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<document ID-DOI="10.11646/megataxa.8.1.1" ID-ISSN="2703-3090" ID-Zenodo-Dep="7526571" approvalRequired="466" approvalRequired_for_bibRefs="26" approvalRequired_for_document="1" approvalRequired_for_illustrations="4" approvalRequired_for_matCits="119" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="233" approvalRequired_for_treatments="77" checkinTime="1673467281002" checkinUser="plazi" docAuthor="Irfan, Muhammad, Zhang, Zhi-Sheng &amp; Peng, Xian-Jin" docDate="2022" docId="03AE87CEBDAFFF6CFC8C3F3EFC50F9A7" docLanguage="en" docName="megataxa.8.1.1.pdf" docOrigin="Megataxa 8 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.3:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="3" docTitle="Ryojius furcatus Irfan &amp; Zhang &amp; Peng 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="256" masterDocId="FF97FFB6BD52FF93FFA63955FFF1FF82" masterDocTitle="Survey of Linyphiidae (Arachnida: Araneae) spiders from Yunnan, China" masterLastPageNumber="292" masterPageNumber="1" pageNumber="254" updateTime="1673469776586" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Survey of Linyphiidae (Arachnida: Araneae) spiders from Yunnan, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Irfan, Muhammad</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-0445-9612</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, 410081, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Zhi-Sheng</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-9304-1789</mods:nameIdentifier>
<mods:affiliation>Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peng, Xian-Jin</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2614-3910</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, 410081, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Megataxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-27</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>8</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>292</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/megataxa.8.1.1</mods:identifier>
<mods:identifier type="ISSN">2703-3090</mods:identifier>
<mods:identifier type="Zenodo-Dep">7526571</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7526764" ID-Zenodo-Dep="7526764" LSID="urn:lsid:plazi:treatment:03AE87CEBDAFFF6CFC8C3F3EFC50F9A7" httpUri="http://treatment.plazi.org/id/03AE87CEBDAFFF6CFC8C3F3EFC50F9A7" lastPageId="255" lastPageNumber="256" pageId="253" pageNumber="254">
<subSubSection box="[810,1262,1642,1669]" pageId="253" pageNumber="254" type="nomenclature">
<paragraph blockId="253.[810,1262,1642,1701]" box="[810,1262,1642,1669]" pageId="253" pageNumber="254">
<heading box="[810,1262,1642,1669]" fontSize="11" level="2" pageId="253" pageNumber="254" reason="2">
<emphasis bold="true" box="[810,1262,1642,1669]" pageId="253" pageNumber="254">
<taxonomicName authority="Irfan &amp; Zhang &amp; Peng, 2022" authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" box="[810,999,1642,1669]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="254" phylum="Arthropoda" rank="species" species="furcatus" status="sp. nov.">
<emphasis bold="true" box="[810,999,1642,1669]" italics="true" pageId="253" pageNumber="254">Ryojius furcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1007,1096,1643,1669]" pageId="253" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
(ĀḦƥŜƦ)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="253" pageNumber="254" type="description">
<paragraph blockId="253.[810,1262,1642,1701]" box="[810,999,1675,1701]" pageId="253" pageNumber="254">
<figureCitation box="[810,999,1675,1701]" captionStart-0="FIGURE 287" captionStart-1="FIGURE 288" captionStart-2="FIGURE 289" captionStart-3="FIGURE 290" captionStartId-0="252.[136,235,1740,1764]" captionStartId-1="253.[136,235,1100,1124]" captionStartId-2="254.[136,235,1868,1892]" captionStartId-3="255.[136,235,812,836]" captionTargetBox-0="[342,1247,167,1726]" captionTargetBox-1="[237,1351,155,1080]" captionTargetBox-2="[264,1323,184,1849]" captionTargetBox-3="[148,765,178,793]" captionTargetId-0="figure-25@252.[340,1247,167,1726]" captionTargetId-1="figure-17@253.[237,1351,155,1080]" captionTargetId-2="figure-25@254.[264,1323,184,1849]" captionTargetId-3="figure-17@255.