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<document id="038FC23314C5D751EBB9C611CD8EBEAC" ID-CLB-Dataset="9608" ID-DOI="10.5252/zoosystema2020v42a28" ID-GBIF-Dataset="c2906488-8302-4ccb-b2cf-e9fe1e782ea5" ID-ISSN="1638-9387" ID-Zenodo-Dep="4265439" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1604967954946" checkinUser="felipe" docAuthor="Ayoro, Halamoussa Joëlle, Segniagbeto, Gabriel Hoinsoudé, Hema, Emmanuel Midibahaye, Penner, Johannes, Oueda, Adama, Dubois, Alain, Rödel, Mark-Oliver, Kabré, Gustave Boureima &amp; Ohler, Annemarie" docDate="2020" docId="038C87CCFFD5FF85FF38FF30FECAF9D7" docLanguage="en" docName="Zoosystema.42.28.547-582.pdf.imf" docOrigin="Zoosystema 42 (28)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Tomopterna milletihorsini" docType="treatment" docVersion="9" lastPageNumber="574" masterDocId="FFB5FFB4FFC8FF98FFDFFFE8FFE3FFE8" masterDocTitle="List of amphibian species (Vertebrata, Tetrapoda) of Burkina Faso" masterLastPageNumber="582" masterPageNumber="547" pageNumber="574" updateTime="1698846188204" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2736F403E7A2835872C5E00885A5621F">List of amphibian species (Vertebrata, Tetrapoda) of Burkina Faso</mods:title>
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<mods:namePart id="900FECD1A66ADFD24037A3159A4B11CC">Ayoro, Halamoussa Joëlle</mods:namePart>
<mods:affiliation id="D31E936BB4C7D54ECD785554A2B0C1A9">Laboratoire de Biologie et Ecologie Animales, UFR / SVT, Université Joseph Ki-Zerbo, 3 BP 7021 Ouagadougou 3 (Burkina Faso) halamoussa. joelle @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="BFDE5114893275247C9C22E7BA3EC5FD">Segniagbeto, Gabriel Hoinsoudé</mods:namePart>
<mods:affiliation id="8A598243FBF08B661AC17D8F1B9CE0EB">Laboratoire d’Écologie et d’Écotoxicologie, Faculté des Sciences, Université de Lomé, BP 1515, Lomé (Togo) gsegniagbeto @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="1B2ADE20EBE3B12660616AA61372F29E" type="email">gsegniagbeto@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EFFF9029302D6F8A63CF5010A318A801">Hema, Emmanuel Midibahaye</mods:namePart>
<mods:affiliation id="06C26219758F0C8908E339A954DDF4EA">Laboratoire de Biologie et Écologie Animales, UFR / SVT, Université Joseph Ki-Zerbo, 3 BP 7021 Ouagadougou 3 (Burkina Faso) hema. emmanuel @ yahoo. fr</mods:affiliation>
<mods:nameIdentifier id="E3303F883E4D795156A1FFB381593ECC" type="email">emmanuel@yahoo.fr</mods:nameIdentifier>
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<mods:namePart id="CED2EE767E239D789B444716F5DFE0B0">Penner, Johannes</mods:namePart>
<mods:affiliation id="194873BCA22EB2AFA0FF7372D6CE58E8">Chair of Wildlife Ecology &amp; Management, University of Freiburg, Tennenbacher Strasse 4, 79106 Freiburg (Germany) johannes. penner @ wildlife. uni-freiburg. de</mods:affiliation>
<mods:nameIdentifier id="8D944B8A4338439131BE897714D83C7B" type="email">penner@wildlife.uni</mods:nameIdentifier>
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<mods:namePart id="5EDE24EBA021F5A3D27CBBABD926A3FF">Oueda, Adama</mods:namePart>
<mods:affiliation id="4372D2C3D6A46637D2F73A0C1A584CCE">Laboratoire de Biologie et Écologie Animales, UFR / SVT, Université Joseph Ki-Zerbo, 3 BP 7021 Ouagadougou 3 (Burkina Faso) oueda 14 @ yahoo. fr</mods:affiliation>
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<mods:namePart id="F0567B55520BA975C9DF7BF924E05F30">Dubois, Alain</mods:namePart>
<mods:affiliation id="524A54B741077689981528409E447EAC">Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d'Histoire naturelle CNRS, Sorbonne Université, EPHE, UA, 57 rue Cuvier, CP 30, F- 75005, Paris (France) alain. dubois @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="AFEFDC3E665F405DEA64DDD512CC8E17" type="email">dubois@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="2D96812F81492A0C4787F5E63FB08B80">Rödel, Mark-Oliver</mods:namePart>
<mods:affiliation id="5CE8EFFEA7CEACA97A9DBEDEC87F90C4">Museum für Naturkunde - Leibniz Institute for Evolution and Biodiversity Science, Invalidenstr. 43, 10115 Berlin (Germany )) Mark-Oliver. Roedel @ mfn. berlin</mods:affiliation>
<mods:nameIdentifier id="E02CA6D6DAE784D275E13218E42B1624" type="email">oedel@mfn.berlin</mods:nameIdentifier>
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<mods:namePart id="9A873B6C7FC584340275B5BA350878A7">Kabré, Gustave Boureima</mods:namePart>
<mods:affiliation id="A3D7328FBF8572520C456641652E7818">Laboratoire de Biologie et Écologie Animales, UFR / SVT, Université Joseph Ki-Zerbo, 3 BP 7021 Ouagadougou 3 (Burkina Faso) gkabre 2 @ gmail. com</mods:affiliation>
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<mods:namePart id="D3925B60A1D41AA82364E4EB5F0C99AC">Ohler, Annemarie</mods:namePart>
<mods:affiliation id="16AD9A0164B6652AB616073C441243B7">Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d'Histoire naturelle CNRS, Sorbonne Université, EPHE, UA, 57 rue Cuvier, CP 30, F- 75005, Paris (France) annemarie. ohler @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="B1B505A247F9BC1DA0624E3AE3CEED8E" type="email">ohler@mnhn.fr</mods:nameIdentifier>
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<mods:typeOfResource id="C06ADDC68898432DF66CF242CA18DF2A">text</mods:typeOfResource>
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<mods:titleInfo id="C96F10FFA52A48F0E033A8DB16772012">
<mods:title id="55B77546DF7733CB20AFBD36E0C0FAD6">Zoosystema</mods:title>
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<mods:part id="49D20D7A783FF58C5085271E25DF68DB">
<mods:date id="9666699DAE19850BD9C7E654DD095858">2020</mods:date>
<mods:detail id="727BA24ABBD75EF7C5D50F2DDD1476B5" type="pubDate">
<mods:number id="EDCD9282BB9D5BB13EE8CCF438CFCA92">2020-11-05</mods:number>
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<mods:detail id="8824E1555AD3A2BB230905B4204A5D1B" type="volume">
<mods:number id="2F545F3CC4835A3958A25D702C66A856">42</mods:number>
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<mods:classification id="16CC6CD20F38C67A2AB2586A22B844A3">journal article</mods:classification>
<mods:identifier id="7C1D041DC4F745ABE904101473B84770" type="CLB-Dataset">9608</mods:identifier>
<mods:identifier id="25F7965308FFDE8058EF335A5ABB96C3" type="DOI">10.5252/zoosystema2020v42a28</mods:identifier>
<mods:identifier id="F54B18E3008DC40111E1DAA2336612E6" type="GBIF-Dataset">c2906488-8302-4ccb-b2cf-e9fe1e782ea5</mods:identifier>
<mods:identifier id="7AEDFFAD16B25E17313930473EB66AFC" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="155D1DE81CCACF8994C32C979D4A31AD" type="Zenodo-Dep">4265439</mods:identifier>
<mods:identifier id="2D26FF4BC6152A79165487416C2176C1" type="ZooBank">5CAF9561-DB13-4E22-BB65-7B771976637C</mods:identifier>
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<treatment id="038C87CCFFD5FF85FF38FF30FECAF9D7" ID-DOI="http://doi.org/10.5281/zenodo.4337058" ID-GBIF-Taxon="168962057" ID-Zenodo-Dep="4337058" LSID="urn:lsid:plazi:treatment:038C87CCFFD5FF85FF38FF30FECAF9D7" httpUri="http://treatment.plazi.org/id/038C87CCFFD5FF85FF38FF30FECAF9D7" lastPageNumber="574" pageId="29" pageNumber="574">
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<taxonomicName id="4C254D59FFD5FF85FF38FF30FD47FF19" ID-CoL="57DRP" authority="(Angel, 1922)" baseAuthorityName="Angel" baseAuthorityYear="1922" box="[231,676,215,242]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="species" species="milletihorsini">
<emphasis id="B951EAC8FFD5FF85FF38FF30FDE5FF1A" bold="true" box="[231,518,215,242]" italics="true" pageId="29" pageNumber="574">Tomopterna milletihorsini</emphasis>
(Angel, 1922)
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</heading>
</paragraph>
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<paragraph id="8B9A36DAFFD5FF85FF5BFEFAFCE9FEB5" blockId="29.[132,778,272,350]" pageId="29" pageNumber="574">
COLLECTION MATERIAL. — There are no voucher specimens available for this species from
<collectingCountry id="F332764AFFD5FF85FE5CFEC3FDE6FEAA" box="[387,517,299,323]" name="Burkina Faso" pageId="29" pageNumber="574">Burkina Faso</collectingCountry>
although its presence was reported in the East and Central-East in this country in the literature.
