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<mods:title id="E694D631DAB15C38294106628EAE0DE4">Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)</mods:title>
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<treatment id="03DA87C4FF8F176FFF4C6E7DA974FDED" ID-DOI="http://doi.org/10.5281/zenodo.6109830" ID-GBIF-Taxon="119596545" ID-Zenodo-Dep="6109830" LSID="urn:lsid:plazi:treatment:03DA87C4FF8F176FFF4C6E7DA974FDED" httpUri="http://treatment.plazi.org/id/03DA87C4FF8F176FFF4C6E7DA974FDED" lastPageId="8" lastPageNumber="119" pageId="7" pageNumber="118">
<subSubSection id="C3696559FF8F1760FF4C6E7DA837FBBB" box="[151,555,1052,1078]" pageId="7" pageNumber="118" type="nomenclature">
<paragraph id="8BCC36D2FF8F1760FF4C6E7DA837FBBB" blockId="7.[151,555,1052,1078]" box="[151,555,1052,1078]" pageId="7" pageNumber="118">
<heading id="D08481BEFF8F1760FF4C6E7DA837FBBB" bold="true" box="[151,555,1052,1078]" fontSize="11" level="1" pageId="7" pageNumber="118" reason="1">
<emphasis id="B907EAC0FF8F1760FF4C6E7DA837FBBB" bold="true" box="[151,555,1052,1078]" pageId="7" pageNumber="118">
Genus
<taxonomicName id="4C734D51FF8F1760FF356E7DA837FBBB" ID-CoL="6QQD" authority="Piersig, 1894" authorityName="Piersig" authorityYear="1894" box="[238,555,1052,1078]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8F1760FF356E7DAB98FBBB" bold="true" box="[238,388,1052,1078]" italics="true" pageId="7" pageNumber="118">Pionacercus</emphasis>
Piersig, 1894
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8F1760FF4C6E02A891FA9F" pageId="7" pageNumber="118" type="reference_group">
<paragraph id="8BCC36D2FF8F1760FF4C6E02A8A7FB1A" blockId="7.[151,955,1122,1298]" pageId="7" pageNumber="118">
<taxonomicName id="4C734D51FF8F1760FF4C6E02ABB0FBF5" ID-CoL="6QQD" box="[151,428,1122,1144]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8F1760FF4C6E02AB09FBF5" box="[151,277,1123,1144]" italics="true" pageId="7" pageNumber="118">Pionacercus</emphasis>
(
<emphasis id="B907EAC0FF8F1760FEF86E02ABBCFBF5" box="[291,416,1123,1144]" italics="true" pageId="7" pageNumber="118">Pionacercus</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="EFE24B23FF8F1760FE6D6E03A827FBF5" author="Cook" box="[438,571,1122,1144]" pageId="7" pageNumber="118" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook, 1974a</bibRefCitation>
, pp. 284285, figs. 1203, 1206, 1207.
<taxonomicName id="4C734D51FF8F1760FF4C6EE3AB09FB1A" ID-CoL="6QQD" box="[151,277,1154,1175]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8F1760FF4C6EE3AB09FB1A" box="[151,277,1154,1175]" italics="true" pageId="7" pageNumber="118">Pionacercus</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8F1760FEA16EE0ABE4FB1A" author="Smith" box="[378,504,1153,1175]" pageId="7" pageNumber="118" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith, 1976</bibRefCitation>
, pp. 4446, 7981.
</paragraph>
<paragraph id="8BCC36D2FF8F1760FF4C6EC0A891FA9F" blockId="7.[151,955,1122,1298]" pageId="7" pageNumber="118">
<taxonomicName id="4C734D51FF8F1760FF4C6EC0AB09FB3B" ID-CoL="6QQD" box="[151,277,1185,1206]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8F1760FF4C6EC0AB09FB3B" box="[151,277,1185,1206]" italics="true" pageId="7" pageNumber="118">Pionacercus</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8F1760FEA16EC1A853FB3B" author="Smith" box="[378,591,1184,1206]" pageId="7" pageNumber="118" refString="Smith, I. M. &amp; Cook, D. R. (1991) Water mites. Chapter 16. In: Thorp, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. Academic Press, San Diego, CA, pp. 523 - 592. [total page number 911 pp.]" type="book chapter" year="1991">Smith &amp; Cook, 1991</bibRefCitation>
, pp. 554, 576, 578.
<taxonomicName id="4C734D51FF8F1760FF4C6EA1AB09FB58" ID-CoL="6QQD" box="[151,277,1216,1237]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8F1760FF4C6EA1AB09FB58" box="[151,277,1216,1237]" italics="true" pageId="7" pageNumber="118">Pionacercus</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8F1760FEA16EDFA82DFB59" author="Smith" box="[378,561,1214,1237]" pageId="7" pageNumber="118" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2001) Water mites and other arachnids. Chapter 16. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 2 nd Edition. Academic Press, Waltham, Massachusetts, pp. 551 - 659. [1056 pp.] http: // dx. doi. org / 10.1016 / B 978 - 012690647 - 9 / 50017 - X" type="book chapter" year="2001">
Smith
<emphasis id="B907EAC0FF8F1760FE666EA1ABF6FB59" box="[445,490,1214,1237]" italics="true" pageId="7" pageNumber="118">et al</emphasis>
., 2001
</bibRefCitation>
, pp. 612, 615, fig. 336.
<taxonomicName id="4C734D51FF8F1760FF4C6EBFAB09FB7E" ID-CoL="6QQD" box="[151,277,1246,1267]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8F1760FF4C6EBFAB09FB7E" box="[151,277,1246,1267]" italics="true" pageId="7" pageNumber="118">Pionacercus</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8F1760FEA16EBCA82DFB7E" author="Smith" box="[378,561,1245,1267]" pageId="7" pageNumber="118" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2010) Water mites and other arachnids. Chapter 15. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 3 rd Edition. Academic Press, Waltham, Massachusetts, pp. 485 - 586. [total page number: 1021 pp.] http: // dx. doi. org / 10.1016 / B 978 - 0 - 12 - 374855 - 3.00015 - 7" type="book chapter" year="2010">
Smith
<emphasis id="B907EAC0FF8F1760FE666EBFABF6FB7E" box="[445,490,1245,1267]" italics="true" pageId="7" pageNumber="118">et al</emphasis>
., 2010
</bibRefCitation>
, pp. 553, 556, fig. 15.337.
<taxonomicName id="4C734D51FF8F1760FF4C6E9CABC9FA9F" authority="Gerecke, 2014" authorityName="Gerecke" authorityYear="2014" box="[151,469,1276,1298]" class="Arachnida" family="Pionidae" genus="Pionacercopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8F1760FF4C6E9CAB2EFA9F" box="[151,306,1277,1298]" italics="true" pageId="7" pageNumber="118">Pionacercopsis</emphasis>
: Gerecke, 2014
</taxonomicName>
, pp. 5960, fig. 2.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8F1760FF4C6F5FAE4BF852" pageId="7" pageNumber="118" type="diagnosis">
<paragraph id="8BCC36D2FF8F1760FF4C6F5FAB56FAF7" blockId="7.[151,1437,1342,2015]" pageId="7" pageNumber="118">
<emphasis id="B907EAC0FF8F1760FF4C6F5FAB7AFADA" bold="true" box="[151,358,1342,1367]" pageId="7" pageNumber="118">
Diagnosis.
<emphasis id="B907EAC0FF8F1760FEC36F5EAB40FADB" bold="true" box="[280,348,1343,1366]" italics="true" pageId="7" pageNumber="118">Larva</emphasis>
:
</emphasis>
Partially described by
<bibRefCitation id="EFE24B23FF8F1760FDB26F5FA935FADA" author="Lundblad" box="[617,809,1342,1367]" pageId="7" pageNumber="118" refString="Lundblad, O. (1927) Die Hydracarinen Schwedens. I. Beitrag zur Systematik, Embryologie, Okologie und Verbreitungsgeschichte der schwedischen Arten. Zoologiska Bidrag frDn Uppsala, 11, 181 - 540." type="journal article" year="1927">Lundblad (1927)</bibRefCitation>
, but insufficiently known to permit listing of diagnostic character states.
