225 lines
32 KiB
XML
225 lines
32 KiB
XML
<document id="56CF5C65F46734A1A62947958F66910D" ID-ISSN="11755334" ID-ZooBank="2412CB4F-4D29-4988-80C1-205D16767678" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629824082200" checkinUser="felipe" docAuthor="Borkent, Christopher J. & Wheeler, Terry A." docDate="2012" docId="CA3487C862799D08EECCF8ADFE93EBF2" docLanguage="en" docName="zt03549p117.pdf" docOrigin="Zootaxa 3549" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leptomorphus crassipilus Borkent 2012, new species" docType="treatment" docUuid="8DB8AB10-8C9D-4B41-A68C-6A72056CF6EC" docUuidSource="ZooBank" docVersion="4" lastPageNumber="27" masterDocId="360DFFB062619D12EE5BFF91FFD9EA01" masterDocTitle="Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549" masterLastPageNumber="117" masterPageNumber="1" pageNumber="25" updateTime="1699117448841" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="23C6FD38B6BFC5CFD7AACF5A14B484B6" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4859724860A2848962E80C052A6A29D8">
|
||
<mods:title id="F800497778480F8CA4DFC934705ACC05">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="2D52AF42176C3317FC11E0996DBA174A" type="personal">
|
||
<mods:role id="207B173A026DD622D7573A316C5CB786">
|
||
<mods:roleTerm id="8E748E4BFB6CC88BACC4F6F4C708917A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F885889EED180EDBAED9B1C2263D294B">Borkent, Christopher J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="AAE4B308A1EC8862F16E9CCD93DECE68" type="personal">
|
||
<mods:role id="A86E8E5BBE06D4CBB2F123CA23697100">
|
||
<mods:roleTerm id="90D0610F9AED2E7CA9F9A5A029CD9983">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="35A25A0075E0873D8A3ECA66A50CD6D7">Wheeler, Terry A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="DA4195581A8895166F76F4D581271C37">text</mods:typeOfResource>
|
||
<mods:relatedItem id="097512CDD529A5667FB4509EDD02F3F4" type="host">
|
||
<mods:titleInfo id="832292CB00A6FC7EE23709364F58207A">
|
||
<mods:title id="03A2BE3AF5FE40661AD246F970CE415F">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="8D79F2F27C09ED8BF55EAADC9DCA5660">
|
||
<mods:date id="6D828D1717A91C7A6FCC22F1F7AC867C">2012</mods:date>
|
||
<mods:detail id="68D58BD6C5ACEFE07CE11C772A6665B4" type="pubDate">
|
||
<mods:number id="62F2FE5CB910DA840CA44A4F60BE1000">2012-11-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="51F5BBF4B1768CEA830170F71E9DD491" type="volume">
|
||
<mods:number id="85EF88955EE8C2CDA83FEB6D6E3CD871">3549</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="1F96ED66113F58AC2A36E6A2EA100C06" unit="page">
|
||
<mods:start id="11E9BAF3A2CF0A0EF86CA355B2C90B29">1</mods:start>
|
||
<mods:end id="3187611D272EF943208D68854DC703B5">117</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="A3C60701682CA9BA53D832CBBDD28AD7">journal article</mods:classification>
|
||
<mods:identifier id="76DD9353A464663AE97C9750E4FF5D7B" type="ISSN">11755334</mods:identifier>
|
||
<mods:identifier id="4B97A40153EA249404FFC33664066490" type="ZooBank">2412CB4F-4D29-4988-80C1-205D16767678</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CA3487C862799D08EECCF8ADFE93EBF2" ID-DOI="http://doi.org/10.5281/zenodo.5258335" ID-Zenodo-Dep="5258335" LSID="urn:lsid:zoobank.org:act:8DB8AB10-8C9D-4B41-A68C-6A72056CF6EC" httpUri="http://treatment.plazi.org/id/CA3487C862799D08EECCF8ADFE93EBF2" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
|
||
<subSubSection id="0A87655562799D0AEECCF8ADFD21ED57" box="[151,760,1852,1878]" pageId="24" pageNumber="25" type="nomenclature">
|
||
<paragraph id="422236DE62799D0AEECCF8ADFD21ED57" blockId="24.[151,760,1852,1911]" box="[151,760,1852,1878]" pageId="24" pageNumber="25">
|
||
<heading id="196A81B262799D0AEECCF8ADFD21ED57" bold="true" box="[151,760,1852,1878]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
|
||
<emphasis id="70E9EACC62799D0AEECCF8ADFD21ED57" bold="true" box="[151,760,1852,1878]" pageId="24" pageNumber="25">
|
||
9.
|
||
<taxonomicName id="859D4D5D62799D0AEEEFF8ADFD82ED57" ID-CoL="8CJ46" authority="Borkent" authorityName="Borkent" authorityYear="2012" box="[180,603,1852,1878]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crassipilus" status="sp. nov.">
|
||
<emphasis id="70E9EACC62799D0AEEEFF8ADFE32ED57" bold="true" box="[180,491,1852,1878]" italics="true" pageId="24" pageNumber="25">Leptomorphus crassipilus</emphasis>
|
||
Borkent
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="6BDA57B762799D0AEC33F8ADFD21ED57" box="[616,760,1852,1878]" pageId="24" pageNumber="25" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562799D0AEECCF8CEFDC5ED76" box="[151,540,1887,1911]" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="422236DE62799D0AEECCF8CEFDC5ED76" blockId="24.[151,760,1852,1911]" box="[151,540,1887,1911]" pageId="24" pageNumber="25">
|
||
(
|
||
<figureCitation id="DAA62A5B62799D0AEEC5F8CEFEC0ED76" box="[158,281,1887,1911]" captionStart="FIGURES 10–15" captionStartId="85.[151,263,1903,1926]" captionTargetBox="[175,1416,182,1881]" captionTargetId="figure-54@85.[173,1417,181,1882]" captionTargetPageId="85" captionText="FIGURES 10–15. Leptomorphus habitus. 10. L. crassipilus, holotype male. 11. L. crosskeyi, female. 12. L. eberhardi, holotype male. 13. L. eberhardi, paratype female. 14. L. fasciculatus, holotype male. 15. L. femoratus, holotype male." pageId="24" pageNumber="25">Figures 10</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62799D0AEF7EF8CEFE9AED76" box="[293,323,1887,1911]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="24" pageNumber="25">50</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62799D0AEF14F8CEFEB4ED76" box="[335,365,1887,1911]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="24" pageNumber="25">98</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62799D0AEF22F8CEFE7CED76" box="[377,421,1887,1911]" captionStart="FIGURE 142–143" captionStartId="110.[151,249,1784,1807]" captionTargetBox="[151,1436,193,1763]" captionTargetId="figure-57@110.[151,1436,193,1763]" captionTargetPageId="110" captionText="FIGURE 142–143. Distribution of Neotropical Leptomorphus. 142. L. amorimi (blue circle), L. crassipilus (red square), L. fasciculatus (black triangle), L. neivai (red star). 143. L. brandiae (purple triangle), L. eberhardi (blue circle), L. femoratus (red star), L. waodani (black square)." pageId="24" pageNumber="25">142</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62799D0AEFEAF8CEFE04ED76" box="[433,477,1887,1911]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values>50% are shown in bold below the branches." pageId="24" pageNumber="25">150</figureCitation>
|
||
,
|
||
<figureCitation id="DAA62A5B62799D0AEFB2F8CEFDCDED76" box="[489,532,1887,1911]" captionStart="FIGURE 155" captionStartId="116.[151,250,858,881]" captionTargetBox="[151,1425,187,838]" captionTargetId="figure-58@116.[151,1436,159,885]" captionTargetPageId="116" captionText="FIGURE 155. One of three equally parsimonious trees showing the relationships between species in the L. furcatus species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="24" pageNumber="25">155</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562799D0BEECCF837FD9BEB86" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="diagnosis">
|
||
<paragraph id="422236DE62799D0BEECCF837FA4EEB1B" blockId="24.[151,1436,1958,2018]" lastBlockId="25.[151,1437,151,2034]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
<emphasis id="70E9EACC62799D0AEECCF837FEEDEDBE" bold="true" box="[151,308,1958,1983]" pageId="24" pageNumber="25">DIAGNOSIS</emphasis>
|
||
: The only extant species of
|
||
<taxonomicName id="859D4D5D62799D0AEC35F836FCC9EDBE" authorityName=": Edwards" authorityYear="1925" box="[622,784,1959,1983]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70E9EACC62799D0AEC35F836FCC9EDBE" box="[622,784,1959,1983]" italics="true" pageId="24" pageNumber="25">Leptomorphus</emphasis>
|
||
</taxonomicName>
|
||
with the following combination of characters: scutum and scutellum evenly covered with fine trichia; male foretibia with a dense row (comb) of short anteroventral bristles; palp segment 5 with thick covering of fine, white, setulae; male tergite 9 posterior margin rounded laterally and concave medially, lateral process apex with 5 rows of tightly spaced short blunt brown bristles; gonocoxite placed basally on, and subequal in length to, tergite 9, gonostylus ~2/3 length of gonocoxite with two apical processes, medial one most pronounced and tapering to a point, the lateral ½ the length of medial, thin and rod-like (
|
||
<figureCitation id="DAA62A5B62789D0BEB6DFE93FA53EB1B" box="[1334,1418,258,282]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="25" pageNumber="26">Fig. 98</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="422236DE62789D0BEE9CFEB7FD9BEB86" blockId="25.[151,1437,151,2034]" pageId="25" pageNumber="26">
|
||
This species is distinguished from most other Neotropical species missing R
|
||
<subScript id="DE19349B62789D0BEA46FEA4FBFCEB42" attach="left" box="[1053,1061,309,323]" fontSize="6" pageId="25" pageNumber="26">4</subScript>
|
||
by the yellow or white scutellum and striped abdomen (
|
||
<figureCitation id="DAA62A5B62789D0BEFC6FEDDFE2CEB65" box="[413,501,332,356]" captionStart="FIGURES 10–15" captionStartId="85.[151,263,1903,1926]" captionTargetBox="[175,1416,182,1881]" captionTargetId="figure-54@85.[173,1417,181,1882]" captionTargetPageId="85" captionText="FIGURES 10–15. Leptomorphus habitus. 10. L. crassipilus, holotype male. 11. L. crosskeyi, female. 12. L. eberhardi, holotype male. 13. L. eberhardi, paratype female. 14. L. fasciculatus, holotype male. 15. L. femoratus, holotype male." pageId="25" pageNumber="26">Fig. 10</figureCitation>
|
||
). It can be distinguished from
|
||
<taxonomicName id="859D4D5D62789D0BED3CFEDCFC2FEB65" authorityName="Borkent" authorityYear="2012" box="[871,1014,332,356]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="eberhardi">
|
||
<emphasis id="70E9EACC62789D0BED3CFEDCFC2FEB65" box="[871,1014,332,356]" italics="true" pageId="25" pageNumber="26">L. eberhardi</emphasis>
|
||
</taxonomicName>
|
||
by the rows of short spines on the posterior margin of tergite 9 (
|
||
<figureCitation id="DAA62A5B62789D0BEFBAFEFEFDECEB86" box="[481,565,367,391]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="25" pageNumber="26">Fig. 98</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562789D08EE9CFE03FE1BEAD5" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
|
||
<paragraph id="422236DE62789D0BEE9CFE03FD5CEDF3" blockId="25.[151,1437,151,2034]" pageId="25" pageNumber="26">
|
||
<emphasis id="70E9EACC62789D0BEE9CFE03FE50EBAA" bold="true" box="[199,393,402,427]" pageId="25" pageNumber="26">DESCRIPTION</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC62789D0BEFC3FE02FE08EBAA" box="[408,465,403,427]" italics="true" pageId="25" pageNumber="26">Male</emphasis>
|
||
. (
|
||
<figureCitation id="DAA62A5B62789D0BEFB2FE02FDE2EBAA" box="[489,571,403,427]" captionStart="FIGURES 10–15" captionStartId="85.[151,263,1903,1926]" captionTargetBox="[175,1416,182,1881]" captionTargetId="figure-54@85.[173,1417,181,1882]" captionTargetPageId="85" captionText="FIGURES 10–15. Leptomorphus habitus. 10. L. crassipilus, holotype male. 11. L. crosskeyi, female. 12. L. eberhardi, holotype male. 13. L. eberhardi, paratype female. 14. L. fasciculatus, holotype male. 15. L. femoratus, holotype male." pageId="25" pageNumber="26">Fig. 10</figureCitation>