</paragraph>
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LITERATURE REPORTS. —
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<collectingCountry id="F332764AFFD5FF85FE5DFE9EFDE5FE65" box="[386,518,373,398]" name="Burkina Faso" pageId="29" pageNumber="574">Burkina Faso</collectingCountry>
</emphasis>
•
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province,
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;
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,
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;
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a.s.l. (Mohneke 2011;
<bibRefCitation id="EFB44B2BFFD5FF85FF5BFE42FEABFE2A" author="MOHNEKE M. &amp; ONADEKO A. B. &amp; RODEL M. - O." box="[132,328,426,450]" pageId="29" pageNumber="574" pagination="78 - 83" refId="ref45109" refString="MOHNEKE M., ONADEKO A. B. &amp; RODEL M. - O. 2011. - Medicinal and dietary uses of amphibians in Burkina Faso. African Journal of Herpetology 60: 78 - 83. https: // doi. org / 10.1080 / 21 564574.2011. 564660" type="journal article" year="2011">
Mohneke
<emphasis id="B951EAC8FFD5FF85FF39FE42FEE8FE2A" box="[230,267,426,450]" italics="true" pageId="29" pageNumber="574">et al</emphasis>
. 2011
</bibRefCitation>
)
</materialsCitation>
•
<materialsCitation id="3B4D3C87FFD5FF85FEBBFE42FECAFE1F" ID-GBIF-Occurrence="2974253742" country="Burkina Faso" elevation="300" latitude="12.2483" location="Zorgho" longLatPrecision="7" longitude="-0.6278" municipality="Ganzourgou" pageId="29" pageNumber="574" specimenCount="1">
<collectingMunicipality id="6BFEACA0FFD5FF85FEBBFE42FE3EFE2A" box="[356,477,426,450]" pageId="29" pageNumber="574">Ganzourgou</collectingMunicipality>
province,
<location id="8EFA6001FFD5FF85FD9FFE42FD68FE2A" LSID="urn:lsid:plazi:treatment:038C87CCFFD5FF85FF38FF30FECAF9D7:8EFA6001FFD5FF85FD9FFE42FD68FE2A" box="[576,651,426,450]" country="Burkina Faso" latitude="12.2483" longLatPrecision="7" longitude="-0.6278" municipality="Ganzourgou" name="Zorgho" pageId="29" pageNumber="574">Zorgho</location>
;
<geoCoordinate id="EE11501DFFD5FF85FD4BFE42FCE0FE2A" box="[660,771,426,450]" degrees="12.2483" direction="north" orientation="latitude" pageId="29" pageNumber="574" precision="5" value="12.2483">12.2483°N</geoCoordinate>
, –
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;
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<elevation id="0008D1E9FFD5FF85FEDEFE2CFEA3FE34" box="[257,320,452,476]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="574" unit="m" value="300.0">300 m</elevation>
</quantity>
a.s.l.; I-III.2008 (
<bibRefCitation id="EFB44B2BFFD5FF85FE31FE2CFD20FE34" author="MOHNEKE M. &amp; ONADEKO A. B. &amp; HIRSCHFELD M. &amp; RODEL M. - O." box="[494,707,452,476]" pageId="29" pageNumber="574" pagination="117 - 128" refId="ref45062" refString="MOHNEKE M., ONADEKO A. B., HIRSCHFELD M. &amp; RODEL M. - O. 2010 b. - Dried or fried: amphibians in local and regional food markets in West Africa. Traffic Bulletin 22: 117 - 128." type="journal article" year="2010">
Mohneke
<emphasis id="B951EAC8FFD5FF85FD8EFE2DFD9CFE34" box="[593,639,452,476]" italics="true" pageId="29" pageNumber="574">et al.</emphasis>
2010b
</bibRefCitation>
;
<bibRefCitation id="EFB44B2BFFD5FF85FD10FE2CFEC3FE1F" author="MOHNEKE M. &amp; ONADEKO A. B. &amp; RODEL M. - O." pageId="29" pageNumber="574" pagination="78 - 83" refId="ref45109" refString="MOHNEKE M., ONADEKO A. B. &amp; RODEL M. - O. 2011. - Medicinal and dietary uses of amphibians in Burkina Faso. African Journal of Herpetology 60: 78 - 83. https: // doi. org / 10.1080 / 21 564574.2011. 564660" type="journal article" year="2011">
Mohneke
<emphasis id="B951EAC8FFD5FF85FF67FE37FF04FE1E" box="[184,231,478,502]" italics="true" pageId="29" pageNumber="574">et al.</emphasis>
2011
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6551FFD5FF85FF5BFDE7FDB8FD2D" pageId="29" pageNumber="574" type="description">
<paragraph id="8B9A36DAFFD5FF85FF5BFDE7FDB8FD2D" blockId="29.[131,777,526,709]" pageId="29" pageNumber="574">
DESCRIPTION.
<emphasis id="B951EAC8FFD5FF85FECFFDE7FECAFDCE" box="[272,297,527,550]" italics="true" pageId="29" pageNumber="574">—</emphasis>
Species with a short head, a horizontal pupil, short and strong limbs. Inner metatarsal tubercles large and shovel-shaped, tarsal tubercles usually present. Frog with a feebly warty back skin and smooth ventral skin. Dorsal colour brown to beige or dark brown. Pale dorsolateral lines often present. A white supratympanic ridge. Numerous dark brown, greenish or reddish spots, often with black borders, distinct on head, back and flanks.
</paragraph>
</subSubSection>
<subSubSection id="C33F6551FFD5FF85FF5BFD35FD7BFB38" pageId="29" pageNumber="574" type="discussion">
<paragraph id="8B9A36DAFFD5FF85FF5BFD35FD7BFB38" blockId="29.[130,777,732,1233]" pageId="29" pageNumber="574">
REMARKS. — The genus
<taxonomicName id="4C254D59FFD5FF85FEADFD35FD3EFD1D" authority="Dumeril &amp; Bibron, 1841" authorityName="Dumeril &amp; Bibron" authorityYear="1841" box="[370,733,732,757]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="genus">
<emphasis id="B951EAC8FFD5FF85FEADFD35FE01FD1C" box="[370,482,733,756]" italics="true" pageId="29" pageNumber="574">Tomopterna</emphasis>
Duméril &amp; Bibron, 1841
</taxonomicName>
still comprises various undescribed species and needs taxonomic revision (
<bibRefCitation id="EFB44B2BFFD5FF85FF64FCF9FE22FCC2" author="PADIAL J. M. &amp; DE LA RIVA I." box="[187,449,785,810]" pageId="29" pageNumber="574" pagination="89 - 99" refId="ref45553" refString="PADIAL J. M. &amp; DE LA RIVA I. 2004. - Annotated checklist of the amphibians of Mauritania (West Africa). Revista Espanola de Herpetologia 18: 89 - 99." type="journal article" year="2004">Padial &amp; De La Riva 2004</bibRefCitation>
;
<bibRefCitation id="EFB44B2BFFD5FF85FE11FCF9FD9BFCC2" author="PADIAL J. M. &amp; CROCHET P. A. &amp; GENIEZ P. &amp; BRITO J. C." box="[462,632,785,810]" pageId="29" pageNumber="574" pagination="11 - 22" refId="ref45500" refString="PADIAL J. M., CROCHET P. A., GENIEZ P. &amp; BRITO J. C. 2013. - Amphibian conservation in Mauritania. Basic and applied Herpetology 27: 11 - 22. http: // dx. doi. org / 10.11160 / bah. 13002" type="journal article" year="2013">
Padial
<emphasis id="B951EAC8FFD5FF85FDCFFCFAFDDCFCC1" box="[528,575,785,809]" italics="true" pageId="29" pageNumber="574">et al.</emphasis>
2013
</bibRefCitation>
). There are 14 species known in this genus and these are also known to be highly cryptic and morphologically similar (
