182 lines
20 KiB
XML
182 lines
20 KiB
XML
<document id="F9FE5FB735B4DA14705657F5681D1EE4" ID-CLB-Dataset="169977" ID-DOI="10.11646/zootaxa.5196.2.3" ID-GBIF-Dataset="78964cd4-d84f-4507-b84c-1b1123848fc3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7224475" ID-ZooBank="659CB97A-6278-4CF6-9583-1A08700DAE1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666167176601" checkinUser="plazi" docAuthor="Zouhair, Laila & Kettani, Kawtar" docDate="2022" docId="03AE87D152442C3D36E6992BAE550BE5" docLanguage="en" docName="zootaxa.5196.2.3.pdf" docOrigin="Zootaxa 5196 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Syndyas subsabinios Chvala 1975" docType="treatment" docVersion="6" lastPageNumber="217" masterDocId="FF97FFA952412C3B36719861AF770808" masterDocTitle="Moroccan Hybotinae (Diptera: Hybotidae): first record of the subfamily and rare genera with an emphasis on their distribution" masterLastPageNumber="222" masterPageNumber="211" pageNumber="216" updateTime="1699408992803" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="651C637E95674AD2303219087BC4F57B">Moroccan Hybotinae (Diptera: Hybotidae): first record of the subfamily and rare genera with an emphasis on their distribution</mods:title>
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<mods:namePart id="54FCFE298489B23333E46955428D7FA6">Zouhair, Laila</mods:namePart>
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<mods:namePart id="9DA03DAD6F1D8759F123DA6411753EE6">Kettani, Kawtar</mods:namePart>
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<mods:title id="DDCF4B7BA6AD94F02EFEF5E19DBA1EE4">Zootaxa</mods:title>
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<mods:part id="4B95EF176DD0B3FB6C73C423B356B480">
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<mods:date id="A4E5B749CD71EB10D1DC4345214FB03E">2022</mods:date>
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<mods:number id="36CA11A30F9848D9EEE2CA51C50A9020">2022-10-19</mods:number>
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<mods:number id="F413DCF9915A677F14C3FF2EB1C3606E">5196</mods:number>
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<treatment id="03AE87D152442C3D36E6992BAE550BE5" ID-DOI="http://doi.org/10.5281/zenodo.7224491" ID-GBIF-Taxon="203119805" ID-Zenodo-Dep="7224491" LSID="urn:lsid:plazi:treatment:03AE87D152442C3D36E6992BAE550BE5" httpUri="http://treatment.plazi.org/id/03AE87D152442C3D36E6992BAE550BE5" lastPageId="6" lastPageNumber="217" pageId="5" pageNumber="216">
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<subSubSection id="C31D654C52442C3E36E6992BAD4D096C" box="[151,570,330,357]" pageId="5" pageNumber="216" type="nomenclature">
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<paragraph id="8BB836C752442C3E36E6992BAD4D096C" blockId="5.[151,570,330,393]" box="[151,570,330,357]" pageId="5" pageNumber="216">
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<heading id="D0F081AB52442C3E36E6992BAD4D096C" bold="true" box="[151,570,330,357]" fontSize="11" level="1" pageId="5" pageNumber="216" reason="1">
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<taxonomicName id="4C074D4452442C3E36E6992BAD4D096C" authority="Chvala, 1975" authorityName="Chvala" authorityYear="1975" box="[151,570,330,357]" class="Insecta" family="Hybotidae" genus="Syndyas" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="216" phylum="Arthropoda" rank="species" species="subsabinios">
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<emphasis id="B973EAD552442C3E36E6992BAD4D096C" bold="true" box="[151,570,330,357]" pageId="5" pageNumber="216">
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<emphasis id="B973EAD552442C3E36E6992BAEE7096D" bold="true" box="[151,400,330,357]" italics="true" pageId="5" pageNumber="216">Syndyas subsabinios</emphasis>
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<bibRefCitation id="EF964B3652442C3E37E6992BAD4D096C" author="Chvala, M." box="[407,570,330,357]" pageId="5" pageNumber="216" pagination="257 - 261" refId="ref5660" refString="Chvala, M. (1975) A new Palaearctic species of Syndyas Loew from Spain, with synonymic notes on the genus Sabinios Jones (Diptera, Empididae). Acta Entomologica Bohemoeslovaca, 72, 257 - 261." type="journal article" year="1975">Chvála, 1975</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D654C52442C3E36E6990EAF9C0981" box="[151,235,367,393]" pageId="5" pageNumber="216" type="description">
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<paragraph id="8BB836C752442C3E36E6990EAF9C0981" blockId="5.[151,570,330,393]" box="[151,235,367,393]" pageId="5" pageNumber="216">
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(
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<figureCitation id="133C2A4252442C3E36EE990EAF940981" box="[159,227,367,393]" captionStart="FIGURE 4" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[151,1436,1010,1920]" captionTargetId="figure-268@5.[151,1436,957,1920]" captionTargetPageId="5" captionText="FIGURE 4. Habitus male of Syndyas subsabinios. Scale: 500 µm" figureDoi="http://doi.org/10.5281/zenodo.7224487" httpUri="https://zenodo.org/record/7224487/files/figure.png" pageId="5" pageNumber="216">Fig. 4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D654C52442C3E36E699D7AE3809FD" pageId="5" pageNumber="216" type="materials_examined">
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<paragraph id="8BB836C752442C3E36E699D7AE3809FD" blockId="5.[151,1437,438,933]" pageId="5" pageNumber="216">
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<emphasis id="B973EAD552442C3E36E699D7AEF709D8" bold="true" box="[151,384,438,464]" pageId="5" pageNumber="216">Material examined.</emphasis>
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<materialsCitation id="3B6F3C9A52442C3E37F699D6AE3C09FD" ID-GBIF-Occurrence="3938602302" collectingDate="2012-09-16" collectionCode="PCLZ" collectorName="K. Kettani" country="Morocco" county="Khemis Anjra" elevation="61" location="Rif" municipality="Oued Khemis" pageId="5" pageNumber="216" specimenCount="1" specimenCount-male="1">
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<emphasis id="B973EAD552442C3E37F699D6AE8009D9" bold="true" box="[391,503,439,465]" pageId="5" pageNumber="216">
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<collectingCountry id="F310765752442C3E37F699D6AE8509D9" box="[391,498,439,465]" name="Morocco" pageId="5" pageNumber="216">Morocco</collectingCountry>
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.
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</emphasis>
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<specimenCount id="9D01FD4E52442C3E378C99D6AD5509D9" box="[509,546,439,465]" pageId="5" pageNumber="216" type="male">1♂</specimenCount>
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,
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<location id="8ED8601C52442C3E345F99D6AD2209D9" LSID="urn:lsid:plazi:treatment:03AE87D152442C3D36E6992BAE550BE5:8ED8601C52442C3E345F99D6AD2209D9" box="[558,597,439,465]" country="Morocco" county="Khemis Anjra" municipality="Oued Khemis" name="Rif" pageId="5" pageNumber="216">Rif</location>
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,
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<collectingMunicipality id="6BDCACBD52442C3E342E99D6AD8B09D9" box="[607,764,439,465]" pageId="5" pageNumber="216">Oued Khemis</collectingMunicipality>
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,
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<collectingCounty id="62D94E4B52442C3E357699D6ACD409D9" box="[775,931,439,465]" pageId="5" pageNumber="216">Khemis Anjra</collectingCounty>
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(site 3),
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<quantity id="4CFF9B2252442C3E327499D6AB3709D9" box="[1029,1088,439,465]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="216" unit="m" value="61.0">
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<elevation id="002AD1F452442C3E327499D6AB3709D9" box="[1029,1088,439,465]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="216" unit="m" value="61.0">61 m</elevation>
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</quantity>
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,
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<date id="FFB9100752442C3E323A99D6ABB209D9" box="[1099,1221,439,465]" pageId="5" pageNumber="216" value="2012-09-16">
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<collectingDate id="EFFDE9EF52442C3E323A99D6ABB209D9" box="[1099,1221,439,465]" pageId="5" pageNumber="216" value="2012-09-16">16.ix.2012</collectingDate>
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</date>
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,
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<collectingMethod id="52464ED052442C3E32A199D6AA3009D9" box="[1232,1351,439,465]" pageId="5" pageNumber="216">Sweep net</collectingMethod>
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, leg.
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<collectorName id="26F2531152442C3E33F099D6AF9E09FD" pageId="5" pageNumber="216">K. Kettani</collectorName>
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(
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<collectionCode id="ED16AE0252442C3E368999BAAE3409FD" box="[248,323,475,501]" pageId="5" pageNumber="216">PCLZ</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D654C52442C3E36B6999FAB230A10" box="[199,1108,510,537]" pageId="5" pageNumber="216" type="distribution">
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<paragraph id="8BB836C752442C3E36B6999FAB230A10" blockId="5.[151,1437,438,933]" box="[199,1108,510,537]" pageId="5" pageNumber="216">
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<emphasis id="B973EAD552442C3E36B6999FAE160A10" bold="true" box="[199,353,510,536]" pageId="5" pageNumber="216">Distribution.</emphasis>
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Known only from Spain.
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<emphasis id="B973EAD552442C3E34F4999FAB230A10" bold="true" box="[645,1108,510,537]" pageId="5" pageNumber="216">
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First record from
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<collectingCountry id="F310765752442C3E352C999EACB10A11" box="[861,966,511,537]" name="Morocco" pageId="5" pageNumber="216">Morocco</collectingCountry>
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and Africa.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D654C52442C3E36B69A43AEBE0AF9" pageId="5" pageNumber="216" type="discussion">
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<paragraph id="8BB836C752442C3E36B69A43AB440A8D" blockId="5.[151,1437,438,933]" pageId="5" pageNumber="216">
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<emphasis id="B973EAD552442C3E36B69A43AE380A34" bold="true" box="[199,335,546,572]" pageId="5" pageNumber="216">Comments.</emphasis>
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<taxonomicName id="4C074D4452442C3E37249A42AD4C0A35" authorityName="Chvala" authorityYear="1975" box="[341,571,547,573]" class="Insecta" family="Hybotidae" genus="Syndyas" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="216" phylum="Arthropoda" rank="species" species="subsabinios">
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<emphasis id="B973EAD552442C3E37249A42AD4C0A35" box="[341,571,547,573]" italics="true" pageId="5" pageNumber="216">Syndyas subsabinios</emphasis>
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</taxonomicName>
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was described from
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<collectingCountry id="F310765752442C3E356E9A42AC150A35" box="[799,866,547,573]" name="Spain" pageId="5" pageNumber="216">Spain</collectingCountry>
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, and no records have since been reported elsewhere (
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<bibRefCitation id="EF964B3652442C3E36EE9A26AE450A69" author="Chvala, M." box="[159,306,582,609]" pageId="5" pageNumber="216" pagination="257 - 261" refId="ref5660" refString="Chvala, M. (1975) A new Palaearctic species of Syndyas Loew from Spain, with synonymic notes on the genus Sabinios Jones (Diptera, Empididae). Acta Entomologica Bohemoeslovaca, 72, 257 - 261." type="journal article" year="1975">Chvála 1975</bibRefCitation>
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). It is worth pointing out that our finding represents the second citation of this species and constitutes the first record of the genus in North Africa and the species in the whole of Africa.
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</paragraph>
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<paragraph id="8BB836C752442C3E36B69AEEAEBE0AF9" blockId="5.[151,1437,438,933]" pageId="5" pageNumber="216">
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In order to compare the North African (Moroccan)
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<taxonomicName id="4C074D4452442C3E35509AEEACB30AA1" authorityName="Chvala" authorityYear="1975" box="[801,964,655,681]" class="Insecta" family="Hybotidae" genus="Syndyas" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="216" phylum="Arthropoda" rank="species" species="subsabinios">
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<emphasis id="B973EAD552442C3E35509AEEACB30AA1" box="[801,964,655,681]" italics="true" pageId="5" pageNumber="216">S. subsabinios</emphasis>
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</taxonomicName>
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with the European (Spanish) population described by
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<bibRefCitation id="EF964B3652442C3E37599AD2AEBE0AC5" author="Chvala, M." box="[296,457,690,717]" pageId="5" pageNumber="216" pagination="257 - 261" refId="ref5660" refString="Chvala, M. (1975) A new Palaearctic species of Syndyas Loew from Spain, with synonymic notes on the genus Sabinios Jones (Diptera, Empididae). Acta Entomologica Bohemoeslovaca, 72, 257 - 261." type="journal article" year="1975">Chvála (1975)</bibRefCitation>
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, we have considered it worthwhile to make a re-description of the species by examination of the Moroccan specimen.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D654C52442C3D36B69A9BAD0B0B89" lastPageId="6" lastPageNumber="217" pageId="5" pageNumber="216" type="diagnosis">
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<paragraph id="8BB836C752442C3E36B69A9BAD9E0B89" blockId="5.[151,1437,438,933]" pageId="5" pageNumber="216">
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<emphasis id="B973EAD552442C3E36B69A9BAE350B1C" bold="true" box="[199,322,762,788]" pageId="5" pageNumber="216">Diagnosis.</emphasis>
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Black species with black legs except the fore tibia, extreme tip of fore femur and fore tarsomeres 1– 4 are yellow, with the apical portion of tarsomere 5 blackish. Wings with a very large and almost spherical blackish brown pterostigma, with blackish brown veins with vein R
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<subScript id="1783348252442C3E356F9B34AC400B6B" attach="left" box="[798,823,853,867]" fontSize="6" pageId="5" pageNumber="216">2+3</subScript>
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abruptly curved around the pterostigma; costal cell and most of the basal cells are bare, lacking microtrichia.
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</paragraph>
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<paragraph id="8BB836C752442C3E36B69BEBADF30BAD" blockId="5.[151,1437,438,933]" box="[199,644,906,933]" pageId="5" pageNumber="216">
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<emphasis id="B973EAD552442C3E36B69BEBAEBE0BAD" bold="true" box="[199,457,906,933]" pageId="5" pageNumber="216">Re-description. Male.</emphasis>
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Length:
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<quantity id="4CFF9B2252442C3E345F9BEBADF30BAD" box="[558,644,906,933]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="216" unit="mm" value="4.5">4.5 mm</quantity>
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</paragraph>
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<caption id="DF78664F52442C3E36E99FF6AC400FB8" ID-DOI="http://doi.org/10.5281/zenodo.7224487" ID-Zenodo-Dep="7224487" box="[152,823,1943,1968]" httpUri="https://zenodo.org/record/7224487/files/figure.png" pageId="5" pageNumber="216" startId="5.[152,255,1943,1967]" targetBox="[151,1436,1010,1920]" targetPageId="5">
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<paragraph id="8BB836C752442C3E36E99FF6AC400FB8" blockId="5.[152,823,1943,1968]" box="[152,823,1943,1968]" pageId="5" pageNumber="216">
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<emphasis id="B973EAD552442C3E36E99FF6AE6F0FA7" bold="true" box="[152,280,1943,1967]" pageId="5" pageNumber="216">FIGURE 4.</emphasis>
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Habitus male of
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<taxonomicName id="4C074D4452442C3E37B69FF9ADEF0FB8" authorityName="Chvala" authorityYear="1975" box="[455,664,1944,1968]" class="Insecta" family="Hybotidae" genus="Syndyas" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="216" phylum="Arthropoda" rank="species" species="subsabinios">
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<emphasis id="B973EAD552442C3E37B69FF9ADEF0FB8" box="[455,664,1944,1968]" italics="true" pageId="5" pageNumber="216">Syndyas subsabinios</emphasis>
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</taxonomicName>
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. Scale: 500 µm
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</paragraph>
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</caption>
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<paragraph id="8BB836C752472C3D36B698F6AC6A0949" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="217">Head black in ground color. Holoptic, eyes meeting on frons, facets more enlarged above. Antenna black, inserted above middle of head in profile; scape very small; pedicel almost as small as scape; postpedicel ovate, slightly elongate; arista-like stylus about twice longer than basal segments with terminal sensillum. Ocellar tubercle prominent, with pair of long black bristles. Mouthparts directed forward, proboscis short blackish brown; palpus slender and shorter than proboscis, with 1 bristle apically.</paragraph>
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<paragraph id="8BB836C752472C3D36B6992AACF109A5" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="217">Thorax black in ground color, strongly arched above, with large thoracic setae brownish; four rows of acrostichalsetae, dorsocentral setae numerous, rows widely separated ending in long prescutellars; pair of long blackish apical scutellars and some shorter marginal hairs on sides.</paragraph>
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<paragraph id="8BB836C752472C3D36B699D6AD280A69" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="217">
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Wings brownish clouded, covered with microtrichia except costal cell and most of basal cells devoid of microtrichia; pterostigma blackish brown and clearly visible, very large and almost spherical; costa running to tip of M
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<subScript id="1783348252472C3D36BF9A70AFA10A17" attach="left" box="[206,214,529,543]" fontSize="6" pageId="6" pageNumber="217">1</subScript>
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; anal cell much longer than basal cells; radial sector very short; R
|
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<subScript id="1783348252472C3D35C19A70ACBE0A17" attach="left" box="[944,969,529,543]" fontSize="6" pageId="6" pageNumber="217">2+3</subScript>
|
||
abruptly curved around pterostigma; vein separating basal cells br and bm faint; veins R
|
||
<subScript id="1783348252472C3D34D19A54ADCD0A4B" attach="left" box="[672,698,565,579]" fontSize="6" pageId="6" pageNumber="217">4+5</subScript>
|
||
and M
|
||
<subScript id="1783348252472C3D357B9A54AC650A4B" attach="left" box="[778,786,565,579]" fontSize="6" pageId="6" pageNumber="217">1</subScript>
|
||
diverge slightly before ending in costa; CuA closing anal cell very slightly recurrent. Halter black.
|
||
</paragraph>
|
||
<paragraph id="8BB836C752472C3D36B69A0AAC2C0B31" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="217">Legs black, with fore and mid tibiae, extreme tips of fore and mid femora and first four tarsal segments of all legs yellow, apical part of last tarsal segment blackish. Anterior two leg pairs slender, hind leg elongated with femora thickened, dilated towards tip; hind tibia conspicuously dilated towards tip; hind metatarsus distinctly dilated, thicker than fore and mid metatarsi. Hind femur with row of strong, long black ventral spine-like bristles (double row on basal third). Fore and mid tibiae with long hairs posteroventrally; mid tibia with shorter pubescence and long bristles above; hind tibia with about 7–8 long bristles.</paragraph>
|
||
<paragraph id="8BB836C752472C3D36B69B22AD0B0B89" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="217">Abdomen blackish, long, covered with scattered pale hairs more dense on two basal tergites and marginal parts of sternites. Genitalia lost during sampling.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D654C52472C3D36B69BEAAE550BE5" pageId="6" pageNumber="217" type="discussion">
|
||
<paragraph id="8BB836C752472C3D36B69BEAAE550BE5" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="217">
|
||
<emphasis id="B973EAD552472C3D36B69BEAAE4C0BAD" bold="true" box="[199,315,907,933]" pageId="6" pageNumber="217">Remarks.</emphasis>
|
||
According to the original description by
|
||
<bibRefCitation id="EF964B3652472C3D354F9BEAAC9E0BAD" author="Chvala, M." box="[830,1001,906,933]" pageId="6" pageNumber="217" pagination="257 - 261" refId="ref5660" refString="Chvala, M. (1975) A new Palaearctic species of Syndyas Loew from Spain, with synonymic notes on the genus Sabinios Jones (Diptera, Empididae). Acta Entomologica Bohemoeslovaca, 72, 257 - 261." type="journal article" year="1975">Chvála (1975)</bibRefCitation>
|
||
, the Moroccan specimen is entirely compatible with the Spanish specimen, with no morphological differences despite the loss of the male genitalia for comparison.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |