317 lines
39 KiB
XML
317 lines
39 KiB
XML
<document id="17A8D61C1EEB0653B0910EF0585D6C64" ID-ISSN="11755334" ID-ZooBank="2412CB4F-4D29-4988-80C1-205D16767678" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629824082200" checkinUser="felipe" docAuthor="Borkent, Christopher J. & Wheeler, Terry A." docDate="2012" docId="CA3487C862209D50EECCFAE1FE5BEDE6" docLanguage="en" docName="zt03549p117.pdf" docOrigin="Zootaxa 3549" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leptomorphus titiwangsensis Borkent 2012, new species" docType="treatment" docUuid="6BB8F467-ABB9-4E29-8B00-A55C34407C43" docUuidSource="ZooBank" docVersion="4" lastPageNumber="67" masterDocId="360DFFB062619D12EE5BFF91FFD9EA01" masterDocTitle="Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549" masterLastPageNumber="117" masterPageNumber="1" pageNumber="66" updateTime="1699117448841" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0ADB8D98A8F0DE6E71BDB9CED5FE113D">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<mods:namePart id="B2915DFC38398632801352FB4CEB0B23">Borkent, Christopher J.</mods:namePart>
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<mods:namePart id="4284857899CD9BEAE5692B8C352C8831">Wheeler, Terry A.</mods:namePart>
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<mods:date id="049B74BA0BCFDAD705524CB2B8C3FDB6">2012</mods:date>
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<mods:number id="645FB43B186CB23CE1E34E217179E24D">3549</mods:number>
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<treatment id="CA3487C862209D50EECCFAE1FE5BEDE6" ID-DOI="http://doi.org/10.5281/zenodo.5258371" ID-Zenodo-Dep="5258371" LSID="urn:lsid:zoobank.org:act:6BB8F467-ABB9-4E29-8B00-A55C34407C43" httpUri="http://treatment.plazi.org/id/CA3487C862209D50EECCFAE1FE5BEDE6" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
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<subSubSection id="0A87655562209D53EECCFAE1FCE9EF8B" box="[151,816,1392,1418]" pageId="65" pageNumber="66" type="nomenclature">
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<paragraph id="422236DE62209D53EECCFAE1FCE9EF8B" blockId="65.[151,816,1392,1450]" box="[151,816,1392,1418]" pageId="65" pageNumber="66">
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<heading id="196A81B262209D53EECCFAE1FCE9EF8B" bold="true" box="[151,816,1392,1418]" fontSize="11" level="1" pageId="65" pageNumber="66" reason="1">
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35.
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<taxonomicName id="859D4D5D62209D53EE98FAE1FD4AEF8B" ID-CoL="8CJ4K" authority="Borkent" authorityName="Borkent" authorityYear="2012" box="[195,659,1392,1418]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="titiwangsensis" status="sp. nov.">
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<emphasis id="70E9EACC62209D53EE98FAE1FDFAEF8B" bold="true" box="[195,547,1392,1418]" italics="true" pageId="65" pageNumber="66">Leptomorphus titiwangsensis</emphasis>
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Borkent
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</taxonomicName>
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,
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<taxonomicNameLabel id="6BDA57B762209D53ECFBFAE1FCE9EF8B" box="[672,816,1392,1418]" pageId="65" pageNumber="66" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562209D53EECCFA03FDF3EFAB" box="[151,554,1426,1450]" pageId="65" pageNumber="66" type="description">
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<paragraph id="422236DE62209D53EECCFA03FDF3EFAB" blockId="65.[151,816,1392,1450]" box="[151,554,1426,1450]" pageId="65" pageNumber="66">
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(
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<figureCitation id="DAA62A5B62209D53EEC5FA03FEC0EFAB" box="[158,281,1426,1450]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="65" pageNumber="66">Figures 38</figureCitation>
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,
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<figureCitation id="DAA62A5B62209D53EF7EFA03FE9AEFAB" box="[293,323,1426,1450]" captionStart="FIGURES 70–77" captionStartId="94.[151,264,1209,1232]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure-136@94.[151,1436,193,1188]" captionTargetPageId="94" captionText="FIGURES 70–77. Leptomorphus wings. 70. L. quadrimaculatus, ventral. 71. L. stigmatus holotype male, dorsal. 72. L. subcaeruleus, ventral. 73. L. tabatius holotype male, dorsal. 74. L. tagbanua holotype male, ventral. 75. L. titiwangsensis paratype female, dorsal. 76. L. walkeri, ventral. 77. L. waodani holotype male, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="65" pageNumber="66">75</figureCitation>
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,
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<figureCitation id="DAA62A5B62209D53EF14FA03FEA2EFAB" box="[335,379,1426,1450]" captionStart="FIGURES 121–124" captionStartId="105.[151,264,1775,1798]" captionTargetBox="[176,1407,186,1696]" captionTargetId="figure-118@105.[161,1446,161,1741]" captionTargetPageId="105" captionText="FIGURES 121–124. Leptomorphus male genitalia. 121. L. talyshensis (reprinted with permission from Zaitzev & Ševčík 2002), A. gonocoxite and gonostylus, ventral. B. dorsal. 122. L. titiwangsensis, ventral. 123. L. walkeri, ventral. 124. L. waodani, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="65" pageNumber="66">122</figureCitation>
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,
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<figureCitation id="DAA62A5B62209D53EFDCFA03FE6AEFAB" box="[391,435,1426,1450]" captionStart="FIGURE 137–139" captionStartId="108.[151,249,1637,1660]" captionTargetBox="[151,1436,193,1616]" captionTargetId="figure-104@108.[151,1436,193,1616]" captionTargetPageId="108" captionText="FIGURE 137–139. Distribution of Oriental and Australasian Leptomorphus. 137. L. chaseni (red circle), L. ornatus (red triangle), L. tabatius (black square), L. tagbanua (blue star), L. titiwangsensis (black circle). 138. Species described by Papp & Ševčík 2011. L. alienus (red triangle), L. ascutellatus (red star), L. baramensis (black triangle), L. gunungmuluensis (purple square), L. longipes (blue circle), L. matilei (yellow star), L. papua (black square), L. utarensis (red circle). 139. Expanded view of northern Borneo showing distribution of L. baramensis, L. gunungmuluensis, and L. longipes." pageId="65" pageNumber="66">137</figureCitation>
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,
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<figureCitation id="DAA62A5B62209D53EFE4FA03FE32EFAB" box="[447,491,1426,1450]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values>50% are shown in bold below the branches." pageId="65" pageNumber="66">150</figureCitation>
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,
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<figureCitation id="DAA62A5B62209D53EFACFA03FDFBEFAB" box="[503,546,1426,1450]" captionStart="FIGURE 152" captionStartId="114.[151,250,1784,1807]" captionTargetBox="[162,1422,1059,1758]" captionTargetId="figure-112@114.[156,1428,1046,1779]" captionTargetPageId="114" captionText="FIGURE 152. One of three equally parsimonious trees showing the relationships between species in the L. ornatus species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="65" pageNumber="66">152</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562209D53EECCFA4BFDE8ED13" pageId="65" pageNumber="66" type="diagnosis">
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<paragraph id="422236DE62209D53EECCFA4BFC9FEC5E" blockId="65.[151,1437,1498,2027]" pageId="65" pageNumber="66">
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<emphasis id="70E9EACC62209D53EECCFA4BFEEDEFF2" bold="true" box="[151,308,1498,1523]" pageId="65" pageNumber="66">DIAGNOSIS</emphasis>
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: The only extant species of
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<taxonomicName id="859D4D5D62209D53ECEAFA4AFC8AEFF2" authorityName=": Edwards" authorityYear="1925" box="[689,851,1499,1523]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
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<emphasis id="70E9EACC62209D53ECEAFA4AFC8AEFF2" box="[689,851,1499,1523]" italics="true" pageId="65" pageNumber="66">Leptomorphus</emphasis>
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</taxonomicName>
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with the following combination of characters: laterotergite and anepisternum brown; wing with pale brown apical wing spot reaching to wing tip, pale medial wing spot absent; male genitalia with gonocoxite as long as tergite 9 with margins distinguishable, apex of gonostylus bifurcated, aedeagal apodemes present (
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<figureCitation id="DAA62A5B62209D53EC8CF9D6FCE0EC5E" box="[727,825,1607,1631]" captionStart="FIGURES 121–124" captionStartId="105.[151,264,1775,1798]" captionTargetBox="[176,1407,186,1696]" captionTargetId="figure-118@105.[161,1446,161,1741]" captionTargetPageId="105" captionText="FIGURES 121–124. Leptomorphus male genitalia. 121. L. talyshensis (reprinted with permission from Zaitzev & Ševčík 2002), A. gonocoxite and gonostylus, ventral. B. dorsal. 122. L. titiwangsensis, ventral. 123. L. walkeri, ventral. 124. L. waodani, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="65" pageNumber="66">Fig. 122</figureCitation>
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).
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</paragraph>
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<paragraph id="422236DE62209D53EE9CF9FBFDE8ED13" blockId="65.[151,1437,1498,2027]" pageId="65" pageNumber="66">
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This species can be distinguished from most other Oriental and eastern Palaearctic species (including those in
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<bibRefCitation id="260C4B2F62209D53EECCF91EFE50ECA6" author="Papp, L. & Sevcik, J." box="[151,393,1679,1703]" pageId="65" pageNumber="66" pagination="139 - 159" refId="ref95170" refString="Papp, L. & Sevcik, J. (2011) Eight new oriental and Australasian species of Leptomorphus (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientarum Hungaricae, 57, 139 - 159." type="journal article" year="2011">Papp & Ševčík 2011</bibRefCitation>
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) based on the yellow katepisternum and metepisternum (
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<figureCitation id="DAA62A5B62209D53EA7CF91EFBA4ECA6" box="[1063,1149,1679,1703]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="65" pageNumber="66">Fig. 38</figureCitation>
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, both brown in all other species in these regions except
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<emphasis id="70E9EACC62209D53EC5DF922FDB7ECCA" box="[518,622,1715,1739]" italics="true" pageId="65" pageNumber="66">
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<taxonomicName id="859D4D5D62209D53EC5DF922FDB0ECCA" authorityName=": Matile" authorityYear="1977" box="[518,617,1715,1739]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="babai">L. babai</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName id="859D4D5D62209D53ECF7F922FCF8ECCB" authorityName=": Matile" authorityYear="1977" box="[684,801,1715,1738]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="ornatus">
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<emphasis id="70E9EACC62209D53ECF7F922FCF8ECCB" box="[684,801,1715,1738]" italics="true" pageId="65" pageNumber="66">L. ornatus</emphasis>
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</taxonomicName>
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). It can be distinguished from
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<taxonomicName id="859D4D5D62209D53EAC2F922FB23ECCA" authorityName=": Matile" authorityYear="1977" box="[1177,1274,1715,1739]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="babai">
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<emphasis id="70E9EACC62209D53EAC2F922FB23ECCA" box="[1177,1274,1715,1739]" italics="true" pageId="65" pageNumber="66">L. babai</emphasis>
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</taxonomicName>
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by its yellow scutellum (brown in
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<taxonomicName id="859D4D5D62209D53EFD9F949FE38ECEE" authorityName=": Matile" authorityYear="1977" box="[386,481,1751,1775]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="babai">
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<emphasis id="70E9EACC62209D53EFD9F949FE38ECEE" box="[386,481,1751,1775]" italics="true" pageId="65" pageNumber="66">L. babai</emphasis>
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</taxonomicName>
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) and from
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<taxonomicName id="859D4D5D62209D53EC3AF949FD0DECEE" authorityName=": Matile" authorityYear="1977" box="[609,724,1752,1775]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="ornatus">
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<emphasis id="70E9EACC62209D53EC3AF949FD0DECEE" box="[609,724,1752,1775]" italics="true" pageId="65" pageNumber="66">L. ornatus</emphasis>
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</taxonomicName>
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by its occurrence only in peninsular
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<collectingCountry id="3A8A764E62209D53EA21F946FB39ECEE" box="[1146,1248,1751,1775]" name="Malaysia" pageId="65" pageNumber="66">Malaysia</collectingCountry>
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and presumably different male genitalic morphology.
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562209D50EE9CF88FFE1BEC35" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="description">
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<paragraph id="422236DE62209D50EE9CF88FFB65EFC9" blockId="65.[151,1437,1498,2027]" lastBlockId="66.[151,1437,151,2024]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
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<emphasis id="70E9EACC62209D53EE9CF88FFE50ED36" bold="true" box="[199,393,1822,1847]" pageId="65" pageNumber="66">DESCRIPTION</emphasis>
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:
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<emphasis id="70E9EACC62209D53EFFBF88EFE00ED36" box="[416,473,1823,1847]" italics="true" pageId="65" pageNumber="66">Male</emphasis>
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.
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<emphasis id="70E9EACC62209D53EFB4F88EFDF6ED36" bold="true" box="[495,559,1823,1847]" pageId="65" pageNumber="66">Head</emphasis>
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: yellow, somewhat dorsoventrally compressed in anterior view. Antenna brown; scape yellow, with brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae present; pedicel yellow, with 1–2 large bristles, few of setae on apicodorsal margin, none ventrally; flagellomere 1 with tapered base brown remainder brown; flagellomere 6 1.1X as long as broad. Clypeus ivory, dorsoventrally elongate oval; bristles on clypeus yellow, 6–8 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 2X as long as face. Face ivory; shape a subequal triangle, bare. Frons yellow; bare, frontal furrow running 3/4-full distance from dorsal margin towards ventral margin, frontal cleft running to lateral ocellus. Palpus yellow; segment 1 hidden behind eye, segments increasing in length, segment 5 ~1.5X length of segment 4, segment 3 spherical with apicolateral depressed patch of fine yellow setae partially encircled by strong dark setae. Labellum yellow. Eye with very few, short inter-ommatidial setulae scattered on surface. Occiput yellow with appressed, anteriorly directed setae. Ocelli in a straight line, space between ocelli 1–1.5X diameter of laterals, lateral ocelli 3X their own diameter from eye margin, ocellar triangle dark brown/black with electric blue green specks.
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<emphasis id="70E9EACC62239D50EEA8FE05FE95EBAD" bold="true" box="[243,332,404,428]" pageId="66" pageNumber="67">Thorax</emphasis>
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: Length
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(
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|
||
, n = 3). Mostly dark brown with anterolateral area yellow. Scutum dark brown/black with blue-green specks, pair of small yellow anterolateral spots; surface of scutum bare; acrostichal setae absent; dorsocentral setae present as fine setae on posterior two thirds; two thirds row of lateral setae present; patch of setae on scutum at wing base present. Scutellum yellow; with 2–4 large bristles and few small bristles. Prescutum brown. Mediotergite dark brown with 2 bristles on posterolateral corners, absent. Laterotergite brown; anterior margin of laterotergite abutting katepisternum. Anepimeron brown. Anepisternum brown. Katepisternum yellow. Antepronotum and proepisternum yellow. Margin of anterior and posterior spiracles yellow with yellow trichia. Metepisternum yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter with basal 1/3 of stem ivory, apical portion and knob dark brown.
|
||
<emphasis id="70E9EACC62239D50ED8BFD25FBD1E8CD" bold="true" box="[976,1032,692,716]" pageId="66" pageNumber="67">Legs</emphasis>
|
||
: principally yellow; hind coxa with brown spot along length of anterior surface; hind femur with apex and basal 1/4 light brown; extreme anteroapical corner light brown on forefemur, on other femora dark brown; tarsi light brown. Midfemur without apical spinelike process. Tibia with covering of brown macrotrichia, foretibia without comb of short setae along length of anteroventral surface, tibial spurs light brown, foretibial spur length 2X apical thickness of foretibia, midtibia with faint, dorsal, bare patch of even thickness for 4/5 of its length, placed basally, shortest midtibial spur 0.6X length of longest, longest midtibial spur 5X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 4X apical thickness of hind tibia. Foreleg first tarsomere 1.8X length of foretibia.
|
||
<emphasis id="70E9EACC62239D50EECCFC42FF01E9ED" bold="true" box="[151,216,979,1004]" pageId="66" pageNumber="67">Wing</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62239D50EEB3FC45FEE7E9ED" box="[232,318,980,1004]" captionStart="FIGURES 70–77" captionStartId="94.[151,264,1209,1232]" captionTargetBox="[151,1436,193,1188]" captionTargetId="figure-136@94.[151,1436,193,1188]" captionTargetPageId="94" captionText="FIGURES 70–77. Leptomorphus wings. 70. L. quadrimaculatus, ventral. 71. L. stigmatus holotype male, dorsal. 72. L. subcaeruleus, ventral. 73. L. tabatius holotype male, dorsal. 74. L. tagbanua holotype male, ventral. 75. L. titiwangsensis paratype female, dorsal. 76. L. walkeri, ventral. 77. L. waodani holotype male, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="66" pageNumber="67">Fig. 75</figureCitation>
|
||
): Length
|
||
<quantity id="85659B3B62239D50EFF0FC45FDDBE9EA" box="[427,514,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="66" pageNumber="67" unit="mm" value="4.5">4.5 mm</quantity>
|
||
(
|
||
<quantity id="85659B3B62239D50EC48FC45FD46E9EA" box="[531,671,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="66" pageNumber="67" unit="mm" value="4.45" valueMax="4.5" valueMin="4.4">4.4–4.5 mm</quantity>
|
||
, n = 3). Hyaline; apical macula light brown, reaching wing tip, but fading towards posterior margin; medial macula absent. Macrotrichia in all cells. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a few short setae. Vein sc-r present (sometimes very light), apical end joining R at 2X its own length prior to origin of Rs. R
|
||
<subScript id="DE19349B62239D50EA5CFBDEFBD6EE5C" attach="left" box="[1031,1039,1103,1117]" fontSize="6" pageId="66" pageNumber="67">4</subScript>
|
||
absent. R
|
||
<subScript id="DE19349B62239D50EA25FBDEFB5FEE5C" attach="left" box="[1150,1158,1103,1117]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
straight, slight posterior turn near tip. M
|
||
<subScript id="DE19349B62239D50EF12FBE7FE88EE85" attach="left" box="[329,337,1142,1156]" fontSize="6" pageId="66" pageNumber="67">1</subScript>
|
||
reaching apex before R
|
||
<subScript id="DE19349B62239D50EC04FBE7FDBEEE85" attach="left" box="[607,615,1142,1156]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
, apices of M veins fading before wing margin. M
|
||
<subScript id="DE19349B62239D50EAC0FBE7FB7AEE85" attach="left" box="[1179,1187,1142,1156]" fontSize="6" pageId="66" pageNumber="67">4</subScript>
|
||
-CuA fork arising just apically of origin of r-m. A
|
||
<subScript id="DE19349B62239D50EF87FB0CFE3DEEAA" attach="left" box="[476,484,1181,1195]" fontSize="6" pageId="66" pageNumber="67">2</subScript>
|
||
absent.
|
||
<emphasis id="70E9EACC62239D50EC13FB1CFD65EEA7" bold="true" box="[584,700,1165,1190]" pageId="66" pageNumber="67">Abdomen</emphasis>
|
||
: Tergites 3–5 with anterior 1/3 yellow, remainder dark brown. Tergite 8 smaller than all other abdominal sclerites, with 4–5 bristles on each apicolateral corner.
|
||
<emphasis id="70E9EACC62239D50EAB4FB25FA84EECC" bold="true" box="[1263,1373,1204,1229]" pageId="66" pageNumber="67">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62239D50EB35FB25FF1CEEF1" captionStart="FIGURES 121–124" captionStartId="105.[151,264,1775,1798]" captionTargetBox="[176,1407,186,1696]" captionTargetId="figure-118@105.[161,1446,161,1741]" captionTargetPageId="105" captionText="FIGURES 121–124. Leptomorphus male genitalia. 121. L. talyshensis (reprinted with permission from Zaitzev & Ševčík 2002), A. gonocoxite and gonostylus, ventral. B. dorsal. 122. L. titiwangsensis, ventral. 123. L. walkeri, ventral. 124. L. waodani, ventral. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="66" pageNumber="67">Fig. 122</figureCitation>
|
||
): yellow. Sternite 9 reduced to thin rectangular sclerite attached to aedeagus. Tergite 9 longer than wide, rounded margins tapering into two short rounded lobes and medial indentation. Gonocoxite placed basally on T9, medial margins not reaching median line, a ventromedial lobe covering base of gonostylus, bearing gonostylus halfway to apex. Gonostylus a single lobe tapering towards middle and then bifurcated into two points and bearing several basal setae, gonocoxite III separate from gonostylus dorsomedially. Aedeagus 2/3 length of gonocoxite, tapering towards apex (slight central swelling), apodemes 1/4 total length. Parameres as a broad lobe with rounded apex and medial bump, apodemes 1.8X length of parameres and strongly bent dorsally at base.
|
||
</paragraph>
|
||
<paragraph id="422236DE62239D50EE9CFA45FC6AEC11" blockId="66.[151,1437,151,2024]" pageId="66" pageNumber="67">
|
||
<emphasis id="70E9EACC62239D50EE9CFA45FE87EFED" box="[199,350,1492,1516]" italics="true" pageId="66" pageNumber="67">Female adult</emphasis>
|
||
. (
|
||
<figureCitation id="DAA62A5B62239D50EF23FA45FE15EFED" box="[376,460,1492,1516]" captionStart="FIGURES 38–41" captionStartId="91.[151,264,1941,1964]" captionTargetBox="[153,1434,258,1918]" captionTargetId="figure-40@91.[151,1436,255,1920]" captionTargetPageId="91" captionText="FIGURES 38–41. Leptomorphus habitus. 38. L. titiwangsensis, paratype female. 39. L. waodani, holotype male. 40. L. tagbanua, holotype male. 41. L. walkeri, male." pageId="66" pageNumber="67">Fig. 38</figureCitation>
|
||
) As for male, except as follows.
|
||
<emphasis id="70E9EACC62239D50ED0EFA45FC74EFED" bold="true" box="[853,941,1492,1516]" pageId="66" pageNumber="67">Thorax</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B62239D50EA42FA45FB59EFED" box="[1049,1152,1492,1516]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="66" pageNumber="67" unit="mm" value="1.29">1.29 mm</quantity>
|
||
(
|
||
<quantity id="85659B3B62239D50EAC8FA45FAE5EFED" box="[1171,1340,1492,1516]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" metricValueMax="1.31" metricValueMin="1.27" pageId="66" pageNumber="67" unit="mm" value="1.29" valueMax="1.31" valueMin="1.27">1.27–1.31 mm</quantity>
|
||
, n = 2).
|
||
<emphasis id="70E9EACC62239D50EECCFA69FF01EC10" bold="true" box="[151,216,1528,1553]" pageId="66" pageNumber="67">Wing</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B62239D50EF66FA69FE4BEC11" box="[317,402,1528,1552]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="66" pageNumber="67" unit="mm" value="5.6">5.6 mm</quantity>
|
||
(
|
||
<quantity id="85659B3B62239D50EFFAFA69FDF2EC11" box="[417,555,1528,1552]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.7" metricValueMin="5.4" pageId="66" pageNumber="67" unit="mm" value="5.55" valueMax="5.7" valueMin="5.4">5.4–5.7 mm</quantity>
|
||
, n = 2).
|
||
<emphasis id="70E9EACC62239D50ECD3FA69FD25EC10" bold="true" box="[648,764,1528,1553]" pageId="66" pageNumber="67">Abdomen</emphasis>
|
||
: Cercus yellow.
|
||
</paragraph>
|
||
<paragraph id="422236DE62239D50EE9CF98CFE1BEC35" blockId="66.[151,1437,151,2024]" box="[199,450,1564,1588]" pageId="66" pageNumber="67">
|
||
<emphasis id="70E9EACC62239D50EE9CF98CFEE7EC35" box="[199,318,1565,1588]" italics="true" pageId="66" pageNumber="67">Immatures</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562239D50EE9CF9D1FE16EC59" box="[199,463,1600,1625]" pageId="66" pageNumber="67" type="biology_ecology">
|
||
<paragraph id="422236DE62239D50EE9CF9D1FE16EC59" blockId="66.[151,1437,151,2024]" box="[199,463,1600,1625]" pageId="66" pageNumber="67">
|
||
<emphasis id="70E9EACC62239D50EE9CF9D1FE93EC58" bold="true" box="[199,330,1600,1625]" pageId="66" pageNumber="67">BIOLOGY</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562239D50EE9CF9F5FC17EC7D" box="[199,974,1636,1661]" pageId="66" pageNumber="67" type="distribution">
|
||
<paragraph id="422236DE62239D50EE9CF9F5FC17EC7D" blockId="66.[151,1437,151,2024]" box="[199,974,1636,1661]" pageId="66" pageNumber="67">
|
||
<emphasis id="70E9EACC62239D50EE9CF9F5FE4FEC7C" bold="true" box="[199,406,1636,1661]" pageId="66" pageNumber="67">DISTRIBUTION</emphasis>
|
||
: Peninsular
|
||
<collectingCountry id="3A8A764E62239D50EC79F9F5FD50EC7D" box="[546,649,1636,1660]" name="Malaysia" pageId="66" pageNumber="67">Malaysia</collectingCountry>
|
||
(
|
||
<figureCitation id="DAA62A5B62239D50ECC3F9F5FD20EC7D" box="[664,761,1636,1660]" captionStart="FIGURE 137–139" captionStartId="108.[151,249,1637,1660]" captionTargetBox="[151,1436,193,1616]" captionTargetId="figure-104@108.[151,1436,193,1616]" captionTargetPageId="108" captionText="FIGURE 137–139. Distribution of Oriental and Australasian Leptomorphus. 137. L. chaseni (red circle), L. ornatus (red triangle), L. tabatius (black square), L. tagbanua (blue star), L. titiwangsensis (black circle). 138. Species described by Papp & Ševčík 2011. L. alienus (red triangle), L. ascutellatus (red star), L. baramensis (black triangle), L. gunungmuluensis (purple square), L. longipes (blue circle), L. matilei (yellow star), L. papua (black square), L. utarensis (red circle). 139. Expanded view of northern Borneo showing distribution of L. baramensis, L. gunungmuluensis, and L. longipes." pageId="66" pageNumber="67">Fig. 137</figureCitation>
|
||
),
|
||
<quantity id="85659B3B62239D50ED56F9F5FC13EC7D" box="[781,970,1636,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.28" metricValueMin="1.22" pageId="66" pageNumber="67" unit="m" value="1250.0" valueMax="1280.0" valueMin="1220.0">1220–1280 masl</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562239D50EE9CF919FA4FECC4" pageId="66" pageNumber="67" type="discussion">
|
||
<paragraph id="422236DE62239D50EE9CF919FA4FECC4" blockId="66.[151,1437,151,2024]" pageId="66" pageNumber="67">
|
||
<emphasis id="70E9EACC62239D50EE9CF919FEABECA0" bold="true" box="[199,370,1672,1697]" pageId="66" pageNumber="67">DISCUSSION</emphasis>
|
||
: This species was compared to the descriptions and figures of the recently described
|
||
<collectingRegion id="8059F83C62239D50EB1BF919FA45ECA1" box="[1344,1436,1672,1696]" country="Republic of the Congo" name="Orientale" pageId="66" pageNumber="67">Oriental</collectingRegion>
|
||
and Australasian species (
|
||
<bibRefCitation id="260C4B2F62239D50EFECF93DFD7BECC5" author="Papp, L. & Sevcik, J." box="[439,674,1708,1732]" pageId="66" pageNumber="67" pagination="139 - 159" refId="ref95170" refString="Papp, L. & Sevcik, J. (2011) Eight new oriental and Australasian species of Leptomorphus (Diptera: Mycetophilidae). Acta Zoologica Academiae Scientarum Hungaricae, 57, 139 - 159." type="journal article" year="2011">Papp & Ševčík 2011</bibRefCitation>
|
||
), and is clearly different from all of them and not a junior synonym.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562239D50EE9DF941FE2DED0D" pageId="66" pageNumber="67" type="etymology">
|
||
<paragraph id="422236DE62239D50EE9DF941FE2DED0D" blockId="66.[151,1437,151,2024]" pageId="66" pageNumber="67">
|
||
<emphasis id="70E9EACC62239D50EE9DF941FE59ECE8" bold="true" box="[198,384,1744,1769]" pageId="66" pageNumber="67">ETYMOLOGY</emphasis>
|
||
:
|
||
<materialsCitation id="F2F53C8362239D50EFCBF941FE29ED0D" country="Malaysia" location="This" municipality="Titiwangsa mountain range" pageId="66" pageNumber="67" specimenCount="1">
|
||
<location id="4742600562239D50EFCBF941FE1BECE9" LSID="urn:lsid:plazi:treatment:CA3487C862209D50EECCFAE1FE5BEDE6:4742600562239D50EFCBF941FE1BECE9" box="[400,450,1744,1768]" country="Malaysia" municipality="Titiwangsa mountain range" name="This" pageId="66" pageNumber="67">This</location>
|
||
species is named for the
|
||
<collectingMunicipality id="A246ACA462239D50ECBEF941FBC1ECE8" box="[741,1048,1744,1769]" pageId="66" pageNumber="67">Titiwangsa mountain range</collectingMunicipality>
|
||
of peninsular
|
||
<collectingCountry id="3A8A764E62239D50EAE5F941FAFCECE9" box="[1214,1317,1744,1768]" name="Malaysia" pageId="66" pageNumber="67">Malaysia</collectingCountry>
|
||
where the
|
||
<typeStatus id="9D26887C62239D50EECCF964FF1EED0C" box="[151,199,1781,1805]" pageId="66" pageNumber="67">type</typeStatus>
|
||
specimens were collected
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562239D50EE9CF889FE5BEDE6" pageId="66" pageNumber="67" type="materials_examined">
|
||
<paragraph id="422236DE62239D50EE9CF889FCF1ED9E" blockId="66.[151,1437,151,2024]" pageId="66" pageNumber="67">
|
||
<materialsCitation id="F2F53C8362239D50EE9CF889FCF1ED9E" collectingDate="1932-05-20" collectionCode="MATERIAL, BMNH" collectorName="H. M. Pendlebury & Det. C. J. Borkent" country="Malaysia" determinerName="Det. C. J. Borkent" location="Fraser's Hill" municipality="Malay Penin" pageId="66" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<emphasis id="70E9EACC62239D50EE9CF889FDDCED30" bold="true" box="[199,517,1816,1841]" pageId="66" pageNumber="67">
|
||
<collectionCode id="248CAE1B62239D50EE9CF889FE86ED30" box="[199,351,1816,1841]" pageId="66" pageNumber="67">MATERIAL</collectionCode>
|
||
EXAMINED
|
||
</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC62239D50EC42F889FD50ED31" box="[537,649,1816,1840]" italics="true" pageId="66" pageNumber="67">
|
||
<typeStatus id="9D26887C62239D50EC42F889FD5AED31" box="[537,643,1816,1840]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
here designated, adult male, pinned on double mount minute with genitalia in plastic vial on main pin, labelled “
|
||
<collectingMunicipality id="A246ACA462239D50ECF8F8ADFCEDED55" box="[675,820,1852,1876]" pageId="66" pageNumber="67">Malay Penin</collectingMunicipality>
|
||
[
|
||
<collectingCountry id="3A8A764E62239D50ED1FF8ADFC39ED55" box="[836,992,1852,1876]" name="Malaysia" pageId="66" pageNumber="67">MALAYSIA</collectingCountry>
|
||
]:/
|
||
<collectingRegion id="8059F83C62239D50EDA2F8ADFB97ED55" box="[1017,1102,1852,1876]" country="Malaysia" name="Pahang" pageId="66" pageNumber="67">Pahang</collectingRegion>
|
||
, F.M.S./
|
||
<location id="4742600562239D50EAEEF8ADFA9BED55" LSID="urn:lsid:plazi:treatment:CA3487C862209D50EECCFAE1FE5BEDE6:4742600562239D50EAEEF8ADFA9BED55" box="[1205,1346,1852,1876]" country="Malaysia" municipality="Malay Penin" name="Fraser's Hill" pageId="66" pageNumber="67" stateProvince="Pahang">Fraser’s Hill</location>
|
||
/ 4200’/
|
||
<date id="3623101E62239D50EECCF8F2FE9AED7A" box="[151,323,1888,1915]" pageId="66" pageNumber="67" value="1932-05-20">
|
||
<collectingDate id="2667E9F662239D50EECCF8F2FE9AED7A" box="[151,323,1888,1915]" pageId="66" pageNumber="67" value="1932-05-20">
|
||
May 20
|
||
<superScript id="B5E89B9662239D50EEA8F8F1FF26ED6F" attach="left" box="[243,255,1888,1902]" fontSize="6" pageId="66" pageNumber="67">th</superScript>
|
||
1932
|
||
</collectingDate>
|
||
</date>
|
||
/ [underside of label]
|
||
<collectorName id="EF68530862239D50EC10F8F2FCCBED7A" box="[587,786,1891,1915]" pageId="66" pageNumber="67">H.M. Pendlebury</collectorName>
|
||
/ F.M.S./ Museums.;
|
||
<typeStatus id="9D26887C62239D50EA54F8F2FB7AED7A" box="[1039,1187,1891,1915]" pageId="66" pageNumber="67" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="549BFD5762239D50EAF4F8F2FB1DED7B" box="[1199,1220,1891,1914]" pageId="66" pageNumber="67" type="male">♂</specimenCount>
|
||
/
|
||
<emphasis id="70E9EACC62239D50EABEF8F2FA45ED7A" box="[1253,1436,1891,1915]" italics="true" pageId="66" pageNumber="67">
|
||
<taxonomicName id="859D4D5D62239D50EABEF8F2FA5EED7A" authorityName=": Edwards" authorityYear="1925" box="[1253,1415,1891,1915]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
|
||
/
|
||
</emphasis>
|
||
<taxonomicName id="859D4D5D62239D50EECCF819FE4FED9E" authority="Borkent" authorityName="Borkent" authorityYear="2012" box="[151,406,1927,1951]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="titiwangsensis">
|
||
<emphasis id="70E9EACC62239D50EECCF819FEEFED9E" box="[151,310,1928,1951]" italics="true" pageId="66" pageNumber="67">titiwangsensis</emphasis>
|
||
Borkent
|
||
</taxonomicName>
|
||
/
|
||
<determinerName id="B7FEDA1A62239D50EFF6F819FDB6ED9E" box="[429,623,1927,1951]" pageId="66" pageNumber="67">
|
||
<collectorName id="EF68530862239D50EFF6F819FDB6ED9E" box="[429,623,1927,1951]" pageId="66" pageNumber="67">Det. C.J. Borkent</collectorName>
|
||
</determinerName>
|
||
2012” [
|
||
<collectionCode id="248CAE1B62239D50EC93F819FCFBED9E" box="[712,802,1928,1951]" pageId="66" pageNumber="67">BMNH</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="422236DE62239D50EE9CF83DFE5BEDE6" blockId="66.[151,1437,151,2024]" pageId="66" pageNumber="67">
|
||
<emphasis id="70E9EACC62239D50EE9CF83DFE98EDC2" box="[199,321,1964,1987]" italics="true" pageId="66" pageNumber="67">
|
||
<typeStatus id="9D26887C62239D50EE9CF83DFEE5EDC2" box="[199,316,1964,1987]" pageId="66" pageNumber="67" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
labelled as for
|
||
<typeStatus id="9D26887C62239D50EFAFF83AFD8FEDC2" box="[500,598,1963,1987]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
except
|
||
<date id="3623101E62239D50ECEBF83AFCC0EDC2" box="[688,793,1963,1987]" pageId="66" pageNumber="67" value="1929-01-29">29.i.1929</date>
|
||
(
|
||
<specimenCount id="549BFD5762239D50ED76F83AFC97EDC2" box="[813,846,1963,1987]" pageId="66" pageNumber="67" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="549BFD5762239D50ED05F83AFC58EDC2" box="[862,897,1963,1987]" pageId="66" pageNumber="67" type="female">1♀</specimenCount>
|
||
);
|
||
<materialsCitation id="F2F53C8362239D50EDC1F83AFB91EDC2" box="[922,1096,1963,1987]" collectingDate="1932-05-29" isTailEnumeration="true" pageId="66" pageNumber="67" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<date id="3623101E62239D50EDC1F83AFBD1EDC2" box="[922,1032,1963,1987]" pageId="66" pageNumber="67" value="1932-05-29">
|
||
<collectingDate id="2667E9F662239D50EDC1F83AFBD1EDC2" box="[922,1032,1963,1987]" pageId="66" pageNumber="67" value="1932-05-29">29.v.1932</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="549BFD5762239D50EA47F83AFBE4EDC2" box="[1052,1085,1963,1987]" pageId="66" pageNumber="67" type="female">1♀</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<date id="3623101E62239D50EA0CF83AFB67EDC2" box="[1111,1214,1963,1987]" pageId="66" pageNumber="67" value="1926-08">viii.1926</date>
|
||
, B. Gater (
|
||
<specimenCount id="549BFD5762239D50EB65F83AFA86EDC2" box="[1342,1375,1963,1987]" pageId="66" pageNumber="67" type="male">1♂</specimenCount>
|
||
).
|
||
<materialsCitation id="F2F53C8362239D50EB23F83AFEA7EDE6" collectionCode="BMNH" pageId="66" pageNumber="67" specimenCount="1" typeStatus="paratype">
|
||
All
|
||
<typeStatus id="9D26887C62239D50EECCF841FEDBEDE9" box="[151,258,2000,2024]" pageId="66" pageNumber="67" type="paratype">paratypes</typeStatus>
|
||
in
|
||
<collectionCode id="248CAE1B62239D50EF7DF841FEA7EDE6" box="[294,382,2000,2023]" pageId="66" pageNumber="67">BMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |