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<mods:title id="6FE9100306D7611A1359FA9BD4B8F7F6">Taxonomic studies on the genus Delphinium (Ranunculaceae) from China (XXVI): Notes on D. altissimum and D. scabriflorum, with description of a new species, D. pseudoscabriflorum</mods:title>
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<mods:namePart id="E18F07F157E2DEDC4F8B9D3EDD9681FC">Yuan, Qiong</mods:namePart>
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<mods:namePart id="C80F2E5D6B60E7138C37260722D30E2F">Yang, Qin-Er</mods:namePart>
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<mods:affiliation id="896472114B3BC9D837765143A1327D3B">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, Guangdong, China. &amp; Center of Conservation Biology, Core Botanical Gardens, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, Guangdong, China.</mods:affiliation>
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<taxonomicName id="4C074D4DFFCBFFEF3F97FF61FD70212C" authority="Q. Yuan &amp; Q.E. Yang" authorityName="Q. Yuan &amp; Q. E. Yang" authorityYear="2023" box="[136,764,244,271]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="29" phylum="Tracheophyta" rank="species" species="pseudoscabriflorum" status="sp. nov.">
<emphasis id="B973EADCFFCBFFEF3F97FF61FD8C212C" bold="true" box="[136,512,244,271]" italics="true" pageId="8" pageNumber="29">Delphinium pseudoscabriflorum</emphasis>
Q. Yuan &amp; Q.E. Yang
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, 
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<figureCitation id="133C2A4BFFCBFFEF3C7BFF61FC64212C" box="[868,1000,244,271]" captionStart="FIGURE 1" captionStartId="1.[136,229,1921,1943]" captionTargetBox="[202,1386,245,1896]" captionTargetId="figure-53@1.[202,1386,245,1896]" captionTargetPageId="1" captionText="FIGURE 1. Selected specimens from Bhutan cited as Delphinium scabriflorum by Munz (1968). A. R.E. Cooper 4114 (BM). B. R.E. Cooper 4114 (E). C. R.E. Cooper 4223 (BM). D. R.E. Cooper 4223 (E). Note that only the two sheets of the collection R.E. Cooper 4114 (A, B) belong to D. scabriflorum while the two sheets of the collection R.E. Cooper 4223 (C, D) belong to D. pseudoscabriflorum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10361855" httpUri="https://zenodo.org/record/10361855/files/figure.png" pageId="8" pageNumber="29">Figs. 1C, D</figureCitation>
, 
<figureCitation id="133C2A4BFFCBFFEF3CECFF61FB8F212D" box="[1011,1027,244,270]" captionStart="FIGURE 2" captionStartId="2.[136,229,1952,1974]" captionTargetBox="[170,1417,190,1928]" captionTargetId="figure-20@2.[170,1417,190,1928]" captionTargetPageId="2" captionText="FIGURE 2. Four specimens from Bhutan cited as Delphinium scabriflorum by Munz (1968).A. F. Ludlow &amp; G. Sherriff 219 (BM). B. F. Ludlow &amp; G. Sherriff 219 (E). C, D. F. Ludlow &amp; G. Sherriff 3342 (BM). Note that they should all belong to D. pseudoscabriflorum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10361861" httpUri="https://zenodo.org/record/10361861/files/figure.png" pageId="8" pageNumber="29">2</figureCitation>
, 
<figureCitation id="133C2A4BFFCBFFEF3B10FF61FBB0212D" box="[1039,1084,244,270]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="4.[136,229,1114,1136]" captionStartId-1="6.[136,229,1952,1974]" captionStartId-2="7.[136,229,1952,1974]" captionStartId-3="8.[136,229,1933,1955]" captionTargetBox-0="[145,1442,190,1089]" captionTargetBox-1="[170,1417,190,1928]" captionTargetBox-2="[170,1417,190,1928]" captionTargetBox-3="[170,1417,436,1909]" captionTargetId-0="figure-20@4.[145,1442,190,1089]" captionTargetId-1="figure-20@6.[170,1417,190,1928]" captionTargetId-2="figure-17@7.[170,1417,190,1928]" captionTargetId-3="figure-91@8.[170,1417,436,1909]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 4. Two specimens from Bhutan identified as Delphinium scabriflorum by Grierson &amp; Long, authors of the Flora of Bhutan, with Grierson as author of the Ranunculaceae. A. J.C. White s.n. (E). B. B.J. Gould 724 (E). Note that both should belong to D. pseudoscabriflorum sp. nov." captionText-1="FIGURE 5. Holotype (A) and isotype (B–D) sheets of Delphinium pseudoscabriflorum sp. nov." captionText-2="FIGURE 6. Isotype (A–D) sheets of Delphinium pseudoscabriflorum sp. nov." captionText-3="FIGURE 7. Delphinium pseudoscabriflorum sp. nov. A. Rootstock. B. Stem (left: proximal part; right: middle part). C. Leaf. D. Portion of leaf (adaxial side). E. Portion of leaf (abaxial side). F. Inflorescence rachis. G. Pedicel. H. Bracteoles. I. Flower (lateral view). J. Flower (close-up). K. Sepals. L. Petals. M. Staminodes. N. Stamens. O, P. Carpels. Q. Follicles." figureDoi-0="http://doi.org/10.5281/zenodo.10361867" figureDoi-1="http://doi.org/10.5281/zenodo.10361871" figureDoi-2="http://doi.org/10.5281/zenodo.10361875" figureDoi-3="http://doi.org/10.5281/zenodo.10361877" httpUri-0="https://zenodo.org/record/10361867/files/figure.png" httpUri-1="https://zenodo.org/record/10361871/files/figure.png" httpUri-2="https://zenodo.org/record/10361875/files/figure.png" httpUri-3="https://zenodo.org/record/10361877/files/figure.png" pageId="8" pageNumber="29">4–7</figureCitation>
, 
<figureCitation id="133C2A4BFFCBFFEF3B58FF61FBEA212D" box="[1095,1126,244,270]" captionStart="FIGURE 17" captionStartId="17.[136,229,1079,1101]" captionTargetBox="[170,1417,190,1055]" captionTargetId="figure-17@17.[170,1417,190,1055]" captionTargetPageId="17" captionText="FIGURE 17. Two specimens from Bhutan identified as Delphinium ludlowii but actually belonging to D. pseudoscabriflorum sp. nov.A. F. Ludlow &amp; G. Sherriff 21303 (BM). B. S.B. Lyon 3480 (BM)." figureDoi="http://doi.org/10.5281/zenodo.10361912" httpUri="https://zenodo.org/record/10361912/files/figure.png" pageId="8" pageNumber="29">17</figureCitation>
.
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Type:— 
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, alt. ca. 
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&amp; 
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WL3157
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<figureCitation id="133C2A4BFFCBFFEF3C93FEF4FC422154" box="[908,974,353,375]" captionStart="FIGURE 5" captionStartId="6.[136,229,1952,1974]" captionTargetBox="[170,1417,190,1928]" captionTargetId="figure-20@6.[170,1417,190,1928]" captionTargetPageId="6" captionText="FIGURE 5. Holotype (A) and isotype (B–D) sheets of Delphinium pseudoscabriflorum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10361871" httpUri="https://zenodo.org/record/10361871/files/figure.png" pageId="8" pageNumber="29">Figs. 5</figureCitation>
, 
<figureCitation id="133C2A4BFFCBFFEF3CC6FEF4FC6B2154" box="[985,999,353,375]" captionStart="FIGURE 6" captionStartId="7.[136,229,1952,1974]" captionTargetBox="[170,1417,190,1928]" captionTargetId="figure-17@7.[170,1417,190,1928]" captionTargetPageId="7" captionText="FIGURE 6. Isotype (A–D) sheets of Delphinium pseudoscabriflorum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10361875" httpUri="https://zenodo.org/record/10361875/files/figure.png" pageId="8" pageNumber="29">6</figureCitation>
.
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<paragraph id="8BB836CEFFCBFFEF3F97FE10FB6B21B9" blockId="8.[136,1451,318,410]" box="[136,1255,388,410]" pageId="8" pageNumber="29">
= 
<taxonomicName id="4C074D4DFFCBFFEF3F83FE11FE0321BA" ID-CoL="34JMK" authorityName=", Hara" authorityYear="1975" box="[156,399,388,409]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="29" phylum="Tracheophyta" rank="species" species="scabriflorum">
<emphasis id="B973EADCFFCBFFEF3F83FE11FE0321BA" box="[156,399,388,409]" italics="true" pageId="8" pageNumber="29">Delphinium scabriflorum</emphasis>
</taxonomicName>
auct. non 
<bibRefCitation id="EF964B3FFFCBFFEF3EECFE10FD1A21B9" author="Don, D." box="[499,662,389,410]" pageId="8" pageNumber="29" pagination="407 - 415" refId="ref5409" refString="Don, D. (1821) Descriptions of several new plants from the Kingdom of Nepaul, taken from specimens preserved in the Herbarium of Aylmer Bourke Lambert, Esq. Memoirs of the Wernerian Natural History Society 3: 407 - 415." type="journal article" year="1821">Don (1821: 412)</bibRefCitation>
: 
<bibRefCitation id="EF964B3FFFCBFFEF3DBFFE10FC9321B9" author="Munz, P. A." box="[672,799,388,410]" pageId="8" pageNumber="29" pagination="73 - 166" refId="ref5784" refString="Munz, P. A. (1968) A synopsis of the Asian species of Delphinium, sensu stricto. Journal of the Arnold Arboretum 49: 73 - 166. https: // doi. org / 10.5962 / p. 185735" type="journal article" year="1968">Munz (1968)</bibRefCitation>
, p.p., excl. 
<emphasis id="B973EADCFFCBFFEF3C91FE10FBB521B9" box="[910,1081,389,410]" italics="true" pageId="8" pageNumber="29">R.E. Cooper 4114</emphasis>
; 
<bibRefCitation id="EF964B3FFFCBFFEF3B59FE11FB6E21B9" author="Grierson, A. J. C." box="[1094,1250,388,410]" pageId="8" pageNumber="29" pagination="309 - 314" refId="ref5474" refString="Grierson, A. J. C. (1984) Delphinium L. In: Grierson, A. J. C. &amp; Long, D. G. (Eds.) Flora of Bhutan, vol. 1, part 2. Royal Botanical Garden, Edinburgh, pp. 309 - 314." type="book chapter" year="1984">Grierson (1984)</bibRefCitation>
.
</paragraph>
<caption id="DF786646FFCBFFEF3F97F818FC5A27C8" ID-DOI="http://doi.org/10.5281/zenodo.10361877" ID-Zenodo-Dep="10361877" httpUri="https://zenodo.org/record/10361877/files/figure.png" pageId="8" pageNumber="29" startId="8.[136,229,1933,1955]" targetBox="[170,1417,436,1909]" targetPageId="8" targetType="figure">
<paragraph id="8BB836CEFFCBFFEF3F97F818FC5A27C8" blockId="8.[136,1452,1933,2027]" pageId="8" pageNumber="29">
<emphasis id="B973EADCFFCBFFEF3F97F818FF722781" bold="true" box="[136,254,1933,1955]" pageId="8" pageNumber="29">FIGURE 7.</emphasis>
<emphasis id="B973EADCFFCBFFEF3E1BF818FD002780" box="[260,652,1933,1955]" italics="true" pageId="8" pageNumber="29">
<taxonomicName id="4C074D4DFFCBFFEF3E1BF818FDB02781" authority="Yuan &amp; Yang, 2023" authorityName="Q. Yuan &amp; Q. E. Yang" authorityYear="2023" box="[260,572,1933,1954]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="29" phylum="Tracheophyta" rank="species" species="pseudoscabriflorum" status="sp. nov.">Delphinium pseudoscabriflorum</taxonomicName>
<taxonomicNameLabel id="A24057A7FFCBFFEF3D5CF81BFD002780" box="[579,652,1934,1955]" pageId="8" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Rootstock. B. Stem (left: proximal part; right: middle part). C. Leaf. D. Portion of leaf (adaxial side). E. Portion of leaf (abaxial side). F. Inflorescence rachis. G. Pedicel. H. Bracteoles. I. Flower (lateral view). J. Flower (close-up). K. Sepals. L. Petals. M. Staminodes. N. Stamens. O, P. Carpels. Q. Follicles.