<bibRefCitation id="EFB44B2BFFD5FF85FE32FCAFFCE2FCB6" author="WASONGA V. D. &amp; CHANNING A." box="[493,769,838,863]" pageId="29" pageNumber="574" pagination="221 - 240" refId="ref47107" refString="WASONGA V. D. &amp; CHANNING A. 2013. - Identification of sand frogs (Anura: Pyxicephalidae: Tomopterna) from Kenya with the description of two new species. Zootaxa 3734: 221 - 240. http: // dx. doi. org / 10.11646 / zootaxa. 3734.2.7" type="journal article" year="2013">Wasonga &amp; Channing 2013</bibRefCitation>
;
<bibRefCitation id="EFB44B2BFFD5FF85FF5BFC89FE6EFC91" author="WILSON L. &amp; CHANNING A." box="[132,397,865,889]" pageId="29" pageNumber="574" pagination="225 - 246" refId="ref47242" refString="WILSON L. &amp; CHANNING A. 2019. - A new sand frog from Namaqualand, South Africa (Pyxicephalidae: Tomopterna). Zootaxa 4609: 225 - 246. http: // dx. doi. org / 10.11646 / zootaxa. 4609.2.2" type="journal article" year="2019">Wilson &amp; Channing 2019</bibRefCitation>
). The commonly used name for West African populations was
<taxonomicName id="4C254D59FFD5FF85FE5EFC94FCE4FC7B" authority="(Boulenger, 1907)" baseAuthorityName="Boulenger" baseAuthorityYear="1907" box="[385,775,891,915]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="species" species="cryptotis">
<emphasis id="B951EAC8FFD5FF85FE5EFC94FDAAFC7B" box="[385,585,892,915]" italics="true" pageId="29" pageNumber="574">Tomopterna cryptotis</emphasis>
(Boulenger, 1907)
</taxonomicName>
(
<bibRefCitation id="EFB44B2BFFD5FF85FF53FC7EFE95FC46" author="LAMOTTE M. &amp; XAVIER F." box="[140,374,918,942]" pageId="29" pageNumber="574" pagination="773 - 816" refId="ref44541" refString="LAMOTTE M. &amp; XAVIER F. 1981. - Les Amphibiens de l'Afrique de l'Ouest, in DURAND J. R. &amp; LEVEQUE C. (eds), Faune et Flore aquatiques de l'Afrique Sahelo-soudanienne, Editions de l'Office de la Recherche Scientifique et Technique d'Outre-Mer 2: 773 - 816." type="journal article" year="1981">Lamotte &amp; Xavier 1981</bibRefCitation>
;
<bibRefCitation id="EFB44B2BFFD5FF85FE5DFC7EFE17FC46" author="RODEL M. - O." box="[386,500,918,942]" pageId="29" pageNumber="574" refId="ref45923" refString="RODEL M. - O. 2000. - Herpetofauna of West Africa: Amphibians of the West African savannah. Edition Chimaira. Vol. 1. Germany, Frankfurt am Main, 300 p." type="book" year="2000">Rödel 2000</bibRefCitation>
;
<bibRefCitation id="EFB44B2BFFD5FF85FDDFFC7EFCE2FC46" author="PADIAL J. M. &amp; DE LA RIVA I." box="[512,769,918,942]" pageId="29" pageNumber="574" pagination="89 - 99" refId="ref45553" refString="PADIAL J. M. &amp; DE LA RIVA I. 2004. - Annotated checklist of the amphibians of Mauritania (West Africa). Revista Espanola de Herpetologia 18: 89 - 99." type="journal article" year="2004">Padial &amp; De La Riva 2004</bibRefCitation>
; Mohneke 2011;
<bibRefCitation id="EFB44B2BFFD5FF85FEF8FC58FE2CFC20" author="PADIAL J. M. &amp; CROCHET P. A. &amp; GENIEZ P. &amp; BRITO J. C." box="[295,463,944,968]" pageId="29" pageNumber="574" pagination="11 - 22" refId="ref45500" refString="PADIAL J. M., CROCHET P. A., GENIEZ P. &amp; BRITO J. C. 2013. - Amphibian conservation in Mauritania. Basic and applied Herpetology 27: 11 - 22. http: // dx. doi. org / 10.11160 / bah. 13002" type="journal article" year="2013">
