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<mods:title id="9A4CB9207B6B2868AEF28DF5472412E9">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<paragraph id="422236DE627B9D08EECCFDD6FD98E860" blockId="26.[151,577,583,642]" box="[151,577,583,609]" pageId="26" pageNumber="27">
<heading id="196A81B2627B9D08EECCFDD6FD98E860" bold="true" box="[151,577,583,609]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="70E9EACC627B9D08EECCFDD6FD98E860" bold="true" box="[151,577,583,609]" pageId="26" pageNumber="27">
10.
<taxonomicName id="859D4D5D627B9D08EE98FDD6FD98E860" ID-CoL="8CJ3J" authority="Matile" authorityName="Matile, 1980:" authorityYear="1229" box="[195,577,583,609]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="crosskeyi">
<emphasis id="70E9EACC627B9D08EE98FDD6FE31E860" bold="true" box="[195,488,583,609]" italics="true" pageId="26" pageNumber="27">Leptomorphus crosskeyi</emphasis>
Matile
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0A876555627B9D08EECCFDFBFDC2E883" box="[151,539,618,642]" pageId="26" pageNumber="27" type="description">
<paragraph id="422236DE627B9D08EECCFDFBFDC2E883" blockId="26.[151,577,583,642]" box="[151,539,618,642]" pageId="26" pageNumber="27">
(
<figureCitation id="DAA62A5B627B9D08EEC5FDFBFEC1E883" box="[158,280,618,642]" captionStart="FIGURES 1015" captionStartId="85.[151,263,1903,1926]" captionTargetBox="[175,1416,182,1881]" captionTargetId="figure-54@85.[173,1417,181,1882]" captionTargetPageId="85" captionText="FIGURES 1015. Leptomorphus habitus. 10. L. crassipilus, holotype male. 11. L. crosskeyi, female. 12. L. eberhardi, holotype male. 13. L. eberhardi, paratype female. 14. L. fasciculatus, holotype male. 15. L. femoratus, holotype male." pageId="26" pageNumber="27">Figures 11</figureCitation>
,
<figureCitation id="DAA62A5B627B9D08EF7FFDFBFE9BE883" box="[292,322,618,642]" captionStart="FIGURES 4255" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 4255. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="26" pageNumber="27">51</figureCitation>
,
<figureCitation id="DAA62A5B627B9D08EF15FDFBFEB5E883" box="[334,364,618,642]" captionStart="FIGURES 99102" captionStartId="99.[151,263,1833,1856]" captionTargetBox="[188,1399,182,1758]" captionTargetId="figure-95@99.[180,1405,176,1767]" captionTargetPageId="99" captionText="FIGURES 99102. Leptomorphus male genitalia, ventral. 99. L. crosskeyi. 100. L. eberhardi. 101. L. fasciculatus. 102. L. femoratus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="26" pageNumber="27">99</figureCitation>
,
<figureCitation id="DAA62A5B627B9D08EF23FDFBFE7DE883" box="[376,420,618,642]" captionStart="FIGURE 134136" captionStartId="107.[151,249,1814,1837]" captionTargetBox="[151,1436,193,1793]" captionTargetId="figure-66@107.[151,1436,193,1793]" captionTargetPageId="107" captionText="FIGURE 134136. Distribution of Afrotropical Leptomorphus. 134. L. grjebinei (red circle), L. mandelai (blue square), L. stigmatus (black triangle). 135. L. aliciae (purple star), L. carnevalei (black square), L. couturieri (black circle), L. gracilis (red triangle). 136. L. crosskeyi (red triangle), L. medleri (blue star), L. obscurus (black circle)." pageId="26" pageNumber="27">136</figureCitation>
,
<figureCitation id="DAA62A5B627B9D08EFEBFDFBFE05E883" box="[432,476,618,642]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values&gt;50% are shown in bold below the branches." pageId="26" pageNumber="27">150</figureCitation>
,
<figureCitation id="DAA62A5B627B9D08EFB3FDFBFDCAE883" box="[488,531,618,642]" captionStart="FIGURE 153" captionStartId="115.[151,250,801,824]" captionTargetBox="[155,1364,186,781]" captionTargetId="figure-100@115.[151,1436,161,836]" captionTargetPageId="115" captionText="FIGURE 153. One of three equally parsimonious trees showing the relationships between species in the L. grjebinei species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="26" pageNumber="27">153</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0A876555627B9D08EECCFD23FCE9E94E" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="422236DE627B9D08EECCFD23FBE4E8C9" blockId="26.[151,1085,690,743]" box="[151,1085,690,712]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="628D11F0627B9D08EECCFD23FBE4E8C9" box="[151,1085,690,712]" pageId="26" pageNumber="27">
<treatmentCitation id="C33C10CF627B9D08EECCFD23FD2CE8C9" author="Matile, L." box="[151,757,690,712]" page="147" pageId="26" pageNumber="27" year="1977">
<taxonomicName id="859D4D5D627B9D08EECCFD23FD2CE8C9" ID-CoL="896PR" authority="Matile, 1977: 147" authorityName="Matile" authorityPageNumber="147" authorityYear="1977" box="[151,757,690,712]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="africanus" subGenus="Gymnoscutum">
<emphasis id="70E9EACC627B9D08EECCFD23FDEEE8C9" box="[151,567,690,712]" italics="true" pageId="26" pageNumber="27">Leptomorphus (Gymnoscutum) africanus</emphasis>
<bibRefCitation id="260C4B2F627B9D08EC66FD23FD2CE8C9" author="Matile, L." box="[573,757,690,712]" pageId="26" pageNumber="27" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile, 1977: 147</bibRefCitation>
</taxonomicName>
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. Preoccupied by Meunier, 1907.