|
||
)
|
||
<emphasis id="70E9EACC62789D0BEC10FE02FD53EBAA" bold="true" box="[587,650,403,427]" pageId="25" pageNumber="26">Head</emphasis>
|
||
: yellow, circular in anterior view. Antenna brown; scape yellow, with yellow setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae present; pedicel yellow, with 2 large bristles, few setae on apicodorsal margin, patch of fine setae apicoventrally; flagellomere 1 with tapered base yellow remainder brown; flagellomere 6 1.9X as long as broad. Clypeus light brown, slightly laterally compressed oval; bristles on clypeus yellow, 6 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 2X as long as face. Face light brown; shape a subequal triangle, bare. Frons light brown; with many bristles covering ventral half, frontal furrow running 1/10 distance from dorsal margin towards ventral margin, frontal cleft just anterior of median ocellus. Palpus yellow; segment 1 small but visible below eye, segments increasing in length, segment 5 not visible, segment 3 without definite patch of fine setae. Labellum brown. Eye with very few, short inter-ommatidial setulae scattered on surface. Occiput yellow with appressed, anteriorly directed setae. Three ocelli; straight line, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle dark brown/black.
|
||
<emphasis id="70E9EACC62789D0BEDB8FCACFBE5E954" bold="true" box="[995,1084,829,853]" pageId="25" pageNumber="26">Thorax</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B62789D0BEAFBFCADFADAE955" box="[1184,1283,828,852]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="25" pageNumber="26" unit="mm" value="1.19">1.19 mm</quantity>
|
||
(n = 1). Dark brown with yellow lateral spots. Scutum dark brown/black with blue-green specks, pair of small yellow posterolateral spots; surface of scutum covered with trichia; acrostichal setae absent; single dorsocentral seta present anteriorly; single row of lateral setae present; patch of setae on scutum at wing base present. Scutellum yellow; without large bristles but with covering of fine trichia. Prescutum yellow. Mediotergite brown with 4 bristles on posterolateral corners, and covering of appressed trichia. Laterotergite brown, with covering of appressed trichia; anterior margin of laterotergite abutting katepisternum. Anepimeron brown with posterodorsal corner yellow. Anepisternum brown. Katepisternum brown with dorsal third yellow. Antepronotum and proepisternum yellow. Margin of anterior and posterior spiracles yellow with yellow trichia. Metepisternum yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter with basal 1/3 of stem ivory, apical portion and knob dark brown.
|
||
<emphasis id="70E9EACC62789D0BEFB4FB0EFDFEEEB6" bold="true" box="[495,551,1183,1207]" pageId="25" pageNumber="26">Legs</emphasis>
|
||
: principally yellow; trochanters light brown; hind femur with basal and apical 1/4 brown; extreme anteroapical corner yellow on forefemur, on other femora dark brown; tarsi brown. Midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia with comb of short setae along length of anteroventral surface, though bristles further apart than in other species, tibial spurs dark brown, foretibial spur length 1.9X apical thickness of foretibia, midtibia with faint, dorsal, bare patch of even thickness for 3/4 of its length, placed basally, shortest midtibial spur 0.8X length of longest, longest midtibial spur 5.3X apical thickness of midtibia, shortest hind tibial spur 0.8X length of longest, longest hind tibial spur 3.5X apical thickness of hind tibia. Foreleg first tarsomere 1.6X length of foretibia.
