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14. 
<taxonomicName id="859D4D5D62419D32EE98FCCAFDAFE975" ID-CoL="8CJ3R" authority="Landrock" authorityName="Landrock" box="[195,630,858,885]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="forcipatus">
<emphasis id="70E9EACC62419D32EE98FCCAFE2DE974" bold="true" box="[195,500,859,885]" italics="true" pageId="32" pageNumber="33">Leptomorphus forcipatus</emphasis>
Landrock
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<subSubSection id="0A87655562419D32EECCFCEFFD8DE997" box="[151,596,894,918]" pageId="32" pageNumber="33" type="description">
<paragraph id="422236DE62419D32EECCFCEFFD8DE997" blockId="32.[151,630,858,918]" box="[151,596,894,918]" pageId="32" pageNumber="33">
(
<figureCitation id="DAA62A5B62419D32EEC5FCEFFEC0E997" box="[158,281,894,918]" captionStart="FIGURES 16–19" captionStartId="86.[151,263,1697,1720]" captionTargetBox="[153,1435,194,1675]" captionTargetId="figure-40@86.[151,1436,193,1676]" captionTargetPageId="86" captionText="FIGURES 16–19. Leptomorphus habitus.16. L. forcipatus, male. 17. L. furcatus, paratype male. 18. L. gracilis, holotype male. 19. L. grjebinei, holotype male." pageId="32" pageNumber="33">Figures 16</figureCitation>
, 
<figureCitation id="DAA62A5B62419D32EF7EFCEFFE9AE997" box="[293,323,894,918]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="32" pageNumber="33">55</figureCitation>
, 
<figureCitation id="DAA62A5B62419D32EF14FCEFFEB4E997" box="[335,365,894,918]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="32" pageNumber="33">85</figureCitation>
, 
<figureCitation id="DAA62A5B62419D32EF22FCEFFE7CE997" box="[377,421,894,918]" captionStart="FIGURES 103–106" captionStartId="100.[151,264,1903,1926]" captionTargetBox="[199,1395,189,1831]" captionTargetId="figure-90@100.[168,1453,169,1891]" captionTargetPageId="100" captionText="FIGURES 103–106. Leptomorphus male genitalia, ventral. 103. L. forcipatus. 104. L. furcatus. 105. L. grjebinei. 106. L. gracilis. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="32" pageNumber="33">103</figureCitation>
, 
<figureCitation id="DAA62A5B62419D32EFEAFCEFFE04E997" box="[433,477,894,918]" captionStart="FIGURE 140–141" captionStartId="109.[151,249,1594,1617]" captionTargetBox="[151,1436,193,1573]" captionTargetId="figure-53@109.[151,1436,193,1573]" captionTargetPageId="109" captionText="FIGURE 140–141. Distribution of Palaearctic Leptomorphus. 140. L. forcipatus (black square), L. subforcipatus (blue triangle), L. talyshensis (black circle), L. walkeri (red star). 141. L. babai (blue triangle), L. panorpiformis (red star), L. quadrimaculatus (black square)." pageId="32" pageNumber="33">140</figureCitation>
, 
<figureCitation id="DAA62A5B62419D32EFB2FCEFFDCCE997" box="[489,533,894,918]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values&gt;50% are shown in bold below the branches." pageId="32" pageNumber="33">150</figureCitation>
, 
<figureCitation id="DAA62A5B62419D32EC7AFCEFFD95E997" box="[545,588,894,918]" captionStart="FIGURE 154" captionStartId="115.[151,250,1776,1799]" captionTargetBox="[151,1431,1079,1755]" captionTargetId="figure-103@115.[151,1436,1055,1790]" captionTargetPageId="115" captionText="FIGURE 154. One of three equally parsimonious trees showing the relationships between species in the L. walkeri species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="32" pageNumber="33">154</figureCitation>
)
</paragraph>
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<paragraph id="422236DE62419D32EECCFC57FD67E9DA" blockId="32.[151,1150,965,1049]" box="[151,702,965,988]" pageId="32" pageNumber="33">
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<treatmentCitation id="C33C10CF62419D32EECCFC57FD62E9DA" author="Landrock, K." box="[151,699,965,988]" page="107" pageId="32" pageNumber="33" year="1918">
<taxonomicName id="859D4D5D62419D32EECCFC57FD62E9DA" authority="Landrock, 1918: 107" authorityName="Landrock" authorityPageNumber="107" authorityYear="1918" box="[151,699,965,988]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="walkeri" subSpecies="forcipata">
<emphasis id="70E9EACC62419D32EECCFC57FE07E9DD" box="[151,478,966,988]" italics="true" pageId="32" pageNumber="33">Leptomorphus walkeri forcipata</emphasis>
<bibRefCitation id="260C4B2F62419D32EFBFFC54FD62E9DA" author="Landrock, K." box="[484,699,965,987]" pageId="32" pageNumber="33" pagination="107 - 120" refId="ref94305" refString="Landrock, K. (1918) Tabellen zum Bestimmen europascher Pilzmucken. II. Wiener Entomologische Zeitung, 37, 107 - 120." type="journal article" year="1918">Landrock, 1918: 107</bibRefCitation>
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.
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<treatmentCitationGroup id="628D11F062419D32EECCFC75FD94E9FB" box="[151,589,996,1018]" pageId="32" pageNumber="33">
<treatmentCitation id="C33C10CF62419D32EECCFC75FD90E9FB" author="Seguy, E." box="[151,585,996,1018]" page="86" pageId="32" pageNumber="33" year="1940">
<taxonomicName id="859D4D5D62419D32EECCFC75FD90E9FB" ID-CoL="8CJ3R" authority=": Seguy, 1940: 86" authorityName="Seguy" authorityPageNumber="86" authorityYear="1940" box="[151,585,996,1018]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="forcipatus">
<emphasis id="70E9EACC62419D32EECCFC75FE4CE9FB" box="[151,405,996,1018]" italics="true" pageId="32" pageNumber="33">Leptomorphus forcipatus</emphasis>
: 
<bibRefCitation id="260C4B2F62419D32EFF8FC75FD90E9FB" author="Seguy, E." box="[419,585,996,1018]" pageId="32" pageNumber="33" pagination="1 - 386" refId="ref95770" refString="Seguy, E. (1940) Dipteres nematoceres (Fungivoridae, Lycoriidae, Hesperinidae, Bibionidae, Scatopsidae, Phrynidae, Pachyneuridae, Blepharoceridae). Faune de France, 36, 1 - 386." type="journal article" year="1940">Séguy, 1940: 86</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="422236DE62419D32EECCFB92FBA7EE18" blockId="32.[151,1150,965,1049]" box="[151,1150,1027,1049]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="628D11F062419D32EECCFB92FBA7EE18" box="[151,1150,1027,1049]" pageId="32" pageNumber="33">
<treatmentCitation id="C33C10CF62419D32EECCFB92FCD8EE18" author="Matile, L." box="[151,769,1027,1049]" page="144" pageId="32" pageNumber="33" year="1977">
<taxonomicName id="859D4D5D62419D32EECCFB92FCD8EE18" ID-CoL="8CJ3R" authority=": Matile, 1977: 144" authorityName=": Matile" authorityPageNumber="144" authorityYear="1977" box="[151,769,1027,1049]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="forcipatus" subGenus="Leptomorphus">
<emphasis id="70E9EACC62419D32EECCFB92FDE6EE18" box="[151,575,1027,1049]" italics="true" pageId="32" pageNumber="33">Leptomorphus (Leptomorphus) forcipatus</emphasis>
: 
<bibRefCitation id="260C4B2F62419D32EC17FB92FCD8EE18" author="Matile, L." box="[588,769,1027,1049]" pageId="32" pageNumber="33" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile, 1977: 144</bibRefCitation>
</taxonomicName>
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(as synonym of 
<taxonomicName id="859D4D5D62419D32EDF2FB95FBB6EE18" ID-CoL="8CJ3T" baseAuthorityName="Matsumura" baseAuthorityYear="1916" box="[937,1135,1027,1049]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="70E9EACC62419D32EDF2FB95FBB6EE18" box="[937,1135,1027,1049]" italics="true" pageId="32" pageNumber="33">L. quadrimaculatus</emphasis>
</taxonomicName>
).
