId="17" pageNumber="18">
<heading id="D0E9741013595324FF4CFDAEFE6DFDED" bold="true" box="[189,491,596,622]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B96A1F6E13595324FF4CFDAEFE6DFDED" bold="true" box="[189,491,596,622]" pageId="17" pageNumber="18">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13595324FF4CFD79FC41FD43" blockId="17.[189,1399,643,705]" pageId="17" pageNumber="18">
Southern end of the Chocó biogeographic region in northwestern
<collectingCountry id="F30983EC13595324FC2CFD79FBC7FD1E" box="[989,1089,643,669]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
,
<quantity id="4CE66E9913595324FBA1FD79FB1BFD1E" box="[1104,1181,643,670]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="18" unit="km" value="75.0">75 km</quantity>
south of the
<typeStatus id="54A57DDE13595324FAB5FD7EFAF1FD1D" box="[1348,1399,644,670]" pageId="17" pageNumber="18">type</typeStatus>
locality of
<taxonomicName id="4C1EB8FF13595324FECDFD5DFDBBFD42" authority="Moret" authorityName="Moret" authorityYear="2020" box="[316,573,679,705]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="incommunis" status="sp. nov.">
<emphasis id="B96A1F6E13595324FECDFD5DFE68FD43" box="[316,494,679,704]" italics="true" pageId="17" pageNumber="18">D. incommunis</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513595324FDB5FD5DFD19FD42" box="[580,671,679,705]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
Probably microendemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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