|
||
<emphasis id="70E9EACC62789D0BED3AFA06FC7BEFB1" bold="true" box="[865,930,1431,1456]" pageId="25" pageNumber="26">Wing</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62789D0BEDE8FA06FBD3EFAE" box="[947,1034,1431,1455]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="25" pageNumber="26">Fig. 50</figureCitation>
|
||
): Length 6.0 mm (n = 1). Hyaline; apical macula dark brown running from anterior to posterior wing margin, beginning halfway along R
|
||
<subScript id="DE19349B62789D0BEB4DFA5BFAC7EFD9" attach="left" box="[1302,1310,1482,1496]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
, but fading towards apex and posterior margin; medial macula extending from Sc to posterior wing margin (fainter on posterior third). Macrotrichia in all cells, though absent from posterobasal margin of cell a. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a few short setae. Vein sc-r present, apical end joining R within its own length prior to origin of Rs. R
|
||
<subScript id="DE19349B62789D0BED70F9CAFCEAEC68" attach="left" box="[811,819,1627,1641]" fontSize="6" pageId="25" pageNumber="26">4</subScript>
|
||
absent. R
|
||
<subScript id="DE19349B62789D0BEDFDF9CAFC77EC68" attach="left" box="[934,942,1627,1641]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
straight, slight posterior turn near tip. M
|
||
<subScript id="DE19349B62789D0BEBCFF9CAFA45EC68" attach="left" box="[1428,1436,1627,1641]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
reaching apex just before R
|
||
<subScript id="DE19349B62789D0BEF8DF910FE07EC8E" attach="left" box="[470,478,1665,1679]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
, apices of M veins clearly reaching wing margin. M
|
||
<subScript id="DE19349B62789D0BEA18F910FB92EC8E" attach="left" box="[1091,1099,1665,1679]" fontSize="6" pageId="25" pageNumber="26">4</subScript>
|
||
-CuA fork arising apically of origin of r-m. A
|
||
<subScript id="DE19349B62789D0BEF10F936FE8AECB4" attach="left" box="[331,339,1703,1717]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
|
||
absent.
|
||
<emphasis id="70E9EACC62789D0BEFEAF906FDFFECB1" bold="true" box="[433,550,1687,1712]" pageId="25" pageNumber="26">Abdomen</emphasis>
|
||
: Tergites principally dark brown, T3–6 with anterior 1/3–1/2 yellow. Tergite 8 smaller than all other abdominal sclerites, with 3–4 bristles on each apicolateral corner.
|
||
<emphasis id="70E9EACC62789D0BEA33F92FFB0FECD6" bold="true" box="[1128,1238,1726,1751]" pageId="25" pageNumber="26">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62789D0BEABDF92FFAE3ECD7" box="[1254,1338,1726,1750]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="25" pageNumber="26">Fig. 98</figureCitation>
|
||
): brown. Sternite 9 not clearly visible due to condition of specimen. Tergite 9 as wide as long, margins ~parallel, ending apically in broad rounded lobes and shallow medial indentation, rounded lobe bearing 5–6 rows of short, blunt, closely spaced bristles on ventroapical surface. Gonocoxite placed basally on T9, medial margin almost reaching medial line, bearing gonostylus on apical 1/3. Gonostylus with two lobes, basolateral lobe shortest and thin, apicomedial lobe broad based and 2X length of basal but tapering to point, bearing setae along medial margin, gonocoxite III fused to dorsolateral margin. Aedeagus 3/4 length of gonostylus, tapering towards apex (slight central swelling), apodemes not observable in specimen. Parameres strongly hooked at apex with apical margin serrated, apodemes 2/5 length of parameres.
