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<document id="7755E2902B8640CC263BCA071352AF44" ID-CLB-Dataset="298350" ID-DOI="10.5194/fr-23-1-2020" ID-GBIF-Dataset="9e9ce0f7-220f-465f-91dc-1de099aae5c1" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965637" IM.bibliography_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1712927515474" checkinUser="felipe" docAuthor="Joyce, Walter G., Rollot, Yann &amp; Cifelli Correspondence, Richard L." docDate="2020" docId="03AE87943006FFD53E64FE1DEA2D342C" docLanguage="en" docName="FossRec.23.1.1-13.pdf" docOrigin="Fossil Record 23 (1)" docSource="http://dx.doi.org/10.5194/fr-23-1-2020" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.2:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="2" docTitle="Lakotemys australodakotensis Joyce &amp; Rollot &amp; Cifelli Correspondence 2020, gen. et sp. nov." docType="treatment" docUuid="00A9D083-7874-4022-AB3C-41B9FF71841E" docUuidSource="ZooBank" docVersion="5" lastPageNumber="7" masterDocId="FF97FFEC3004FFD33D0DFF95EB6F3155" masterDocTitle="A new species of baenid turtle from the Early Cretaceous Lakota Formation of South Dakota" masterLastPageNumber="13" masterPageNumber="1" pageNumber="3" updateTime="1718070541761" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9694EF004E641F9232FE144CA198A2C5">A new species of baenid turtle from the Early Cretaceous Lakota Formation of South Dakota</mods:title>
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<mods:namePart id="77C66CA2E60EF73A8BCEBF15047ABDEB">Joyce, Walter G.</mods:namePart>
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<mods:namePart id="2AE44E8C9E0F6A1C3707C71AD5C64A41">Rollot, Yann</mods:namePart>
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<mods:namePart id="8E67C489646E364161C29A68D7EDA8F4">Cifelli Correspondence, Richard L.</mods:namePart>
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<mods:title id="10F0863E637CEE5A4032DFDA1FDE0F46">Fossil Record</mods:title>
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<treatment id="03AE87943006FFD53E64FE1DEA2D342C" ID-DOI="http://doi.org/10.5281/zenodo.11557476" ID-GBIF-Taxon="232727668" ID-Zenodo-Dep="11557476" LSID="urn:lsid:zoobank.org:act:00A9D083-7874-4022-AB3C-41B9FF71841E" httpUri="http://treatment.plazi.org/id/03AE87943006FFD53E64FE1DEA2D342C" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
<subSubSection id="C31D65093006FFD13E64FE1DEE2030CA" box="[873,1359,392,416]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BB836823006FFD13E64FE1DEE2030CA" blockId="2.[873,1359,392,416]" box="[873,1359,392,416]" pageId="2" pageNumber="3">
<heading id="D0F081EE3006FFD13E64FE1DEE2030CA" box="[873,1359,392,416]" fontSize="10" level="2" pageId="2" pageNumber="3" reason="5">
<taxonomicName id="4C074D013006FFD13E64FE1DEFC630F5" authority="Joyce &amp; Rollot &amp; Cifelli Correspondence, 2020" authorityName="Joyce &amp; Rollot &amp; Cifelli Correspondence" authorityYear="2020" box="[873,1193,392,416]" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="australodakotensis" status="gen. et sp. nov.">
<emphasis id="B973EA903006FFD13E64FE1DEFC630F5" box="[873,1193,392,416]" italics="true" pageId="2" pageNumber="3">Lakotemys australodakotensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057EB3006FFD139A2FE1DEE2030CA" box="[1199,1359,392,415]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C31D65093006FFD13E3EFE55EF973347" pageId="2" pageNumber="3" type="type_taxon">
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<emphasis id="B973EA903006FFD13E3EFE55E8B4308D" bold="true" box="[819,987,448,472]" pageId="2" pageNumber="3">