</treatmentCitationGroup>
</paragraph>
<paragraph id="422236DE627B9D08EECCFD40FC3FE8E6" blockId="26.[151,1085,690,743]" box="[151,998,721,743]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="628D11F0627B9D08EECCFD40FC3FE8E6" box="[151,998,721,743]" pageId="26" pageNumber="27">
<treatmentCitation id="C33C10CF627B9D08EECCFD40FD8CE8E6" author="Matile, L." box="[151,597,721,743]" page="1229" pageId="26" pageNumber="27" year="1980">
<taxonomicName id="859D4D5D627B9D08EECCFD40FD8CE8E6" ID-CoL="8CJ3J" authority="Matile, 1980: 1229" authorityName="Matile, 1980:" authorityPageNumber="1229" authorityYear="1229" box="[151,597,721,743]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="crosskeyi">
<emphasis id="70E9EACC627B9D08EECCFD40FE55E8E6" box="[151,396,721,743]" italics="true" pageId="26" pageNumber="27">Leptomorphus crosskeyi</emphasis>
<bibRefCitation id="260C4B2F627B9D08EFC8FD40FD8CE8E6" author="Matile, L." box="[403,597,721,743]" pageId="26" pageNumber="27" pagination="216 - 230" refId="ref94631" refString="Matile, L. (1980) 15. Family Mycetophilidae. In: Crosskey, R. W. (ed.). Catalogue of the Diptera of the Afrotropical region. British Museum of Natural History, London, pp. 216 - 230." type="book chapter" year="1980">Matile, 1980: 1229</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(new replacement name for
<taxonomicName id="859D4D5D627B9D08ED2DFD40FC0FE8E6" ID-CoL="896PR" authorityName="Matile" authorityYear="1977" box="[886,982,721,743]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="70E9EACC627B9D08ED2DFD40FC0FE8E6" box="[886,982,721,743]" italics="true" pageId="26" pageNumber="27">africanus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="422236DE627B9D08EECCFC82FCE9E94E" blockId="26.[151,1437,787,2036]" pageId="26" pageNumber="27">
<emphasis id="70E9EACC627B9D08EECCFC82FEC0E92D" bold="true" box="[151,281,787,812]" pageId="26" pageNumber="27">References</emphasis>
:
<bibRefCitation id="260C4B2F627B9D08EF72FC85FDC1E92D" author="Crosskey, R. W." box="[297,536,788,812]" pageId="26" pageNumber="27" refId="ref92665" refString="Crosskey, R. W. (ed.) (1980) Catalogue of the Diptera of the Afrotropical Region. British Museum of Natural History, London, 1437 pp." type="book" year="1980">Crosskey 1980: 1221</bibRefCitation>
(catalogue appendix as
<taxonomicName id="859D4D5D627B9D08ED70FC84FC6AE92D" ID-CoL="896PR" authorityName="Matile" authorityYear="1977" box="[811,947,788,812]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="70E9EACC627B9D08ED70FC84FC6AE92D" box="[811,947,788,812]" italics="true" pageId="26" pageNumber="27">L. africanus</emphasis>
</taxonomicName>
);
<bibRefCitation id="260C4B2F627B9D08ED97FC85FB49E92D" author="Matile, L." box="[972,1168,788,812]" pageId="26" pageNumber="27" pagination="143 - 150" refId="ref94722" refString="Matile, L. (1997) Nouvelles donnees sur les Leptomorphus Afrotropicaux (Diptera, Mycetophilidae). Revue Francaise d'Entomologie (Nouvelle Serie) 1996, 18 (4), 143 - 150." type="journal article" year="1997">Matile 1997: 143</bibRefCitation>
, 147150 (figures, new records, morphological variation, key,
<emphasis id="70E9EACC627B9D08EC13FCA9FD56E951" box="[584,655,824,848]" italics="true" pageId="26" pageNumber="27">lapsus</emphasis>
as
<emphasis id="70E9EACC627B9D08ECEFFCA8FCF8E951" box="[692,801,825,848]" italics="true" pageId="26" pageNumber="27">nom. nov.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="0A876555627B9D08EE9CFCCAFB4EEE6E" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="422236DE627B9D08EE9CFCCAFE26E9DE" blockId="26.[151,1437,787,2036]" pageId="26" pageNumber="27">
<emphasis id="70E9EACC627B9D08EE9CFCCAFEBDE975" bold="true" box="[199,356,859,884]" pageId="26" pageNumber="27">DIAGNOSIS</emphasis>
: The only extant species of
<taxonomicName id="859D4D5D627B9D08EC92FCCDFCB2E975" authorityName=": Edwards" authorityYear="1925" box="[713,875,860,884]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="70E9EACC627B9D08EC92FCCDFCB2E975" box="[713,875,860,884]" italics="true" pageId="26" pageNumber="27">Leptomorphus</emphasis>
</taxonomicName>
with the following combination of characters: laterotergite, anepisternum and anepimeron yellow; scutellum brown; male genitalia with posterolateral projection of tergite 9 (tergal evagination) bearing distinct thin point on medial corner, gonostylus with shortest lobe gradually tapering to thick point (
<figureCitation id="DAA62A5B627B9D08EFC5FC56FE2AE9DE" box="[414,499,967,991]" captionStart="FIGURES 99102" captionStartId="99.[151,263,1833,1856]" captionTargetBox="[188,1399,182,1758]" captionTargetId="figure-95@99.[180,1405,176,1767]" captionTargetPageId="99" captionText="FIGURES 99102. Leptomorphus male genitalia, ventral. 99. L. crosskeyi. 100. L. eberhardi. 101. L. fasciculatus. 102. L. femoratus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="26" pageNumber="27">Fig. 99</figureCitation>
).
</paragraph>
<paragraph id="422236DE627B9D08EE9CFC7DFB4EEE6E" blockId="26.[151,1437,787,2036]" pageId="26" pageNumber="27">
This species can be distinguished from most other Afrotropical species by the brown scutellum and lack of small setae covering the scutum (except along dorsocentral lines). It can be separated from
<taxonomicName id="859D4D5D627B9D08EACEFB80FAD1EE29" box="[1173,1288,1040,1064]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="70E9EACC627B9D08EACEFB80FAD1EE29" box="[1173,1288,1040,1064]" italics="true" pageId="26" pageNumber="27">L. gracilis</emphasis>
</taxonomicName>
by the larger width of the posteromedial concavity of tergite 9 and thin point on the medial corner of the tergal evagination and from
<taxonomicName id="859D4D5D627B9D08EE8FFBC8FE8EEE71" box="[212,343,1112,1136]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="70E9EACC627B9D08EE8FFBC8FE8EEE71" box="[212,343,1112,1136]" italics="true" pageId="26" pageNumber="27">L. obscurus</emphasis>
</taxonomicName>
by the almost square posterior margin of the tergal evaginations (
<figureCitation id="DAA62A5B627B9D08EA6CFBC6FB52EE6E" box="[1079,1163,1111,1135]" captionStart="FIGURES 99102" captionStartId="99.[151,263,1833,1856]" captionTargetBox="[188,1399,182,1758]" captionTargetId="figure-95@99.[180,1405,176,1767]" captionTargetPageId="99" captionText="FIGURES 99102. Leptomorphus male genitalia, ventral. 99. L. crosskeyi. 100. L. eberhardi. 101. L. fasciculatus. 102. L. femoratus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="26" pageNumber="27">Fig. 99</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0A876555627B9D09EE9CFBEAFE1BEE14" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="422236DE627B9D09EE9CFBEAFDEBE984" blockId="26.[151,1437,787,2036]" lastBlockId="27.[151,1436,151,1549]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="70E9EACC627B9D08EE9CFBEAFE52EE95" bold="true" box="[199,395,1147,1172]" pageId="26" pageNumber="27">DESCRIPTION</emphasis>
:
<emphasis id="70E9EACC627B9D08EFC6FBEDFE0EEE95" box="[413,471,1148,1172]" italics="true" pageId="26" pageNumber="27">Male</emphasis>
.
<emphasis id="70E9EACC627B9D08EFB2FBEDFDF0EE95" bold="true" box="[489,553,1148,1172]" pageId="26" pageNumber="27">Head</emphasis>
: yellow with some brown spots, circular in anterior view. Antenna brown; scape dark yellow, with brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae present; pedicel yellow, with 2 large bristles, several setae on apicodorsal margin, number of fine setae on apicoventral margin; flagellomere 1 with tapered base brown remainder brown; flagellomere 6 1.8X as long as broad. Clypeus yellow, dorsoventrally elongate oval; bristles on clypeus brown, 68 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 2X as long as face. Face yellow; shape a subequal triangle, with few bristles ventrolaterally. Frons ventral half brown rest yellow; with few bristles medioventrally, frontal furrow running full distance from dorsal margin towards ventral margin, frontal cleft just anterior of median ocellus. Palpus with segments 12 yellow, 35 light brown; segment 1 hidden behind eye, segments increasing in length, segment 5 2.5X length of segment 4 with central half thinner than base and apex, segment 3 with apicolateral patch of fine yellow setae encircled by strong dark setae. Labellum yellow. Eye with interommatidial setulae absent. Occiput yellow with some adventitious brown spotting anteriorly, with appressed, anteriorly directed setae. Ocelli in a straight line, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle dark brown.
<emphasis id="70E9EACC627B9D08EDF3F9E5FBD8EC8D" bold="true" box="[936,1025,1652,1676]" pageId="26" pageNumber="27">Thorax</emphasis>
: Length
<quantity id="85659B3B627B9D08EA3CF9E5FB12EC8A" box="[1127,1227,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="26" pageNumber="27" unit="mm" value="1.19">1.19 mm</quantity>
(
<quantity id="85659B3B627B9D08EA81F9E5FA5AEC8A" box="[1242,1411,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" metricValueMax="1.23" metricValueMin="1.15" pageId="26" pageNumber="27" unit="mm" value="1.19" valueMax="1.23" valueMin="1.15">1.151.23 mm</quantity>
, n = 4). Dark brown dorsally, yellow laterally. Scutum dark brown/black with blue-green specks, pair of yellow mediolateral and posterolateral spots; surface of scutum bare; acrostichal setae absent; dorsocentral setae present as fine setae for most of length; multiple rows of lateral setae present; patch of setae on scutum at wing base present. Scutellum brown; with 810 large bristles and few small bristles. Prescutum yellow. Mediotergite brown, lighter posteriorly with 68 bristles on posterolateral corners, anteromedial patch of small bristles. Laterotergite yellow; anterior margin of laterotergite not reaching katepisternum. Anepimeron yellow. Anepisternum yellow. Katepisternum yellow. Antepronotum and proepisternum yellow. Margin of anterior and posterior spiracles yellow with light brown and yellow trichia respectively. Metepisternum yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter with basal 1/3 of stem ivory, apical portion and knob dark brown.
<emphasis id="70E9EACC627B9D08EB00F829FA4AEDD1" bold="true" box="[1371,1427,1976,2000]" pageId="26" pageNumber="27">Legs</emphasis>
: principally yellow; extreme anteroapical corner dark brown on all femora; midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia without comb of short setae along length of anteroventral surface, hind tibial spur yellow, remainder brown, foretibial spur length 2X apical thickness of foretibia, midtibia with strong, dorsal, bare patch of even thickness for 3/4 of its length, placed centrally, shortest midtibial spur 0.78X length of longest, longest midtibial spur 4.5X apical thickness of midtibia, shortest hind tibial spur 0.8X length of longest, longest hind tibial spur 6.7X apical thickness of hind tibia. Foreleg first tarsomere 1.8X length of foretibia.
<emphasis id="70E9EACC627A9D09EC70FEDAFDB5EB65" bold="true" box="[555,620,331,356]" pageId="27" pageNumber="28">Wing</emphasis>
(
<figureCitation id="DAA62A5B627A9D09EC26FEDDFD0AEB65" box="[637,723,332,356]" captionStart="FIGURES 4255" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 4255. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="27" pageNumber="28">Fig. 51</figureCitation>
): Length
<quantity id="85659B3B627A9D09ED1BFEDDFC41EB62" box="[832,920,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="27" pageNumber="28" unit="mm" value="5.5">5.5 mm</quantity>
(
<quantity id="85659B3B627A9D09EDFCFEDDFBECEB62" box="[935,1077,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.7" metricValueMin="5.3" pageId="27" pageNumber="28" unit="mm" value="5.5" valueMax="5.7" valueMin="5.3">5.35.7 mm</quantity>
, n = 4). Hyaline; apical macula absent or, if present, very light, restricted to apical 1/4 of cell r1; medial macula absent. Macrotrichia in all cells, though absent from posterobasal margin of cell a. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with group of macrotrichia. Vein sc-r present, apical end joining R at 12X its own length prior to origin of Rs. R
<subScript id="DE19349B627A9D09EC7FFE7AFDF5EBF8" attach="left" box="[548,556,491,505]" fontSize="6" pageId="27" pageNumber="28">4</subScript>
absent. R
<subScript id="DE19349B627A9D09ECC1FE7AFD7BEBF8" attach="left" box="[666,674,491,505]" fontSize="6" pageId="27" pageNumber="28">5</subScript>
straight, slight posterior turn near tip. M
<subScript id="DE19349B627A9D09EA2FFE7AFBA5EBF8" attach="left" box="[1140,1148,491,505]" fontSize="6" pageId="27" pageNumber="28">1</subScript>
reaching apex before R
<subScript id="DE19349B627A9D09EBD6FE7AFA4CEBF8" attach="left" box="[1421,1429,491,505]" fontSize="6" pageId="27" pageNumber="28">5</subScript>
, apices of M veins thinning towards margin. M
<subScript id="DE19349B627A9D09ECFCFD83FD76E821" attach="left" box="[679,687,530,544]" fontSize="6" pageId="27" pageNumber="28">4</subScript>
-CuA fork arising just apically of origin of r-m. A
<subScript id="DE19349B627A9D09EAB2FD83FB28E821" attach="left" box="[1257,1265,530,544]" fontSize="6" pageId="27" pageNumber="28">2</subScript>
faintly present as crease.
<emphasis id="70E9EACC627A9D09EF4CFDB8FE55E843" bold="true" box="[279,396,553,578]" pageId="27" pageNumber="28">Abdomen</emphasis>
: Tergites 12 yellow with posterodorsal brown triangle, 36 with anterior 1/2 yellow remainder dark brown, T7 yellow. Tergite 8 smaller than all other abdominal sclerites, with 3 bristles on each apicolateral corner.
<emphasis id="70E9EACC627A9D09EF27FDE0FE35E88B" bold="true" box="[380,492,625,650]" pageId="27" pageNumber="28">Genitalia</emphasis>
(
<figureCitation id="DAA62A5B627A9D09EFA6FDE0FD8DE888" box="[509,596,625,649]" captionStart="FIGURES 99102" captionStartId="99.[151,263,1833,1856]" captionTargetBox="[188,1399,182,1758]" captionTargetId="figure-95@99.[180,1405,176,1767]" captionTargetPageId="99" captionText="FIGURES 99102. Leptomorphus male genitalia, ventral. 99. L. crosskeyi. 100. L. eberhardi. 101. L. fasciculatus. 102. L. femoratus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="27" pageNumber="28">Fig. 99</figureCitation>
): yellow. Sternite 9 sclerotized, circular, 1/3 width of genitalia at widest point, overlapping medial margin of gonocoxite. Tergite 9 longer than wide, margins parallel on apical 2/3 ending in lateral squared off lobe with medially directed spine and medial U-shaped indentation, one thick and one thin, ventrally extending point at base of posterior lobe. Gonocoxite placed basally on T9, medial margin not reaching medial line, bearing gonostylus apically. Gonostylus with two lobes, dorsal lobe a broad based point, ventral lobe half the width of dorsal but 2.5X length and barely tapering until apex, gonocoxite III fused to dorsolateral margin. Aedeagus 2/3 length of T9, tapering towards apex, apodemes ~1/4 total length. Parameres a simple taper, apodemes ~1/2 length of parameres.
</paragraph>
<paragraph id="422236DE627A9D09EE9CFC00FCA3E9F0" blockId="27.[151,1436,151,1549]" pageId="27" pageNumber="28">
<emphasis id="70E9EACC627A9D09EE9CFC00FEC2E9A8" box="[199,283,913,937]" italics="true" pageId="27" pageNumber="28">Female</emphasis>
. (
<figureCitation id="DAA62A5B627A9D09EF6FFC00FE5EE9A8" box="[308,391,913,937]" captionStart="FIGURES 1015" captionStartId="85.[151,263,1903,1926]" captionTargetBox="[175,1416,182,1881]" captionTargetId="figure-54@85.[173,1417,181,1882]" captionTargetPageId="85" captionText="FIGURES 1015. Leptomorphus habitus. 10. L. crassipilus, holotype male. 11. L. crosskeyi, female. 12. L. eberhardi, holotype male. 13. L. eberhardi, paratype female. 14. L. fasciculatus, holotype male. 15. L. femoratus, holotype male." pageId="27" pageNumber="28">Fig. 11</figureCitation>
) As for male, except as follows.
<emphasis id="70E9EACC627A9D09ED4AFC00FCB0E9A8" bold="true" box="[785,873,913,937]" pageId="27" pageNumber="28">Thorax</emphasis>
: Length
<quantity id="85659B3B627A9D09ED8EFC00FBF4E9A8" box="[981,1069,913,937]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="28" unit="mm" value="1.5">1.5 mm</quantity>
(
<quantity id="85659B3B627A9D09EA1BFC00FB30E9A8" box="[1088,1257,913,937]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.56" metricValueMin="1.44" pageId="27" pageNumber="28" unit="mm" value="1.5" valueMax="1.56" valueMin="1.44">1.441.56 mm</quantity>
, n = 2).
<emphasis id="70E9EACC627A9D09EB08FC00FA4DE9AB" bold="true" box="[1363,1428,913,938]" pageId="27" pageNumber="28">Wing</emphasis>
: Length
<quantity id="85659B3B627A9D09EEB5FC24FE9DE9CC" box="[238,324,949,973]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="27" pageNumber="28" unit="mm" value="6.5">6.5 mm</quantity>
(
<quantity id="85659B3B627A9D09EF0EFC24FE39E9CC" box="[341,480,949,973]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="6.6" metricValueMin="6.4" pageId="27" pageNumber="28" unit="mm" value="6.5" valueMax="6.6" valueMin="6.4">6.46.6 mm</quantity>
, n = 2).
<emphasis id="70E9EACC627A9D09EC1AFC24FD6CE9CF" bold="true" box="[577,693,949,974]" pageId="27" pageNumber="28">Abdomen</emphasis>
: Tergites 16 with anterior 1/3 yellow remainder dark brown, T7 dark brown with pair of small lateral yellow spots. Cercus yellow.
</paragraph>
<paragraph id="422236DE627A9D09EE9CFC6FFE1BEE14" blockId="27.[151,1436,151,1549]" box="[199,450,1021,1045]" pageId="27" pageNumber="28">
<emphasis id="70E9EACC627A9D09EE9CFC6FFEE7EE14" box="[199,318,1022,1045]" italics="true" pageId="27" pageNumber="28">Immatures</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0A876555627A9D09EE9CFBB0FE16EE38" box="[199,463,1057,1082]" pageId="27" pageNumber="28" type="biology_ecology">
<paragraph id="422236DE627A9D09EE9CFBB0FE16EE38" blockId="27.[151,1436,151,1549]" box="[199,463,1057,1082]" pageId="27" pageNumber="28">
<emphasis id="70E9EACC627A9D09EE9CFBB0FE93EE3B" bold="true" box="[199,330,1057,1082]" pageId="27" pageNumber="28">BIOLOGY</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0A876555627A9D09EE9CFBD4FB3AEE5C" box="[199,1251,1093,1118]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="422236DE627A9D09EE9CFBD4FB3AEE5C" blockId="27.[151,1436,151,1549]" box="[199,1251,1093,1118]" pageId="27" pageNumber="28">
<emphasis id="70E9EACC627A9D09EE9CFBD4FE4FEE5F" bold="true" box="[199,406,1093,1118]" pageId="27" pageNumber="28">DISTRIBUTION</emphasis>
:
<collectingCountry id="3A8A764E627A9D09EFFFFBD4FE29EE5C" box="[420,496,1093,1117]" name="Ghana" pageId="27" pageNumber="28">Ghana</collectingCountry>
,
<collectingCountry id="3A8A764E627A9D09EFA7FBD4FCC3EE5C" box="[508,794,1093,1117]" name="Central African Republic" pageId="27" pageNumber="28">Central African Republic</collectingCountry>
, and
<collectingCountry id="3A8A764E627A9D09ED0FFBD4FC72EE5C" box="[852,939,1093,1117]" name="Uganda" pageId="27" pageNumber="28">Uganda</collectingCountry>
(
<figureCitation id="DAA62A5B627A9D09EDE1FBD4FBC5EE5C" box="[954,1052,1093,1117]" captionStart="FIGURE 134136" captionStartId="107.[151,249,1814,1837]" captionTargetBox="[151,1436,193,1793]" captionTargetId="figure-66@107.[151,1436,193,1793]" captionTargetPageId="107" captionText="FIGURE 134136. Distribution of Afrotropical Leptomorphus. 134. L. grjebinei (red circle), L. mandelai (blue square), L. stigmatus (black triangle). 135. L. aliciae (purple star), L. carnevalei (black square), L. couturieri (black circle), L. gracilis (red triangle). 136. L. crosskeyi (red triangle), L. medleri (blue star), L. obscurus (black circle)." pageId="27" pageNumber="28">Fig. 136</figureCitation>
),
<quantity id="85659B3B627A9D09EA6BFBD4FB06EE5C" box="[1072,1247,1093,1117]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="12.5" metricValueMin="1.8" pageId="27" pageNumber="28" unit="m" value="715.0" valueMax="1250.0" valueMin="180.0">1801250 masl</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0A876555627A9D09EE9CFBF8FCE4EEC8" pageId="27" pageNumber="28" type="discussion">
<paragraph id="422236DE627A9D09EE9CFBF8FCE4EEC8" blockId="27.[151,1436,151,1549]" pageId="27" pageNumber="28">
<emphasis id="70E9EACC627A9D09EE9CFBF8FEA2EE83" bold="true" box="[199,379,1129,1154]" pageId="27" pageNumber="28">DISCUSSION:</emphasis>
As discussed below in the phylogeny section, the placement of
<taxonomicName id="859D4D5D627A9D09EA27FBF8FAA2EE80" box="[1148,1403,1129,1153]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="70E9EACC627A9D09EA27FBF8FAC7EE80" box="[1148,1310,1129,1153]" italics="true" pageId="27" pageNumber="28">Leptomorphus</emphasis>
species
</taxonomicName>
in subgenera (
<bibRefCitation id="260C4B2F627A9D09EF43FB1CFE7DEEA4" author="Matile, L." box="[280,420,1165,1189]" pageId="27" pageNumber="28" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977</bibRefCitation>
) is not supported by our phylogenetic results. This species is therefore removed from the subgenus
<taxonomicName id="859D4D5D627A9D09EF5DFB20FE7FEEC8" box="[262,422,1201,1225]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subGenus" subGenus="Gymnoscutum">
<emphasis id="70E9EACC627A9D09EF5DFB20FE7FEEC8" box="[262,422,1201,1225]" italics="true" pageId="27" pageNumber="28">Gymnoscutum</emphasis>
</taxonomicName>
and placed solely in
<emphasis id="70E9EACC627A9D09ECC8FB20FCE4EEC8" box="[659,829,1201,1225]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="859D4D5D627A9D09ECC8FB20FCE0EEC8" authorityName=": Edwards" authorityYear="1925" box="[659,825,1201,1225]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0A876555627A9D09EE9CFB44FD19EC0C" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="422236DE627A9D09EE9CFB44FC43EF58" blockId="27.[151,1436,151,1549]" pageId="27" pageNumber="28">
<materialsCitation id="F2F53C83627A9D09EE9CFB44FC43EF58" collectingDate="1970-09-29" collectionCode="MATERIAL" collectorName="Det. C. J. Borkent" determinerName="L. Matile &amp; Det. C. J. Borkent" location="Matile" pageId="27" pageNumber="28" specimenCount="2" specimenCount-male="2" stateProvince="Monaghan" typeStatus="holotype">
<emphasis id="70E9EACC627A9D09EE9CFB44FDDBEEEF" bold="true" box="[199,514,1237,1262]" pageId="27" pageNumber="28">
<collectionCode id="248CAE1B627A9D09EE9CFB44FE86EEEF" box="[199,351,1237,1262]" pageId="27" pageNumber="28">MATERIAL</collectionCode>
EXAMINED
</emphasis>
:
<emphasis id="70E9EACC627A9D09EC4BFB44FD59EEEC" box="[528,640,1237,1261]" italics="true" pageId="27" pageNumber="28">
<typeStatus id="9D26887C627A9D09EC4BFB44FDA3EEEC" box="[528,634,1237,1261]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
:
</emphasis>
adult male, pinned on double mount minuten, genitalia in glass vial on pin, labelled “[Blue label] REP. CENTRAFRIC./ LA MABOKE/
<date id="3623101E627A9D09EDF0FB68FBF0EF10" box="[939,1065,1273,1297]" pageId="27" pageNumber="28" value="1970-09-29">
<collectingDate id="2667E9F6627A9D09EDF0FB68FBF0EF10" box="[939,1065,1273,1297]" pageId="27" pageNumber="28" value="1970-09-29">29.IX.1970</collectingDate>
</date>
/ L. MATILE rec.; [Red label]
<typeStatus id="9D26887C627A9D09EECCFA8CFEF7EF34" box="[151,302,1309,1333]" pageId="27" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
;
<taxonomicName id="859D4D5D627A9D09EF64FA8CFE38EF34" authorityName=": Edwards" authorityYear="1925" box="[319,481,1309,1333]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
/ (Afroleptomorphus)/africanus
<taxonomicNameLabel id="6BDA57B7627A9D09ED08FA8FFC4CEF34" box="[851,917,1310,1333]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
<specimenCount id="549BFD57627A9D09EDF9FA8FFC6EEF34" box="[930,951,1310,1333]" pageId="27" pageNumber="28" type="male"></specimenCount>
ht/
<determinerName id="B7FEDA1A627A9D09EDBAFA8FFB96EF34" box="[993,1103,1309,1333]" pageId="27" pageNumber="28">L. Matile</determinerName>
det. 1974;
<typeStatus id="9D26887C627A9D09EA84FA8CFAAAEF34" box="[1247,1395,1309,1333]" pageId="27" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="549BFD57627A9D09EB24FA8FFA4DEF34" box="[1407,1428,1310,1333]" pageId="27" pageNumber="28" type="male"></specimenCount>
/
<taxonomicName id="859D4D5D627A9D09EECCFAD0FE7FEF58" authorityName="Matile, 1980:" authorityYear="1229" box="[151,422,1345,1369]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="crosskeyi">
<emphasis id="70E9EACC627A9D09EECCFAD0FE7FEF58" box="[151,422,1345,1369]" italics="true" pageId="27" pageNumber="28">Leptomorphus crosskeyi</emphasis>
</taxonomicName>
/
<location id="47426005627A9D09EFEEFAD0FE24EF58" LSID="urn:lsid:plazi:treatment:CA3487C8627B9D09EECCFDD6FD19EC0C:47426005627A9D09EFEEFAD0FE24EF58" box="[437,509,1345,1369]" name="Matile" pageId="27" pageNumber="28" stateProvince="Monaghan">Matile</location>
/
<determinerName id="B7FEDA1A627A9D09EC56FAD3FD0AEF58" box="[525,723,1345,1369]" pageId="27" pageNumber="28">
<collectorName id="EF685308627A9D09EC56FAD3FD0AEF58" box="[525,723,1345,1369]" pageId="27" pageNumber="28">Det. C.J. Borkent</collectorName>
</determinerName>
, 2012” [
<collectingRegion id="8059F83C627A9D09ED6BFAD3FC56EF58" box="[816,911,1346,1369]" country="Ireland" name="Monaghan" pageId="27" pageNumber="28">MNHN</collectingRegion>
].
</materialsCitation>
</paragraph>
<paragraph id="422236DE627A9D09EE9CFAF7FAF2EFA0" blockId="27.[151,1436,151,1549]" pageId="27" pageNumber="28">
<materialsCitation id="F2F53C83627A9D09EE9CFAF7FBDEEFA0" collectingDate="1967-08-20" collectingDateMax="1972-01-09" collectingDateMin="1967-08-20" collectionCode="MNHN, R" collectorName="Labelled" country="Uganda" location="Forest" pageId="27" pageNumber="28" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Kibale" typeStatus="paratype">
<emphasis id="70E9EACC627A9D09EE9CFAF7FE98EF7C" box="[199,321,1382,1405]" italics="true" pageId="27" pageNumber="28">
<typeStatus id="9D26887C627A9D09EE9CFAF7FEE5EF7C" box="[199,316,1382,1405]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectorName id="EF685308627A9D09EF15FAF4FE69EF7C" box="[334,432,1381,1405]" pageId="27" pageNumber="28">Labelled</collectorName>
as for
<typeStatus id="9D26887C627A9D09EC54FAF4FDA8EF7C" box="[527,625,1381,1405]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(
<specimenCount id="549BFD57627A9D09ECD2FAF4FD73EF7C" box="[649,682,1381,1405]" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
, labelled as
<typeStatus id="9D26887C627A9D09ED1CFAF4FC75EF7C" box="[839,940,1381,1405]" pageId="27" pageNumber="28" type="allotype">Allotype</typeStatus>
,
<collectionCode id="248CAE1B627A9D09EDE6FAF4FBC1EF7C" box="[957,1048,1381,1405]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="28">MNHN</collectionCode>
); except
<date id="3623101E627A9D09EADCFAF4FB31EF7C" box="[1159,1256,1381,1405]" pageId="27" pageNumber="28" value="1970-10-02">
<collectingDate id="2667E9F6627A9D09EADCFAF4FB31EF7C" box="[1159,1256,1381,1405]" pageId="27" pageNumber="28" value="1970-10-02">2.x.1970</collectingDate>
</date>
(
<specimenCount id="549BFD57627A9D09EB5BFAF4FAF8EF7C" box="[1280,1313,1381,1405]" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
,
<collectionCode id="248CAE1B627A9D09EB6EFAF4FA49EF7C" box="[1333,1424,1381,1405]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="28">MNHN</collectionCode>
);
<date id="3623101E627A9D09EECCFA18FEC7EFA0" box="[151,286,1417,1441]" pageId="27" pageNumber="28" value="1967-08-20">
<collectingDate id="2667E9F6627A9D09EECCFA18FEC7EFA0" box="[151,286,1417,1441]" pageId="27" pageNumber="28" value="1967-08-20">20.viii.1967</collectingDate>
</date>
(
<specimenCount id="549BFD57627A9D09EF6BFA18FE88EFA0" box="[304,337,1417,1441]" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B627A9D09EF04FA18FE63EFA0" box="[351,442,1417,1441]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="28">MNHN</collectionCode>
);
<collectingCountry id="3A8A764E627A9D09EF96FA18FD90EFA0" box="[461,585,1417,1441]" name="Uganda" pageId="27" pageNumber="28">UGANDA</collectingCountry>
,
<collectingRegion id="8059F83C627A9D09EC0EFA18FD46EFA0" box="[597,671,1417,1441]" country="Uganda" name="Kibale" pageId="27" pageNumber="28">Kibale</collectingRegion>
<location id="47426005627A9D09ECFDFA1BFD36EFA0" LSID="urn:lsid:plazi:treatment:CA3487C8627B9D09EECCFDD6FD19EC0C:47426005627A9D09ECFDFA1BFD36EFA0" box="[678,751,1418,1441]" country="Uganda" name="Forest" pageId="27" pageNumber="28" stateProvince="Kibale">Forest</location>
,
<date id="3623101E627A9D09ECA1FA18FCA3EFA0" box="[762,890,1417,1441]" pageId="27" pageNumber="28" value="1971-12-12">
<collectingDate id="2667E9F6627A9D09ECA1FA18FCA3EFA0" box="[762,890,1417,1441]" pageId="27" pageNumber="28" value="1971-12-12">12.xii.1971</collectingDate>
</date>
<date id="3623101E627A9D09EDD0FA18FC3FEFA0" box="[907,998,1417,1441]" pageId="27" pageNumber="28" value="1972-01-09">
<collectingDate id="2667E9F6627A9D09EDD0FA18FC3FEFA0" box="[907,998,1417,1441]" pageId="27" pageNumber="28" value="1972-01-09">9.i.1972</collectingDate>
</date>
,
<collectionCode id="248CAE1B627A9D09EDAAFA1BFBDEEFA0" box="[1009,1031,1418,1441]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="28">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="F2F53C83627A9D09EA50FA1BFAFEEFA0" box="[1035,1319,1417,1441]" collectionCode="BMNH" collectorName="L. Mason" country="Uganda" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Kibale" typeStatus="paratype">
<collectorName id="EF685308627A9D09EA50FA1BFBAFEFA0" box="[1035,1142,1418,1441]" pageId="27" pageNumber="28">L. Mason</collectorName>
(1#m,
<collectionCode id="248CAE1B627A9D09EA9DFA18FAC6EFA0" box="[1222,1311,1417,1441]" pageId="27" pageNumber="28">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="422236DE627A9D09EE9CFA3CFD19EC0C" blockId="27.[151,1436,151,1549]" pageId="27" pageNumber="28">
<emphasis id="70E9EACC627A9D09EE9CFA3CFEAEEFC4" box="[199,375,1453,1477]" italics="true" pageId="27" pageNumber="28">Other material:</emphasis>
<materialsCitation id="F2F53C83627A9D09EF26FA3CFA88EFC4" box="[381,1361,1453,1477]" collectingDate="1967-08-20" collectionCode="MNHN" collectorName="L. Matile." country="Central African Republic" location="La Maboke" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1">
<collectingCountry id="3A8A764E627A9D09EF26FA3CFCC8EFC4" box="[381,785,1453,1477]" name="Central African Republic" pageId="27" pageNumber="28">CENTRAL AFRICAN REPUBLIC</collectingCountry>
,
<location id="47426005627A9D09ED40FA3FFC47EFC4" LSID="urn:lsid:plazi:treatment:CA3487C8627B9D09EECCFDD6FD19EC0C:47426005627A9D09ED40FA3FFC47EFC4" box="[795,926,1453,1477]" country="Central African Republic" name="La Maboke" pageId="27" pageNumber="28">La Maboke</location>
,
<date id="3623101E627A9D09EDF2FA3CFBEAEFC4" box="[937,1075,1453,1477]" pageId="27" pageNumber="28" value="1967-08-20">
<collectingDate id="2667E9F6627A9D09EDF2FA3CFBEAEFC4" box="[937,1075,1453,1477]" pageId="27" pageNumber="28" value="1967-08-20">20.viii.1967</collectingDate>
</date>
,
<collectorName id="EF685308627A9D09EA65FA3FFB74EFC4" box="[1086,1197,1453,1477]" pageId="27" pageNumber="28">L. Matile.</collectorName>
(
<specimenCount id="549BFD57627A9D09EAE4FA3CFB39EFC4" box="[1215,1248,1453,1477]" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B627A9D09EAB5FA3CFA90EFC4" box="[1262,1353,1453,1477]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="28">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C83627A9D09EB00FA3FFDD9EFE8" collectingDate="1970-10-02" collectionCode="MNHN" country="Central African Republic" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1">
same, except
<date id="3623101E627A9D09EEB0FA40FE89EFE8" box="[235,336,1489,1513]" pageId="27" pageNumber="28" value="1970-10-02">
<collectingDate id="2667E9F6627A9D09EEB0FA40FE89EFE8" box="[235,336,1489,1513]" pageId="27" pageNumber="28" value="1970-10-02">2.x.1970</collectingDate>
</date>
. (
<specimenCount id="549BFD57627A9D09EF31FA40FE52EFE8" box="[362,395,1489,1513]" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
,
<collectionCode id="248CAE1B627A9D09EFC6FA40FE21EFE8" box="[413,504,1489,1513]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="28">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C83627A9D09EC55FA40FB86EFE8" box="[526,1119,1489,1513]" collectingDate="1994-10-08" collectingDateMax="1994-10-15" collectingDateMin="1994-10-08" collectionCode="ZMUN" country="Ghana" location="Kakum" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1">
<collectingCountry id="3A8A764E627A9D09EC55FA40FDAFEFE8" box="[526,630,1489,1513]" name="Ghana" pageId="27" pageNumber="28">GHANA</collectingCountry>
,
<location id="47426005627A9D09ECDDFA40FD01EFE8" LSID="urn:lsid:plazi:treatment:CA3487C8627B9D09EECCFDD6FD19EC0C:47426005627A9D09ECDDFA40FD01EFE8" box="[646,728,1489,1513]" country="Ghana" name="Kakum" pageId="27" pageNumber="28">Kakum</location>
N.P.,
<date id="3623101E627A9D09ED7FFA40FC6BEFE8" box="[804,946,1489,1513]" pageId="27" pageNumber="28" value="1994-10-08" valueMax="1994-10-15" valueMin="1994-10-08">
<collectingDate id="2667E9F6627A9D09ED7FFA40FC6BEFE8" box="[804,946,1489,1513]" pageId="27" pageNumber="28" value="1994-10-08" valueMax="1994-10-15" valueMin="1994-10-08">815.x.1994</collectingDate>
</date>
. (
<specimenCount id="549BFD57627A9D09ED97FA40FC37EFE8" box="[972,1006,1489,1513]" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B627A9D09EA5BFA40FB81EFE8" box="[1024,1112,1489,1513]" pageId="27" pageNumber="28">ZMUN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F2F53C83627A9D09EA35FA40FE7DEC0C" collectingDate="1971-12-12" collectingDateMax="1972-01-09" collectingDateMin="1971-12-12" collectionCode="R" country="Uganda" location="Forest" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Kibale">
<collectingCountry id="3A8A764E627A9D09EA35FA40FB33EFE8" box="[1134,1258,1489,1513]" name="Uganda" pageId="27" pageNumber="28">UGANDA</collectingCountry>
,
<collectingRegion id="8059F83C627A9D09EAA2FA40FA9DEFE8" box="[1273,1348,1489,1513]" country="Uganda" name="Kibale" pageId="27" pageNumber="28">Kibale</collectingRegion>
<location id="47426005627A9D09EB14FA43FA41EFE8" LSID="urn:lsid:plazi:treatment:CA3487C8627B9D09EECCFDD6FD19EC0C:47426005627A9D09EB14FA43FA41EFE8" box="[1359,1432,1490,1513]" country="Uganda" name="Forest" pageId="27" pageNumber="28" stateProvince="Kibale">Forest</location>
,
<date id="3623101E627A9D09EECCFA64FECEEC0C" box="[151,279,1525,1549]" pageId="27" pageNumber="28" value="1971-12-12">
<collectingDate id="2667E9F6627A9D09EECCFA64FECEEC0C" box="[151,279,1525,1549]" pageId="27" pageNumber="28" value="1971-12-12">12.xii.1971</collectingDate>
</date>
<date id="3623101E627A9D09EF7CFA64FE5AEC0C" box="[295,387,1525,1549]" pageId="27" pageNumber="28" value="1972-01-09">
<collectingDate id="2667E9F6627A9D09EF7CFA64FE5AEC0C" box="[295,387,1525,1549]" pageId="27" pageNumber="28" value="1972-01-09">9.i.1972</collectingDate>
</date>
,
<collectionCode id="248CAE1B627A9D09EFD5FA67FE7DEC0C" box="[398,420,1526,1549]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="28">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="F2F53C83627A9D09EFF3FA67FD65EC0C" box="[424,700,1525,1549]" collectionCode="BMNH" collectorName="Masou." country="Uganda" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1">
L.
<collectorName id="EF685308627A9D09EF9CFA67FDC2EC0C" box="[455,539,1526,1549]" pageId="27" pageNumber="28">Masou.</collectorName>
(
<specimenCount id="549BFD57627A9D09EC77FA64FD94EC0C" box="[556,589,1525,1549]" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
,
<collectionCode id="248CAE1B627A9D09EC00FA64FD6DEC0C" box="[603,692,1525,1549]" pageId="27" pageNumber="28">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>