[148,765,178,793]" captionTargetPageId-0="252" captionTargetPageId-1="253" captionTargetPageId-2="254" captionTargetPageId-3="255" captionText-0="FIGURE 287. Ryojius furcatus sp. nov., male palp of holotype. A prolateral view B retrolateral view C dorsal view D ventral view." captionText-1="FIGURE 288. Ryojius furcatus sp. nov., female (one of paratypes). A, C Epigyne, ventral view B Epigyne, lateral view D Vulva, dorsal view." captionText-2="FIGURE 289. Ryojius furcatus sp. nov., male holotype (A, B) and female (one of paratypes, C, D). A, C Habitus, dorsal view B, D Habitus, ventral view." captionText-3="FIGURE 290. Distribution of Ryojius furcatus sp. nov.." figureDoi-0="http://doi.org/10.5281/zenodo.7527211" figureDoi-1="http://doi.org/10.5281/zenodo.7527213" figureDoi-2="http://doi.org/10.5281/zenodo.7527215" figureDoi-3="http://doi.org/10.5281/zenodo.7527217" httpUri-0="https://zenodo.org/record/7527211/files/figure.png" httpUri-1="https://zenodo.org/record/7527213/files/figure.png" httpUri-2="https://zenodo.org/record/7527215/files/figure.png" httpUri-3="https://zenodo.org/record/7527217/files/figure.png" pageId="253" pageNumber="254">Figures 287290</figureCitation>
</paragraph>
<paragraph blockId="253.[810,1452,1738,2085]" pageId="253" pageNumber="254">
<emphasis bold="true" box="[810,886,1739,1765]" pageId="253" pageNumber="254">Types.</emphasis>
<materialsCitation collectingDate="2004-05-04" collectionCode="CGY" collectorName="Charles Griswold &amp; D. H. Kavanaugh" country="China" county="Fugong County" elevation="2563" latitude="27.16636" location="Lumadeng Township" longLatPrecision="1" longitude="98.77667" pageId="253" pageNumber="254" specimenCode="CGY28" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus box="[896,1000,1739,1765]" pageId="253" pageNumber="254">Holotype</typeStatus>
♁,
<emphasis bold="true" box="[1048,1251,1738,1765]" pageId="253" pageNumber="254">
<collectingCountry box="[1048,1143,1738,1764]" name="China" pageId="253" pageNumber="254">CHINA</collectingCountry>
,
<collectingRegion box="[1157,1251,1739,1765]" country="China" name="Yunnan" pageId="253" pageNumber="254">Yunnan</collectingRegion>
</emphasis>
,
<collectingCounty box="[1268,1448,1739,1765]" pageId="253" pageNumber="254">Fugong County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:03AE87CEBDAFFF6CFC8C3F3EFC50F9A7:8ED86003BDAFFF6EFC8C3FBEFBD0F887" box="[810,1057,1771,1797]" country="China" county="Fugong County" latitude="27.16636" longLatPrecision="1" longitude="98.77667" name="Lumadeng Township" pageId="253" pageNumber="254" stateProvince="Yunnan">Lumadeng Township</location>
,
<collectingMethod box="[1076,1383,1771,1797]" pageId="253" pageNumber="254">Shibali forest station trap</collectingMethod>
area,
<geoCoordinate box="[810,949,1803,1829]" degrees="27.16636" direction="north" orientation="latitude" pageId="253" pageNumber="254" precision="1" value="27.16636">27.16636°N</geoCoordinate>
,
<geoCoordinate box="[971,1108,1803,1829]" degrees="98.77667" direction="east" orientation="longitude" pageId="253" pageNumber="254" precision="1" value="98.77667">98.77667°E</geoCoordinate>
, alt.
<quantity box="[1183,1264,1803,1830]" metricMagnitude="3" metricUnit="m" metricValue="2.563" pageId="253" pageNumber="254" unit="m" value="2563.0">
<elevation box="[1183,1264,1803,1830]" metricMagnitude="3" metricUnit="m" metricValue="2.563" pageId="253" pageNumber="254" unit="m" value="2563.0">2563m</elevation>
</quantity>
,
<date box="[1286,1447,1803,1829]" pageId="253" pageNumber="254" value="2004-05-04">
<collectingDate box="[1286,1447,1803,1829]" pageId="253" pageNumber="254" value="2004-05-04">4 May 2004</collectingDate>
</date>
,
<collectorName box="[810,1010,1835,1861]" pageId="253" pageNumber="254">Charles Griswold</collectorName>
and
<collectorName box="[1072,1276,1835,1861]" pageId="253" pageNumber="254">D. H. Kavanaugh</collectorName>
leg. (
<specimenCode box="[1347,1440,1835,1861]" collectionCode="CGY" pageId="253" pageNumber="254">CGY28</specimenCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="2004-05-04" collectionCode="CGY" collectorName="Charles Griswold &amp; D. H. Kavanaugh" country="China" county="Fugong County" elevation="2563" latitude="27.16636" location="Lumadeng Township" longLatPrecision="1" longitude="98.77667" pageId="253" pageNumber="254" specimenCode="CGY28" specimenCount="10" specimenCount-female="10" stateProvince="Yunnan" typeStatus="paratype">
<emphasis bold="true" box="[810,939,1867,1893]" pageId="253" pageNumber="254">
<typeStatus box="[810,934,1867,1893]" pageId="253" pageNumber="254">Paratypes</typeStatus>
:
</emphasis>
1♁
<specimenCount box="[993,1047,1868,1895]" pageId="253" pageNumber="254" type="female">10♀</specimenCount>
, same data as holotype male (