Padial
<emphasis id="B951EAC8FFD5FF85FEB7FC59FE74FC20" box="[360,407,944,968]" italics="true" pageId="29" pageNumber="574">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFB44B2BFFD5FF85FE04FC58FD92FC20" author="SOW A. S. &amp; GONCALVES D. V. &amp; SOUSA F. V. &amp; FREIRIA F. &amp; SANTAREM F. &amp; ANTON G. &amp; DIENG H. &amp; CAMPOS J. C. &amp; DIAGNE S. K. &amp; BORATYNSKI Z. &amp; BRITO J. C." box="[475,625,944,968]" pageId="29" pageNumber="574" pagination="101 - 116" refId="ref46751" refString="SOW A. S., GONCALVES D. V., SOUSA F. V., MARTINEZ- FREIRIA F., SANTAREM F., VELO- ANTON G., DIENG H., CAMPOS J. C., DIAGNE S. K., BORATYNSKI Z. &amp; BRITO J. C. 2017. - Atlas of the distribution of amphibians and reptiles in the Diawling National Park, Mauritania. Basic and applied Herpetology 31: 101 - 116. http: // dx. doi. org / 10.11160 / bah. 87" type="journal article" year="2017">
Sow
<emphasis id="B951EAC8FFD5FF85FDD5FC59FDDAFC20" box="[522,569,944,968]" italics="true" pageId="29" pageNumber="574">et al.</emphasis>
2017
</bibRefCitation>
).
<bibRefCitation id="EFB44B2BFFD5FF85FD5AFC58FF09FC0B" author="OHLER A. &amp; FRETEY T." pageId="29" pageNumber="574" pagination="173 - 175" refId="ref45385" refString="OHLER A. &amp; FRETEY T. 2008. - Statut du nom Arthroleptis milletihorsini, Angel, 1922 (Amphibia, Anura). Alytes 25: 173 - 175." type="journal article" year="2008">Ohler &amp; Frétey (2008)</bibRefCitation>
investigated the status of
<taxonomicName id="4C254D59FFD5FF85FE3CFC23FF0AFC15" authority="(Angel, 1922)" baseAuthorityName="Angel" baseAuthorityYear="1922" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="species" species="milletihorsini">
<emphasis id="B951EAC8FFD5FF85FE3CFC23FD30FC0B" box="[483,723,971,995]" italics="true" pageId="29" pageNumber="574">Arthroleptis milletihorsini</emphasis>
(Angel, 1922)
</taxonomicName>
, the type of which was deposited in MNHN collection. According to their examination, the characters did not correspond to the morphological characters of the genus
<taxonomicName id="4C254D59FFD5FF85FDCAFBF2FCE0FBDA" authority="Smith, 1849" authorityName="Smith" authorityYear="1849" box="[533,771,1050,1074]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="genus">
<emphasis id="B951EAC8FFD5FF85FDCAFBF2FD62FBDA" box="[533,641,1050,1074]" italics="true" pageId="29" pageNumber="574">Arthroleptis</emphasis>
Smith, 1849
</taxonomicName>
, but rather to the characters of the genus
<taxonomicName id="4C254D59FFD5FF85FDFFFBDDFEEEFB8F" authority="Dumeril &amp; Bibron, 1841" authorityName="Dumeril &amp; Bibron" authorityYear="1841" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="genus">
<emphasis id="B951EAC8FFD5FF85FDFFFBDDFD73FBA4" box="[544,656,1077,1100]" italics="true" pageId="29" pageNumber="574">Tomopterna</emphasis>
Duméril &amp; Bibron, 1841
</taxonomicName>
. The
<typeStatus id="549E8878FFD5FF85FE9CFBA7FE7AFB8F" box="[323,409,1103,1127]" pageId="29" pageNumber="574" type="holotype">holotype</typeStatus>
is a young specimen of
<quantity id="4CDD9B3FFFD5FF85FD58FBA7FD33FB8E" box="[647,720,1103,1127]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="574" unit="mm" value="18.0">18 mm</quantity>
SVL.