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</paragraph>
<paragraph id="422236DE62419D32EECCFBD4FBF9EF10" blockId="32.[151,1437,1093,2018]" pageId="32" pageNumber="33">
<emphasis id="70E9EACC62419D32EECCFBD4FEC0EE5F" bold="true" box="[151,281,1093,1118]" pageId="32" pageNumber="33">References</emphasis>
: 
<bibRefCitation id="260C4B2F62419D32EF77FBD7FE2FEE5F" author="Okada, I." box="[300,502,1094,1118]" pageId="32" pageNumber="33" pagination="99 - 103" refId="ref94924" refString="Okada, I. (1936) Einige Nematocera aus den Sud-Kurilen (Diptera). Insecta Matsumurana, 10, 99 - 103." type="journal article" year="1936">Okada 1936: 100</bibRefCitation>
(synonymized with 
<taxonomicName id="859D4D5D62419D32ECB1FBD6FC13EE5F" ID-CoL="8CJ3T" baseAuthorityName="Matsumura" baseAuthorityYear="1916" box="[746,970,1094,1118]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="70E9EACC62419D32ECB1FBD6FC13EE5F" box="[746,970,1094,1118]" italics="true" pageId="32" pageNumber="33">L. quadrimaculatus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="260C4B2F62419D32EDBCFBD7FB79EE5F" author="Seguy, E." box="[999,1184,1094,1118]" pageId="32" pageNumber="33" pagination="1 - 386" refId="ref95770" refString="Seguy, E. (1940) Dipteres nematoceres (Fungivoridae, Lycoriidae, Hesperinidae, Bibionidae, Scatopsidae, Phrynidae, Pachyneuridae, Blepharoceridae). Faune de France, 36, 1 - 386." type="journal article" year="1940">Séguy 1940: 86</bibRefCitation>
(distribution); 
<bibRefCitation id="260C4B2F62419D32EB08FBD7FEDEEE80" author="Matile, L." pageId="32" pageNumber="33" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977: 144</bibRefCitation>
(subgeneric placement), 1988: 234 (catalogue); 
<bibRefCitation id="260C4B2F62419D32ED7EFBF8FB2EEE80" author="Ostroverkhova, G. P. &amp; Shtakel'berg A. A." box="[805,1271,1129,1153]" pageId="32" pageNumber="33" pagination="404 - 487" refId="ref95062" refString="Ostroverkhova, G. P. &amp; Shtakel'berg A. A. (1988) 24. Family Mycetophilidae (Fungivoridae). In: Bei-Bienko G., (Ed.) Keys to the Insects of the European Part of the USSR. Volume 5: Diptera and Siphonaptera Part 1. Smithsonian Institution Libraries &amp; National Research Foundation, Washington D. C., pp. 404 - 487." type="book chapter" year="1988">Ostroverkhova &amp; Shtakel’berg 1988: 416</bibRefCitation>
, 418 (genitalia figure and key reference); 
<bibRefCitation id="260C4B2F62419D32EF9BFB1FFD57EEA7" author="Zaitzev, A. I." box="[448,654,1166,1190]" pageId="32" pageNumber="33" refId="ref96742" refString="Zaitzev, A. I. (1994) Fungus gnats of the fauna of Russia and adjacent regions. Part 1. Nauka, Moscow, 288 pp. [in Russian]" type="book" year="1994">Zaitzev 1994: 157</bibRefCitation>
, 161 (key, re-description, male genitalia figure); 
<bibRefCitation id="260C4B2F62419D32EAE8FB1FFA4EEEA7" author="Yakovlev, E. B." box="[1203,1431,1166,1190]" pageId="32" pageNumber="33" pagination="335 - 362" refId="ref96606" refString="Yakovlev, E. B. (1995) Species diversity and abundance of fungivorous Diptera in forests and city parks of Russian Karelia. An International Journal of Dipterological Research, 6, 335 - 362." type="journal article" year="1995">Yakovlev 1995: 351</bibRefCitation>
, 356 (rearing record); 
<bibRefCitation id="260C4B2F62419D32EFDDFB20FD00EEC8" author="Zaitzev, A. I. &amp; Sevcik J." box="[390,729,1201,1226]" pageId="32" pageNumber="33" pagination="203 - 211" refId="ref96838" refString="Zaitzev, A. I. &amp; Sevcik J. (2002) A review of the Palaearctic species of the Leptomorphus quadrimaculatus (Matsumura) group (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientiarum Hungaricae, 48, 203 - 211." type="journal article" year="2002">Zaitzev and Ševčík, 2002: 204</bibRefCitation>
(removed from synonymy with 
<taxonomicName id="859D4D5D62419D32EA1AFB23FAC5EECB" ID-CoL="8CJ3T" baseAuthorityName="Matsumura" baseAuthorityYear="1916" box="[1089,1308,1202,1226]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="70E9EACC62419D32EA1AFB23FAC5EECB" box="[1089,1308,1202,1226]" italics="true" pageId="32" pageNumber="33">L. quadrimaculatus</emphasis>
</taxonomicName>
); 
<bibRefCitation id="260C4B2F62419D32EB6FFB20FE91EEEF" author="Sevcik, J. &amp; Papp, L." pageId="32" pageNumber="33" pagination="285 - 295" refId="ref95873" refString="Sevcik, J. &amp; Papp, L. (2003) New Mycetophilidae (Diptera) and additions to the Hungarian checklist. Folia Entomologica Hungarica, 64, 285 - 295." type="journal article" year="2003">Ševčík &amp; Papp 2003: 288</bibRefCitation>
(
<collectingCountry id="3A8A764E62419D32EF03FB47FE19EEEF" box="[344,448,1238,1262]" name="Hungary" pageId="32" pageNumber="33">Hungary</collectingCountry>
); 
<bibRefCitation id="260C4B2F62419D32EF8EFB47FCC3EEEF" author="Gammelmo, O. &amp; Soli, G." box="[469,794,1238,1262]" pageId="32" pageNumber="33" pagination="57 - 69" refId="ref93311" refString="Gammelmo, O. &amp; Soli, G. (2006) Norwegian fungus gnats of the family Mycetophilidae (Diptera, Nematocera). Norwegian Journal of Entomology, 53, 57 - 69." type="journal article" year="2006">Gammelmo &amp; Søli 2006: 60</bibRefCitation>
, (
<collectingCountry id="3A8A764E62419D32ED74FB47FC56EEEF" box="[815,911,1238,1262]" name="Norway" pageId="32" pageNumber="33">Norway</collectingCountry>
); 
<bibRefCitation id="260C4B2F62419D32EDFFFB47FB82EEEF" author="Sevcik, J." box="[932,1115,1238,1262]" pageId="32" pageNumber="33" pagination="1 - 84" refId="ref95807" refString="Sevcik, J. (2006) Diptera associated with fungi in the Czech and Slovak Republics. C asopis Slezskeho Zemskeho Muzea Serie A V e dy P r irodni, 55 (supplement 2), 1 - 84." type="journal article" year="2006">Ševčík 2006: 14</bibRefCitation>