</paragraph>
<paragraph id="8BCC36D2FF8F1760FF1C6FE7AE4BF852" blockId="7.[151,1437,1342,2015]" pageId="7" pageNumber="118">
<emphasis id="B907EAC0FF8F1760FF1C6FE7AB0EFA12" bold="true" box="[199,274,1414,1439]" italics="true" pageId="7" pageNumber="118">Adults</emphasis>
(modified from
<bibRefCitation id="EFE24B23FF8F1760FE316FE7A86BFA12" author="Smith" box="[490,631,1414,1439]" pageId="7" pageNumber="118" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith 1976</bibRefCitation>
): Character states of subfamily Foreliinae. Idiosoma with dorsal integument mostly soft and bearing only tiny platelets associated with glandularia and idiosomal setae, or mostly covered by a dorsal shield that extends anteriorly to and including level of postocular setae in males; mostly soft and bearing only tiny platelets associated with glandularia and idiosomal setae in females. Venter with coxal plates and genital field incorporated into a ventral shield in males, but surrounded by soft integument bearing only tiny sclerites associated with glandularia and idiosomal setae in females. Fourth coxal plate with medial edge relatively long in males, short but not reduced to angles in females. Genital field with three pairs of relatively small acetabula borne on ventral shield flanking gonopore in males and on triangular acetabular plates flanking gonopore in females, extending laterally to but not beyond posterolateral angles of fourth coxal plates. Pedipalp segments moderately long and slender and with tibia bearing two slender setae on small projections ventrally and a sessile, peg-like seta distomedially. Male third leg with tarsus bearing slightly reduced claw sockets that are terminal in position and small but only slightly modified claws; fourth leg with genu short, moderately expanded, and bearing a row of moderately long, slightly thickened setae ventrally, a patch of slender swimming setae anteriorly, and patches of slender swimming setae and thick, blade-like setae distoventrally, lacking a prominent distoventral projection, tibia relatively long and slender and bearing a row of slender swimming setae ventrally and several swimming setae distally, tarsus curved dorsally, bearing a row of thick, peg-like setae that extends both proximal and distal to deepest part of curved portion, and bearing thick, highly modified claws.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF80176FFF1C6AF6A8BFFEE8" pageId="8" pageNumber="119" type="materials_examined">
<paragraph id="8BCC36D2FF80176FFF1C6AF6A8A7FF3D" blockId="8.[151,1436,151,608]" box="[199,699,151,176]" pageId="8" pageNumber="119">
<emphasis id="B907EAC0FF80176FFF1C6AF6AB7EFF3D" bold="true" box="[199,354,151,176]" pageId="8" pageNumber="119">
<typeStatus id="54C88870FF80176FFF1C6AF6AB1EFF3D" box="[199,258,151,176]" pageId="8" pageNumber="119">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C734D51FF80176FFEB26AF8A8A7FF3D" authority="Piersig." authorityName="Piersig." box="[361,699,151,176]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="leuckarti">
<emphasis id="B907EAC0FF80176FFEB26AF8A842FF3D" box="[361,606,151,176]" italics="true" pageId="8" pageNumber="119">Pionacercus leuckarti</emphasis>
Piersig.
</taxonomicName>
</paragraph>
<paragraph id="8BCC36D2FF80176FFF1C6ADDA8BFFEE8" blockId="8.[151,1436,151,608]" pageId="8" pageNumber="119">
<emphasis id="B907EAC0FF80176FFF1C6ADDAB8BFF58" bold="true" box="[199,407,188,213]" pageId="8" pageNumber="119">Species included.</emphasis>
<taxonomicName id="4C734D51FF80176FFE796ADCA97BFF59" authority="Piersig" authorityName="Piersig" box="[418,871,188,213]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="leuckarti">
<emphasis id="B907EAC0FF80176FFE796ADCA880FF58" box="[418,668,188,213]" italics="true" pageId="8" pageNumber="119">Pionacercus leuckarti</emphasis>
Piersig (Europe)
</taxonomicName>
,
<taxonomicName id="4C734D51FF80176FFCAD6ADCAEEEFF59" authority="(Koenike) (Europe)" authorityName="Koenike" baseAuthorityName="Koenike" box="[886,1266,188,213]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="uncinatus">
<emphasis id="B907EAC0FF80176FFCAD6ADCAE1FFF59" box="[886,1027,189,212]" italics="true" pageId="8" pageNumber="119">P. uncinatus</emphasis>
(Koenike) (Europe)
</taxonomicName>
,
<taxonomicName id="4C734D51FF80176FFADA6ADCAB5CFF75" authority="Thor" authorityName="Thor" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="norvegicus">
<emphasis id="B907EAC0FF80176FFADA6ADCAF80FF59" box="[1281,1436,189,212]" italics="true" pageId="8" pageNumber="119">P. norvegicus</emphasis>
Thor (Europe)
</taxonomicName>
,
<taxonomicName id="4C734D51FF80176FFE8A6A80A8BBFF75" authority="Imamura" authorityName="Imamura" box="[337,679,224,248]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B907EAC0FF80176FFE8A6A80ABC8FF75" box="[337,468,225,248]" italics="true" pageId="8" pageNumber="119">P. japonica</emphasis>
Imamura (
<collectingCountry id="F3647642FF80176FFD816A81A8BCFF75" box="[602,672,224,248]" name="Japan" pageId="8" pageNumber="119">Japan</collectingCountry>
)
</taxonomicName>
. There are five undescribed species of
<taxonomicName id="4C734D51FF80176FFB5B6A80AF17FF75" box="[1152,1291,225,248]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFB5B6A80AF17FF75" box="[1152,1291,225,248]" italics="true" pageId="8" pageNumber="119">Pionacercus</emphasis>
</taxonomicName>
from North
<collectingCountry id="F3647642FF80176FFF4C6B65AAEBFE90" box="[151,247,260,285]" name="United States of America" pageId="8" pageNumber="119">America</collectingCountry>
in the Canadian National Collection of Insects and Arachnids and larvae have been associated with adults by rearing for one of them. One of these species is also represented in the Habeeb Collection and was incorrectly reported as
<taxonomicName id="4C734D51FF80176FFEC26B2CA882FEE8" authority="Piersig (Habeeb 1958)" authorityName="Piersig (Habeeb" authorityYear="1958" box="[281,670,332,357]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="leuckarti">
<emphasis id="B907EAC0FF80176FFEC26B2CAB85FEE8" box="[281,409,332,357]" italics="true" pageId="8" pageNumber="119">P. leuckarti</emphasis>
Piersig (
<bibRefCitation id="EFE24B23FF80176FFE276B2DA88AFEE8" author="Habeeb" box="[508,662,332,357]" pageId="8" pageNumber="119" refString="Habeeb, H. (1958) New mites from New Brunswick. Leaflets of Acadian Biology, 18, 1 - 4." type="journal article" year="1958">Habeeb 1958</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF80176FFF1C6B0EA8E7FE05" box="[199,763,367,392]" pageId="8" pageNumber="119" type="distribution">
<paragraph id="8BCC36D2FF80176FFF1C6B0EA8E7FE05" blockId="8.[151,1436,151,608]" box="[199,763,367,392]" pageId="8" pageNumber="119">
<emphasis id="B907EAC0FF80176FFF1C6B0EAB7CFE05" bold="true" box="[199,352,367,392]" pageId="8" pageNumber="119">Distribution.</emphasis>
Holarctic (Eurasia, North
<collectingCountry id="F3647642FF80176FFD516B0EA8ECFE05" box="[650,752,367,392]" name="United States of America" pageId="8" pageNumber="119">America</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF80176FFF1C6BF5A974FDED" pageId="8" pageNumber="119" type="discussion">
<paragraph id="8BCC36D2FF80176FFF1C6BF5A974FDED" blockId="8.[151,1436,151,608]" pageId="8" pageNumber="119">
<emphasis id="B907EAC0FF80176FFF1C6BF5AB50FE20" bold="true" box="[199,332,404,429]" pageId="8" pageNumber="119">Discussion.</emphasis>
<bibRefCitation id="EFE24B23FF80176FFE836BF5ABE5FE21" author="Cook" box="[344,505,404,429]" pageId="8" pageNumber="119" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook (1974a)</bibRefCitation>
treated
<taxonomicName id="4C734D51FF80176FFD856BF4A8F5FE21" box="[606,745,405,428]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFD856BF4A8F5FE21" box="[606,745,405,428]" italics="true" pageId="8" pageNumber="119">Pionacercus</emphasis>
</taxonomicName>
(including the subgenus
<taxonomicName id="4C734D51FF80176FFBC26BF4AED9FE21" box="[1049,1221,405,428]" class="Arachnida" family="Pionidae" genus="Pionacercopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFBC26BF4AED9FE21" box="[1049,1221,405,428]" italics="true" pageId="8" pageNumber="119">Pionacercopsis</emphasis>
</taxonomicName>
) in the subfamily Tiphyinae.
<bibRefCitation id="EFE24B23FF80176FFECE6BD6ABB5FE5D" author="Smith" box="[277,425,439,464]" pageId="8" pageNumber="119" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith (1976)</bibRefCitation>
placed
<taxonomicName id="4C734D51FF80176FFE246BD8A896FE5D" box="[511,650,441,464]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFE246BD8A896FE5D" box="[511,650,441,464]" italics="true" pageId="8" pageNumber="119">Pionacercus</emphasis>
</taxonomicName>
(including
<taxonomicName id="4C734D51FF80176FFCD16BD8A9AAFE5D" box="[778,950,441,464]" class="Arachnida" family="Pionidae" genus="Pionacercopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFCD16BD8A9AAFE5D" box="[778,950,441,464]" italics="true" pageId="8" pageNumber="119">Pionacercopsis</emphasis>
</taxonomicName>
) in the tribe Foreliini along with the genera
<taxonomicName id="4C734D51FF80176FFF4C6BBDAAF7FE78" box="[151,235,476,501]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFF4C6BBDAAF7FE78" box="[151,235,476,501]" italics="true" pageId="8" pageNumber="119">Forelia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D51FF80176FFEFC6BBDABA2FE78" box="[295,446,476,501]" class="Arachnida" family="Pionidae" genus="Pseudofeltria" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFEFC6BBDABA2FE78" box="[295,446,476,501]" italics="true" pageId="8" pageNumber="119">Pseudofeltria</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE24B23FF80176FFE0B6BBDA8D4FE79" author="Smith" box="[464,712,476,501]" pageId="8" pageNumber="119" refString="Smith, I. M. &amp; Cook, D. R. (1991) Water mites. Chapter 16. In: Thorp, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. Academic Press, San Diego, CA, pp. 523 - 592. [total page number 911 pp.]" type="book chapter" year="1991">Smith &amp; Cook (1991</bibRefCitation>
,
<bibRefCitation id="EFE24B23FF80176FFD0C6BBDA90CFE79" author="Smith" box="[727,784,476,500]" pageId="8" pageNumber="119" refString="Smith, I. M. &amp; Cook, D. R. (1999) An assessment of global distribution patterns in water mites (Acari: Hydrachnida). In: Needham, G. R., Mitchell, R. D., Horn, D. J. &amp; Welbourn, W. C. (Eds.), Acarology IX. Vol. 2. Symposia. Ohio Biological Survey, Columbus, pp. 109 - 124." type="book" year="1999">1999</bibRefCitation>
) and
<bibRefCitation id="EFE24B23FF80176FFC8D6BBDAE32FE79" author="Smith" box="[854,1070,476,501]" pageId="8" pageNumber="119" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2001) Water mites and other arachnids. Chapter 16. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 2 nd Edition. Academic Press, Waltham, Massachusetts, pp. 551 - 659. [1056 pp.] http: // dx. doi. org / 10.1016 / B 978 - 012690647 - 9 / 50017 - X" type="book chapter" year="2001">
Smith
<emphasis id="B907EAC0FF80176FFC7F6BBCA9CBFE78" box="[932,983,476,501]" italics="true" pageId="8" pageNumber="119">et al</emphasis>
. (2001
</bibRefCitation>
, 2010) treated this clade as the subfamily Foreliinae. Gerecke (2014) proposed placing
<taxonomicName id="4C734D51FF80176FFCE56860A9D5FD95" box="[830,969,513,536]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFCE56860A9D5FD95" box="[830,969,513,536]" italics="true" pageId="8" pageNumber="119">Pionacercus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D51FF80176FFBCA6860AEA1FD95" box="[1041,1213,513,536]" class="Arachnida" family="Pionidae" genus="Pionacercopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFBCA6860AEA1FD95" box="[1041,1213,513,536]" italics="true" pageId="8" pageNumber="119">Pionacercopsis</emphasis>
</taxonomicName>
in the Foreliinae, apparently unaware that
<bibRefCitation id="EFE24B23FF80176FFE6B6845A85AFDB0" author="Smith" box="[432,582,548,573]" pageId="8" pageNumber="119" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith (1976)</bibRefCitation>
had already assigned them to Foreliini. Here we follow Gerecke (2014) in considering
<taxonomicName id="4C734D51FF80176FFEFB6828ABB7FDED" box="[288,427,585,608]" class="Arachnida" family="Pionidae" genus="Pionacercus" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFEFB6828ABB7FDED" box="[288,427,585,608]" italics="true" pageId="8" pageNumber="119">Pionacercus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D51FF80176FFE396828A892FDED" box="[482,654,585,608]" class="Arachnida" family="Pionidae" genus="Pionacercopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF80176FFE396828A892FDED" box="[482,654,585,608]" italics="true" pageId="8" pageNumber="119">Pionacercopsis</emphasis>
</taxonomicName>
as separate genera.
</paragraph>
</subSubSection>
</treatment>
</document>