<bibRefCitation id="EFB44B2BFFD5FF85FF5AFB81FEBAFB69" author="OHLER A. &amp; FRETEY T." box="[133,345,1129,1153]" pageId="29" pageNumber="574" pagination="173 - 175" refId="ref45385" refString="OHLER A. &amp; FRETEY T. 2008. - Statut du nom Arthroleptis milletihorsini, Angel, 1922 (Amphibia, Anura). Alytes 25: 173 - 175." type="journal article" year="2008">Ohler &amp; Frétey (2008)</bibRefCitation>
allocated the nomen
<taxonomicName id="4C254D59FFD5FF85FDC1FB81FCE6FB69" baseAuthorityName="Angel" baseAuthorityYear="1922" box="[542,773,1129,1153]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="species" species="milletihorsini">
<emphasis id="B951EAC8FFD5FF85FDC1FB81FCE6FB69" box="[542,773,1129,1153]" italics="true" pageId="29" pageNumber="574">Arthroleptis milletihorsini</emphasis>
</taxonomicName>
to
<taxonomicName id="4C254D59FFD5FF85FF41FB6DFEEDFB74" authorityName="Dumeril &amp; Bibron" authorityYear="1841" box="[158,270,1157,1180]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="genus">
<emphasis id="B951EAC8FFD5FF85FF41FB6DFEEDFB74" box="[158,270,1157,1180]" italics="true" pageId="29" pageNumber="574">Tomopterna</emphasis>
</taxonomicName>
, as
<taxonomicName id="4C254D59FFD5FF85FEEEFB6DFD52FB74" authority="(Angel, 1922)" baseAuthorityName="Angel" baseAuthorityYear="1922" box="[305,689,1156,1180]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="species" species="milletihorsini">
<emphasis id="B951EAC8FFD5FF85FEEEFB6DFDC0FB74" box="[305,547,1156,1180]" italics="true" pageId="29" pageNumber="574">Tomopterna milletihorsini</emphasis>
(Angel, 1922)
</taxonomicName>
with the type locality being Kati
<quantity id="4CDD9B3FFFD5FF85FEABFB76FE55FB5E" box="[372,438,1182,1206]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="574" unit="km" value="12.0">12 km</quantity>
north to Bamako from
<collectingCountry id="F332764AFFD5FF85FD75FB76FD38FB5E" box="[682,731,1182,1206]" name="Serbia" pageId="29" pageNumber="574">Mali</collectingCountry>
. We tentatively use this name for West African
<taxonomicName id="4C254D59FFD5FF85FDFDFB51FD71FB38" authorityName="Dumeril &amp; Bibron" authorityYear="1841" box="[546,658,1209,1232]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="genus">
<emphasis id="B951EAC8FFD5FF85FDFDFB51FD71FB38" box="[546,658,1209,1232]" italics="true" pageId="29" pageNumber="574">Tomopterna</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6551FFD5FF85FF5BFB02FECAF9D7" pageId="29" pageNumber="574" type="biology_ecology">
<paragraph id="8B9A36DAFFD5FF85FF5BFB02FECAF9D7" blockId="29.[129,777,1258,1599]" pageId="29" pageNumber="574">
HABITAT. — During our field sampling, the presence of
<taxonomicName id="4C254D59FFD5FF85FD48FB03FF1EFAF4" baseAuthorityName="Angel" baseAuthorityYear="1922" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="species" species="milletihorsini">
<emphasis id="B951EAC8FFD5FF85FD48FB03FF1EFAF4" italics="true" pageId="29" pageNumber="574">Tomopterna milletihorsini</emphasis>
</taxonomicName>
has been reported from Dori and Yakouta zones where gardeners and fishermen reported a frog species which inflate their body as
<taxonomicName id="4C254D59FFD5FF85FF03FAD1FE5FFAB9" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[220,444,1337,1361]" class="Amphibia" family="Pyxicephalidae" genus="Pyxicephalus" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="species" species="maltzanii">
<emphasis id="B951EAC8FFD5FF85FF03FAD1FE5FFAB9" box="[220,444,1337,1361]" italics="true" pageId="29" pageNumber="574">Pyxicephalus maltzanii</emphasis>