(biology, photo of adults en copula), 2010: 17 (fungal association); 
<bibRefCitation id="260C4B2F62419D32EC14FB68FCA4EF10" author="Kjaerandsen, J. &amp; Hedmark, K. &amp; Kurina, O. &amp; Polevoi, A. &amp; Okland, B. &amp; Gotmark, F." box="[591,893,1273,1298]" pageId="32" pageNumber="33" pagination="1 - 128" refId="ref93903" refString="Kjaerandsen, J., Hedmark, K., Kurina, O., Polevoi, A., Okland, B. &amp; Gotmark, F. (2007) Annotated checklist of fungus gnats from Sweden (Diptera: Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae and Mycetophilidae). Insect Systematics and Evolution, Supplement, 65, 1 - 128." type="journal article" year="2007">
Kjaerandsen 
<emphasis id="70E9EACC62419D32EC87FB6BFCCDEF13" box="[732,788,1274,1298]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2007: 35
</bibRefCitation>
(distribution).
</paragraph>
</subSubSection>
<subSubSection id="0A87655562419D32EE9CFA8CFDFDEC57" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="422236DE62419D32EE9CFA8CFDF6EFA0" blockId="32.[151,1437,1093,2018]" pageId="32" pageNumber="33">
<emphasis id="70E9EACC62419D32EE9CFA8CFEBDEF37" bold="true" box="[199,356,1309,1334]" pageId="32" pageNumber="33">DIAGNOSIS</emphasis>
: The only extant species of 
<taxonomicName id="859D4D5D62419D32EC92FA8FFCB2EF37" authorityName=": Edwards" authorityYear="1925" box="[713,875,1310,1334]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="70E9EACC62419D32EC92FA8FFCB2EF37" box="[713,875,1310,1334]" italics="true" pageId="32" pageNumber="33">Leptomorphus</emphasis>
</taxonomicName>
with the following combination of characters: foretibia with dense row (comb) of short anteroventral bristles; wing with dark apical and medial spots present; male genitalia with gonocoxite smoothly curved along entire length, not swollen at apex and without triangular medial protrusion at base (
<figureCitation id="DAA62A5B62419D32EF9BFA18FDFBEFA0" box="[448,546,1417,1441]" captionStart="FIGURES 103–106" captionStartId="100.[151,264,1903,1926]" captionTargetBox="[199,1395,189,1831]" captionTargetId="figure-90@100.[168,1453,169,1891]" captionTargetPageId="100" captionText="FIGURES 103–106. Leptomorphus male genitalia, ventral. 103. L. forcipatus. 104. L. furcatus. 105. L. grjebinei. 106. L. gracilis. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="32" pageNumber="33">Fig. 103</figureCitation>
).
</paragraph>
<paragraph id="422236DE62419D32EE9CFA3FFDFDEC57" blockId="32.[151,1437,1093,2018]" pageId="32" pageNumber="33">
This species can be distinguished from other Palaearctic species by the long gonocoxites (more than half total length of male genitalia, 
<figureCitation id="DAA62A5B62419D32EFEAFA40FDCBEFE8" box="[433,530,1489,1513]" captionStart="FIGURES 103–106" captionStartId="100.[151,264,1903,1926]" captionTargetBox="[199,1395,189,1831]" captionTargetId="figure-90@100.[168,1453,169,1891]" captionTargetPageId="100" captionText="FIGURES 103–106. Leptomorphus male genitalia, ventral. 103. L. forcipatus. 104. L. furcatus. 105. L. grjebinei. 106. L. gracilis. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="32" pageNumber="33">Fig. 103</figureCitation>
), almost completely dark yellow abdomen (tergite 7 and posterior half of tergite 6, brown, 
<figureCitation id="DAA62A5B62419D32EF5CFA67FE8EEC0F" box="[263,343,1526,1550]" captionStart="FIGURES 16–19" captionStartId="86.[151,263,1697,1720]" captionTargetBox="[153,1435,194,1675]" captionTargetId="figure-40@86.[151,1436,193,1676]" captionTargetPageId="86" captionText="FIGURES 16–19. Leptomorphus habitus.16. L. forcipatus, male. 17. L. furcatus, paratype male. 18. L. gracilis, holotype male. 19. L. grjebinei, holotype male." pageId="32" pageNumber="33">Fig. 16</figureCitation>
) and scutum evenly covered with small setae. It is the only species with long gonocoxites that has the gonocoxite both smoothly curved (not strongly bent as in 
<emphasis id="70E9EACC62419D32ED3DF98BFBD1EC33" box="[870,1032,1562,1586]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="859D4D5D62419D32ED3DF98BFBDCEC33" box="[870,1029,1562,1586]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="talyshensis">L. talyshensis</taxonomicName>
,
</emphasis>
<figureCitation id="DAA62A5B62419D32EA48F988FBADEC30" box="[1043,1140,1561,1585]" captionStart="FIGURES 121–124" captionStartId="105.[151,264,1775,1798]" captionTargetBox="[176,1407,186,1696]" captionTargetId="figure-118@105.[161,1446,161,1741]" captionTargetPageId="105" captionText="FIGURES 121–124. Leptomorphus male genitalia. 121. L. talyshensis (reprinted with permission from Zaitzev &amp; Ševčík 2002), A. gonocoxite and gonostylus, ventral. B. dorsal. 122. L. titiwangsensis, ventral. 123. L. walkeri, ventral. 124. L. waodani, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="32" pageNumber="33">Fig. 121</figureCitation>
) and without a triangular protrusion mediobasally (
<figureCitation id="DAA62A5B62419D32EFEDF9AFFDCEEC57" box="[438,535,1598,1622]" captionStart="FIGURES 103–106" captionStartId="100.[151,264,1903,1926]" captionTargetBox="[199,1395,189,1831]" captionTargetId="figure-90@100.[168,1453,169,1891]" captionTargetPageId="100" captionText="FIGURES 103–106. Leptomorphus male genitalia, ventral. 103. L. forcipatus. 104. L. furcatus. 105. L. grjebinei. 106. L. gracilis. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="32" pageNumber="33">Fig. 103</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0A87655562419D33EE9CF9F0FE3AEC12" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="422236DE62419D33EE9CF9F0FD65EFA6" blockId="32.[151,1437,1093,2018]" lastBlockId="33.[151,1437,151,2024]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="70E9EACC62419D32EE9CF9F0FE50EC7B" bold="true" box="[199,393,1633,1658]" pageId="32" pageNumber="33">DESCRIPTION</emphasis>
: 
<emphasis id="70E9EACC62419D32EFFFF9F3FE04EC7B" box="[420,477,1634,1658]" italics="true" pageId="32" pageNumber="33">Male</emphasis>
. (
<figureCitation id="DAA62A5B62419D32EC5BF9F0FD84EC78" box="[512,605,1633,1657]" captionStart="FIGURES 16–19" captionStartId="86.[151,263,1697,1720]" captionTargetBox="[153,1435,194,1675]" captionTargetId="figure-40@86.[151,1436,193,1676]" captionTargetPageId="86" captionText="FIGURES 16–19. Leptomorphus habitus.16. L. forcipatus, male. 17. L. furcatus, paratype male. 18. L. gracilis, holotype male. 19. L. grjebinei, holotype male." pageId="32" pageNumber="33">Fig. 16</figureCitation>
) 
<emphasis id="70E9EACC62419D32EC23F9F3FD6EEC7B" bold="true" box="[632,695,1634,1658]" pageId="32" pageNumber="33">Head</emphasis>
: brown dorsally yellow ventrally, somewhat dorsoventrally compressed in anterior view. Antenna brown; scape yellow, with light brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae present; pedicel light brown/yellow, with 3 large bristles, several setae on apicodorsal margin, a few fine setae on apicoventral margin; flagellomere 1 with tapered base brown remainder brown; flagellomere 6 2.0X as long as broad. Clypeus light brown, strongly protruding, slightly laterally compressed oval; bristles on clypeus light brown, both strong, smaller bristles on entire surface, all directed ventrally, clypeus 2X as long as face. Face brown; shape a slightly wider than tall triangle, with fine bristles covering face. Frons brown; with few bristles medioventrally, frontal furrow running 3/4–full distance from dorsal margin towards ventral margin, frontal cleft just anterior of median ocellus. Palpus yellow; segment 1 small but visible below eye, segments increasing in length, segment 5 1.5X length of segment 4 with even width from base to apex, segment 3 with apicolateral patch of fine yellow setae encircled by strong dark setae. Labellum yellow. Eye with long interommatidial setulae in ~1/4 of the ommatidial junctions, scattered on all but medial margin. Occiput brown with appressed, anteriorly directed setae. Ocelli with median slightly in front of laterals, space between ocelli less than diameter of laterals, lateral ocelli 1.5X their own diameter from eye margin, ocellar triangle dark brown. 
<emphasis id="70E9EACC62409D33EB67FE95FA4DEB1D" bold="true" box="[1340,1428,260,284]" pageId="33" pageNumber="34">Thorax</emphasis>
: Length 
<quantity id="85659B3B62409D33EEAAFEB6FE81EB3E" box="[241,344,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="33" pageNumber="34" unit="mm" value="1.54">1.54 mm</quantity>
(
<quantity id="85659B3B62409D33EF31FEB6FDCDEB3E" box="[362,532,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="1.515" metricValueMax="1.64" metricValueMin="1.39" pageId="33" pageNumber="34" unit="mm" value="1.515" valueMax="1.64" valueMin="1.39">1.39–1.64 mm</quantity>
, n = 3). Dorsally brown, laterally yellow. Scutum brown on disc with lighter dorsocentral lines, yellow on anterior corners, lateral and posterior margins except for dark spot at wing base; surface of scutum covered with small setae; acrostichal setae absent; dorsocentral setae probably present but not clearly distinguishable from other setae; multiple rows of lateral setae present; patch of setae on scutum at wing base small. Scutellum yellow to light brown; with 6–8 large bristles and many small bristles. Prescutum brown. Mediotergite yellow, darker posteriorly with 8–16 bristles on posterolateral corners, small bristles covering. Laterotergite yellow; anterior margin of laterotergite abutting katepisternum. Anepimeron yellow. Anepisternum yellow. Katepisternum yellow with anteroventral corner brown. Antepronotum and proepisternum light brown to yellow. Margin of anterior and posterior spiracles yellow with yellow trichia. Metepisternum yellow. Anapleural suture straight and clear. Halter stem yellow, knob light brown. 
<emphasis id="70E9EACC62409D33EDD9FDFDFC63E885" bold="true" box="[898,954,620,644]" pageId="33" pageNumber="34">Legs</emphasis>
: principally yellow; extreme anteroapical corner dark brown on all femora; foretarsi brown. Midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia with weak comb of short setae along length of anteroventral surface (
<figureCitation id="DAA62A5B62409D33EB63FD25FA57E8CD" box="[1336,1422,692,716]" captionStart="FIGURES 82–90" captionStartId="96.[151,263,1444,1467]" captionTargetBox="[151,1436,211,1413]" captionTargetId="figure-84@96.[151,1436,211,1413]" captionTargetPageId="96" captionText="FIGURES 82–90. Leptomorphus morphology. 82. Head of L. brandiae, anterolateral, showing swollen palpus segment 5 covered in fine white trichia. 83–84. Male midfemur, anterior, showing apicoventral process (arrow). 83. L. nebulosus. 84. L. subcaeruleus. 85–90. Male foretibia, posterior, showing presence (arrow) or absence of dense comb of ventral bristles. 85. L. forcipatus. 86. L. magnificus. 87. L. mandelai. 88. L. tagbanua. 89. L. walkeri. 90. L. waodani." pageId="33" pageNumber="34">Fig. 85</figureCitation>
), tibial spurs yellow, foretibial spur length 2X apical thickness of foretibia, midtibia with strong, dorsal, bare patch of even thickness for 3/4 of its length, placed basally, shortest midtibial spur 0.7X length of longest, longest midtibial spur 4.5X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 4.5X apical thickness of hind tibia. Foreleg first tarsomere 1.7X length of foretibia. 