</paragraph>
</caption>
<paragraph id="8BB836CEFFCAFFEE3F97FF35FAA920F4" blockId="9.[136,1452,159,286]" pageId="9" pageNumber="30">
= 
<taxonomicName id="4C074D4DFFCAFFEE3F84FF0AFC7D2096" ID-CoL="5NHFY" authority="(Bruhl ex Huth 1895: 419) Wang &amp; Warnock" authorityName="Wang &amp; Warnock" baseAuthorityName="Bruhl ex Huth" baseAuthorityPageNumber="419" baseAuthorityYear="1895" box="[155,1009,159,181]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="9" pageNumber="30" phylum="Tracheophyta" rank="variety" species="umbrosum" variety="drepanocentrum">
<emphasis id="B973EADCFFCAFFEE3F84FF0AFEF92097" box="[155,373,159,180]" italics="true" pageId="9" pageNumber="30">Delphinium umbrosum</emphasis>
var. 
<emphasis id="B973EADCFFCAFFEE3EBCFF0AFDCD2097" box="[419,577,159,180]" italics="true" pageId="9" pageNumber="30">drepanocentrum</emphasis>
(Brühl ex 
<bibRefCitation id="EF964B3FFFCAFFEE3DBAFF0AFCB72097" author="Huth, E." box="[677,827,159,181]" pageId="9" pageNumber="30" pagination="322 - 499" refId="ref5726" refString="Huth, E. (1895) Monographie der Gattung Delphinium. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 20: 322 - 499. https: // doi. org / 10.5962 / bhl. title. 15558" type="journal article" year="1895">Huth 1895: 419</bibRefCitation>
) Wang &amp; Warnock
</taxonomicName>
in 
<bibRefCitation id="EF964B3FFFCAFFEE3B12FF35FB7C2097" author="Wang, W. T. &amp; Warnock, M. &amp; Zhu, G." box="[1037,1264,159,181]" pageId="9" pageNumber="30" pagination="382 - 388" refId="ref6231" refString="Wang, W. T., Warnock, M. &amp; Zhu, G. (1995) Notulae de la Ranunculaceis sinensibus (XX). Phytologia 79: 382 - 388." type="journal article" year="1995">
Wang 
<emphasis id="B973EADCFFCAFFEE3B55FF35FBFF2097" box="[1098,1139,159,181]" italics="true" pageId="9" pageNumber="30">et al</emphasis>
. (1995: 384)
</bibRefCitation>
; 
<bibRefCitation id="EF964B3FFFCAFFEE3BE6FF35FA2B2097" author="Wang, W. T." box="[1273,1447,159,181]" pageId="9" pageNumber="30" pagination="1 - 254" refId="ref6074" refString="Wang, W. T. (2020) A revision of the genus Delphinium (Ranunculaceae) of China (II). Guihaia 40 (Suppl.): 1 - 254. https: // doi. org / 10.3847 / 1538 - 4365 / aba 190" type="journal article" year="2020">Wang (2020: 147)</bibRefCitation>
, p.p., quoad 
<emphasis id="B973EADCFFCAFFEE3E31FF57FD2C20F4" box="[302,672,194,216]" italics="true" pageId="9" pageNumber="30">
<collectingRegion id="49C3F82CFFCAFFEE3E31FF57FEF120F4" box="[302,381,194,215]" country="China" name="Qinghai" pageId="9" pageNumber="30">Qinghai</collectingRegion>
- 
<collectingRegion id="49C3F82CFFCAFFEE3E9AFF56FE4420FB" box="[389,456,195,216]" country="China" name="Xizang" pageId="9" pageNumber="30">Xizang</collectingRegion>
Suppl. Exped. 751642
</emphasis>
, 
<emphasis id="B973EADCFFCAFFEE3DB3FF57FB9220F4" box="[684,1054,194,216]" italics="true" pageId="9" pageNumber="30">
<collectingRegion id="49C3F82CFFCAFFEE3DB3FF57FD7720F4" box="[684,763,194,215]" country="China" name="Qinghai" pageId="9" pageNumber="30">Qinghai</collectingRegion>
- 
<collectingRegion id="49C3F82CFFCAFFEE3C1CFF56FCCA20FB" box="[771,838,195,216]" country="China" name="Xizang" pageId="9" pageNumber="30">Xizang</collectingRegion>
Suppl. Exped. 751720
</emphasis>
[see 
<bibRefCitation id="EF964B3FFFCAFFEE3B4FFF56FA9720F4" author="Yuan, Q. &amp; Yang, Q. E." box="[1104,1307,194,216]" pageId="9" pageNumber="30" pagination="243 - 258" refId="ref6317" refString="Yuan, Q. &amp; Yang, Q. E. (2022) Taxonomic studies on the genus Delphinium (Ranunculaceae) from China (XXII): Clarifying morphological distinction between D. drepanocentrum and D. umbrosum and synonymizing D. umbrosoides with D. drepanocentrum. Phytotaxa 572: 243 - 258. https: // doi. org / 10.11646 / phytotaxa. 572.3.3" type="journal article" year="2022">Yuan &amp; Yang (2022)</bibRefCitation>
].
</paragraph>
<paragraph id="8BB836CEFFCAFFEE3F97FF73FEC8213D" blockId="9.[136,1452,159,286]" pageId="9" pageNumber="30">
<treatmentCitationGroup id="AB1711E0FFCAFFEE3F97FF73FEC8213D" pageId="9" pageNumber="30">
= 
<taxonomicName id="4C074D4DFFCAFFEE3F84FF70FD7120D8" ID-CoL="34J6H" authority="Wang in Wang &amp; Li (1987: 34)" box="[155,765,229,251]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="9" pageNumber="30" phylum="Tracheophyta" rank="species" species="conaense">
<emphasis id="B973EADCFFCAFFEE3F84FF70FEE020D8" box="[155,364,229,251]" italics="true" pageId="9" pageNumber="30">Delphinium conaense</emphasis>
auct. non Wang in 
<treatmentCitation id="0AA610DFFFCAFFEE3D3BFF73FD7120D8" author="Wang, W. T. &amp; Li, L. Q." box="[548,765,229,251]" page="34" pageId="9" pageNumber="30" year="1987">
<bibRefCitation id="EF964B3FFFCAFFEE3D3BFF73FD7120D8" author="Wang, W. T. &amp; Li, L. Q." box="[548,765,229,251]" pageId="9" pageNumber="30" pagination="24 - 38" refId="ref6124" refString="Wang, W. T. &amp; Li, L. Q. (1987) Notulae de la Ranunculaceis sinensibus (X). Acta Phytotaxonomica Sinica 25: 24 - 38." type="journal article" year="1987">Wang &amp; Li (1987: 34)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
: 
<treatmentCitation id="0AA610DFFFCAFFEE3C19FF73FC3920D8" author="Wang, W. T." box="[774,949,229,251]" page="165" pageId="9" pageNumber="30" year="2020">
<bibRefCitation id="EF964B3FFFCAFFEE3C19FF73FC3920D8" author="Wang, W. T." box="[774,949,229,251]" pageId="9" pageNumber="30" pagination="1 - 254" refId="ref6074" refString="Wang, W. T. (2020) A revision of the genus Delphinium (Ranunculaceae) of China (II). Guihaia 40 (Suppl.): 1 - 254. https: // doi. org / 10.3847 / 1538 - 4365 / aba 190" type="journal article" year="2020">Wang (2020: 165)</bibRefCitation>
</treatmentCitation>
, p.p., quoad 
<emphasis id="B973EADCFFCAFFEE3B30FF73FABD20D9" box="[1071,1329,229,251]" italics="true" pageId="9" pageNumber="30">
PE 
<collectingRegion id="49C3F82CFFCAFFEE3B4CFF73FB1A20D8" box="[1107,1174,230,251]" country="China" name="Xizang" pageId="9" pageNumber="30">Xizang</collectingRegion>
Exped. PE6212
</emphasis>
[see 
<bibRefCitation id="EF964B3FFFCAFFEE3A7DFF73FEB6213D" author="Yuan, Q. &amp; Yang, Q. E." pageId="9" pageNumber="30" pagination="291 - 300" refId="ref6386" refString="Yuan, Q. &amp; Yang, Q. E. (2023) Taxonomic studies on the genus Delphinium (Ranunculaceae) from China (XXIII): Reduction of D. conaense to the synonymy D. bhutanicum, a species newly reported for China. Phytotaxa 599: 291 - 300. https: // doi. org / 10.11646 / phytotaxa. 599.5.3" type="journal article" year="2023">Yuan &amp; Yang (2023)</bibRefCitation>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF786646FFCAFFEE3F97FB41FD70252D" ID-DOI="http://doi.org/10.5281/zenodo.10361880" ID-Zenodo-Dep="10361880" httpUri="https://zenodo.org/record/10361880/files/figure.png" pageId="9" pageNumber="30" startId="9.[136,229,1236,1258]" targetBox="[145,1442,312,1211]" targetPageId="9" targetType="figure">
<paragraph id="8BB836CEFFCAFFEE3F97FB41FD70252D" blockId="9.[136,1452,1236,1294]" pageId="9" pageNumber="30">
<emphasis id="B973EADCFFCAFFEE3F97FB41FF7424CA" bold="true" box="[136,248,1236,1258]" pageId="9" pageNumber="30">FIGURE 8</emphasis>
. Two specimens from Nepal identified or cited as 
<taxonomicName id="4C074D4DFFCAFFEE3DFCFB40FC5B24C9" authorityName=", Hara" authorityYear="1975" box="[739,983,1237,1258]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="9" pageNumber="30" phylum="Tracheophyta" rank="species" species="scabriflorum">
<emphasis id="B973EADCFFCAFFEE3DFCFB40FC5B24C9" box="[739,983,1237,1258]" italics="true" pageId="9" pageNumber="30">Delphinium scabriflorum</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF964B3FFFCAFFEE3CE3FB40FBFE24C9" author="Hara, H." box="[1020,1138,1236,1258]" pageId="9" pageNumber="30" pagination="263 - 271" refId="ref5532" refString="Hara, H. (1975) New or noteworthy flowering plants from Eastern Himalaya (16). The Journal of Japanese Botany 50: 263 - 271." type="journal article" year="1975">Hara (1975)</bibRefCitation>
and Yonekura (2018). A. 
<emphasis id="B973EADCFFCAFFEE3A70FB40FEC8252D" italics="true" pageId="9" pageNumber="30">J.D.A. Stainton et al. 7521</emphasis>
(BM). B. 
<emphasis id="B973EADCFFCAFFEE3EB8FB6CFD13252E" box="[423,671,1272,1294]" italics="true" pageId="9" pageNumber="30">M. Mikage et al. 9682513</emphasis>
(KATH).
</paragraph>
</caption>
<subSubSection id="C31D6545FFCAFFEE3F97FAAEFAF82746" pageId="9" pageNumber="30" type="description">
<paragraph id="8BB836CEFFCAFFEE3F97FAAEFAF82746" blockId="9.[136,1452,1339,1894]" pageId="9" pageNumber="30">
<emphasis id="B973EADCFFCAFFEE3F97FAAEFE9E2576" bold="true" box="[136,274,1339,1365]" pageId="9" pageNumber="30">Description</emphasis>
:—Perennial herbs. Rootstock thickened, woody. Stems 
<quantity id="4CFF9B2BFFCAFFEE3C8AFAAEFB852575" box="[917,1033,1339,1366]" metricMagnitude="-1" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="9" pageNumber="30" unit="m" value="0.85" valueMax="1.2" valueMin="0.5">0.5–1.2 m</quantity>
or more tall, more or less spreading puberulent throughout or at least above, scattered-leafy; leaves pentagonal, the blades of the lower 
<quantity id="4CFF9B2BFFCAFFEE3BE7FACBFAD7255A" box="[1272,1371,1374,1401]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="9" pageNumber="30" unit="cm" value="10.0" valueMax="15.0" valueMin="5.0">5–15 cm</quantity>
broad, open-cordate, 5-fid to near the base or 
<quantity id="4CFF9B2BFFCAFFEE3D5DFA17FD0325BF" box="[578,655,1410,1437]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="30" unit="cm" value="0.5">0.5 cm</quantity>
above, the segments cuneate below, stiff-pubescent on both sides, but especially on nerves beneath, the segments incised-lobed then coarsely toothed. the lobules oblong-ovate, 
<quantity id="4CFF9B2BFFCAFFEE3A53FA30FA20259C" box="[1356,1452,1445,1472]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="9" pageNumber="30" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
or more wide; petioles more or less spreading puberulent, 
<quantity id="4CFF9B2BFFCAFFEE3C32FA5DFC1325C0" box="[813,927,1480,1507]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="30" unit="cm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 cm</quantity>
long, scarcely vaginate at base; upper leaves gradually reduced. Inflorescence often amply paniculate, central raceme open, 8–12-flowered, with fewer-flowered branches from below; rachis densely more or less spreading puberulent, some of the hairs glandular; lower bracts foliose, divided, to 
<quantity id="4CFF9B2BFFCAFFEE3E7CF9A7FE17266F" box="[355,411,1586,1612]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="30" unit="cm" value="3.0">3 cm</quantity>
long, the upper simple, entire, linear to lanceolate, 
<quantity id="4CFF9B2BFFCAFFEE3CFCF9A7FBBB266F" box="[995,1079,1586,1612]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="30" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
long; pedicels arched-spreading, 
<quantity id="4CFF9B2BFFCAFFEE3F97F9C0FF50264C" box="[136,220,1621,1648]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="9" pageNumber="30" unit="cm" value="3.0" valueMax="5.0" valueMin="1.0">1–5 cm</quantity>
long, densely more or less spreading puberulent, some of the hairs glandular; bracteoles linear to lanceolate, ca. 
<quantity id="4CFF9B2BFFCAFFEE3FACF9EDFF6126B0" box="[179,237,1656,1683]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="30" unit="cm" value="1.0">1 cm</quantity>
long, abaxially stiff-puberulent, closely appressed to the flower. Flowers blue to rich blue-violet; sepals densely stiff-puberulent, the upper sepal round-ovate, ca. 
<quantity id="4CFF9B2BFFCAFFEE3C0AF90EFCEE2695" box="[789,866,1691,1718]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="30" unit="cm" value="1.5">1.5 cm</quantity>
long, ca. 
<quantity id="4CFF9B2BFFCAFFEE3CCBF90EFB802695" box="[980,1036,1691,1718]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="30" unit="cm" value="1.0">1 cm</quantity>
broad, subscute, the spur ca. 
<quantity id="4CFF9B2BFFCAFFEE3A41F90EFA272695" box="[1374,1451,1691,1718]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="30" unit="cm" value="1.8">1.8 cm</quantity>
long, ca. 
<quantity id="4CFF9B2BFFCAFFEE3FEFF92BFEBE26FA" box="[240,306,1726,1753]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="30" unit="mm" value="5.0">5 mm</quantity>
broad at base, lateral sepals oblong-obovate, ca. 
<quantity id="4CFF9B2BFFCAFFEE3C43F92AFC2426FA" box="[860,936,1727,1753]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="30" unit="cm" value="1.8">1.8 cm</quantity>
long, ca. 
<quantity id="4CFF9B2BFFCAFFEE3B07F92BFBD526FA" box="[1048,1113,1726,1753]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="30" unit="mm" value="5.0">5 mm</quantity>
broad, subacute, lower sepals asymmetrically obovate, ca. 
<quantity id="4CFF9B2BFFCAFFEE3EDAF977FD8326DF" box="[453,527,1762,1788]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="30" unit="cm" value="1.8">1.8 cm</quantity>
long, ca. 
<quantity id="4CFF9B2BFFCAFFEE3D66F977FD3426DF" box="[633,696,1762,1789]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="30" unit="mm" value="5.0">5 mm</quantity>
broad, subacute; petals blue, the laminae oblique, ca. 
<quantity id="4CFF9B2BFFCAFFEE3A16F977FAC426DF" box="[1289,1352,1762,1788]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="30" unit="mm" value="8.0">8 mm</quantity>
long, ca. 
<quantity id="4CFF9B2BFFCAFFEE3F97F890FF44273C" box="[136,200,1797,1823]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="30" unit="mm" value="3.0">3 mm</quantity>
wide, glabrous, bidentate, the spur ca. 
<quantity id="4CFF9B2BFFCAFFEE3D68F890FD4F273C" box="[631,707,1797,1824]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="30" unit="cm" value="1.5">1.5 cm</quantity>
long; staminodes oblong, ca. 
<quantity id="4CFF9B2BFFCAFFEE3B11F890FBC8273C" box="[1038,1092,1797,1823]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="30" unit="cm" value="1.0">1 cm</quantity>
long, 
<quantity id="4CFF9B2BFFCAFFEE3B96F890FB52273C" box="[1161,1246,1797,1824]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="30" unit="mm" value="3.5">3.5 mm</quantity>
broad, bifid into 2 unequal lanceolate-oblong lobes, white ciliate, the claw ca. 
<quantity id="4CFF9B2BFFCAFFEE3C01F8BDFCD22760" box="[798,862,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="30" unit="mm" value="6.0">6 mm</quantity>
long, white bearded; stamens ca. 
<quantity id="4CFF9B2BFFCAFFEE3BCAF8BDFA992760" box="[1237,1301,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="30" unit="mm" value="8.0">8 mm</quantity>
long; anthers elongate, 
<quantity id="4CFF9B2BFFCAFFEE3FEAF8DEFEC72745" box="[245,331,1867,1894]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="30" unit="mm" value="1.5">1.5 mm</quantity>
long; carpels 3, ventrally puberulent, more densely so on styles. Follicles ventrally puberulent.
</paragraph>
</subSubSection>
<caption id="DF786646FFC9FFED3F97F835FCF127F9" ID-DOI="http://doi.org/10.5281/zenodo.10361884" ID-Zenodo-Dep="10361884" httpUri="https://zenodo.org/record/10361884/files/figure.png" pageId="10" pageNumber="31" startId="10.[136,229,1952,1974]" targetBox="[170,1417,190,1928]" targetPageId="10" targetType="figure">
<paragraph id="8BB836CEFFC9FFED3F97F835FCF127F9" blockId="10.[136,1451,1952,2010]" pageId="10" pageNumber="31">
<emphasis id="B973EADCFFC9FFED3F97F835FF7A2796" bold="true" box="[136,246,1952,1974]" pageId="10" pageNumber="31">FIGURE 9</emphasis>
. Four specimens from Bhutan cited as 
<taxonomicName id="4C074D4DFFC9FFED3D78F834FCCA2795" authorityName="Wallich" authorityYear="1831" box="[615,838,1953,1974]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="10" pageNumber="31" phylum="Tracheophyta" rank="species" species="altissimum">
<emphasis id="B973EADCFFC9FFED3D78F834FCCA2795" box="[615,838,1953,1974]" italics="true" pageId="10" pageNumber="31">Delphinium altissimum</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF964B3FFFC9FFED3C77F834FC6B2794" author="Munz, P. A." box="[872,999,1953,1975]" pageId="10" pageNumber="31" pagination="73 - 166" refId="ref5784" refString="Munz, P. A. (1968) A synopsis of the Asian species of Delphinium, sensu stricto. Journal of the Arnold Arboretum 49: 73 - 166. https: // doi. org / 10.5962 / p. 185735" type="journal article" year="1968">Munz (1968)</bibRefCitation>
. A. 
<emphasis id="B973EADCFFC9FFED3B14F834FB3A2795" box="[1035,1206,1953,1974]" italics="true" pageId="10" pageNumber="31">R.E. Cooper 2008</emphasis>
(E). B. 
<emphasis id="B973EADCFFC9FFED3A1FF834FA272795" box="[1280,1451,1953,1974]" italics="true" pageId="10" pageNumber="31">R.E. Cooper 2556</emphasis>
(E). C. 
<emphasis id="B973EADCFFC9FFED3FD0F850FE5227F9" box="[207,478,1989,2010]" italics="true" pageId="10" pageNumber="31">F. Ludlow &amp; G. Sherriff 298</emphasis>
(BM). D. 
<emphasis id="B973EADCFFC9FFED3D5DF850FCDD27F9" box="[578,849,1989,2010]" italics="true" pageId="10" pageNumber="31">F. Ludlow &amp; G. Sherriff 298</emphasis>
(E).
</paragraph>
</caption>
<caption id="DF786646FFC8FFEC3F97F834FCDF27F9" ID-DOI="http://doi.org/10.5281/zenodo.10361892" ID-Zenodo-Dep="10361892" httpUri="https://zenodo.org/record/10361892/files/figure.png" pageId="11" pageNumber="32" startId="11.[136,229,1953,1975]" targetBox="[170,1417,190,1929]" targetPageId="11" targetType="figure">
<paragraph id="8BB836CEFFC8FFEC3F97F834FCDF27F9" blockId="11.[136,1453,1953,2011]" pageId="11" pageNumber="32">
<emphasis id="B973EADCFFC8FFEC3F97F834FE8F2795" bold="true" box="[136,259,1953,1975]" pageId="11" pageNumber="32">FIGURE 10</emphasis>
. Four specimens from Bhutan cited as 
<taxonomicName id="4C074D4DFFC8FFEC3D67F834FCDB2795" authorityName="Wallich" authorityYear="1831" box="[632,855,1953,1974]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="11" pageNumber="32" phylum="Tracheophyta" rank="species" species="altissimum">
<emphasis id="B973EADCFFC8FFEC3D67F834FCDB2795" box="[632,855,1953,1974]" italics="true" pageId="11" pageNumber="32">Delphinium altissimum</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF964B3FFFC8FFEC3C65F837FC762794" author="Munz, P. A." box="[890,1018,1953,1975]" pageId="11" pageNumber="32" pagination="73 - 166" refId="ref5784" refString="Munz, P. A. (1968) A synopsis of the Asian species of Delphinium, sensu stricto. Journal of the Arnold Arboretum 49: 73 - 166. https: // doi. org / 10.5962 / p. 185735" type="journal article" year="1968">Munz (1968)</bibRefCitation>
. A. 
<emphasis id="B973EADCFFC8FFEC3B00F837FAB42795" box="[1055,1336,1953,1975]" italics="true" pageId="11" pageNumber="32">F. Ludlow &amp; G. Sherriff 3496</emphasis>
(BM). B. 
<emphasis id="B973EADCFFC8FFEC3A85F837FE0727F9" italics="true" pageId="11" pageNumber="32">F. Ludlow &amp; G. Sherriff 3496</emphasis>
(E). C, D. 
<emphasis id="B973EADCFFC8FFEC3EEAF853FC9C27F9" box="[501,784,1989,2011]" italics="true" pageId="11" pageNumber="32">F. Ludlow &amp; G. Sherriff 3558</emphasis>
(BM).
</paragraph>
</caption>
<caption id="DF786646FFCFFFEB3F97FBD1FB84245E" ID-DOI="http://doi.org/10.5281/zenodo.10361896" ID-Zenodo-Dep="10361896" httpUri="https://zenodo.org/record/10361896/files/figure.png" pageId="12" pageNumber="33" startId="12.[136,229,1092,1114]" targetBox="[201,1386,190,1067]" targetPageId="12" targetType="figure">
<paragraph id="8BB836CEFFCFFFEB3F97FBD1FB84245E" blockId="12.[136,1452,1092,1150]" pageId="12" pageNumber="33">
<emphasis id="B973EADCFFCFFFEB3F97FBD1FE8E247A" bold="true" box="[136,258,1092,1114]" pageId="12" pageNumber="33">FIGURE 11</emphasis>
. Two specimens from Bhutan identified as 
<taxonomicName id="4C074D4DFFCFFFEB3DBFFBD1FCF3247A" authorityName="Wallich" authorityYear="1831" box="[672,895,1092,1113]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="12" pageNumber="33" phylum="Tracheophyta" rank="species" species="altissimum">
<emphasis id="B973EADCFFCFFFEB3DBFFBD1FCF3247A" box="[672,895,1092,1113]" italics="true" pageId="12" pageNumber="33">Delphinium altissimum</emphasis>
</taxonomicName>
by Grierson &amp; Long, authors of the 
<emphasis id="B973EADCFFCFFFEB3BC4FBD1FAF9247A" box="[1243,1397,1092,1113]" italics="true" pageId="12" pageNumber="33">Flora of Bhutan</emphasis>
, with Grierson as author of the 
<taxonomicName id="4C074D4DFFCFFFEB3E62FBFDFD9D245D" box="[381,529,1128,1150]" class="Magnoliopsida" family="Ranunculaceae" kingdom="Plantae" order="Ranunculales" pageId="12" pageNumber="33" phylum="Tracheophyta" rank="family">Ranunculaceae</taxonomicName>
. A. 
<emphasis id="B973EADCFFCFFFEB3D29FBFDFD6F245E" box="[566,739,1128,1149]" italics="true" pageId="12" pageNumber="33">R.E. Cooper 4186</emphasis>
(E). B. 
<emphasis id="B973EADCFFCFFFEB3C2FFBFDFC51245E" box="[816,989,1128,1149]" italics="true" pageId="12" pageNumber="33">R.E. Cooper 4360</emphasis>
(E).
</paragraph>
</caption>
<caption id="DF786646FFCFFFEB3F97F873FDAE283C" ID-DOI="http://doi.org/10.5281/zenodo.10361898" ID-Zenodo-Dep="10361898" httpUri="https://zenodo.org/record/10361898/files/figure.png" pageId="12" pageNumber="33" startId="12.[136,229,2022,2044]" targetBox="[201,1386,1175,1996]" targetPageId="12" targetType="figure">
<paragraph id="8BB836CEFFCFFFEB3F97F873FDAE283C" blockId="12.[136,1451,2022,2079]" pageId="12" pageNumber="33">
<emphasis id="B973EADCFFCFFFEB3F97F873FE8827D8" bold="true" box="[136,260,2022,2044]" pageId="12" pageNumber="33">FIGURE 12</emphasis>
. A collection from Gyirong in southern Xizang, China cited as 
<taxonomicName id="4C074D4DFFCFFFEB3C79F873FBC927D8" authorityName="Wallich" authorityYear="1831" box="[870,1093,2022,2043]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="12" pageNumber="33" phylum="Tracheophyta" rank="species" species="altissimum">
<emphasis id="B973EADCFFCFFFEB3C79F873FBC927D8" box="[870,1093,2022,2043]" italics="true" pageId="12" pageNumber="33">Delphinium altissimum</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF964B3FFFCFFFEB3B76F873FB6D27D8" author="Wang, W. T." box="[1129,1249,2022,2043]" pageId="12" pageNumber="33" pagination="153 - 163" refId="ref6047" refString="Wang, W. T. (1983) Notulae de la Ranunculaceis sinensibus (VI). Acta Botanica Yunnanica 5: 153 - 163." type="journal article" year="1983">Wang (1983</bibRefCitation>
, 
<bibRefCitation id="EF964B3FFFCFFFEB3BF4F873FA9327D8" author="Wang, W. T." box="[1259,1311,2022,2043]" pageId="12" pageNumber="33" pagination="1 - 254" refId="ref6074" refString="Wang, W. T. (2020) A revision of the genus Delphinium (Ranunculaceae) of China (II). Guihaia 40 (Suppl.): 1 - 254. https: // doi. org / 10.3847 / 1538 - 4365 / aba 190" type="journal article" year="2020">2020</bibRefCitation>
). A. 
<emphasis id="B973EADCFFCFFFEB3A54F873FF5B283C" italics="true" pageId="12" pageNumber="33">C.C. Ni et al. 2305</emphasis>
(PE). B. 
<emphasis id="B973EADCFFCFFFEB3E2EF79FFE69283C" box="[305,485,2058,2079]" italics="true" pageId="12" pageNumber="33">C.C. Ni et al. 2305</emphasis>
(XZ).
</paragraph>
</caption>
<caption id="DF786646FFCEFFEA3F97FB83FCD8240F" ID-DOI="http://doi.org/10.5281/zenodo.10361900" ID-Zenodo-Dep="10361900" box="[136,852,1046,1068]" httpUri="https://zenodo.org/record/10361900/files/figure.png" pageId="13" pageNumber="34" startId="13.[136,229,1046,1068]" targetBox="[145,1443,190,1022]" targetPageId="13" targetType="figure">
<paragraph id="8BB836CEFFCEFFEA3F97FB83FCD8240F" blockId="13.[136,852,1046,1068]" box="[136,852,1046,1068]" pageId="13" pageNumber="34">
<emphasis id="B973EADCFFCEFFEA3F97FB83FE8F2408" bold="true" box="[136,259,1046,1068]" pageId="13" pageNumber="34">FIGURE 13</emphasis>
. Distribution of 
<emphasis id="B973EADCFFCEFFEA3EBCFB83FCA6240F" box="[419,810,1046,1068]" italics="true" pageId="13" pageNumber="34">
<taxonomicName id="4C074D4DFFCEFFEA3EBCFB83FD572408" authority="Yuan &amp; Yang, 2023" authorityName="Q. Yuan &amp; Q. E. Yang" authorityYear="2023" box="[419,731,1046,1067]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="13" pageNumber="34" phylum="Tracheophyta" rank="species" species="pseudoscabriflorum" status="sp. nov.">Delphinium pseudoscabriflorum</taxonomicName>
<taxonomicNameLabel id="A24057A7FFCEFFEA3DFEFB82FCA6240F" box="[737,810,1047,1068]" pageId="13" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(●).
</paragraph>
</caption>
<subSubSection id="C31D6545FFCEFFEA3FA2FBCCFEB72499" pageId="13" pageNumber="34" type="distribution">
<paragraph id="8BB836CEFFCEFFEA3FA2FBCCFEB72499" blockId="13.[136,1452,1113,1844]" pageId="13" pageNumber="34">
<emphasis id="B973EADCFFCEFFEA3FA2FBCCFE6A2450" bold="true" box="[189,486,1113,1139]" pageId="13" pageNumber="34">Distribution and habitat</emphasis>
:— 
<taxonomicName id="4C074D4DFFCEFFEA3D16FBCCFCFB2450" authorityName="Q. Yuan &amp; Q. E. Yang" authorityYear="2023" box="[521,887,1113,1139]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="13" pageNumber="34" phylum="Tracheophyta" rank="species" species="pseudoscabriflorum">
<emphasis id="B973EADCFFCEFFEA3D16FBCCFCFB2450" box="[521,887,1113,1139]" italics="true" pageId="13" pageNumber="34">Delphinium pseudoscabriflorum</emphasis>
</taxonomicName>
is distributed in 
<collectingCountry id="F310765EFFCEFFEA3B22FBCCFB022450" box="[1085,1166,1113,1139]" name="Bhutan" pageId="13" pageNumber="34">Bhutan</collectingCountry>
and the closely adjacent Cona county in southern 
<collectingRegion id="49C3F82CFFCEFFEA3EBDFBE9FE7A24B5" box="[418,502,1148,1174]" country="China" name="Xizang" pageId="13" pageNumber="34">Xizang</collectingRegion>
, 
<collectingCountry id="F310765EFFCEFFEA3D1EFBE9FDC824B5" box="[513,580,1148,1174]" name="China" pageId="13" pageNumber="34">China</collectingCountry>
(
<figureCitation id="133C2A4BFFCEFFEA3D4BFBE9FD2B24B4" box="[596,679,1148,1175]" captionStart="FIGURE 13" captionStartId="13.[136,229,1046,1068]" captionTargetBox="[145,1443,190,1022]" captionTargetId="figure-17@13.[145,1443,190,1022]" captionTargetPageId="13" captionText="FIGURE 13. Distribution of Delphinium pseudoscabriflorum sp. nov. (●)." figureDoi="http://doi.org/10.5281/zenodo.10361900" httpUri="https://zenodo.org/record/10361900/files/figure.png" pageId="13" pageNumber="34">Fig. 13</figureCitation>
). It grows in forests or at forest margins at altitudes of 
<quantity id="4CFF9B2BFFCEFFEA3A0FFBE9FA2024B4" box="[1296,1452,1148,1175]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="13" pageNumber="34" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">2400–2600 m</quantity>
above sea level.
</paragraph>
</subSubSection>
<subSubSection id="C31D6545FFCEFFEA3FA2FB56FB0E24FE" box="[189,1154,1219,1245]" pageId="13" pageNumber="34" type="biology_ecology">
<paragraph id="8BB836CEFFCEFFEA3FA2FB56FB0E24FE" blockId="13.[136,1452,1113,1844]" box="[189,1154,1219,1245]" pageId="13" pageNumber="34">
<emphasis id="B973EADCFFCEFFEA3FA2FB56FEB524FE" bold="true" box="[189,313,1219,1245]" pageId="13" pageNumber="34">Phenology</emphasis>
:—Flowering from July to September; fruiting from September to October.
</paragraph>
</subSubSection>
<subSubSection id="C31D6545FFCEFFEA3FA2FB73FE0C2639" pageId="13" pageNumber="34" type="materials_examined">
<paragraph id="8BB836CEFFCEFFEA3FA2FB73FA8A2564" blockId="13.[136,1452,1113,1844]" pageId="13" pageNumber="34">
<materialsCitation id="3B6F3C93FFCEFFEA3FA2FB73FEBF2500" collectionCode="E, BM" collectorName="B. J. Gould &amp; F. Ludlow &amp; G. Sherriff" country="Bhutan" location="BHUTAN" pageId="13" pageNumber="34" specimenCount="1" typeStatus="paratype">
<emphasis id="B973EADCFFCEFFEA3FA2FB73FD402523" bold="true" box="[189,716,1254,1280]" pageId="13" pageNumber="34">
Additional specimens examined (
<typeStatus id="54BC886CFFCEFFEA3D54FB73FD492523" box="[587,709,1254,1280]" pageId="13" pageNumber="34" type="paratype">paratypes</typeStatus>
)
</emphasis>
:— 
<collectingCountry id="F310765EFFCEFFEA3DF2FB73FCEA2523" box="[749,870,1254,1280]" name="Bhutan" pageId="13" pageNumber="34">BHUTAN</collectingCountry>
. 