</taxonomicName>
and is living under the sand. In Koti women indicated its presence as common in their groundnut fields after the harvesting (mostly from November to December). Women reported that the species was found by young shepherds who search it in sand in dried backwaters or in holes of groundnut fields after harvesting. In West Africa,
<taxonomicName id="4C254D59FFD5FF85FE2FFA56FD83FA3D" authorityName="Dumeril &amp; Bibron" authorityYear="1841" box="[496,608,1470,1493]" class="Amphibia" family="Pyxicephalidae" genus="Tomopterna" kingdom="Animalia" order="Anura" pageId="29" pageNumber="574" phylum="Chordata" rank="genus">
<emphasis id="B951EAC8FFD5FF85FE2FFA56FD83FA3D" box="[496,608,1470,1493]" italics="true" pageId="29" pageNumber="574">Tomopterna</emphasis>
</taxonomicName>
is found in semiarid to arid habitats on sandy soils in the Sahel zone (
<bibRefCitation id="EFB44B2BFFD5FF85FD51FA30FCE2FA18" author="RODEL M. - O." box="[654,769,1496,1520]" pageId="29" pageNumber="574" refId="ref45923" refString="RODEL M. - O. 2000. - Herpetofauna of West Africa: Amphibians of the West African savannah. Edition Chimaira. Vol. 1. Germany, Frankfurt am Main, 300 p." type="book" year="2000">Rödel 2000</bibRefCitation>
;
<bibRefCitation id="EFB44B2BFFD5FF85FF5BFA1AFE6BF9E3" author="PADIAL J. M. &amp; DE LA RIVA I." box="[132,392,1522,1547]" pageId="29" pageNumber="574" pagination="89 - 99" refId="ref45553" refString="PADIAL J. M. &amp; DE LA RIVA I. 2004. - Annotated checklist of the amphibians of Mauritania (West Africa). Revista Espanola de Herpetologia 18: 89 - 99." type="journal article" year="2004">Padial &amp; De La Riva 2004</bibRefCitation>
; Mohneke 2011). The species has been also found in mountains zones from
<collectingCountry id="F332764AFFD5FF85FE3DF9E5FDACF9CD" box="[482,591,1549,1573]" name="Mauritania" pageId="29" pageNumber="574">Mauritania</collectingCountry>
(
<bibRefCitation id="EFB44B2BFFD5FF85FD82F9E5FCE2F9CD" author="PADIAL J. M. &amp; CROCHET P. A. &amp; GENIEZ P. &amp; BRITO J. C." box="[605,769,1549,1573]" pageId="29" pageNumber="574" pagination="11 - 22" refId="ref45500" refString="PADIAL J. M., CROCHET P. A., GENIEZ P. &amp; BRITO J. C. 2013. - Amphibian conservation in Mauritania. Basic and applied Herpetology 27: 11 - 22. http: // dx. doi. org / 10.11160 / bah. 13002" type="journal article" year="2013">
Padial
<emphasis id="B951EAC8FFD5FF85FD43F9E6FD29F9CD" box="[668,714,1549,1573]" italics="true" pageId="29" pageNumber="574">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFB44B2BFFD5FF85FF5BF9CFFEF8F9D7" author="SOW A. S. &amp; GONCALVES D. V. &amp; SOUSA F. V. &amp; FREIRIA F. &amp; SANTAREM F. &amp; ANTON G. &amp; DIENG H. &amp; CAMPOS J. C. &amp; DIAGNE S. K. &amp; BORATYNSKI Z. &amp; BRITO J. C." box="[132,283,1575,1599]" pageId="29" pageNumber="574" pagination="101 - 116" refId="ref46751" refString="SOW A. S., GONCALVES D. V., SOUSA F. V., MARTINEZ- FREIRIA F., SANTAREM F., VELO- ANTON G., DIENG H., CAMPOS J. C., DIAGNE S. K., BORATYNSKI Z. &amp; BRITO J. C. 2017. - Atlas of the distribution of amphibians and reptiles in the Diawling National Park, Mauritania. Basic and applied Herpetology 31: 101 - 116. http: // dx. doi. org / 10.11160 / bah. 87" type="journal article" year="2017">
Sow
<emphasis id="B951EAC8FFD5FF85FF6CF9C0FF01F9D7" box="[179,226,1575,1599]" italics="true" pageId="29" pageNumber="574">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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