<typeStatus id="54BC88203006FFD13E3EFE55E804308D" box="[819,875,448,472]" pageId="2" pageNumber="3">Type</typeStatus>
specimen
</emphasis>
</paragraph>
<paragraph id="8BB836823006FFD13E3EFE6EEF973347" blockId="2.[819,1272,507,530]" box="[819,1272,507,530]" pageId="2" pageNumber="3">
<materialsCitation id="3B6F3CDF3006FFD13E3EFE6EEF9B3347" box="[819,1268,507,530]" collectionCode="OMNH" pageId="2" pageNumber="3" specimenCode="OMNH 67133" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBA19EF93006FFD13E3EFE6EE8BF3347" box="[819,976,507,530]" collectionCode="OMNH" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="2" pageNumber="3" type="Museum">OMNH 67133</specimenCode>
, a partial shell (
<figureCitation id="133C2A073006FFD13974FE6EEFAB3347" box="[1145,1220,507,530]" captionStart="Figure 2" captionStartId="3.[124,191,1660,1682]" captionTargetBox="[157,1434,295,1628]" captionTargetId="figure-18@3.[150,1435,293,1628]" captionTargetPageId="3" captionText="Figure 2. OMNH 67133, a partial shell, holotype, Lakotemys australodakotensis gen. et. sp. nov., BerriasianBarremian Lakota Formation, Fall River County, South Dakota, USA. Photographs and illustrations in (a) dorsal and (b) ventral view. Dashed lines connote sutures observed in 3-D-rendered CT scans (see Fig. 4). Abbreviations are as follows: Ab: abdominal scute; An: anal scute; co: costal; Fe: femoral scute; Hu: humeral scute; hyo: hyoplastron; hyp: hypoplastron; Ma: marginal scutes; mes: mesoplastron; ne: neural; Pe: pectoral scutes; per: peripheral; Pl: pleural scutes; sp: suprapygal; Ve: vertebral scute; xi: xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.10965641" httpUri="https://zenodo.org/record/10965641/files/figure.png" pageId="2" pageNumber="3">Figs. 2</figureCitation>
,
<figureCitation id="133C2A073006FFD139C2FE6EEF843347" box="[1231,1259,507,530]" captionStart="Figure 4" captionStartId="5.[124,191,1771,1793]" captionTargetBox="[151,1435,213,1740]" captionTargetId="figure-18@5.[150,1435,212,1740]" captionTargetPageId="5" captionText="Figure 4. The 3-D-rendered CT images of shells, Lakotemys australodakotensis gen. et. sp. nov., BerriasianBarremian Lakota Formation, Fall River County, South Dakota, USA.Dorsal and ventral views of (a) OMNH 67133, the holotype, and (b) OMNH 63615.Abbreviations are as follows: co: costal; ent: entoplastron; hyo: hyoplastron; hyp: hypoplastron; mes: mesoplastron; ne: neural; per: peripheral; sp: suprapygal; xi: xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.10965645" httpUri="https://zenodo.org/record/10965645/files/figure.png" pageId="2" pageNumber="3">4a</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D65093006FFD13E3EFDA1EEE833B3" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BB836823006FFD13E3EFDA1EF393319" blockId="2.[819,1110,564,588]" box="[819,1110,564,588]" pageId="2" pageNumber="3">
<emphasis id="B973EA903006FFD13E3EFDA1EF393319" bold="true" box="[819,1110,564,588]" pageId="2" pageNumber="3">
<typeStatus id="54BC88203006FFD13E3EFDA1E8043319" box="[819,875,564,588]" pageId="2" pageNumber="3">Type</typeStatus>
locality and stratum
</emphasis>
</paragraph>
<paragraph id="8BB836823006FFD13E3EFDE5EEE833B3" blockId="2.[819,1462,624,742]" pageId="2" pageNumber="3">
<collectionCode id="ED16AE473006FFD13E3EFDE5E8EB33D2" box="[819,900,624,647]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="2" pageNumber="3" type="Museum">OMNH</collectionCode>
locality
<materialsCitation id="3B6F3CDF3006FFD13EE6FDE5EEEC33B3" collectionCode="OMNH" county="Fall River County" location="Unit" municipality="Dick Canyon" pageId="2" pageNumber="3" specimenCode="V1332" specimenCount="1" stateProvince="South Dakota" typeStatus="holotype">