<emphasis id="B973EADCFFCEFFEA3C6EFB72FB922523" box="[881,1054,1254,1280]" italics="true" pageId="13" pageNumber="34">
<collectorName id="26F25318FFCEFFEA3C6EFB72FC602523" box="[881,1004,1254,1280]" pageId="13" pageNumber="34">B.J. Gould</collectorName>
724
</emphasis>
(
<collectionCode id="ED16AE0BFFCEFFEA3B30FB73FBCF2523" box="[1071,1091,1254,1280]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="13" pageNumber="34" type="Herbarium">E</collectionCode>
), 
<emphasis id="B973EADCFFCEFFEA3B45FB72FA272523" box="[1114,1451,1254,1281]" italics="true" pageId="13" pageNumber="34">
<collectorName id="26F25318FFCEFFEA3B45FB72FB472523" box="[1114,1227,1254,1280]" pageId="13" pageNumber="34">F. Ludlow</collectorName>
&amp; 
<collectorName id="26F25318FFCEFFEA3BEDFB72FAE72523" box="[1266,1387,1254,1281]" pageId="13" pageNumber="34">G. Sherriff</collectorName>
3342
</emphasis>
(
<collectionCode id="ED16AE0BFFCEFFEA3F8FFA9CFF332500" box="[144,191,1289,1315]" country="United Kingdom" name="Bristol Museum" pageId="13" pageNumber="34">BM</collectionCode>
, 2 sheets)
</materialsCitation>
; 
<materialsCitation id="3B6F3C93FFCEFFEA3E5FFA9CFB132500" box="[320,1183,1289,1316]" collectionCode="BM, E" collectorName="R. E. Cooper &amp; F. Ludlow &amp; G. Sherriff" country="Bhutan" location="Lhuentse" pageId="13" pageNumber="34" specimenCount="1" stateProvince="Lhuentse">
<collectingRegion id="49C3F82CFFCEFFEA3E5FFA9CFE262500" box="[320,426,1289,1315]" country="Bhutan" name="Lhuntshi" pageId="13" pageNumber="34">Lhuentse</collectingRegion>
, 
<emphasis id="B973EADCFFCEFFEA3EA9FA9FFD0E2500" box="[438,642,1289,1316]" italics="true" pageId="13" pageNumber="34">
<collectorName id="26F25318FFCEFFEA3EA9FA9FFDCE2507" box="[438,578,1290,1316]" pageId="13" pageNumber="34">R.E. Cooper</collectorName>
4223
</emphasis>
(
<collectionCode id="ED16AE0BFFCEFFEA3D8DFA9CFD4D2500" box="[658,705,1289,1315]" country="United Kingdom" name="Bristol Museum" pageId="13" pageNumber="34">BM</collectionCode>
, 
<collectionCode id="ED16AE0BFFCEFFEA3DD1FA9CFD6E2500" box="[718,738,1289,1315]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="13" pageNumber="34" type="Herbarium">E</collectionCode>
), 
<emphasis id="B973EADCFFCEFFEA3DE7FA9FFBD92500" box="[760,1109,1289,1316]" italics="true" pageId="13" pageNumber="34">
<collectorName id="26F25318FFCEFFEA3DE7FA9FFCE42500" box="[760,872,1289,1315]" pageId="13" pageNumber="34">F. Ludlow</collectorName>
&amp; 
<collectorName id="26F25318FFCEFFEA3C91FA9FFB842500" box="[910,1032,1289,1316]" pageId="13" pageNumber="34">G. Sherriff</collectorName>
21303
</emphasis>
(
<collectionCode id="ED16AE0BFFCEFFEA3B79FA9CFB1B2500" box="[1126,1175,1289,1315]" country="United Kingdom" name="Bristol Museum" pageId="13" pageNumber="34">BM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B6F3C93FFCEFFEA3BB4FA9CFC812564" collectionCode="BM, E" collectorName="F. Ludlow &amp; G. Sherriff &amp; S. B. Lyon" country="Bhutan" location="Wangdue Phodrang" municipality="Ludlow" pageId="13" pageNumber="34" specimenCount="1" stateProvince="Wangdue Phodrang">
<collectingRegion id="49C3F82CFFCEFFEA3BB4FA9CFA062500" box="[1195,1418,1289,1315]" country="Bhutan" name="Wangdi Phodrang" pageId="13" pageNumber="34">Wangdue Phodrang</collectingRegion>
, 
<emphasis id="B973EADCFFCEFFEA3A88FA9FFE252564" italics="true" pageId="13" pageNumber="34">
<collectorName id="26F25318FFCEFFEA3A88FA9FFF502565" pageId="13" pageNumber="34">F. Ludlow</collectorName>
&amp; 
<collectorName id="26F25318FFCEFFEA3E1FFAB8FEF42565" box="[256,376,1324,1351]" pageId="13" pageNumber="34">G. Sherriff</collectorName>
219
</emphasis>
(
<collectionCode id="ED16AE0BFFCEFFEA3EA7FAB8FE6B2564" box="[440,487,1325,1351]" country="United Kingdom" name="Bristol Museum" pageId="13" pageNumber="34">BM</collectionCode>
, 
<collectionCode id="ED16AE0BFFCEFFEA3EECFAB8FD8B2564" box="[499,519,1325,1351]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="13" pageNumber="34" type="Herbarium">E</collectionCode>
), 
<emphasis id="B973EADCFFCEFFEA3D04FAB8FD492564" box="[539,709,1325,1351]" italics="true" pageId="13" pageNumber="34">
<collectorName id="26F25318FFCEFFEA3D04FAB8FD0A2565" box="[539,646,1325,1351]" pageId="13" pageNumber="34">S.B. Lyon</collectorName>
3480
</emphasis>
(
<collectionCode id="ED16AE0BFFCEFFEA3DCAFAB8FC8A2564" box="[725,774,1325,1351]" country="United Kingdom" name="Bristol Museum" pageId="13" pageNumber="34">BM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B6F3C93FFCEFFEA3C06FAB9FA8E2564" box="[793,1282,1324,1351]" collectionCode="E" collectorName="J. C. White" country="Bhutan" location="Without" pageId="13" pageNumber="34" specimenCount="1">
Without precise locality, 
<emphasis id="B973EADCFFCEFFEA3B2EFAB8FB582565" box="[1073,1236,1324,1351]" italics="true" pageId="13" pageNumber="34">
<collectorName id="26F25318FFCEFFEA3B2EFAB8FB2A2565" box="[1073,1190,1324,1351]" pageId="13" pageNumber="34">J.C. White</collectorName>
s.n.
</emphasis>
(
<collectionCode id="ED16AE0BFFCEFFEA3BFBFAB8FB752564" box="[1252,1273,1325,1351]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="13" pageNumber="34" type="Herbarium">E</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB836CEFFCEFFEA3FA2FAC5FE0C2639" blockId="13.[136,1452,1113,1844]" pageId="13" pageNumber="34">
<collectingCountry id="F310765EFFCEFFEA3FA2FAC5FE952549" box="[189,281,1360,1386]" name="China" pageId="13" pageNumber="34">CHINA</collectingCountry>
. 
<collectingRegion id="49C3F82CFFCEFFEA3E37FAC5FEF12549" box="[296,381,1360,1386]" country="China" name="Xizang" pageId="13" pageNumber="34">Xizang</collectingRegion>
: Cona, 
<emphasis id="B973EADCFFCEFFEA3EC6FAC5FD482549" box="[473,708,1360,1386]" italics="true" pageId="13" pageNumber="34">Anonymous 75-1841</emphasis>
(PE00580621, PE00580622, PE00580623), 
<emphasis id="B973EADCFFCEFFEA3BD5FAC5FF6925AE" italics="true" pageId="13" pageNumber="34">
FLPH 
<collectingRegion id="49C3F82CFFCEFFEA3A04FAC5FADF2549" box="[1307,1363,1360,1386]" country="China" name="Xizang" pageId="13" pageNumber="34">Tibet</collectingRegion>
Exped. 12-0849
</emphasis>
(PE01966670, PE01966671), 
<emphasis id="B973EADCFFCEFFEA3D50FAE1FCE725AE" box="[591,875,1395,1421]" italics="true" pageId="13" pageNumber="34">
PE 
<collectingRegion id="49C3F82CFFCEFFEA3D61FAE1FD4025AE" box="[638,716,1396,1421]" country="China" name="Xizang" pageId="13" pageNumber="34">Xizang</collectingRegion>
Exped. 6212
</emphasis>
(PE02347446, PE02347447), 
<emphasis id="B973EADCFFCEFFEA3BCBFAE1FF4C2593" italics="true" pageId="13" pageNumber="34">
PE 
<collectingRegion id="49C3F82CFFCEFFEA3A1CFAE1FADD25AE" box="[1283,1361,1396,1421]" country="China" name="Xizang" pageId="13" pageNumber="34">Xizang</collectingRegion>
Exped. 6282
</emphasis>
(PE02309341, PE02309342, PE02309399, PE02309400), 
<emphasis id="B973EADCFFCEFFEA3C6CFA03FAB22593" box="[883,1342,1430,1457]" italics="true" pageId="13" pageNumber="34">
<collectingRegion id="49C3F82CFFCEFFEA3C6CFA03FC432593" box="[883,975,1430,1456]" country="China" name="Qinghai" pageId="13" pageNumber="34">Qinghai</collectingRegion>
- 
<collectingRegion id="49C3F82CFFCEFFEA3CC7FA02FBAA2593" box="[984,1062,1431,1456]" country="China" name="Xizang" pageId="13" pageNumber="34">Xizang</collectingRegion>
Suppl. Exped. 75-1642
</emphasis>
(HNWP, KUN0685490, KUN0685491, PE00500068, PE00500070), 
<emphasis id="B973EADCFFCEFFEA3C45FA2CFAB525F0" box="[858,1337,1465,1492]" italics="true" pageId="13" pageNumber="34">
<collectingRegion id="49C3F82CFFCEFFEA3C45FA2CFC3A25F0" box="[858,950,1465,1491]" country="China" name="Qinghai" pageId="13" pageNumber="34">Qinghai</collectingRegion>
- 
<collectingRegion id="49C3F82CFFCEFFEA3CA0FA2FFB8125F0" box="[959,1037,1466,1491]" country="China" name="Xizang" pageId="13" pageNumber="34">Xizang</collectingRegion>
Suppl. Exped. 75-1720
</emphasis>
(HNWP, KUN0685488, KUN0685489, PE00500067), 
<emphasis id="B973EADCFFCEFFEA3D90FA49FB9C25D4" box="[655,1040,1500,1527]" italics="true" pageId="13" pageNumber="34">Qinghai-Xizang Veg. Exped. 2471</emphasis>
(PE00048368), 
<emphasis id="B973EADCFFCEFFEA3BD3FA48FF4C2639" italics="true" pageId="13" pageNumber="34">Z. Zhou et al. ZJW-3923</emphasis>
(KUN1274266).
</paragraph>
</subSubSection>
<subSubSection id="C31D6545FFCEFFE83FA2F9B6FC9027E8" lastPageId="15" lastPageNumber="36" pageId="13" pageNumber="34" type="discussion">
<paragraph id="8BB836CEFFCEFFEA3FA2F9B6FA832710" blockId="13.[136,1452,1113,1844]" pageId="13" pageNumber="34">
<emphasis id="B973EADCFFCEFFEA3FA2F9B6FE8C261E" bold="true" box="[189,256,1571,1597]" pageId="13" pageNumber="34">Notes</emphasis>
:— 
<taxonomicName id="4C074D4DFFCEFFEA3E3BF9B6FD1F261E" authorityName="Q. Yuan &amp; Q. E. Yang" authorityYear="2023" box="[292,659,1571,1597]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="13" pageNumber="34" phylum="Tracheophyta" rank="species" species="pseudoscabriflorum">
<emphasis id="B973EADCFFCEFFEA3E3BF9B6FD1F261E" box="[292,659,1571,1597]" italics="true" pageId="13" pageNumber="34">Delphinium pseudoscabriflorum</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C074D4DFFCEFFEA3C2DF9B1FB1C261D" authority="Munz (1968: 105)" authorityName="Munz" authorityPageNumber="105" authorityYear="1968" box="[818,1168,1571,1598]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="13" pageNumber="34" phylum="Tracheophyta" rank="species" species="ludlowii">
<emphasis id="B973EADCFFCEFFEA3C2DF9B1FCCA261E" box="[818,838,1572,1597]" italics="true" pageId="13" pageNumber="34">D</emphasis>
. 
<emphasis id="B973EADCFFCEFFEA3C47F9B6FC38261E" box="[856,948,1571,1597]" italics="true" pageId="13" pageNumber="34">ludlowii</emphasis>
<bibRefCitation id="EF964B3FFFCEFFEA3CA1F9B6FB1C261D" author="Munz, P. A." box="[958,1168,1571,1598]" pageId="13" pageNumber="34" pagination="73 - 166" refId="ref5784" refString="Munz, P. A. (1968) A synopsis of the Asian species of Delphinium, sensu stricto. Journal of the Arnold Arboretum 49: 73 - 166. https: // doi. org / 10.5962 / p. 185735" type="journal article" year="1968">Munz (1968: 105)</bibRefCitation>
</taxonomicName>
in habit and position of bracteoles on the pedicel, but differs by having leaves 5-fid to near the base or about 
<quantity id="4CFF9B2BFFCEFFEA3B5BF9D3FB1C2642" box="[1092,1168,1606,1633]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="34" unit="cm" value="0.5">0.5 cm</quantity>
above (vs. 5-fid to about 
<quantity id="4CFF9B2BFFCEFFEA3F97F9FCFF4C26A7" box="[136,192,1641,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="34" unit="cm" value="1.0">1 cm</quantity>
from the base), pedicels densely more or less spreading puberulent (vs. appressed strigulose), bracteoles linear to lanceolate (vs. subspathulate), sepaline spurs slightly curved downward (vs. slightly curved upward), and follicles ventrally puberulent (vs. glabrous). Our new species is also somewhat similar to 
<taxonomicName id="4C074D4DFFCEFFEA3B07F925FB4026E9" authorityName=", Hara" authorityYear="1975" box="[1048,1228,1712,1738]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="13" pageNumber="34" phylum="Tracheophyta" rank="species" species="scabriflorum">
<emphasis id="B973EADCFFCEFFEA3B07F925FB4026E9" box="[1048,1228,1712,1738]" italics="true" pageId="13" pageNumber="34">D. scabriflorum</emphasis>
</taxonomicName>
in habit, but differs by having leaves 3-fid to near the base or about 
<quantity id="4CFF9B2BFFCEFFEA3D88F946FD6E26CE" box="[663,738,1747,1774]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="34" unit="cm" value="0.5">0.5 cm</quantity>
above (vs. moderately 5-lobed), pedicels more or less spreading puberulent (vs. appressed strigulose), bracteoles closely appressed to the flower (vs. below middle of the pedicel), sepaline spurs slightly curved downward (vs. straight), and follicles ventrally puberulent (vs. glabrous).
</paragraph>
<caption id="DF786646FFCDFFE93F97F834FD3427F9" ID-DOI="http://doi.org/10.5281/zenodo.10361904" ID-Zenodo-Dep="10361904" httpUri="https://zenodo.org/record/10361904/files/figure.png" pageId="14" pageNumber="35" startId="14.[136,229,1953,1975]" targetBox="[170,1417,190,1929]" targetPageId="14" targetType="figure">
<paragraph id="8BB836CEFFCDFFE93F97F834FD3427F9" blockId="14.[136,1452,1953,2011]" pageId="14" pageNumber="35">
<emphasis id="B973EADCFFCDFFE93F97F834FE8F2795" bold="true" box="[136,259,1953,1975]" pageId="14" pageNumber="35">FIGURE 14</emphasis>
. Paratype sheets of 
<taxonomicName id="4C074D4DFFCDFFE93EDBF834FD062795" authorityName="Munz" authorityYear="1968" box="[452,650,1953,1974]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="14" pageNumber="35" phylum="Tracheophyta" rank="species" species="ludlowii">
<emphasis id="B973EADCFFCDFFE93EDBF834FD062795" box="[452,650,1953,1974]" italics="true" pageId="14" pageNumber="35">Delphinium ludlowii</emphasis>
</taxonomicName>
. A. 
<emphasis id="B973EADCFFCDFFE93DAEF837FCD32795" box="[689,863,1953,1975]" italics="true" pageId="14" pageNumber="35">R.E. Cooper 4322</emphasis>
(BM). B. 
<emphasis id="B973EADCFFCDFFE93CDDF837FB5C2795" box="[962,1232,1953,1975]" italics="true" pageId="14" pageNumber="35">F. Ludlow &amp; G. Sherriff 368</emphasis>
(BM). C. 
<emphasis id="B973EADCFFCDFFE93A2CF837FE9527F9" italics="true" pageId="14" pageNumber="35">F. Ludlow &amp; G. Sherriff 470</emphasis>
(BM). D. 
<emphasis id="B973EADCFFCDFFE93E62F853FD0027F9" box="[381,652,1989,2011]" italics="true" pageId="14" pageNumber="35">F. Ludlow &amp; G. Sherriff 470</emphasis>
(E).
</paragraph>
</caption>
<caption id="DF786646FFCCFFE83F97FBA2FEAB2452" ID-DOI="http://doi.org/10.5281/zenodo.10361906" ID-Zenodo-Dep="10361906" httpUri="https://zenodo.org/record/10361906/files/figure.png" pageId="15" pageNumber="36" startId="15.[136,229,1079,1101]" targetBox="[170,1417,190,1055]" targetPageId="15" targetType="figure">
<paragraph id="8BB836CEFFCCFFE83F97FBA2FEAB2452" blockId="15.[136,1452,1079,1137]" pageId="15" pageNumber="36">
<emphasis id="B973EADCFFCCFFE83F97FBA2FE89246E" bold="true" box="[136,261,1079,1101]" pageId="15" pageNumber="36">FIGURE 15</emphasis>
. Holotype (A) and isotype (B) sheets of 
<taxonomicName id="4C074D4DFFCCFFE83D84FBADFCE8246E" authorityName="Munz" authorityYear="1968" box="[667,868,1080,1101]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="ludlowii">
<emphasis id="B973EADCFFCCFFE83D84FBADFCE8246E" box="[667,868,1080,1101]" italics="true" pageId="15" pageNumber="36">Delphinium ludlowii</emphasis>
</taxonomicName>
. A. 
<emphasis id="B973EADCFFCCFFE83C90FBADFB2B246E" box="[911,1191,1080,1101]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 894</emphasis>
(BM). B. 
<emphasis id="B973EADCFFCCFFE83A0FFBADFF772452" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 894</emphasis>
(E).
</paragraph>
</caption>
<paragraph id="8BB836CEFFCCFFE83FA2FB35FD1125D4" blockId="15.[136,1452,1183,1995]" pageId="15" pageNumber="36">
<bibRefCitation id="EF964B3FFFCCFFE83FA2FB35FEDE249A" author="Munz, P. A." box="[189,338,1183,1210]" pageId="15" pageNumber="36" pagination="73 - 166" refId="ref5784" refString="Munz, P. A. (1968) A synopsis of the Asian species of Delphinium, sensu stricto. Journal of the Arnold Arboretum 49: 73 - 166. https: // doi. org / 10.5962 / p. 185735" type="journal article" year="1968">Munz (1968)</bibRefCitation>
described 
<taxonomicName id="4C074D4DFFCCFFE83ED0FB0AFD34249A" authorityName="Munz" authorityYear="1968" box="[463,696,1183,1209]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="ludlowii">
<emphasis id="B973EADCFFCCFFE83ED0FB0AFD34249A" box="[463,696,1183,1209]" italics="true" pageId="15" pageNumber="36">Delphinium ludlowii</emphasis>
</taxonomicName>
on the basis of eight collections from 
<collectingCountry id="F310765EFFCCFFE83B6BFB0AFB45249A" box="[1140,1225,1183,1209]" name="Bhutan" pageId="15" pageNumber="36">Bhutan</collectingCountry>
. These include 
<emphasis id="B973EADCFFCCFFE83A64FB35FE9024FE" italics="true" pageId="15" pageNumber="36">R.E. Cooper 4322</emphasis>
(BM; 
<figureCitation id="133C2A4BFFCCFFE83E76FB57FE5D24FE" box="[361,465,1218,1245]" captionStart="FIGURE 14" captionStartId="14.[136,229,1953,1975]" captionTargetBox="[170,1417,190,1929]" captionTargetId="figure-20@14.[170,1417,190,1929]" captionTargetPageId="14" captionText="FIGURE 14. Paratype sheets of Delphinium ludlowii. A. R.E. Cooper 4322 (BM). B. F. Ludlow &amp; G. Sherriff 368 (BM). C. F. Ludlow &amp; G. Sherriff 470 (BM). D. F. Ludlow &amp; G. Sherriff 470 (E)." figureDoi="http://doi.org/10.5281/zenodo.10361904" httpUri="https://zenodo.org/record/10361904/files/figure.png" pageId="15" pageNumber="36">Fig. 14A</figureCitation>
), 
<emphasis id="B973EADCFFCCFFE83EF9FB56FCAB24FF" box="[486,807,1218,1245]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 368</emphasis>
(BM, E; the BM sheet shown in 
<figureCitation id="133C2A4BFFCCFFE83BBCFB57FAA924FE" box="[1187,1317,1218,1245]" captionStart="FIGURE 14" captionStartId="14.[136,229,1953,1975]" captionTargetBox="[170,1417,190,1929]" captionTargetId="figure-20@14.[170,1417,190,1929]" captionTargetPageId="14" captionText="FIGURE 14. Paratype sheets of Delphinium ludlowii. A. R.E. Cooper 4322 (BM). B. F. Ludlow &amp; G. Sherriff 368 (BM). C. F. Ludlow &amp; G. Sherriff 470 (BM). D. F. Ludlow &amp; G. Sherriff 470 (E)." figureDoi="http://doi.org/10.5281/zenodo.10361904" httpUri="https://zenodo.org/record/10361904/files/figure.png" pageId="15" pageNumber="36">Figure 14B</figureCitation>
), 
<emphasis id="B973EADCFFCCFFE83A24FB56FEC22523" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 470</emphasis>
(BM, E; 
<figureCitation id="133C2A4BFFCCFFE83EA7FB73FDB32523" box="[440,575,1254,1280]" captionStart="FIGURE 14" captionStartId="14.[136,229,1953,1975]" captionTargetBox="[170,1417,190,1929]" captionTargetId="figure-20@14.[170,1417,190,1929]" captionTargetPageId="14" captionText="FIGURE 14. Paratype sheets of Delphinium ludlowii. A. R.E. Cooper 4322 (BM). B. F. Ludlow &amp; G. Sherriff 368 (BM). C. F. Ludlow &amp; G. Sherriff 470 (BM). D. F. Ludlow &amp; G. Sherriff 470 (E)." figureDoi="http://doi.org/10.5281/zenodo.10361904" httpUri="https://zenodo.org/record/10361904/files/figure.png" pageId="15" pageNumber="36">Fig. 14C, D</figureCitation>
), 
<emphasis id="B973EADCFFCCFFE83D4BFB73FC1C2523" box="[596,912,1254,1280]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 894</emphasis>
(BM, E; 
<figureCitation id="133C2A4BFFCCFFE83CE5FB73FBC02523" box="[1018,1100,1253,1280]" captionStart="FIGURE 15" captionStartId="15.[136,229,1079,1101]" captionTargetBox="[170,1417,190,1055]" captionTargetId="figure-17@15.[170,1417,190,1055]" captionTargetPageId="15" captionText="FIGURE 15. Holotype (A) and isotype (B) sheets of Delphinium ludlowii. A. F. Ludlow &amp; G. Sherriff 894 (BM). B. F. Ludlow &amp; G. Sherriff 894 (E)." figureDoi="http://doi.org/10.5281/zenodo.10361906" httpUri="https://zenodo.org/record/10361906/files/figure.png" pageId="15" pageNumber="36">Fig. 15</figureCitation>
), 
<emphasis id="B973EADCFFCCFFE83B7EFB73FA272523" box="[1121,1451,1254,1280]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 1008</emphasis>
(BM; 
<figureCitation id="133C2A4BFFCCFFE83FD1FA9CFEB42500" box="[206,312,1289,1315]" captionStart="FIGURE 16" captionStartId="16.[136,229,1953,1975]" captionTargetBox="[170,1417,190,1929]" captionTargetId="figure-20@16.[170,1417,190,1929]" captionTargetPageId="16" captionText="FIGURE 16. Paratype sheets of Delphinium ludlowii. A. F. Ludlow &amp; G. Sherriff 1008 (BM). B. F. Ludlow &amp; G. Sherriff 16972 (BM). C. F. Ludlow &amp; G. Sherriff 19485 (BM). D. F. Ludlow &amp; G. Sherriff 20978 (BM)." figureDoi="http://doi.org/10.5281/zenodo.10361908" httpUri="https://zenodo.org/record/10361908/files/figure.png" pageId="15" pageNumber="36">Fig. 16A</figureCitation>
), 
<emphasis id="B973EADCFFCCFFE83E51FA9FFD3F2500" box="[334,691,1289,1315]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 16972</emphasis>
(BM; 
<figureCitation id="133C2A4BFFCCFFE83C1CFA9CFCE72500" box="[771,875,1289,1315]" captionStart="FIGURE 16" captionStartId="16.[136,229,1953,1975]" captionTargetBox="[170,1417,190,1929]" captionTargetId="figure-20@16.[170,1417,190,1929]" captionTargetPageId="16" captionText="FIGURE 16. Paratype sheets of Delphinium ludlowii. A. F. Ludlow &amp; G. Sherriff 1008 (BM). B. F. Ludlow &amp; G. Sherriff 16972 (BM). C. F. Ludlow &amp; G. Sherriff 19485 (BM). D. F. Ludlow &amp; G. Sherriff 20978 (BM)." figureDoi="http://doi.org/10.5281/zenodo.10361908" httpUri="https://zenodo.org/record/10361908/files/figure.png" pageId="15" pageNumber="36">Fig. 16B</figureCitation>
), 
<emphasis id="B973EADCFFCCFFE83C9DFA9FFB6B2500" box="[898,1255,1289,1315]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 19485</emphasis>
(BM; 
<figureCitation id="133C2A4BFFCCFFE83A29FA9CFA122500" box="[1334,1438,1289,1315]" captionStart="FIGURE 16" captionStartId="16.[136,229,1953,1975]" captionTargetBox="[170,1417,190,1929]" captionTargetId="figure-20@16.[170,1417,190,1929]" captionTargetPageId="16" captionText="FIGURE 16. Paratype sheets of Delphinium ludlowii. A. F. Ludlow &amp; G. Sherriff 1008 (BM). B. F. Ludlow &amp; G. Sherriff 16972 (BM). C. F. Ludlow &amp; G. Sherriff 19485 (BM). D. F. Ludlow &amp; G. Sherriff 20978 (BM)." figureDoi="http://doi.org/10.5281/zenodo.10361908" httpUri="https://zenodo.org/record/10361908/files/figure.png" pageId="15" pageNumber="36">Fig. 16C</figureCitation>
), and 
<emphasis id="B973EADCFFCCFFE83FA7FAB8FD9E2565" box="[184,530,1324,1350]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 20978</emphasis>
(BM; 
<figureCitation id="133C2A4BFFCCFFE83D42FAB9FD482565" box="[605,708,1324,1350]" captionStart="FIGURE 16" captionStartId="16.[136,229,1953,1975]" captionTargetBox="[170,1417,190,1929]" captionTargetId="figure-20@16.[170,1417,190,1929]" captionTargetPageId="16" captionText="FIGURE 16. Paratype sheets of Delphinium ludlowii. A. F. Ludlow &amp; G. Sherriff 1008 (BM). B. F. Ludlow &amp; G. Sherriff 16972 (BM). C. F. Ludlow &amp; G. Sherriff 19485 (BM). D. F. Ludlow &amp; G. Sherriff 20978 (BM)." figureDoi="http://doi.org/10.5281/zenodo.10361908" httpUri="https://zenodo.org/record/10361908/files/figure.png" pageId="15" pageNumber="36">Fig. 16D</figureCitation>
), with the BM sheet of the collection 
<emphasis id="B973EADCFFCCFFE83B72FAB8FA272565" box="[1133,1451,1324,1350]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 894</emphasis>
designated as 
<typeStatus id="54BC886CFFCCFFE83E3BFADAFE0A254A" box="[292,390,1359,1385]" pageId="15" pageNumber="36" type="holotype">holotype</typeStatus>
(
<figureCitation id="133C2A4BFFCCFFE83E8AFADAFE712549" box="[405,509,1359,1386]" captionStart="FIGURE 15" captionStartId="15.[136,229,1079,1101]" captionTargetBox="[170,1417,190,1055]" captionTargetId="figure-17@15.[170,1417,190,1055]" captionTargetPageId="15" captionText="FIGURE 15. Holotype (A) and isotype (B) sheets of Delphinium ludlowii. A. F. Ludlow &amp; G. Sherriff 894 (BM). B. F. Ludlow &amp; G. Sherriff 894 (E)." figureDoi="http://doi.org/10.5281/zenodo.10361906" httpUri="https://zenodo.org/record/10361906/files/figure.png" pageId="15" pageNumber="36">Fig. 15A</figureCitation>
). In the protologue, the author stated that this species seemed close to 
<taxonomicName id="4C074D4DFFCCFFE83A16FAC5FA28254A" authorityName="Wallich" authorityYear="1831" box="[1289,1444,1359,1385]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="altissimum">
<emphasis id="B973EADCFFCCFFE83A16FAC5FA28254A" box="[1289,1444,1359,1385]" italics="true" pageId="15" pageNumber="36">D. altissimum</emphasis>
</taxonomicName>
, but had more evenly toothed leaves, bracteoles nearer the flower, and sepals more minutely pubescent. Since its description, 
<taxonomicName id="4C074D4DFFCCFFE83E0BFA03FE182593" authorityName="Munz" authorityYear="1968" box="[276,404,1430,1456]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="ludlowii">
<emphasis id="B973EADCFFCCFFE83E0BFA03FE182593" box="[276,404,1430,1456]" italics="true" pageId="15" pageNumber="36">D. ludlowii</emphasis>
</taxonomicName>
has been recognized by 
<bibRefCitation id="EF964B3FFFCCFFE83DADFA03FCB22593" author="Hara, H." box="[690,830,1430,1456]" pageId="15" pageNumber="36" pagination="9 - 23" refId="ref5562" refString="Hara, H. (1979) Ranunculaceae. In: Hara, H. &amp; Williams, L. H. J. (Eds.) An Enumeration of the Flowering Plants of Nepal, vol. 2. Trustees of British Museum (Natural History), London, pp. 9 - 23." type="book chapter" year="1979">Hara (1979)</bibRefCitation>
, 
<bibRefCitation id="EF964B3FFFCCFFE83C53FA03FB8F2593" author="Grierson, A. J. C." box="[844,1027,1430,1456]" pageId="15" pageNumber="36" pagination="309 - 314" refId="ref5474" refString="Grierson, A. J. C. (1984) Delphinium L. In: Grierson, A. J. C. &amp; Long, D. G. (Eds.) Flora of Bhutan, vol. 1, part 2. Royal Botanical Garden, Edinburgh, pp. 309 - 314." type="book chapter" year="1984">Grierson (1984)</bibRefCitation>
, 
<bibRefCitation id="EF964B3FFFCCFFE83B0EFA03FB182593" author="Rau, M. A." box="[1041,1172,1430,1456]" pageId="15" pageNumber="36" pagination="84 - 102" refId="ref5877" refString="Rau, M. A. (1993) Delphinium L. In: Sharma, B. D., Balakrishnan, N. P. &amp; Rao, R. R. (Eds.) Flora of India, vol. 1. Botanical Survey of India, Calcutta, pp. 84 - 102." type="book chapter" year="1993">Rau (1993)</bibRefCitation>
, and 
<bibRefCitation id="EF964B3FFFCCFFE83BCCFA03FA2B2593" author="Press, J. R. &amp; Shrestha, K. K. &amp; Sutton, D. A." box="[1235,1447,1430,1456]" pageId="15" pageNumber="36" pagination="248 - 258" refId="ref5828" refString="Press, J. R., Shrestha, K. K. &amp; Sutton, D. A. (2000) Ranunculaceae. In: Annotated Checklist of the Flowering Plants of Nepal. The Natural History Museum, London, pp. 248 - 258." type="book chapter" year="2000">
Press 
<emphasis id="B973EADCFFCCFFE83A08FA03FAC62593" box="[1303,1354,1430,1456]" italics="true" pageId="15" pageNumber="36">et al</emphasis>
. (2000)
</bibRefCitation>
, with its geographical range extended to include 
<collectingCountry id="F310765EFFCCFFE83DBAFA2CFD6425F0" box="[677,744,1465,1491]" name="Nepal" pageId="15" pageNumber="36">Nepal</collectingCountry>
and 
<collectingCountry id="F310765EFFCCFFE83C3FFA2CFCD625F0" box="[800,858,1465,1491]" name="India" pageId="15" pageNumber="36">India</collectingCountry>
(
<collectingRegion id="49C3F82CFFCCFFE83C76FA2CFC4C25F0" box="[873,960,1465,1491]" country="India" name="Sikkim" pageId="15" pageNumber="36">Sikkim</collectingRegion>
). However, we have been unable to see any material of 
<taxonomicName id="4C074D4DFFCCFFE83E16FA4BFE0B25D4" authorityName="Munz" authorityYear="1968" box="[265,391,1501,1527]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="ludlowii">
<emphasis id="B973EADCFFCCFFE83E16FA4BFE0B25D4" box="[265,391,1501,1527]" italics="true" pageId="15" pageNumber="36">D. ludlowii</emphasis>
</taxonomicName>
from 
<collectingCountry id="F310765EFFCCFFE83ED3FA48FD8325D4" box="[460,527,1501,1527]" name="Nepal" pageId="15" pageNumber="36">Nepal</collectingCountry>
and Sikkim.
</paragraph>
<paragraph id="8BB836CEFFCCFFE83FA2F994FC3B26D0" blockId="15.[136,1452,1183,1995]" pageId="15" pageNumber="36">
From the type material, 
<taxonomicName id="4C074D4DFFCCFFE83EC8F994FD4D2638" authorityName="Munz" authorityYear="1968" box="[471,705,1537,1563]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="ludlowii">
<emphasis id="B973EADCFFCCFFE83EC8F994FD4D2638" box="[471,705,1537,1563]" italics="true" pageId="15" pageNumber="36">Delphinium ludlowii</emphasis>
</taxonomicName>
is characterized by having stems up to 
<quantity id="4CFF9B2BFFCCFFE83B8CF994FB4D2638" box="[1171,1217,1537,1563]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="36" unit="m" value="1.0">1 m</quantity>
tall, pubescent with deflexed hairs (minutely strigulose) especially above, leaves 5-fid to about 
<quantity id="4CFF9B2BFFCCFFE83CD1F9B0FB88261C" box="[974,1028,1573,1599]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="36" unit="cm" value="1.0">1 cm</quantity>
from base, inflorescence mostly a lax 6–12-flowered raceme, inflorescence axis and pedicels densely pubescent with deflexed hairs (minutely strigulose), bracteoles linear-spathulate and closely appressed to sepals (subtending the flower), sepaline spurs slightly curved upward, and glabrous follicles. It is noteworthy that the degree of leaf division is somewhat variable in this species. In the 
<typeStatus id="54BC886CFFCCFFE83FAEF920FE9F26EC" box="[177,275,1717,1743]" pageId="15" pageNumber="36" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54BC886CFFCCFFE83E56F920FE1626EC" box="[329,410,1717,1743]" pageId="15" pageNumber="36" type="isotype">isotype</typeStatus>
the leaves are divided to about 
<quantity id="4CFF9B2BFFCCFFE83DE5F920FCBD26EC" box="[762,817,1717,1743]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="36" unit="cm" value="1.0">1 cm</quantity>
from the base (
<figureCitation id="133C2A4BFFCCFFE83CC1F920FBBC26F3" box="[990,1072,1717,1744]" captionStart="FIGURE 15" captionStartId="15.[136,229,1079,1101]" captionTargetBox="[170,1417,190,1055]" captionTargetId="figure-17@15.[170,1417,190,1055]" captionTargetPageId="15" captionText="FIGURE 15. Holotype (A) and isotype (B) sheets of Delphinium ludlowii. A. F. Ludlow &amp; G. Sherriff 894 (BM). B. F. Ludlow &amp; G. Sherriff 894 (E)." figureDoi="http://doi.org/10.5281/zenodo.10361906" httpUri="https://zenodo.org/record/10361906/files/figure.png" pageId="15" pageNumber="36">Fig. 15</figureCitation>
), but in 
<specimenCount id="9D01FD47FFCCFFE83B93F920FABA26EC" box="[1164,1334,1717,1743]" count="3" pageId="15" pageNumber="36" type="generic" typeStatus="paratype">three paratypes</specimenCount>
the leaves are often more deeply divided to less than 
<quantity id="4CFF9B2BFFCCFFE83D7BF94CFD1726D0" box="[612,667,1753,1779]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="36" unit="cm" value="1.0">1 cm</quantity>
from base (
<figureCitation id="133C2A4BFFCCFFE83C3EF94CFC2626D0" box="[801,938,1753,1779]" captionStart="FIGURE 16" captionStartId="16.[136,229,1953,1975]" captionTargetBox="[170,1417,190,1929]" captionTargetId="figure-20@16.[170,1417,190,1929]" captionTargetPageId="16" captionText="FIGURE 16. Paratype sheets of Delphinium ludlowii. A. F. Ludlow &amp; G. Sherriff 1008 (BM). B. F. Ludlow &amp; G. Sherriff 16972 (BM). C. F. Ludlow &amp; G. Sherriff 19485 (BM). D. F. Ludlow &amp; G. Sherriff 20978 (BM)." figureDoi="http://doi.org/10.5281/zenodo.10361908" httpUri="https://zenodo.org/record/10361908/files/figure.png" pageId="15" pageNumber="36">Fig. 16B–D</figureCitation>
).
</paragraph>
<paragraph id="8BB836CEFFCCFFE83FA2F968FC9027E8" blockId="15.[136,1452,1183,1995]" pageId="15" pageNumber="36">
Of the collections cited as 
<taxonomicName id="4C074D4DFFCCFFE83EF1F968FC872734" authorityName=", Hara" authorityYear="1975" box="[494,779,1789,1815]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="scabriflorum">
<emphasis id="B973EADCFFCCFFE83EF1F968FC872734" box="[494,779,1789,1815]" italics="true" pageId="15" pageNumber="36">Delphinium scabriflorum</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF964B3FFFCCFFE83C28F968FC422734" author="Munz, P. A." box="[823,974,1789,1815]" pageId="15" pageNumber="36" pagination="73 - 166" refId="ref5784" refString="Munz, P. A. (1968) A synopsis of the Asian species of Delphinium, sensu stricto. Journal of the Arnold Arboretum 49: 73 - 166. https: // doi. org / 10.5962 / p. 185735" type="journal article" year="1968">Munz (1968)</bibRefCitation>
, only 
<emphasis id="B973EADCFFCCFFE83B0AF96BFB6C2734" box="[1045,1248,1789,1815]" italics="true" pageId="15" pageNumber="36">R.E. Cooper 4114</emphasis>
(BM, E; 
<figureCitation id="133C2A4BFFCCFFE83A52F968FF172718" captionStart="FIGURE 1" captionStartId="1.[136,229,1921,1943]" captionTargetBox="[202,1386,245,1896]" captionTargetId="figure-53@1.[202,1386,245,1896]" captionTargetPageId="1" captionText="FIGURE 1. Selected specimens from Bhutan cited as Delphinium scabriflorum by Munz (1968). A. R.E. Cooper 4114 (BM). B. R.E. Cooper 4114 (E). C. R.E. Cooper 4223 (BM). D. R.E. Cooper 4223 (E). Note that only the two sheets of the collection R.E. Cooper 4114 (A, B) belong to D. scabriflorum while the two sheets of the collection R.E. Cooper 4223 (C, D) belong to D. pseudoscabriflorum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10361855" httpUri="https://zenodo.org/record/10361855/files/figure.png" pageId="15" pageNumber="36">Fig. 1A, B</figureCitation>
) belongs to this species. The remaining collections should belong to our new species 
<taxonomicName id="4C074D4DFFCCFFE83B77F8B7FAE72718" authorityName="Q. Yuan &amp; Q. E. Yang" authorityYear="2023" box="[1128,1387,1825,1851]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="pseudoscabriflorum">
<emphasis id="B973EADCFFCCFFE83B77F8B7FAE72718" box="[1128,1387,1825,1851]" italics="true" pageId="15" pageNumber="36">D. pseudoscabriflorum</emphasis>
</taxonomicName>
. 
<specimenCount id="9D01FD47FFCCFFE83A65F8B4FF71277C" count="2" pageId="15" pageNumber="36" type="generic">Two specimens</specimenCount>
from 
<collectingCountry id="F310765EFFCCFFE83E5AF8D0FE15277C" box="[325,409,1861,1887]" name="Bhutan" pageId="15" pageNumber="36">Bhutan</collectingCountry>
, 
<emphasis id="B973EADCFFCCFFE83EBAF8D3FC8A277C" box="[421,774,1861,1887]" italics="true" pageId="15" pageNumber="36">F. Ludlow &amp; G. Sherriff 21303</emphasis>
(BM; 
<figureCitation id="133C2A4BFFCCFFE83C4BF8D0FC36277C" box="[852,954,1861,1887]" captionStart="FIGURE 17" captionStartId="17.[136,229,1079,1101]" captionTargetBox="[170,1417,190,1055]" captionTargetId="figure-17@17.[170,1417,190,1055]" captionTargetPageId="17" captionText="FIGURE 17. Two specimens from Bhutan identified as Delphinium ludlowii but actually belonging to D. pseudoscabriflorum sp. nov.A. F. Ludlow &amp; G. Sherriff 21303 (BM). B. S.B. Lyon 3480 (BM)." figureDoi="http://doi.org/10.5281/zenodo.10361912" httpUri="https://zenodo.org/record/10361912/files/figure.png" pageId="15" pageNumber="36">Fig. 17A</figureCitation>
) and 
<emphasis id="B973EADCFFCCFFE83CE3F8D0FB26277C" box="[1020,1194,1861,1887]" italics="true" pageId="15" pageNumber="36">S.B. Lyon 3480</emphasis>
(BM; 
<figureCitation id="133C2A4BFFCCFFE83BE7F8D0FAD2277C" box="[1272,1374,1861,1887]" captionStart="FIGURE 17" captionStartId="17.[136,229,1079,1101]" captionTargetBox="[170,1417,190,1055]" captionTargetId="figure-17@17.[170,1417,190,1055]" captionTargetPageId="17" captionText="FIGURE 17. Two specimens from Bhutan identified as Delphinium ludlowii but actually belonging to D. pseudoscabriflorum sp. nov.A. F. Ludlow &amp; G. Sherriff 21303 (BM). B. S.B. Lyon 3480 (BM)." figureDoi="http://doi.org/10.5281/zenodo.10361912" httpUri="https://zenodo.org/record/10361912/files/figure.png" pageId="15" pageNumber="36">Fig. 17B</figureCitation>
), were identified as 
<taxonomicName id="4C074D4DFFCCFFE83E07F8FFFE1927A0" authorityName="Munz" authorityYear="1968" box="[280,405,1897,1923]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="ludlowii">
<emphasis id="B973EADCFFCCFFE83E07F8FFFE1927A0" box="[280,405,1897,1923]" italics="true" pageId="15" pageNumber="36">D. ludlowii</emphasis>
</taxonomicName>
on the determination slips but not cited under this species by 
<bibRefCitation id="EF964B3FFFCCFFE83B5BF8FCFB5627A0" author="Munz, P. A." box="[1092,1242,1897,1923]" pageId="15" pageNumber="36" pagination="73 - 166" refId="ref5784" refString="Munz, P. A. (1968) A synopsis of the Asian species of Delphinium, sensu stricto. Journal of the Arnold Arboretum 49: 73 - 166. https: // doi. org / 10.5962 / p. 185735" type="journal article" year="1968">Munz (1968)</bibRefCitation>
. From their degree of leaf division, shape of bracteoles and their position on the pedicel, and shape of sepaline spurs, both can also be readily referred to our new species 
<taxonomicName id="4C074D4DFFCCFFE83D0CF827FC9927E8" authorityName="Q. Yuan &amp; Q. E. Yang" authorityYear="2023" box="[531,789,1969,1995]" class="Magnoliopsida" family="Ranunculaceae" genus="Delphinium" kingdom="Plantae" order="Ranunculales" pageId="15" pageNumber="36" phylum="Tracheophyta" rank="species" species="pseudoscabriflorum">
<emphasis id="B973EADCFFCCFFE83D0CF827FC9927E8" box="[531,789,1969,1995]" italics="true" pageId="15" pageNumber="36">D. pseudoscabriflorum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>