<specimenCode box="[818,911,1899,1925]" collectionCode="CGY" pageId="253" pageNumber="254">CGY28</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2004-05-08" collectionCode="CGY" collectorName="Charles Griswold &amp; D. H. Kavanaugh &amp; Heng-mei Yan" country="China" elevation="2475" latitude="27.16337" location="Forest" longLatPrecision="1" longitude="98.78208" municipality="Yamu river" pageId="253" pageNumber="254" specimenCode="CGY37" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount box="[933,967,1899,1925]" pageId="253" pageNumber="254" type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03AE87CEBDAFFF6CFC8C3F3EFC50F9A7:8ED86003BDAFFF6EFC7E3E3EFBEFF807" box="[984,1054,1899,1925]" country="China" latitude="27.16337" longLatPrecision="1" longitude="98.78208" municipality="Yamu river" name="Forest" pageId="253" pageNumber="254" stateProvince="Yunnan">Forest</location>
along shelf above
<collectingMunicipality box="[1273,1403,1899,1925]" pageId="253" pageNumber="254">Yamu river</collectingMunicipality>
,
<locationDeviation location="Shibali" pageId="253" pageNumber="254">
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="253" pageNumber="254" unit="km" value="0.4">0.4 km</quantity>
SE Shibali
</locationDeviation>
,
<geoCoordinate box="[994,1134,1931,1957]" degrees="27.16337" direction="north" orientation="latitude" pageId="253" pageNumber="254" precision="1" value="27.16337">27.16337°N</geoCoordinate>
,
<geoCoordinate box="[1148,1284,1931,1957]" degrees="98.78208" direction="east" orientation="longitude" pageId="253" pageNumber="254" precision="1" value="98.78208">98.78208°E</geoCoordinate>
, alt.
<quantity box="[1343,1423,1931,1958]" metricMagnitude="3" metricUnit="m" metricValue="2.475" pageId="253" pageNumber="254" unit="m" value="2475.0">
<elevation box="[1343,1423,1931,1958]" metricMagnitude="3" metricUnit="m" metricValue="2.475" pageId="253" pageNumber="254" unit="m" value="2475.0">2475m</elevation>
</quantity>
,
<date pageId="253" pageNumber="254" value="2004-05-08">
<collectingDate pageId="253" pageNumber="254" value="2004-05-08">8 May 2004</collectingDate>
</date>
,
<collectorName box="[943,1142,1963,1989]" pageId="253" pageNumber="254">Charles Griswold</collectorName>
and
<collectorName box="[1201,1402,1963,1989]" pageId="253" pageNumber="254">D. H. Kavanaugh</collectorName>
and
<collectorName box="[810,972,1995,2021]" pageId="253" pageNumber="254">Heng-mei Yan</collectorName>
leg. (
<specimenCode box="[1035,1128,1995,2021]" collectionCode="CGY" pageId="253" pageNumber="254">CGY37</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="255" lastPageNumber="256" pageId="253" pageNumber="254" type="etymology">
<paragraph blockId="253.[810,1452,1738,2085]" lastBlockId="255.[136,778,907,2085]" lastPageId="255" lastPageNumber="256" pageId="253" pageNumber="254">
<emphasis bold="true" box="[858,992,2027,2053]" pageId="253" pageNumber="254">Etymology.</emphasis>
This epithet derives from the Latin adjective “
<taxonomicName authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" box="[934,1025,2059,2085]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="253" pageNumber="254" phylum="Arthropoda" rank="species" species="furcatus">
<emphasis box="[934,1025,2059,2085]" italics="true" pageId="253" pageNumber="254">furcatus</emphasis>
</taxonomicName>
”, meaning “forked”, referring to the forked tip of the terminal apophysis in ventral view in the male palp.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7527215" ID-Zenodo-Dep="7527215" httpUri="https://zenodo.org/record/7527215/files/figure.png" pageId="254" pageNumber="255" startId="254.[136,235,1868,1892]" targetBox="[264,1323,184,1849]" targetPageId="254">
<paragraph blockId="254.[136,1451,1868,1925]" pageId="254" pageNumber="255">
<emphasis bold="true" box="[136,286,1868,1892]" pageId="254" pageNumber="255">FIGURE 289.</emphasis>
<taxonomicName authority="Irfan &amp; Zhang &amp; Peng, 2022" authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" box="[292,456,1869,1893]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="254" pageNumber="255" phylum="Arthropoda" rank="species" species="furcatus" status="sp. nov.">
<emphasis box="[292,456,1869,1893]" italics="true" pageId="254" pageNumber="255">Ryojius furcatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[463,543,1869,1893]" pageId="254" pageNumber="255">
<taxonomicNameLabel box="[463,543,1869,1893]" pageId="254" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (A, B) and female (one of paratypes, C, D).
<emphasis bold="true" box="[1156,1205,1868,1892]" pageId="254" pageNumber="255">A, C</emphasis>
Habitus, dorsal view
<emphasis bold="true" pageId="254" pageNumber="255">B, D</emphasis>
Habitus, ventral view.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7527217" ID-Zenodo-Dep="7527217" box="[136,705,812,837]" httpUri="https://zenodo.org/record/7527217/files/figure.png" pageId="255" pageNumber="256" startId="255.[136,235,812,836]" targetBox="[148,765,178,793]" targetPageId="255">
<paragraph blockId="255.[136,705,812,837]" box="[136,705,812,837]" pageId="255" pageNumber="256">
<emphasis bold="true" box="[136,286,812,836]" pageId="255" pageNumber="256">FIGURE 290.</emphasis>
Distribution of
<taxonomicName authority="Irfan &amp; Zhang &amp; Peng, 2022" authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" box="[448,612,813,837]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="furcatus" status="sp. nov.">
<emphasis box="[448,612,813,837]" italics="true" pageId="255" pageNumber="256">Ryojius furcatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[618,699,813,837]" pageId="255" pageNumber="256">
<taxonomicNameLabel box="[618,699,813,837]" pageId="255" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</caption>
<subSubSection pageId="255" pageNumber="256" type="diagnosis">
<paragraph blockId="255.[136,778,907,2085]" pageId="255" pageNumber="256">
<emphasis bold="true" box="[184,305,971,997]" pageId="255" pageNumber="256">Diagnosis.</emphasis>
The new species resembles
<taxonomicName authority="(Chen &amp; Yin, 2000)" baseAuthorityName="Chen &amp; Yin" baseAuthorityYear="2000" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="nanyuensis">
<emphasis italics="true" pageId="255" pageNumber="256">Ryojius nanyuensis</emphasis>
(
<bibRefCitation author="Chen, J. &amp; Yin, C. M." box="[283,518,1003,1029]" pageId="255" pageNumber="256" pagination="86 - 93" refId="ref86737" refString="Chen, J. &amp; Yin, C. M. (2000) On five species of linyphiid spiders from Hunan, China (Araneae, Linyphiidae) Acta Arachnologica Sinica, 9, 86 - 93." type="journal article" year="2000">Chen &amp; Yin, 2000</bibRefCitation>
)
</taxonomicName>
in having a similar paracymbium and scape of epigyne (
<figureCitation box="[606,774,1035,1061]" captionStart="FIGURE 287" captionStartId="252.[136,235,1740,1764]" captionTargetBox="[342,1247,167,1726]" captionTargetId="figure-25@252.[340,1247,167,1726]" captionTargetPageId="252" captionText="FIGURE 287. Ryojius furcatus sp. nov., male palp of holotype. A prolateral view B retrolateral view C dorsal view D ventral view." figureDoi="http://doi.org/10.5281/zenodo.7527211" httpUri="https://zenodo.org/record/7527211/files/figure.png" pageId="255" pageNumber="256">Figs 287AD</figureCitation>
,
<figureCitation box="[136,235,1067,1093]" captionStart="FIGURE 288" captionStartId="253.[136,235,1100,1124]" captionTargetBox="[237,1351,155,1080]" captionTargetId="figure-17@253.[237,1351,155,1080]" captionTargetPageId="253" captionText="FIGURE 288. Ryojius furcatus sp. nov., female (one of paratypes). A, C Epigyne, ventral view B Epigyne, lateral view D Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7527213" httpUri="https://zenodo.org/record/7527213/files/figure.png" pageId="255" pageNumber="256">288AD</figureCitation>
;
<bibRefCitation author="Saaristo, M. I. &amp; Tu, L. H. &amp; Li, S. Q." box="[247,511,1067,1093]" pageId="255" pageNumber="256" pagination="383 - 401" refId="ref89901" refString="Saaristo, M. I., Tu, L. H. &amp; Li, S. Q. (2006) A review of Chinese micronetine species (Araneae, Linyphiidae) Part I, species of ex- Arcuphantes and ex- Centromerus. Animal Biology, 56 (3), 383 - 401. https: // doi. org / 10.1163 / 157075606778441886" type="journal article" year="2006">Tu, Saaristo &amp; Li, 2006</bibRefCitation>
figs 1523), but can be distinguished by the distal tip of lamella characteristca webbed feet-shaped in
<taxonomicName authority="Irfan &amp; Zhang &amp; Peng, 2022" authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" box="[389,509,1131,1157]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="furcatus" status="sp. nov.">
<emphasis box="[389,509,1131,1157]" italics="true" pageId="255" pageNumber="256">R. furcatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[515,602,1131,1157]" pageId="255" pageNumber="256">
<taxonomicNameLabel box="[515,602,1131,1157]" pageId="255" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[616,764,1131,1157]" captionStart="FIGURE 287" captionStartId="252.[136,235,1740,1764]" captionTargetBox="[342,1247,167,1726]" captionTargetId="figure-25@252.[340,1247,167,1726]" captionTargetPageId="252" captionText="FIGURE 287. Ryojius furcatus sp. nov., male palp of holotype. A prolateral view B retrolateral view C dorsal view D ventral view." figureDoi="http://doi.org/10.5281/zenodo.7527211" httpUri="https://zenodo.org/record/7527211/files/figure.png" pageId="255" pageNumber="256">Fig. 287A, D</figureCitation>
), whereas flower-shaped in
<taxonomicName baseAuthorityName="Chen &amp; Yin" baseAuthorityYear="2000" box="[435,592,1164,1189]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="nanyuensis">
<emphasis box="[435,592,1164,1189]" italics="true" pageId="255" pageNumber="256">R. nanyuensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Saaristo, M. I. &amp; Tu, L. H. &amp; Li, S. Q." pageId="255" pageNumber="256" pagination="383 - 401" refId="ref89901" refString="Saaristo, M. I., Tu, L. H. &amp; Li, S. Q. (2006) A review of Chinese micronetine species (Araneae, Linyphiidae) Part I, species of ex- Arcuphantes and ex- Centromerus. Animal Biology, 56 (3), 383 - 401. https: // doi. org / 10.1163 / 157075606778441886" type="journal article" year="2006">Tu, Saaristo &amp; Li, 2006</bibRefCitation>
figs 15, 16). Terminal apophysis tip forked in
<taxonomicName authority="Irfan &amp; Zhang &amp; Peng, 2022" authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="furcatus" status="sp. nov.">
<emphasis italics="true" pageId="255" pageNumber="256">R. furcatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[230,316,1227,1253]" pageId="255" pageNumber="256">
<taxonomicNameLabel box="[230,316,1227,1253]" pageId="255" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[328,442,1227,1253]" captionStart="FIGURE 287" captionStartId="252.[136,235,1740,1764]" captionTargetBox="[342,1247,167,1726]" captionTargetId="figure-25@252.[340,1247,167,1726]" captionTargetPageId="252" captionText="FIGURE 287. Ryojius furcatus sp. nov., male palp of holotype. A prolateral view B retrolateral view C dorsal view D ventral view." figureDoi="http://doi.org/10.5281/zenodo.7527211" httpUri="https://zenodo.org/record/7527211/files/figure.png" pageId="255" pageNumber="256">Fig.287D</figureCitation>
), whereas relatively broad with round end in
<taxonomicName baseAuthorityName="Chen &amp; Yin" baseAuthorityYear="2000" box="[288,444,1260,1285]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="nanyuensis">
<emphasis box="[288,444,1260,1285]" italics="true" pageId="255" pageNumber="256">R. nanyuensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Saaristo, M. I. &amp; Tu, L. H. &amp; Li, S. Q." box="[461,731,1259,1285]" pageId="255" pageNumber="256" pagination="383 - 401" refId="ref89901" refString="Saaristo, M. I., Tu, L. H. &amp; Li, S. Q. (2006) A review of Chinese micronetine species (Araneae, Linyphiidae) Part I, species of ex- Arcuphantes and ex- Centromerus. Animal Biology, 56 (3), 383 - 401. https: // doi. org / 10.1163 / 157075606778441886" type="journal article" year="2006">Tu, Saaristo &amp; Li, 2006</bibRefCitation>
fig. 18). Apex of embolus finger-shaped in
<taxonomicName authority="Irfan &amp; Zhang &amp; Peng, 2022" authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" box="[565,684,1291,1317]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="furcatus" status="sp. nov.">
<emphasis box="[565,684,1291,1317]" italics="true" pageId="255" pageNumber="256">R. furcatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[690,778,1291,1317]" pageId="255" pageNumber="256">
<taxonomicNameLabel box="[690,778,1291,1317]" pageId="255" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[144,299,1323,1349]" captionStart="FIGURE 287" captionStartId="252.[136,235,1740,1764]" captionTargetBox="[342,1247,167,1726]" captionTargetId="figure-25@252.[340,1247,167,1726]" captionTargetPageId="252" captionText="FIGURE 287. Ryojius furcatus sp. nov., male palp of holotype. A prolateral view B retrolateral view C dorsal view D ventral view." figureDoi="http://doi.org/10.5281/zenodo.7527211" httpUri="https://zenodo.org/record/7527211/files/figure.png" pageId="255" pageNumber="256">Fig. 287A, D</figureCitation>
), whereas tongue-shaped in
<taxonomicName baseAuthorityName="Chen &amp; Yin" baseAuthorityYear="2000" box="[622,778,1324,1349]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="nanyuensis">
<emphasis box="[622,778,1324,1349]" italics="true" pageId="255" pageNumber="256">R. nanyuensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Saaristo, M. I. &amp; Tu, L. H. &amp; Li, S. Q." box="[144,403,1355,1381]" pageId="255" pageNumber="256" pagination="383 - 401" refId="ref89901" refString="Saaristo, M. I., Tu, L. H. &amp; Li, S. Q. (2006) A review of Chinese micronetine species (Araneae, Linyphiidae) Part I, species of ex- Arcuphantes and ex- Centromerus. Animal Biology, 56 (3), 383 - 401. https: // doi. org / 10.1163 / 157075606778441886" type="journal article" year="2006">Tu, Saaristo &amp; Li, 2006</bibRefCitation>
fig. 18). Posterior median plate of epigyne posteriorly with two tooth-shaped apophyses in
<taxonomicName authority="Irfan &amp; Zhang &amp; Peng, 2022" authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="furcatus" status="sp. nov.">
<emphasis italics="true" pageId="255" pageNumber="256">R. furcatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[233,323,1419,1445]" pageId="255" pageNumber="256">
<taxonomicNameLabel box="[233,323,1419,1445]" pageId="255" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[338,456,1419,1445]" captionStart="FIGURE 288" captionStartId="253.[136,235,1100,1124]" captionTargetBox="[237,1351,155,1080]" captionTargetId="figure-17@253.[237,1351,155,1080]" captionTargetPageId="253" captionText="FIGURE 288. Ryojius furcatus sp. nov., female (one of paratypes). A, C Epigyne, ventral view B Epigyne, lateral view D Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7527213" httpUri="https://zenodo.org/record/7527213/files/figure.png" pageId="255" pageNumber="256">Fig. 288D</figureCitation>
), whereas without apophysis in
<taxonomicName baseAuthorityName="Chen &amp; Yin" baseAuthorityYear="2000" box="[169,327,1452,1477]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="nanyuensis">
<emphasis box="[169,327,1452,1477]" italics="true" pageId="255" pageNumber="256">R. nanyuensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Saaristo, M. I. &amp; Tu, L. H. &amp; Li, S. Q." box="[347,627,1451,1477]" pageId="255" pageNumber="256" pagination="383 - 401" refId="ref89901" refString="Saaristo, M. I., Tu, L. H. &amp; Li, S. Q. (2006) A review of Chinese micronetine species (Araneae, Linyphiidae) Part I, species of ex- Arcuphantes and ex- Centromerus. Animal Biology, 56 (3), 383 - 401. https: // doi. org / 10.1163 / 157075606778441886" type="journal article" year="2006">Tu, Saaristo &amp; Li, 2006</bibRefCitation>
figs 2022). Dorsal part of scape as wide as long in
<taxonomicName authority="Irfan &amp; Zhang &amp; Peng, 2022" authorityName="Irfan &amp; Zhang &amp; Peng" authorityYear="2022" box="[564,684,1483,1509]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="furcatus" status="sp. nov.">
<emphasis box="[564,684,1483,1509]" italics="true" pageId="255" pageNumber="256">R. furcatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[690,778,1483,1509]" pageId="255" pageNumber="256">
<taxonomicNameLabel box="[690,778,1483,1509]" pageId="255" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[144,291,1515,1541]" captionStart="FIGURE 288" captionStartId="253.[136,235,1100,1124]" captionTargetBox="[237,1351,155,1080]" captionTargetId="figure-17@253.[237,1351,155,1080]" captionTargetPageId="253" captionText="FIGURE 288. Ryojius furcatus sp. nov., female (one of paratypes). A, C Epigyne, ventral view B Epigyne, lateral view D Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7527213" httpUri="https://zenodo.org/record/7527213/files/figure.png" pageId="255" pageNumber="256">Fig. 288A, C</figureCitation>
), whereas longer than wide in
<taxonomicName baseAuthorityName="Chen &amp; Yin" baseAuthorityYear="2000" box="[625,778,1516,1541]" class="Arachnida" family="Linyphiidae" genus="Ryojius" kingdom="Animalia" order="Araneae" pageId="255" pageNumber="256" phylum="Arthropoda" rank="species" species="nanyuensis">
<emphasis box="[625,778,1516,1541]" italics="true" pageId="255" pageNumber="256">R. nanyuensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Saaristo, M. I. &amp; Tu, L. H. &amp; Li, S. Q." box="[144,407,1547,1573]" pageId="255" pageNumber="256" pagination="383 - 401" refId="ref89901" refString="Saaristo, M. I., Tu, L. H. &amp; Li, S. Q. (2006) A review of Chinese micronetine species (Araneae, Linyphiidae) Part I, species of ex- Arcuphantes and ex- Centromerus. Animal Biology, 56 (3), 383 - 401. https: // doi. org / 10.1163 / 157075606778441886" type="journal article" year="2006">Tu, Saaristo &amp; Li, 2006</bibRefCitation>
figs 2022)
</paragraph>
</subSubSection>
<subSubSection pageId="255" pageNumber="256" type="description">
<paragraph blockId="255.[136,778,907,2085]" lastBlockId="255.[810,1452,171,1573]" pageId="255" pageNumber="256">
<emphasis bold="true" box="[184,402,1579,1605]" pageId="255" pageNumber="256">Description. Male</emphasis>
(
<typeStatus box="[420,522,1579,1605]" pageId="255" pageNumber="256">holotype</typeStatus>
,
<figureCitation box="[538,696,1579,1605]" captionStart="FIGURE 289" captionStartId="254.[136,235,1868,1892]" captionTargetBox="[264,1323,184,1849]" captionTargetId="figure-25@254.[264,1323,184,1849]" captionTargetPageId="254" captionText="FIGURE 289. Ryojius furcatus sp. nov., male holotype (A, B) and female (one of paratypes, C, D). A, C Habitus, dorsal view B, D Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7527215" httpUri="https://zenodo.org/record/7527215/files/figure.png" pageId="255" pageNumber="256">Fig. 289A, B</figureCitation>
): Total length: 1.87. Carapace 0.99 long, 0.77 wide, brown; fovea, cervical and radial grooves distinct. Clypeus 0.22 high. Sternum longer than wide, dark yellow,, sparsely covered with microsetae. Labium wider than long. Maxillae long, distal end broad with scopulae. Chelicerae with four promarginal and five retromarginal teeth. AER recurved, PER straight, slightly wider. Eye sizes and interdistances: AME 0.04, ALE 0.06, PME 0.05, PLE 0.06, AMEAME 0.02, PMEPME 0.05, AMEALE, 0.03, PMEPLE 0.07, AMEPME 0.04, ALEALE 0.28, PLEPLE 0.33, ALEPLE 0.01. Length of legs: I 2.47 (0.71, 0.85, 0.52, 0.39), II 2.14 (0.66, 0.75, 0.47, 0.36), III 1.70 (0.53, 0.56, 0.32, 0.29), IV 2.22 (0.70, 0.79, 0.39, 0.34). Leg formula I-IV-II-III. Tm I 0.28 and Tm IV 0.17. Tibial spine formula: 2-2-1-1. Abdomen 1.03 long, 0.69 wide, oval, greenish-grey, dorsally with two pairs of sigillae; ventral side greenish-grey.
</paragraph>
<paragraph blockId="255.[810,1452,171,1573]" pageId="255" pageNumber="256">
Palp (
<figureCitation box="[926,1084,235,261]" captionStart="FIGURE 287" captionStartId="252.[136,235,1740,1764]" captionTargetBox="[342,1247,167,1726]" captionTargetId="figure-25@252.[340,1247,167,1726]" captionTargetPageId="252" captionText="FIGURE 287. Ryojius furcatus sp. nov., male palp of holotype. A prolateral view B retrolateral view C dorsal view D ventral view." figureDoi="http://doi.org/10.5281/zenodo.7527211" httpUri="https://zenodo.org/record/7527211/files/figure.png" pageId="255" pageNumber="256">Fig. 287AD</figureCitation>
): Patella as long as tibia with a long dorsal spine; tibia with strongly sclerotized dorsal apophysis pointing laterally in dorsal view; with one retrolateral and one dorsal trichobothria. Cymbium with shallow depression at base. Paracymbium strongly scleortized, distal arm with V-shaped apex. The distal suprategular apophysis small, covered by the tegulum with narrow hook-shaped dorsal tip. Radix C-shaped. Lamella characteristca with two branches; posterior one hornshaped, anterior one long, extending forward, tip webbed feet-shaped. Terminal apophysis longer than wide, tip forked. Embolus conspicuously large, posteriorly curved about half circle, apex strongly sclerotized, finger-shaped with blunt end; thumb short, covered by the lamella characteristca; embolus proper with pointed end.
</paragraph>
<paragraph blockId="255.[810,1452,171,1573]" pageId="255" pageNumber="256">
<emphasis bold="true" box="[858,945,715,741]" pageId="255" pageNumber="256">Female</emphasis>
(one of the
<typeStatus box="[1092,1202,715,741]" pageId="255" pageNumber="256">paratypes</typeStatus>
,
<figureCitation box="[1216,1372,715,741]" captionStart="FIGURE 289" captionStartId="254.[136,235,1868,1892]" captionTargetBox="[264,1323,184,1849]" captionTargetId="figure-25@254.[264,1323,184,1849]" captionTargetPageId="254" captionText="FIGURE 289. Ryojius furcatus sp. nov., male holotype (A, B) and female (one of paratypes, C, D). A, C Habitus, dorsal view B, D Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7527215" httpUri="https://zenodo.org/record/7527215/files/figure.png" pageId="255" pageNumber="256">Fig. 289C, D</figureCitation>
): Total length: 2.15. Carapace 0.88 long, 0.67 wide, brown; fovea, cervical and radial grooves distinct. Clypeus 0.22 high. Sternum longer than wide, yellow,, sparsely covered with microsetae. Labium wider than long. Maxillae long, distal end broad with scopulae. Chelicerae with four promarginal and five retromarginal teeth. AER recurved, PER straight, slightly wider. Eye sizes and interdistances: AME 0.04, ALE 0.07, PME 0.05, PLE 0.07, AMEAME 0.02, PMEPME 0.06, AMEALE, 0.03, PMEPLE 0.05, AMEPME 0.05, ALEALE 0.28, PLEPLE 0.31, ALEPLE 0.01. Length of legs: I 2.27 (0.68, 0.78, 0.44, 0.37), II 1.98 (0.59, 0.66, 0.39, 0.34), III 1.68 (0.51, 0.53, 0.35, 0.29), IV 2.19 (0.68, 0.76, 0.41, 0.34). Leg formula I-IV-II-III. Tm I 0.23 and Tm IV 0.13. Tibial spine formula: 2-2-1-1. Abdomen 1.39 long, 1.03 wide, oval, pale, posteriorly with two greenish-grey chevrons; ventral side pale greenish-grey.
</paragraph>
<paragraph blockId="255.[810,1452,171,1573]" pageId="255" pageNumber="256">
Epigyne (
<figureCitation box="[974,1133,1291,1317]" captionStart="FIGURE 288" captionStartId="253.[136,235,1100,1124]" captionTargetBox="[237,1351,155,1080]" captionTargetId="figure-17@253.[237,1351,155,1080]" captionTargetPageId="253" captionText="FIGURE 288. Ryojius furcatus sp. nov., female (one of paratypes). A, C Epigyne, ventral view B Epigyne, lateral view D Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7527213" httpUri="https://zenodo.org/record/7527213/files/figure.png" pageId="255" pageNumber="256">Fig. 288AD</figureCitation>
) protruding; basal part of scape wide, posteriorly projecting, dorsally with a narrow tongue-shaped stretcher. Distal part of scape longer than wide with small lateral pockets. Posterior median plate posteriorly with two tooth-shaped apophyses. Spermathecae globular, situated at the anterior lateral sides of dorsal plate apices.
</paragraph>
</subSubSection>
<subSubSection pageId="255" pageNumber="256" type="distribution">
<paragraph blockId="255.[810,1452,171,1573]" pageId="255" pageNumber="256">
<emphasis bold="true" box="[858,1011,1515,1541]" pageId="255" pageNumber="256">Distribution.</emphasis>
Known only from the
<typeStatus box="[1305,1353,1515,1541]" pageId="255" pageNumber="256">type</typeStatus>
locality (
<figureCitation box="[818,917,1547,1573]" captionStart="FIGURE 290" captionStartId="255.[136,235,812,836]" captionTargetBox="[148,765,178,793]" captionTargetId="figure-17@255.[148,765,178,793]" captionTargetPageId="255" captionText="FIGURE 290. Distribution of Ryojius furcatus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.7527217" httpUri="https://zenodo.org/record/7527217/files/figure.png" pageId="255" pageNumber="256">Fig. 290</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>