<date id="FFB910423006FFD13EE6FDE5EF5933D2" box="[1003,1078,624,647]" pageId="2" pageNumber="3" value="1332-05">
<specimenCode id="DBA19EF93006FFD13EE6FDE5EF5933D2" box="[1003,1078,624,647]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="3" type="Museum">V1332</specimenCode>
</date>
,
<collectingMunicipality id="6BDCACF83006FFD1394BFDE5EFB533D2" box="[1094,1242,624,647]" pageId="2" pageNumber="3">Dick Canyon</collectingMunicipality>
,
<collectingCounty id="62D94E0E3006FFD139E4FDE5EEDD33D2" box="[1257,1458,624,647]" pageId="2" pageNumber="3">Fall River County</collectingCounty>
,
<collectingRegion id="49C3F8603006FFD13E3EFD05E8A833F2" box="[819,967,656,679]" country="United States of America" name="South Dakota" pageId="2" pageNumber="3">South Dakota</collectingRegion>
(more detailed locality information is on file at
<collectionCode id="ED16AE473006FFD13E5EFD3AE8C63393" box="[851,937,687,710]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="2" pageNumber="3" type="Museum">OMNH</collectionCode>
);
<location id="8ED860593006FFD13ECFFD3AE89E3393" LSID="urn:lsid:plazi:treatment:03AE87943006FFD53E64FE1DEA2D342C:8ED860593006FFD13ECFFD3AE89E3393" box="[962,1009,687,710]" county="Fall River County" municipality="Dick Canyon" name="Unit" pageId="2" pageNumber="3" stateProvince="South Dakota">Unit</location>
2,
<location id="8ED860593006FFD13912FD3AEFB53393" LSID="urn:lsid:plazi:treatment:03AE87943006FFD53E64FE1DEA2D342C:8ED860593006FFD13912FD3AEFB53393" box="[1055,1242,687,710]" county="Fall River County" municipality="Dick Canyon" name="Chilson Member" pageId="2" pageNumber="3" stateProvince="South Dakota">Chilson Member</location>
,
<taxonomicName id="4C074D013006FFD139E6FD3AE8F033B3" authority="Formation, Berriasian" authorityName="Formation, Berriasian" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Lakota Formation, Berriasian</taxonomicName>
<location id="8ED860593006FFD13EA0FD5AEF4233B3" LSID="urn:lsid:plazi:treatment:03AE87943006FFD53E64FE1DEA2D342C:8ED860593006FFD13EA0FD5AEF4233B3" box="[941,1069,719,742]" county="Fall River County" municipality="Dick Canyon" name="Valanginian" pageId="2" pageNumber="3" stateProvince="South Dakota">Valanginian</location>
(see Geological settings above)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D65093006FFD13E3EFC9DE88A350F" pageId="2" pageNumber="3" type="description">
<paragraph id="8BB836823006FFD13E3EFC9DEF7F3275" blockId="2.[819,1040,776,800]" box="[819,1040,776,800]" pageId="2" pageNumber="3">
<emphasis id="B973EA903006FFD13E3EFC9DEF7F3275" bold="true" box="[819,1040,776,800]" pageId="2" pageNumber="3">Nomenclatural acts</emphasis>
</paragraph>
<paragraph id="8BB836823006FFD13E3EFCD1E88A350F" blockId="2.[819,1462,836,1114]" pageId="2" pageNumber="3">
This publication and its nomenclatural acts were registered at ZooBank on
<date id="FFB910423006FFD13985FCF1EE07322E" box="[1160,1384,868,891]" pageId="2" pageNumber="3" value="2020-02-12">12 February 2020</date>
, prior to publication. The
<collectionCode id="ED16AE473006FFD1394AFC11EFED32CE" box="[1095,1154,900,923]" pageId="2" pageNumber="3">LSID</collectionCode>
of the publication is
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urn:lsid:zoobank.org:pub:
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, that of the new genus
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urn:lsid:zoobank.org:act:
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, and that of the new species
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urn:lsid:zoobank.org:act:
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.
</paragraph>
</subSubSection>
<subSubSection id="C31D65093006FFD13E3EFBE9EFA437F8" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BB836823006FFD13E3EFBE9E8CF35C1" blockId="2.[819,928,1148,1172]" box="[819,928,1148,1172]" pageId="2" pageNumber="3">
<emphasis id="B973EA903006FFD13E3EFBE9E8CF35C1" bold="true" box="[819,928,1148,1172]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BB836823006FFD13E3EFB22EFA437F8" blockId="2.[819,1462,1207,1709]" pageId="2" pageNumber="3">
<taxonomicName id="4C074D013006FFD13E3EFB22EF19359A" authority="Joyce &amp; Rollot &amp; Cifelli Correspondence, 2020" authorityName="Joyce &amp; Rollot &amp; Cifelli Correspondence" authorityYear="2020" box="[819,1142,1207,1231]" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="australodakotensis" status="gen. et sp. nov.">
<emphasis id="B973EA903006FFD13E3EFB22EF19359A" box="[819,1142,1207,1231]" italics="true" pageId="2" pageNumber="3">Lakotemys australodakotensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057EB3006FFD13972FB22EE48359B" box="[1151,1319,1207,1230]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
can be diagnosed as a representative of Paracryptodira by the presence of a finely sculptured skull and shell and the likely location of the foramen posterius canalis carotici interni midline along the contact of the basisphenoid and pterygoid and as a representative of
<taxonomicName id="4C074D013006FFD13EB9FAC2EF78343B" authorityName="Cope" authorityYear="1873" box="[948,1047,1367,1390]" family="Baenidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="family">Baenidae</taxonomicName>
by the absence of epiplastral processes and the development of well-developed axillary and inguinal buttresses. Among named Early Cretaceous baenids,
<taxonomicName id="4C074D013006FFD13868FA03EF5D349B" authorityName="Joyce &amp; Rollot &amp; Cifelli Correspondence" authorityYear="2020" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="australodakotensis">
<emphasis id="B973EA903006FFD13868FA03EF5D349B" italics="true" pageId="2" pageNumber="3">Lakotemys australodakotensis</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C074D013006FFD1387FFA23EF7A34BB" baseAuthorityName="Gilmore" baseAuthorityYear="1919" family="Baenidae" genus="Protobaena" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="wyomingensis">
<emphasis id="B973EA903006FFD1387FFA23EF7A34BB" italics="true" pageId="2" pageNumber="3">Protobaena wyomingensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C074D013006FFD1395CFA43EE7634BB" authorityName="Gaffney" authorityYear="1972" box="[1105,1305,1494,1518]" family="Baenidae" genus="Trinitichelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="hiatti">
<emphasis id="B973EA903006FFD1395CFA43EE7634BB" box="[1105,1305,1494,1518]" italics="true" pageId="2" pageNumber="3">Trinitichelys hiatti</emphasis>
</taxonomicName>
by having an irregularly shaped vertebral
<collectionCode id="ED16AE473006FFD13967FA63EF123758" box="[1130,1149,1526,1549]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="3" type="Museum">V</collectionCode>
that does not lap onto neural VIII and that forms two anterolateral processes that partially hinder vertebral IV from contacting pleural IV and can be differentiated from
<taxonomicName id="4C074D013006FFD13912F9C0EE6E3738" box="[1055,1281,1621,1645]" family="Pleurosternidae" genus="Arundelemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="dardeni">
<emphasis id="B973EA903006FFD13912F9C0EE6E3738" box="[1055,1281,1621,1645]" italics="true" pageId="2" pageNumber="3">Arundelemys dardeni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C074D013006FFD1383CF9C0E80437D8" authorityName="Gaffney" authorityYear="1972" family="Baenidae" genus="Trinitichelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="hiatti">
<emphasis id="B973EA903006FFD1383CF9C0E80437D8" italics="true" pageId="2" pageNumber="3">Trinitichelys hiatti</emphasis>
</taxonomicName>
by having a broader skull and triturating surfaces (see Discussion for additional differences).
</paragraph>
</subSubSection>
<subSubSection id="C31D65093006FFD53E3EF95AE99E35D0" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BB836823006FFD13E3EF95AE8C137B2" blockId="2.[819,942,1743,1767]" box="[819,942,1743,1767]" pageId="2" pageNumber="3">
<emphasis id="B973EA903006FFD13E3EF95AE8C137B2" bold="true" box="[819,942,1743,1767]" pageId="2" pageNumber="3">Etymology</emphasis>
</paragraph>
<paragraph id="8BB836823006FFD53E3EF89FE99E35D0" blockId="2.[819,1462,1802,1921]" lastBlockId="6.[124,767,1038,1158]" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
The compound word
<taxonomicName id="4C074D013006FFD13911F89FEFE23677" authorityName="Joyce, Rollot &amp; Cifelli" authorityYear="2020" box="[1052,1165,1802,1826]" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
<emphasis id="B973EA903006FFD13911F89FEFE23677" box="[1052,1165,1802,1826]" italics="true" pageId="2" pageNumber="3">Lakotemys</emphasis>
</taxonomicName>
is derived from the combination of the word
<taxonomicName id="4C074D013006FFD13903F8BFEF373617" box="[1038,1112,1834,1858]" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B973EA903006FFD13903F8BFEF373617" box="[1038,1112,1834,1858]" italics="true" pageId="2" pageNumber="3">Lakota</emphasis>
</taxonomicName>
, in reference to the indigenous
<taxonomicName id="4C074D013006FFD13E3EF8DFE8123634" box="[819,893,1866,1889]" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Lakota</taxonomicName>
people of the greater Black Hills region, and the fossil-bearing
<taxonomicName id="4C074D013006FFD13EDCF8FFEFF936D4" authority="Formation" authorityName="Formation" box="[977,1174,1898,1921]" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Lakota Formation</taxonomicName>
, and
<emphasis id="B973EA903006FFD139DBF8FCEE6436D4" box="[1238,1291,1897,1921]" italics="true" pageId="2" pageNumber="3">emys</emphasis>
, ancient Greek for turtles. The species epithet
<taxonomicName id="4C074D013002FFD53CC4FB9BE9FE3573" authorityName="Joyce &amp; Rollot &amp; Cifelli Correspondence" authorityYear="2020" box="[457,657,1038,1062]" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="australodakotensis">
<emphasis id="B973EA903002FFD53CC4FB9BE9FE3573" box="[457,657,1038,1062]" italics="true" pageId="6" pageNumber="7">australodakotensis</emphasis>
</taxonomicName>
is formed in allusion to the state of
<collectingRegion id="49C3F8603002FFD53C88FBBAE97A3513" box="[389,533,1071,1094]" country="United States of America" name="South Dakota" pageId="6" pageNumber="7">South Dakota</collectingRegion>
through the combination of the word
<emphasis id="B973EA903002FFD53C3DFBDBEA1D3533" box="[304,370,1102,1126]" italics="true" pageId="6" pageNumber="7">auster</emphasis>
with the suffix -
<emphasis id="B973EA903002FFD53F29FBDBE9253533" box="[548,586,1102,1126]" italics="true" pageId="6" pageNumber="7">alis</emphasis>
, Latin for south, and
<emphasis id="B973EA903002FFD53DA4FBFBEB9835D3" box="[169,247,1134,1158]" italics="true" pageId="6" pageNumber="7">Dakota</emphasis>
. The suffix -
<emphasis id="B973EA903002FFD53C8DFBFBEADA35D3" box="[384,437,1134,1158]" italics="true" pageId="6" pageNumber="7">ensis</emphasis>
is used to indicate a location.
</paragraph>
</subSubSection>
<caption id="DF78660A3007FFD03D71F9E9E8FC3652" ID-DOI="http://doi.org/10.5281/zenodo.10965641" ID-Zenodo-Dep="10965641" httpUri="https://zenodo.org/record/10965641/files/figure.png" pageId="3" pageNumber="4" startId="3.[124,191,1660,1682]" targetBox="[157,1434,295,1628]" targetPageId="3" targetType="figure">
<paragraph id="8BB836823007FFD03D71F9E9E8FC3652" blockId="3.[124,1463,1660,1799]" pageId="3" pageNumber="4">
<emphasis id="B973EA903007FFD03D71F9E9EBB737C7" bold="true" box="[124,216,1660,1682]" pageId="3" pageNumber="4">Figure 2.</emphasis>
OMNH 67133, a partial shell, holotype,
<taxonomicName id="4C074D013007FFD03F6FF9E8E8EC37C7" authorityName="Joyce &amp; Rollot &amp; Cifelli Correspondence" authorityYear="2020" box="[610,899,1661,1682]" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="australodakotensis">
<emphasis id="B973EA903007FFD03F6FF9E8E8EC37C7" box="[610,899,1661,1682]" italics="true" pageId="3" pageNumber="4">Lakotemys australodakotensis</emphasis>
</taxonomicName>
gen. et. sp. nov., BerriasianBarremian
<taxonomicName id="4C074D013007FFD03809F9E8EA4037FA" authority="Formation, Fall River County" authorityName="Formation, Fall River County" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Lakota Formation, Fall River County</taxonomicName>
, South Dakota, USA. Photographs and illustrations in
<emphasis id="B973EA903007FFD03E5EF90CE80037FA" bold="true" box="[851,879,1689,1711]" pageId="3" pageNumber="4">(a)</emphasis>
dorsal and
<emphasis id="B973EA903007FFD03EE5F90CEF6A37FA" bold="true" box="[1000,1029,1689,1711]" pageId="3" pageNumber="4">(b)</emphasis>
ventral view. Dashed lines connote sutures observed in 3-D-rendered CT scans (see Fig. 4). Abbreviations are as follows: Ab: abdominal scute; An: anal scute; co: costal; Fe: femoral scute; Hu: humeral scute; hyo: hyoplastron; hyp: hypoplastron; Ma: marginal scutes; mes: mesoplastron; ne: neural; Pe: pectoral scutes; per: peripheral; Pl: pleural scutes; sp: suprapygal; Ve: vertebral scute; xi: xiphiplastron.
</paragraph>
</caption>
<caption id="DF78660A3000FFD73D71F925EE6F366E" ID-DOI="http://doi.org/10.5281/zenodo.10965643" ID-Zenodo-Dep="10965643" httpUri="https://zenodo.org/record/10965643/files/figure.png" pageId="4" pageNumber="5" startId="4.[124,191,1712,1734]" targetBox="[150,1433,242,1681]" targetPageId="4" targetType="figure">
<paragraph id="8BB836823000FFD73D71F925EE6F366E" blockId="4.[124,1463,1712,1851]" pageId="4" pageNumber="5">
<emphasis id="B973EA903000FFD73D71F925EBB83793" bold="true" box="[124,215,1712,1734]" pageId="4" pageNumber="5">Figure 3.</emphasis>
OMNH 63615, a partial shell,
<taxonomicName id="4C074D013000FFD73F0DF925E84F3790" authorityName="Joyce &amp; Rollot &amp; Cifelli Correspondence" authorityYear="2020" box="[512,800,1712,1733]" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="australodakotensis">
<emphasis id="B973EA903000FFD73F0DF925E84F3790" box="[512,800,1712,1733]" italics="true" pageId="4" pageNumber="5">Lakotemys australodakotensis</emphasis>
</taxonomicName>
gen. et. sp. nov., BerriasianBarremian
<taxonomicName id="4C074D013000FFD739ADF924EBAC37B6" authority="Formation, Fall River County" authorityName="Formation, Fall River County" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Lakota Formation, Fall River County</taxonomicName>
, South Dakota, USA. Photographs and illustrations in
<emphasis id="B973EA903000FFD73FECF958E99337B6" bold="true" box="[737,764,1741,1763]" pageId="4" pageNumber="5">(a)</emphasis>
dorsal and
<emphasis id="B973EA903000FFD73E7EF958E8FF37B6" bold="true" box="[883,912,1741,1763]" pageId="4" pageNumber="5">(b)</emphasis>
ventral view. Dashed lines connote sutures observed in 3-D-rendered CT scans (see Fig. 4). Abbreviations are as follows: Ab: abdominal scute; co: costal; ent: entoplastron; epi: epiplastron; Ex: extragular scute; Fe: femoral scute; Gu: gular scute; Hu: humeral scute; hyo: hyoplastron; hyp: hypoplastron; IM: inframarginal scute; Ma: marginal scutes; mes: mesoplastron; ne: neural; nu: nuchal scute; Pe: pectoral scutes; per: peripheral; Ve: vertebral scute.
</paragraph>
</caption>
<caption id="DF78660A3001FFD63D71F97EEA4E360C" ID-DOI="http://doi.org/10.5281/zenodo.10965645" ID-Zenodo-Dep="10965645" httpUri="https://zenodo.org/record/10965645/files/figure.png" pageId="5" pageNumber="6" startId="5.[124,191,1771,1793]" targetBox="[151,1435,213,1740]" targetPageId="5" targetType="figure">
<paragraph id="8BB836823001FFD63D71F97EEA4E360C" blockId="5.[124,1462,1771,1881]" pageId="5" pageNumber="6">
<emphasis id="B973EA903001FFD63D71F97EEBB73654" bold="true" box="[124,216,1771,1793]" pageId="5" pageNumber="6">Figure 4.</emphasis>
The 3-D-rendered CT images of shells,
<taxonomicName id="4C074D013001FFD63F52F979E8EF3654" authorityName="Joyce &amp; Rollot &amp; Cifelli Correspondence" authorityYear="2020" box="[607,896,1772,1793]" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="australodakotensis">
<emphasis id="B973EA903001FFD63F52F979E8EF3654" box="[607,896,1772,1793]" italics="true" pageId="5" pageNumber="6">Lakotemys australodakotensis</emphasis>
</taxonomicName>
gen. et. sp. nov., BerriasianBarremian
<taxonomicName id="4C074D013001FFD63809F979EA4A364B" authority="Formation, Fall River County" authorityName="Formation, Fall River County" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Lakota Formation, Fall River County</taxonomicName>
, South Dakota, USA. Dorsal and ventral views of
<emphasis id="B973EA903001FFD63FF6F89DE878364B" bold="true" box="[763,791,1800,1822]" pageId="5" pageNumber="6">(a)</emphasis>
OMNH 67133, the holotype, and
<emphasis id="B973EA903001FFD6395AF89DEF1B364B" bold="true" box="[1111,1140,1800,1822]" pageId="5" pageNumber="6">(b)</emphasis>
OMNH 63615. Abbreviations are as follows: co: costal; ent: entoplastron; hyo: hyoplastron; hyp: hypoplastron; mes: mesoplastron; ne: neural; per: peripheral; sp: suprapygal; xi: xiphiplastron.
</paragraph>
</caption>
<caption id="DF78660A3002FFD53D71FCEEE820329E" ID-DOI="http://doi.org/10.5281/zenodo.10965647" ID-Zenodo-Dep="10965647" httpUri="https://zenodo.org/record/10965647/files/figure.png" pageId="6" pageNumber="7" startId="6.[124,191,891,913]" targetBox="[151,1435,177,859]" targetPageId="6" targetType="figure">
<paragraph id="8BB836823002FFD53D71FCEEE820329E" blockId="6.[124,1462,891,971]" pageId="6" pageNumber="7">
<emphasis id="B973EA903002FFD53D71FCEEEBB832C4" bold="true" box="[124,215,891,913]" pageId="6" pageNumber="7">Figure 5.</emphasis>
The 3-D-rendered CT images of a skull,
<taxonomicName id="4C074D013002FFD53F6EFCEEE8EB32C5" authorityName="Joyce &amp; Rollot &amp; Cifelli Correspondence" authorityYear="2020" box="[611,900,891,912]" family="Baenidae" genus="Lakotemys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="australodakotensis">
<emphasis id="B973EA903002FFD53F6EFCEEE8EB32C5" box="[611,900,891,912]" italics="true" pageId="6" pageNumber="7">Lakotemys australodakotensis</emphasis>
</taxonomicName>
gen. et. sp. nov., BerriasianBarremian
<taxonomicName id="4C074D013002FFD53809FCE9EA4832FB" authority="Formation, Fall River County" authorityName="Formation, Fall River County" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Lakota Formation, Fall River County</taxonomicName>
, South Dakota, USA. OMNH 66106 in
<emphasis id="B973EA903002FFD53FAEFC0DE9D032FB" bold="true" box="[675,703,920,942]" pageId="6" pageNumber="7">(a)</emphasis>
dorsal,
<emphasis id="B973EA903002FFD53E06FC0DE84732FB" bold="true" box="[779,808,920,942]" pageId="6" pageNumber="7">(b)</emphasis>
ventral,
<emphasis id="B973EA903002FFD53E71FC0DE8F932FB" bold="true" box="[892,918,920,942]" pageId="6" pageNumber="7">(c)</emphasis>
anterior,
<emphasis id="B973EA903002FFD53EFCFC0DEF6132FB" bold="true" box="[1009,1038,920,942]" pageId="6" pageNumber="7">(d)</emphasis>
posterior,
<emphasis id="B973EA903002FFD53979FC0DEFE132FB" bold="true" box="[1140,1166,920,942]" pageId="6" pageNumber="7">(e)</emphasis>
right lateral, and
<emphasis id="B973EA903002FFD5383AFC0DEE2032FB" bold="true" box="[1335,1359,920,942]" pageId="6" pageNumber="7">(f)</emphasis>
left lateral views. Abbreviations are as follows: fr: frontal; pa: parietal; po: postorbital.
</paragraph>
</caption>
<subSubSection id="C31D65093002FFD53D71FB32EA2D342C" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BB836823002FFD53D71FB32EA2735EA" blockId="6.[124,328,1191,1215]" box="[124,328,1191,1215]" pageId="6" pageNumber="7">
<emphasis id="B973EA903002FFD53D71FB32EA2735EA" bold="true" box="[124,328,1191,1215]" pageId="6" pageNumber="7">Referred material</emphasis>
</paragraph>
<paragraph id="8BB836823002FFD53D71FB77EA2D342C" blockId="6.[124,766,1250,1401]" pageId="6" pageNumber="7">
<materialsCitation id="3B6F3CDF3002FFD53D71FB77EA3A344C" collectionCode="OMNH" pageId="6" pageNumber="7" specimenCode="OMNH 66106" specimenCount="1">
<specimenCode id="DBA19EF93002FFD53D71FB77EA7435AC" box="[124,283,1250,1273]" collectionCode="OMNH" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="6" pageNumber="7" type="Museum">OMNH 66106</specimenCode>
, a poorly preserved skull collected from the
<typeStatus id="54BC88203002FFD53D71FA97EBC5344C" box="[124,170,1282,1305]" pageId="6" pageNumber="7">type</typeStatus>
locality (
<figureCitation id="133C2A073002FFD53C01FA97EA24344C" box="[268,331,1282,1305]" captionStart="Figure 3" captionStartId="4.[124,191,1712,1734]" captionTargetBox="[150,1433,242,1681]" captionTargetId="figure-18@4.[150,1435,242,1681]" captionTargetPageId="4" captionText="Figure 3. OMNH 63615, a partial shell, Lakotemys australodakotensis gen. et. sp. nov., BerriasianBarremian Lakota Formation, Fall River County, South Dakota, USA. Photographs and illustrations in (a) dorsal and (b) ventral view. Dashed lines connote sutures observed in 3-D-rendered CT scans (see Fig. 4). Abbreviations are as follows: Ab: abdominal scute; co: costal; ent: entoplastron; epi: epiplastron; Ex: extragular scute; Fe: femoral scute; Gu: gular scute; Hu: humeral scute; hyo: hyoplastron; hyp: hypoplastron; IM: inframarginal scute; Ma: marginal scutes; mes: mesoplastron; ne: neural; nu: nuchal scute; Pe: pectoral scutes; per: peripheral; Ve: vertebral scute." figureDoi="http://doi.org/10.5281/zenodo.10965643" httpUri="https://zenodo.org/record/10965643/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
)
</materialsCitation>
;
<materialsCitation id="3B6F3CDF3002FFD53C52FA97EA78346C" collectionCode="OMNH" pageId="6" pageNumber="7" specimenCode="OMNH 63615" specimenCount="1">
<specimenCode id="DBA19EF93002FFD53C52FA97EA94344C" box="[351,507,1282,1305]" collectionCode="OMNH" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="6" pageNumber="7" type="Museum">OMNH 63615</specimenCode>
, a poorly preserved shell collected from
</materialsCitation>
<collectionCode id="ED16AE473002FFD53C13FAB7EA00346C" box="[286,367,1314,1337]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="6" pageNumber="7" type="Museum">OMNH</collectionCode>
site
<materialsCitation id="3B6F3CDF3002FFD53CAEFAB7EA51342C" collectionCode="V" pageId="6" pageNumber="7" specimenCode="V1382" specimenCount="1">
<date id="FFB910423002FFD53CAEFAB7EA84346C" box="[419,491,1314,1337]" pageId="6" pageNumber="7" value="1382-05">
<specimenCode id="DBA19EF93002FFD53CAEFAB7EA84346C" box="[419,491,1314,1337]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="7" type="Museum">V1382</specimenCode>
</date>
(
<figureCitation id="133C2A073002FFD53CF7FAB7E952346C" box="[506,573,1314,1337]" captionStart="Figure 4" captionStartId="5.[124,191,1771,1793]" captionTargetBox="[151,1435,213,1740]" captionTargetId="figure-18@5.[150,1435,212,1740]" captionTargetPageId="5" captionText="Figure 4. The 3-D-rendered CT images of shells, Lakotemys australodakotensis gen. et. sp. nov., BerriasianBarremian Lakota Formation, Fall River County, South Dakota, USA.Dorsal and ventral views of (a) OMNH 67133, the holotype, and (b) OMNH 63615.Abbreviations are as follows: co: costal; ent: entoplastron; hyo: hyoplastron; hyp: hypoplastron; mes: mesoplastron; ne: neural; per: peripheral; sp: suprapygal; xi: xiphiplastron." figureDoi="http://doi.org/10.5281/zenodo.10965645" httpUri="https://zenodo.org/record/10965645/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
). All referred material originates from Unit 2 of the Chilson Member of the
<taxonomicName id="4C074D013002FFD53D71FAF7EA2D342C" authority="Formation." authorityName="Formation." box="[124,322,1378,1401]" class="Malacostraca" family="Tryphosidae" genus="Lakota" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Lakota Formation.</taxonomicName>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>