<emphasis id="70E9EACC62409D33EAD4FCD2FB09E95D" bold="true" box="[1167,1232,835,860]" pageId="33" pageNumber="34">Wing</emphasis>
(
<figureCitation id="DAA62A5B62409D33EABAFCD5FAEEE95D" box="[1249,1335,836,860]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="33" pageNumber="34">Fig. 55</figureCitation>
): Length 
<quantity id="85659B3B62409D33EECCFCF6FF35E97E" box="[151,236,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="33" pageNumber="34" unit="mm" value="7.2">7.2 mm</quantity>
(
<quantity id="85659B3B62409D33EEA7FCF6FE5EE97E" box="[252,391,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.5" metricValueMin="6.6" pageId="33" pageNumber="34" unit="mm" value="7.05" valueMax="7.5" valueMin="6.6">6.6–7.5 mm</quantity>
, n = 3). Hyaline; apical macula not reaching wing tip, extending faintly along posterior wing margin into apex of cell cua 
<subScript id="DE19349B62409D33EF8AFC0AFE00E9A8" attach="left" box="[465,473,923,937]" fontSize="6" pageId="33" pageNumber="34">1</subScript>
and cup though not joining with medial macula; medial macula extending from Sc to stem of M 
<subScript id="DE19349B62409D33EF57FC53FEFDE9D1" attach="left" box="[268,292,962,976]" fontSize="6" pageId="33" pageNumber="34">1+2</subScript>
. Macrotrichia in all cells. Setae on basal posterior margin of wing (along base of cell a) all the same length. Calypter with a few short setae. Vein sc-r present, apical end joining R within its own length prior to origin of Rs. R 
<subScript id="DE19349B62409D33EEA0FB9CFEDAEE1A" attach="left" box="[251,259,1037,1051]" fontSize="6" pageId="33" pageNumber="34">4</subScript>
absent. R 
<subScript id="DE19349B62409D33EF2FFB9CFEA5EE1A" attach="left" box="[372,380,1037,1051]" fontSize="6" pageId="33" pageNumber="34">5</subScript>
straight, slight posterior turn near tip. M 
<subScript id="DE19349B62409D33ED06FB9CFCBCEE1A" attach="left" box="[861,869,1037,1051]" fontSize="6" pageId="33" pageNumber="34">1</subScript>
reaching apex just before R 
<subScript id="DE19349B62409D33EAE9FB9CFB63EE1A" attach="left" box="[1202,1210,1037,1051]" fontSize="6" pageId="33" pageNumber="34">5</subScript>
, apices of M veins thinning towards wing margin. M 
<subScript id="DE19349B62409D33EC7CFBA2FDF6EE40" attach="left" box="[551,559,1075,1089]" fontSize="6" pageId="33" pageNumber="34">4</subScript>
-CuA fork arising well before origin of r-m. A 
<subScript id="DE19349B62409D33EA38FBA2FBB2EE40" attach="left" box="[1123,1131,1075,1089]" fontSize="6" pageId="33" pageNumber="34">2</subScript>
faintly present as crease. 
<emphasis id="70E9EACC62409D33EECCFBDBFED5EE62" bold="true" box="[151,268,1098,1123]" pageId="33" pageNumber="34">Abdomen</emphasis>
: Tergites T1–6 yellow, T6 brown posteriorly, T7 brown. Tergite 8 smaller than all other abdominal sclerites, without bristles. 
<emphasis id="70E9EACC62409D33EF93FBFFFDEEEE86" bold="true" box="[456,567,1134,1159]" pageId="33" pageNumber="34">Genitalia</emphasis>
(
<figureCitation id="DAA62A5B62409D33EC11FBFEFD69EE86" box="[586,688,1135,1159]" captionStart="FIGURES 103–106" captionStartId="100.[151,264,1903,1926]" captionTargetBox="[199,1395,189,1831]" captionTargetId="figure-90@100.[168,1453,169,1891]" captionTargetPageId="100" captionText="FIGURES 103–106. Leptomorphus male genitalia, ventral. 103. L. forcipatus. 104. L. furcatus. 105. L. grjebinei. 106. L. gracilis. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="33" pageNumber="34">Fig. 103</figureCitation>
): light yellow. Sternite 9 lightly sclerotized, posteriorly directed triangle 1/2 length of T9 but as wide as genitalia. Tergite 9 subcircular though apical margin somewhat flattened. Gonocoxite placed centrally on T9, much longer than length of T9, medial margin not reaching median line, bearing gonostylus basally. Gonostylus a single lobe tapering to a point apically, strongly bent at halfway point and with several setae, gonocoxite III associated with dorsal margin of gonostylus but not fused to it. Aedeagus 3/4 length of T9, tapering slightly towards apex for basal 3/4 and then strongly indented and remainder squarish, apodemes ~1/2 total length. Parameres consisting of two lobes, lateral lobe a broad based spine curving laterally, medial lobe 2X length of lateral and very thin, apodemes 1/2 length of parameres, strongly united with gonocoxal apodemes and with a dorsal pointing small hook.
</paragraph>
<paragraph id="422236DE62409D33EE9CFA22FE6DEFEE" blockId="33.[151,1437,151,2024]" pageId="33" pageNumber="34">
<emphasis id="70E9EACC62409D33EE9CFA22FEC2EFCA" box="[199,283,1459,1483]" italics="true" pageId="33" pageNumber="34">Female</emphasis>
. No specimens examined, but coloration apparently similar to male with female terminalia yellow (
<bibRefCitation id="260C4B2F62409D33EEC5FA46FE7EEFEE" author="Zaitzev, A. I. &amp; Sevcik J." box="[158,423,1495,1519]" pageId="33" pageNumber="34" pagination="203 - 211" refId="ref96838" refString="Zaitzev, A. I. &amp; Sevcik J. (2002) A review of the Palaearctic species of the Leptomorphus quadrimaculatus (Matsumura) group (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientiarum Hungaricae, 48, 203 - 211." type="journal article" year="2002">Zaitzev &amp; Ševčík 2002</bibRefCitation>
).
</paragraph>
<paragraph id="422236DE62409D33EE9CFA6DFE3AEC12" blockId="33.[151,1437,151,2024]" box="[199,483,1531,1555]" pageId="33" pageNumber="34">
<emphasis id="70E9EACC62409D33EE9CFA6DFE9EEC12" box="[199,327,1532,1555]" italics="true" pageId="33" pageNumber="34">Immatures:</emphasis>
Undescribed.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562409D33EE9CF98FFCD8EC7E" pageId="33" pageNumber="34" type="biology_ecology">
<paragraph id="422236DE62409D33EE9CF98FFCD8EC7E" blockId="33.[151,1437,151,2024]" pageId="33" pageNumber="34">
<emphasis id="70E9EACC62409D33EE9CF98FFE93EC36" bold="true" box="[199,330,1566,1591]" pageId="33" pageNumber="34">BIOLOGY</emphasis>
: Larvae are known to feed on the spores of 
<taxonomicName id="859D4D5D62409D33ED6FF98EFC57EC36" box="[820,910,1567,1591]" class="Agaricomycetes" family="Stereaceae" genus="Stereum" kingdom="Fungi" order="Russulales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="genus">
<emphasis id="70E9EACC62409D33ED6FF98EFC57EC36" box="[820,910,1567,1591]" italics="true" pageId="33" pageNumber="34">Stereum</emphasis>
</taxonomicName>
(
<taxonomicName id="859D4D5D62409D33EDC5F98EFBB1EC36" box="[926,1128,1567,1591]" class="Agaricomycetes" family="Stereaceae" genus="Stereum" kingdom="Fungi" order="Russulales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="subtomentosum">
<emphasis id="70E9EACC62409D33EDC5F98EFBB1EC36" box="[926,1128,1567,1591]" italics="true" pageId="33" pageNumber="34">S. subtomentosum</emphasis>
</taxonomicName>
and 
<taxonomicName id="859D4D5D62409D33EAC6F98EFAC5EC36" box="[1181,1308,1567,1591]" class="Agaricomycetes" family="Stereaceae" genus="Stereum" kingdom="Fungi" order="Russulales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="species" species="hirsutum">
<emphasis id="70E9EACC62409D33EAC6F98EFAC5EC36" box="[1181,1308,1567,1591]" italics="true" pageId="33" pageNumber="34">S. hirsutum</emphasis>
</taxonomicName>
; 
<bibRefCitation id="260C4B2F62409D33EB70F98EFEFEEC5A" author="Zaitzev, A. I. &amp; Sevcik J." pageId="33" pageNumber="34" pagination="203 - 211" refId="ref96838" refString="Zaitzev, A. I. &amp; Sevcik J. (2002) A review of the Palaearctic species of the Leptomorphus quadrimaculatus (Matsumura) group (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientiarum Hungaricae, 48, 203 - 211." type="journal article" year="2002">Zaitzev &amp; Ševčík 2002</bibRefCitation>
) and 
<taxonomicName id="859D4D5D62409D33EF30F9D2FE35EC5A" box="[363,492,1603,1627]" class="Agaricomycetes" family="Schizoporaceae" genus="Trichaptum" higherTaxonomySource="CoL" kingdom="Fungi" order="Hymenochaetales" pageId="33" pageNumber="34" phylum="Basidiomycota" rank="genus">
<emphasis id="70E9EACC62409D33EF30F9D2FE35EC5A" box="[363,492,1603,1627]" italics="true" pageId="33" pageNumber="34">Trichaptum</emphasis>
</taxonomicName>
(
<bibRefCitation id="260C4B2F62409D33EFA5F9D2FD77EC5A" author="Yakovlev, E. B." box="[510,686,1603,1627]" pageId="33" pageNumber="34" pagination="335 - 362" refId="ref96606" refString="Yakovlev, E. B. (1995) Species diversity and abundance of fungivorous Diptera in forests and city parks of Russian Karelia. An International Journal of Dipterological Research, 6, 335 - 362." type="journal article" year="1995">Yakovlev 1995</bibRefCitation>
, 
<bibRefCitation id="260C4B2F62409D33ECE6F9D2FCBFEC5A" author="Jakovlev, J." box="[701,870,1603,1627]" pageId="33" pageNumber="34" pagination="157 - 189" refId="ref93492" refString="Jakovlev, J. (2011) Fungus gnats (Diptera: Sciaroidea) associated with dead wood and wood growing fungi: new rearing data from Finland and Russian Karelia and general analysis of known larval microhabitats in Europe Entomologica Fennica, 22, 157 - 189." type="journal article" year="2011">Jakovlev 2011</bibRefCitation>
). Adults copulate soon after emergence from the pupa, and copulation lasts several hours (
<bibRefCitation id="260C4B2F62409D33EC3FF9F6FD2FEC7E" author="Sevcik, J." box="[612,758,1639,1663]" pageId="33" pageNumber="34" pagination="1 - 84" refId="ref95807" refString="Sevcik, J. (2006) Diptera associated with fungi in the Czech and Slovak Republics. C asopis Slezskeho Zemskeho Muzea Serie A V e dy P r irodni, 55 (supplement 2), 1 - 84." type="journal article" year="2006">Ševčík 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0A87655562409D33EE9CF91BFE75ECC6" pageId="33" pageNumber="34" type="distribution">
<paragraph id="422236DE62409D33EE9CF91BFE75ECC6" blockId="33.[151,1437,151,2024]" pageId="33" pageNumber="34">
<emphasis id="70E9EACC62409D33EE9CF91BFE4FECA2" bold="true" box="[199,406,1674,1699]" pageId="33" pageNumber="34">DISTRIBUTION</emphasis>
: 
<collectingCountry id="3A8A764E62409D33EFFEF91AFE24ECA2" box="[421,509,1675,1699]" name="Finland" pageId="33" pageNumber="34">Finland</collectingCountry>
, 
<collectingCountry id="3A8A764E62409D33EC52F91AFD64ECA2" box="[521,701,1675,1699]" name="Czech Republic" pageId="33" pageNumber="34">Czech Republic</collectingCountry>
, 
<collectingCountry id="3A8A764E62409D33EC91F91AFCF6ECA2" box="[714,815,1675,1699]" name="Slovakia" pageId="33" pageNumber="34">Slovakia</collectingCountry>
, 
<collectingCountry id="3A8A764E62409D33ED61F91AFC4CECA2" box="[826,917,1675,1699]" name="Sweden" pageId="33" pageNumber="34">Sweden</collectingCountry>
, 
<collectingCountry id="3A8A764E62409D33EDFAF91AFC25ECA2" box="[929,1020,1675,1699]" name="Norway" pageId="33" pageNumber="34">Norway</collectingCountry>
, 
<collectingCountry id="3A8A764E62409D33EA53F91AFBB2ECA2" box="[1032,1131,1675,1699]" name="Hungary" pageId="33" pageNumber="34">Hungary</collectingCountry>
, 
<collectingCountry id="3A8A764E62409D33EA2CF91AFB06ECA2" box="[1143,1247,1675,1699]" name="Germany" pageId="33" pageNumber="34">Germany</collectingCountry>
and 
<collectingCountry id="3A8A764E62409D33EB4DF91AFA45ECA2" box="[1302,1436,1675,1699]" name="Switzerland" pageId="33" pageNumber="34">Switzerland</collectingCountry>
(
<figureCitation id="DAA62A5B62409D33EEC4F93EFED8ECC6" box="[159,257,1711,1735]" captionStart="FIGURE 140–141" captionStartId="109.[151,249,1594,1617]" captionTargetBox="[151,1436,193,1573]" captionTargetId="figure-53@109.[151,1436,193,1573]" captionTargetPageId="109" captionText="FIGURE 140–141. Distribution of Palaearctic Leptomorphus. 140. L. forcipatus (black square), L. subforcipatus (blue triangle), L. talyshensis (black circle), L. walkeri (red star). 141. L. babai (blue triangle), L. panorpiformis (red star), L. quadrimaculatus (black square)." pageId="33" pageNumber="34">Fig. 140</figureCitation>
), 
<quantity id="85659B3B62409D33EF4EF93EFE7EECC6" box="[277,423,1711,1735]" metricMagnitude="2" metricUnit="m" metricValue="4.075" metricValueMax="7.8" metricValueMin="0.35" pageId="33" pageNumber="34" unit="m" value="407.5" valueMax="780.0" valueMin="35.0">35–780 masl</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0A87655562409D33EE9CF943FC99EDE6" pageId="33" pageNumber="34" type="discussion">
<paragraph id="422236DE62409D33EE9CF943FAA0ED7A" blockId="33.[151,1437,151,2024]" pageId="33" pageNumber="34">
<emphasis id="70E9EACC62409D33EE9CF943FEABECEA" bold="true" box="[199,370,1746,1771]" pageId="33" pageNumber="34">DISCUSSION</emphasis>
: This species was originally described as a subspecies of 
<taxonomicName id="859D4D5D62409D33EDA9F945FBBAECEA" box="[1010,1123,1747,1771]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="walkeri">
<emphasis id="70E9EACC62409D33EDA9F945FBBAECEA" box="[1010,1123,1747,1771]" italics="true" pageId="33" pageNumber="34">L. walkeri</emphasis>
</taxonomicName>
, and was later thought to be a junior synonym of 
<taxonomicName id="859D4D5D62409D33EFCEF969FDAFED0E" baseAuthorityName="Matsumura" baseAuthorityYear="1916" box="[405,630,1783,1807]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="70E9EACC62409D33EFCEF969FDAFED0E" box="[405,630,1783,1807]" italics="true" pageId="33" pageNumber="34">L. quadrimaculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="260C4B2F62409D33ECD2F966FCC2ED0E" author="Okada, I." box="[649,795,1783,1807]" pageId="33" pageNumber="34" pagination="99 - 103" refId="ref94924" refString="Okada, I. (1936) Einige Nematocera aus den Sud-Kurilen (Diptera). Insecta Matsumurana, 10, 99 - 103." type="journal article" year="1936">Okada 1936</bibRefCitation>
, 
<bibRefCitation id="260C4B2F62409D33ED77F966FC64ED0E" author="Matile, L." box="[812,957,1783,1807]" pageId="33" pageNumber="34" pagination="231 - 241" refId="ref94679" refString="Matile, L. (1988) Tribe Sciophilini. In: Soos, A. and Papp, L. Catalogue of Palaearctic Diptera: Volume 3: Ceratopogonidae - Mycetophilidae. Elsevier, Amsterdam, pp. 231 - 241." type="book chapter" year="1988">Matile 1988</bibRefCitation>
). 
<bibRefCitation id="260C4B2F62409D33ED8EF966FB22ED0E" author="Zaitzev, A. I. &amp; Sevcik J." box="[981,1275,1783,1807]" pageId="33" pageNumber="34" pagination="203 - 211" refId="ref96838" refString="Zaitzev, A. I. &amp; Sevcik J. (2002) A review of the Palaearctic species of the Leptomorphus quadrimaculatus (Matsumura) group (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientiarum Hungaricae, 48, 203 - 211." type="journal article" year="2002">Zaitzev &amp; Ševčík (2002)</bibRefCitation>
considered it significantly different from the latter, reinstated it and designated a 
<typeStatus id="9D26887C62409D33EDC0F88AFBD8ED32" box="[923,1025,1819,1843]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
from the single type specimen they found. 
<typeStatus id="9D26887C62409D33EEB3F8D1FE81ED59" box="[232,344,1856,1880]" pageId="33" pageNumber="34" type="lectotype">Lectotype</typeStatus>
information from 
<bibRefCitation id="260C4B2F62409D33EC76F8AEFC94ED56" author="Zaitzev, A. I. &amp; Sevcik J." box="[557,845,1855,1879]" pageId="33" pageNumber="34" pagination="203 - 211" refId="ref96838" refString="Zaitzev, A. I. &amp; Sevcik J. (2002) A review of the Palaearctic species of the Leptomorphus quadrimaculatus (Matsumura) group (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientiarum Hungaricae, 48, 203 - 211." type="journal article" year="2002">Zaitzev &amp; Ševčík (2002)</bibRefCitation>
: adult male, pinned, labelled “Árvaváralja, Kertész, 
<date id="3623101E62409D33EECCF8F2FEC8ED7A" box="[151,273,1891,1915]" pageId="33" pageNumber="34" value="1914-06-24">24.vi.1914</date>
, [underside of label] Collectio K. Landrock, K Czižek, D. Jacentkovský” [type depository: MMBC].
</paragraph>
<paragraph id="422236DE62409D33EE9CF816FC99EDE6" blockId="33.[151,1437,151,2024]" pageId="33" pageNumber="34">
As discussed below in the phylogeny section, the placement of 
<taxonomicName id="859D4D5D62409D33EDF3F816FB7CED9E" box="[936,1189,1927,1951]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="70E9EACC62409D33EDF3F816FB93ED9E" box="[936,1098,1927,1951]" italics="true" pageId="33" pageNumber="34">Leptomorphus</emphasis>
species
</taxonomicName>
in subgenera (
<bibRefCitation id="260C4B2F62409D33EB0AF816FF09EDC2" author="Matile, L." pageId="33" pageNumber="34" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977</bibRefCitation>
) is not supported by our phylogenetic results. This species is therefore removed from the subgenus 
<taxonomicName id="859D4D5D62409D33EECCF85EFEE0EDE6" authorityName=": Edwards" authorityYear="1925" box="[151,313,1999,2023]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="70E9EACC62409D33EECCF85EFEE0EDE6" box="[151,313,1999,2023]" italics="true" pageId="33" pageNumber="34">Leptomorphus</emphasis>
</taxonomicName>
and placed solely in the genus 
<emphasis id="70E9EACC62409D33ECCCF85EFC99EDE6" box="[663,832,1999,2023]" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="859D4D5D62409D33ECCCF85EFCE5EDE6" authorityName=": Edwards" authorityYear="1925" box="[663,828,1999,2023]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0A87655562439D30EE9CFF06FB6BEB1D" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="422236DE62439D30EE9CFF06FB6BEB1D" blockId="34.[151,1437,151,284]" pageId="34" pageNumber="35">
<collectionCode id="248CAE1B62439D30EE9CFF06FE86EAB1" box="[199,351,151,176]" pageId="34" pageNumber="35">
<emphasis id="70E9EACC62439D30EE9CFF06FE86EAB1" bold="true" box="[199,351,151,176]" pageId="34" pageNumber="35">MATERIAL</emphasis>
</collectionCode>
<materialsCitation id="F2F53C8362439D30EF36FF06FB77EB1D" collectingDate="1989-06-22" collectingDateMax="2007-07-11" collectingDateMin="1989-06-22" collectionCode="LEM, MTD, ZMUN" collectorName="J. Preisler &amp; P. Vonicka. &amp; Sachsische Schweiz N. P. &amp; U. Kallweit. &amp; E. Rindal &amp; L. Aarvik. &amp; J. Rohacek &amp; J. Sevcik." country="Czech Republic" county="Josefuv Dul" location="Polana Biosphere Reserve" municipality="Jedlovy" pageId="34" pageNumber="35" specimenCount="4" specimenCount-male="4" stateProvince="Jihocesky">
<emphasis id="70E9EACC62439D30EF36FF06FDD1EAB1" bold="true" box="[365,520,151,176]" pageId="34" pageNumber="35">EXAMINED</emphasis>
: 
<collectingCountry id="3A8A764E62439D30EC45FF06FCCBEAAE" box="[542,786,151,175]" name="Czech Republic" pageId="34" pageNumber="35">CZECH REPUBLIC</collectingCountry>
, 
<collectingRegion id="8059F83C62439D30ED7EFF06FC54EAAE" box="[805,909,151,175]" country="Czech Republic" name="Jihocesky" pageId="34" pageNumber="35">Bohemia</collectingRegion>
, 
<collectingCounty id="AB434E5262439D30EDC4FF06FBF6EAAE" box="[927,1071,151,175]" pageId="34" pageNumber="35">Josefuv Dul</collectingCounty>
, 
<collectingMunicipality id="A246ACA462439D30EA18FF06FB43EAAE" box="[1091,1178,151,175]" pageId="34" pageNumber="35">Jedlový</collectingMunicipality>
dul, 
<date id="3623101E62439D30EABAFF06FAAEEAAE" box="[1249,1399,151,175]" pageId="34" pageNumber="35" value="2005-09-01" valueMax="2005-09-22" valueMin="2005-09-01">
<collectingDate id="2667E9F662439D30EABAFF06FAAEEAAE" box="[1249,1399,151,175]" pageId="34" pageNumber="35" value="2005-09-01" valueMax="2005-09-22" valueMin="2005-09-01">1–22.ix.2005</collectingDate>
</date>
, 
<collectorName id="EF68530862439D30EBD1FF09FF35EAD5" pageId="34" pageNumber="35">J. Preisler</collectorName>
&amp; 
<collectorName id="EF68530862439D30EF4DFF2DFE4CEAD5" box="[278,405,188,212]" pageId="34" pageNumber="35">P. Vonička.</collectorName>
(
<specimenCount id="549BFD5762439D30EFF1FF2DFE12EAD2" box="[426,459,188,212]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
, 
<collectionCode id="248CAE1B62439D30EF87FF2DFDC5EAD5" box="[476,540,188,212]" pageId="34" pageNumber="35">LEM</collectionCode>
); 
<collectingCountry id="3A8A764E62439D30EC69FF2DFD66EAD5" box="[562,703,188,212]" name="Germany" pageId="34" pageNumber="35">GERMANY</collectingCountry>
, 
<collectingRegion id="8059F83C62439D30EC96FF2DFCFBEAD5" box="[717,802,188,212]" country="Germany" name="Sachsen" pageId="34" pageNumber="35">Saxony</collectingRegion>
, 
<collectorName id="EF68530862439D30ED6BFF2DFB97EAD2" box="[816,1102,188,212]" pageId="34" pageNumber="35">Sachsische Schweiz N.P.</collectorName>
, 
<date id="3623101E62439D30EA06FF2DFAD8EAD5" box="[1117,1281,188,212]" pageId="34" pageNumber="35" value="1989-06-22" valueMax="1989-06-23" valueMin="1989-06-22">
<collectingDate id="2667E9F662439D30EA06FF2DFAD8EAD5" box="[1117,1281,188,212]" pageId="34" pageNumber="35" value="1989-06-22" valueMax="1989-06-23" valueMin="1989-06-22">22–23.vi.1989</collectingDate>
</date>
, 
<collectorName id="EF68530862439D30EB54FF2DFA44EAD5" box="[1295,1437,188,212]" pageId="34" pageNumber="35">U. Kallweit.</collectorName>
(
<specimenCount id="549BFD5762439D30EEF9FF4EFF1AEAF6" box="[162,195,223,247]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
, 
<collectionCode id="248CAE1B62439D30EE83FF4EFEC2EAF6" box="[216,283,223,247]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="34" pageNumber="35">MTD</collectionCode>
); 
<collectingCountry id="3A8A764E62439D30EF6FFF4EFE72EAF6" box="[308,427,223,247]" name="Norway" pageId="34" pageNumber="35">NORWAY</collectingCountry>
, AK, As; Arungen, Syverud, 
<date id="3623101E62439D30ED4DFF4EFC01EAF6" box="[790,984,223,247]" pageId="34" pageNumber="35" value="2003-08-15" valueMax="2003-09-03" valueMin="2003-08-15">
<collectingDate id="2667E9F662439D30ED4DFF4EFC01EAF6" box="[790,984,223,247]" pageId="34" pageNumber="35" value="2003-08-15" valueMax="2003-09-03" valueMin="2003-08-15">15.viii–3.ix.2003</collectingDate>
</date>
, 
<collectorName id="EF68530862439D30EDB0FF71FB84EAF6" box="[1003,1117,223,247]" pageId="34" pageNumber="35">E. Rindal</collectorName>
, 
<collectorName id="EF68530862439D30EA2BFF71FB32EAF6" box="[1136,1259,223,247]" pageId="34" pageNumber="35">L. Aarvik.</collectorName>
(
<specimenCount id="549BFD5762439D30EB58FF4EFAFDEAF6" box="[1283,1316,223,247]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
, 
<collectionCode id="248CAE1B62439D30EB63FF4EFA49EAF6" box="[1336,1424,223,247]" pageId="34" pageNumber="35">ZMUN</collectionCode>
); 
<collectingCountry id="3A8A764E62439D30EECCFE95FEFCEB1D" box="[151,293,260,284]" name="Slovakia" pageId="34" pageNumber="35">SLOVAKIA</collectingCountry>
, 
<location id="4742600562439D30EF6BFE95FD80EB1A" LSID="urn:lsid:plazi:treatment:CA3487C862419D30EECCFCCBFB6BEB1D:4742600562439D30EF6BFE95FD80EB1A" box="[304,601,260,284]" country="Czech Republic" county="Josefuv Dul" municipality="Jedlovy" name="Polana Biosphere Reserve" pageId="34" pageNumber="35" stateProvince="Jihocesky">Polana Biosphere Reserve</location>
, 
<date id="3623101E62439D30EC3FFE95FCFDEB1D" box="[612,804,260,284]" pageId="34" pageNumber="35" value="2007-05-24" valueMax="2007-07-11" valueMin="2007-05-24">
<collectingDate id="2667E9F662439D30EC3FFE95FCFDEB1D" box="[612,804,260,284]" pageId="34" pageNumber="35" value="2007-05-24" valueMax="2007-07-11" valueMin="2007-05-24">24.v–11.vii.2007</collectingDate>
</date>
, 
<collectorName id="EF68530862439D30ED74FE95FC74EB1D" box="[815,941,260,284]" pageId="34" pageNumber="35">J. Roháček</collectorName>
, 
<collectorName id="EF68530862439D30EDE3FE95FBFCEB1D" box="[952,1061,260,284]" pageId="34" pageNumber="35">J. Ševčík.</collectorName>
(
<specimenCount id="549BFD5762439D30EA6CFE95FB81EB1A" box="[1079,1112,260,284]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
, 
<collectionCode id="248CAE1B62439D30EA3DFE95FB7FEB1D" box="[1126,1190,260,284]" pageId="34" pageNumber="35">LEM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>