|
||
</paragraph>
|
||
<paragraph id="422236DE627B9D08EE9CFF09FE47EAAE" blockId="26.[151,1436,151,500]" box="[199,414,151,176]" pageId="26" pageNumber="27">
|
||
<emphasis id="70E9EACC627B9D08EE9CFF09FEC2EAB1" box="[199,283,152,176]" italics="true" pageId="26" pageNumber="27">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="422236DE627B9D08EE9CFF2CFE1BEAD5" blockId="26.[151,1436,151,500]" box="[199,450,188,212]" pageId="26" pageNumber="27">
|
||
<emphasis id="70E9EACC627B9D08EE9CFF2CFEE7EAD5" box="[199,318,189,212]" italics="true" pageId="26" pageNumber="27">Immatures</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555627B9D08EE9CFF4EFE16EAF6" box="[199,463,223,248]" pageId="26" pageNumber="27" type="biology_ecology">
|
||
<paragraph id="422236DE627B9D08EE9CFF4EFE16EAF6" blockId="26.[151,1436,151,500]" box="[199,463,223,248]" pageId="26" pageNumber="27">
|
||
<emphasis id="70E9EACC627B9D08EE9CFF4EFE93EAF9" bold="true" box="[199,330,223,248]" pageId="26" pageNumber="27">BIOLOGY</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555627B9D08EE9CFE92FCABEB1D" box="[199,882,259,284]" pageId="26" pageNumber="27" type="distribution">
|
||
<paragraph id="422236DE627B9D08EE9CFE92FCABEB1D" blockId="26.[151,1436,151,500]" box="[199,882,259,284]" pageId="26" pageNumber="27">
|
||
<emphasis id="70E9EACC627B9D08EE9CFE92FE4FEB1D" bold="true" box="[199,406,259,284]" pageId="26" pageNumber="27">DISTRIBUTION</emphasis>
|
||
: Northern
|
||
<collectingCountry id="3A8A764E627B9D08EC4AFE95FD59EB1D" box="[529,640,260,284]" name="Argentina" pageId="26" pageNumber="27">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="DAA62A5B627B9D08ECD4FE95FD29EB1D" box="[655,752,260,284]" captionStart="FIGURE 142–143" captionStartId="110.[151,249,1784,1807]" captionTargetBox="[151,1436,193,1763]" captionTargetId="figure-57@110.[151,1436,193,1763]" captionTargetPageId="110" captionText="FIGURE 142–143. Distribution of Neotropical Leptomorphus. 142. L. amorimi (blue circle), L. crassipilus (red square), L. fasciculatus (black triangle), L. neivai (red star). 143. L. brandiae (purple triangle), L. eberhardi (blue circle), L. femoratus (red star), L. waodani (black square)." pageId="26" pageNumber="27">Fig. 142</figureCitation>
|
||
),
|
||
<quantity id="85659B3B627B9D08ED5EFE95FCB7EB1D" box="[773,878,260,284]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="26" pageNumber="27" unit="m" value="980.0">980 masl</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555627B9D08EE9CFEB6FD2FEB62" pageId="26" pageNumber="27" type="etymology">
|
||
<paragraph id="422236DE627B9D08EE9CFEB6FD2FEB62" blockId="26.[151,1436,151,500]" pageId="26" pageNumber="27">
|
||
<emphasis id="70E9EACC627B9D08EE9CFEB6FE58EB41" bold="true" box="[199,385,295,320]" pageId="26" pageNumber="27">ETYMOLOGY</emphasis>
|
||
: The species name refers to the rows of thick (
|
||
<emphasis id="70E9EACC627B9D08EDCCFEB9FC3AEB3E" box="[919,995,296,319]" italics="true" pageId="26" pageNumber="27">crassi-</emphasis>
|
||
) setae (
|
||
<emphasis id="70E9EACC627B9D08EA66FEB9FBA4EB41" box="[1085,1149,296,320]" italics="true" pageId="26" pageNumber="27">-pilus</emphasis>
|
||
) on the apex of tergite 9, which is a unique characteristic within
|
||
<taxonomicName id="859D4D5D627B9D08EC16FEDDFD36EB65" authorityName=": Edwards" authorityYear="1925" box="[589,751,332,356]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70E9EACC627B9D08EC16FEDDFD36EB65" box="[589,751,332,356]" italics="true" pageId="26" pageNumber="27">Leptomorphus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A876555627B9D08EE9CFEFEFE93EBF2" pageId="26" pageNumber="27" type="materials_examined">
|
||
<paragraph id="422236DE627B9D08EE9CFEFEFE93EBF2" blockId="26.[151,1436,151,500]" pageId="26" pageNumber="27">
|
||
<materialsCitation id="F2F53C83627B9D08EE9CFEFEFE93EBF2" collectionCode="MATERIAL, MNHN" collectorName="S. Javier & Col. Duret & Det. C. J. Borkent" country="Argentina" determinerName="Det. C. J. Borkent" latitude="-26.783" location="Tucuman" longLatPrecision="74" longitude="-65.383" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman" typeStatus="holotype">
|
||
<emphasis id="70E9EACC627B9D08EE9CFEFEFDD8EB89" bold="true" box="[199,513,367,392]" pageId="26" pageNumber="27">
|
||
<collectionCode id="248CAE1B627B9D08EE9CFEFEFE86EB89" box="[199,351,367,392]" pageId="26" pageNumber="27">MATERIAL</collectionCode>
|
||
EXAMINED
|
||
</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC627B9D08EC4BFEE1FD59EB89" box="[528,640,368,392]" italics="true" pageId="26" pageNumber="27">
|
||
<typeStatus id="9D26887C627B9D08EC4BFEE1FDA3EB89" box="[528,634,368,392]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
here designated, adult male, pinned, labelled “
|
||
<collectingCountry id="3A8A764E627B9D08EAD6FEFEFB26EB86" box="[1165,1279,367,391]" name="Argentina" pageId="26" pageNumber="27">Argentina</collectingCountry>
|
||
/
|
||
<collectingRegion id="8059F83C627B9D08EB56FEFEFAACEB86" box="[1293,1397,367,391]" country="Argentina" name="Tucuman" pageId="26" pageNumber="27">Tucuman</collectingRegion>
|
||
/
|
||
<collectorName id="EF685308627B9D08EBDEFEFEFF03EBAD" pageId="26" pageNumber="27">S. Javier</collectorName>
|
||
[
|
||
<geoCoordinate id="27A95019627B9D08EEB2FE05FE8CEBAD" box="[233,341,404,428]" degrees="26.783" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="55" value="-26.783">26.783°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="27A95019627B9D08EF38FE05FE0EEBAD" box="[355,471,404,428]" degrees="65.383" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="55" value="-65.383">65.383°W</geoCoordinate>
|
||
]/
|
||
<collectorName id="EF685308627B9D08EFB4FE05FDBFEBAD" box="[495,614,404,428]" pageId="26" pageNumber="27">Col. Duret</collectorName>
|
||
/ [vertically on margin] IV 44; [pink label with genitalia in between two coverslips attached]; 20779;
|
||
<typeStatus id="9D26887C627B9D08EF87FE26FDB6EBCE" box="[476,623,439,463]" pageId="26" pageNumber="27" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="549BFD57627B9D08EC22FE29FD57EBCE" box="[633,654,440,463]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
|
||
/
|
||
<taxonomicName id="859D4D5D627B9D08ECC7FE29FC66EBD1" authorityName="Borkent" authorityYear="2012" box="[668,959,440,464]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="crassipilus">
|
||
<emphasis id="70E9EACC627B9D08ECC7FE29FC66EBD1" box="[668,959,440,464]" italics="true" pageId="26" pageNumber="27">Leptomorphus crassipilus</emphasis>
|
||
</taxonomicName>
|
||
/ Borkent,
|
||
<taxonomicNameLabel id="6BDA57B7627B9D08EA63FE29FB19EBCE" box="[1080,1216,439,463]" pageId="26" pageNumber="27" rank="species">new species</taxonomicNameLabel>
|
||
/
|
||
<determinerName id="B7FEDA1A627B9D08EA8CFE29FA42EBCE" box="[1239,1435,439,463]" pageId="26" pageNumber="27">
|
||
<collectorName id="EF685308627B9D08EA8CFE29FA42EBCE" box="[1239,1435,439,463]" pageId="26" pageNumber="27">Det. C.J. Borkent</collectorName>
|
||
</determinerName>
|
||
2012” [
|
||
<collectionCode id="248CAE1B627B9D08EEB3FE4DFE9AEBF2" box="[232,323,476,499]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="27">MNHN</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |