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<document id="D8C8002F2A77F4C43400B888AC66DDE8" ID="10.11646/zootaxa.4178.2.1" ID-DOI="10.11646/zootaxa.4178.2.1" ID-GBIF-Dataset="0a00d470-cfa7-48f5-9fb3-c11fa79b5f35" ID-ISSN="1175-5326" ID-Zenodo-Dep="163174" ID-ZooBank="C533EE2A-5831-49A2-A4ED-2E7CD94EC663" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477378507121" checkinUser="plazi" docAuthor="Delgado-Blas, Víctor Hugo &amp; Díaz-Díaz, Oscar" docDate="2016" docId="CA1187ADCA44E859FF30A738DEA1FD12" docLanguage="en" docName="zootaxa.4178.2.1.pdf" docOrigin="Zootaxa 4178 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dispio bescanzae Delgado-Blas &amp; Díaz-Díaz, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="166" masterDocId="3628FFD5CA48E856FFA7A47BDD22FF96" masterDocTitle="Redescription of two species and five new species of Dispio Hartman, 1951 (Spionidae: Polychaeta) from the eastern Pacific Coast and Caribbean Sea, with a review of the genus" masterLastPageNumber="181" masterPageNumber="151" pageNumber="163" updateTime="1698390455347" updateUser="plazi">
<mods:mods id="5948ACEEDAEAAE7A9D8828962AC78DF9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="283F5A4BD1F0EAC9070DC8EF0F7AE323">
<mods:title id="C84DEC8E3DA463732E48224DFA80D27F">Redescription of two species and five new species of Dispio Hartman, 1951 (Spionidae: Polychaeta) from the eastern Pacific Coast and Caribbean Sea, with a review of the genus</mods:title>
</mods:titleInfo>
<mods:name id="D265503AB92166758226566FD7A8300C" type="personal">
<mods:role id="FD18712A8570D82553568B8BAB5DEB7A">
<mods:roleTerm id="1DB51130EDB334C2DC4B6DBE3EE00AB1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02CD413243B4FABB3DC2867E9F96DD8D">Delgado-Blas, Víctor Hugo</mods:namePart>
</mods:name>
<mods:name id="BCBEEBEFB9140D1F563C5A70705C3060" type="personal">
<mods:role id="0658D5E4D8A84C080BEE362C3EFE9520">
<mods:roleTerm id="04B6B2028AC8D6FA3A1552BF6B443566">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="82EC6F275EA32B7CF3B68302B03F01A2">Díaz-Díaz, Oscar</mods:namePart>
</mods:name>
<mods:typeOfResource id="231BAE8649E098673739EB190BBB7C1E">text</mods:typeOfResource>
<mods:relatedItem id="BC0B677B2F0C4CBC7F625BA29304D0BA" type="host">
<mods:titleInfo id="3F67197242B09EEBD797CA75BF696478">
<mods:title id="122790DFD60907E43A7B0AF9E01F8A50">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B5DC306725B7B36A0B435868AA7A94D0">
<mods:date id="FED5E3C389E375656B637C94549B0E85">2016</mods:date>
<mods:detail id="D8605DFCCCB56A41075B5DFD42067D3C" type="volume">
<mods:number id="C59608FB7B12868FB6D288440CC4B688">4178</mods:number>
</mods:detail>
<mods:detail id="E271189BBAFCE078F54CD25BA0FB762F" type="issue">
<mods:number id="263087B692108BB7CA1A9DF8342A9951">2</mods:number>
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<mods:start id="9DA2A07B6FC9D90769446AE1EF8CB53F">151</mods:start>
<mods:end id="F0560F41D2CD5DDA07A8D089DC74C505">181</mods:end>
</mods:extent>
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<mods:classification id="02A51E28C5D87AF37E0350E2E5C1FF9E">journal article</mods:classification>
<mods:identifier id="9DAE45E0F66BFD6B5635F9617E483D14" type="DOI">10.11646/zootaxa.4178.2.1</mods:identifier>
<mods:identifier id="8AC5A5F35B612B490476C9350C909B0C" type="GBIF-Dataset">0a00d470-cfa7-48f5-9fb3-c11fa79b5f35</mods:identifier>
<mods:identifier id="230AD0FE448793934BAAAAAB05171712" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="850C4F7653B0F7F0F2C60B5F9E7FB5C3" type="Zenodo-Dep">163174</mods:identifier>
<mods:identifier id="60C13DA02AFE90F44519D1179E1E45E0" type="ZooBank">C533EE2A-5831-49A2-A4ED-2E7CD94EC663</mods:identifier>
</mods:mods>
<treatment id="CA1187ADCA44E859FF30A738DEA1FD12" ID-DOI="http://doi.org/10.5281/zenodo.5661065" ID-GBIF-Taxon="124557970" ID-Zenodo-Dep="5661065" LSID="urn:lsid:plazi:treatment:CA1187ADCA44E859FF30A738DEA1FD12" httpUri="http://treatment.plazi.org/id/CA1187ADCA44E859FF30A738DEA1FD12" lastPageId="15" lastPageNumber="166" pageId="12" pageNumber="163">
<subSubSection id="0AA26530CA44E85AFF30A738DC19FCE9" pageId="12" pageNumber="163" type="nomenclature">
<paragraph id="420736BBCA44E85AFF30A738DCE9FCC8" blockId="12.[151,459,835,895]" box="[151,459,835,862]" pageId="12" pageNumber="163">
<heading id="194F81D7CA44E85AFF30A738DCE9FCC8" bold="true" box="[151,459,835,862]" fontSize="11" level="1" pageId="12" pageNumber="163" reason="1">
<emphasis id="70CCEAA9CA44E85AFF30A738DCE9FCC8" bold="true" box="[151,459,835,862]" pageId="12" pageNumber="163">
<taxonomicName id="85B84D38CA44E85AFF30A738DC44FCCB" ID-CoL="36VMR" box="[151,358,835,861]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="bescanzae" status="sp. nov.">
<emphasis id="70CCEAA9CA44E85AFF30A738DC44FCCB" bold="true" box="[151,358,835,861]" italics="true" pageId="12" pageNumber="163">Dispio bescanzae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BFF57D2CA44E85AFEC9A73FDCE9FCC8" box="[366,459,836,862]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="420736BBCA44E85AFF30A71DDC19FCE9" blockId="12.[151,459,835,895]" box="[151,315,870,895]" pageId="12" pageNumber="163">
(
<figureCitation id="DA832A3ECA44E85AFF39A71DDDDEFCE9" box="[158,252,870,895]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Figure 5</figureCitation>
AZ)
</paragraph>
</subSubSection>
<subSubSection id="0AA26530CA44E85AFF30A7D5DF2EFB12" pageId="12" pageNumber="163" type="materials_examined">
<paragraph id="420736BBCA44E85AFF30A7D5DF2EFB12" blockId="12.[151,1437,942,2020]" pageId="12" pageNumber="163">
<materialsCitation id="F2D03CE6CA44E85AFF30A7D5D8B5FB87" ID-GBIF-Occurrence="1324019885" collectionCode="ECOSUR, MOBR" collectorName="Oscar Diaz-Diaz &amp; March &amp; El Penon" country="Venezuela" county="Material" elevation="2" latitude="10.566112" location="Guiria" longLatPrecision="21" longitude="-62.739166" municipality="Atlantic" pageId="12" pageNumber="163" specimenCode="ECOSUR 0178, ECOSUR 0179" specimenCount="12" stateProvince="Sucre" typeStatus="holotype">
<emphasis id="70CCEAA9CA44E85AFF30A7D5DCA3FC51" bold="true" box="[151,385,942,967]" pageId="12" pageNumber="163">
<collectingCounty id="AB664E37CA44E85AFF30A7D5DDDDFC51" box="[151,255,942,967]" pageId="12" pageNumber="163">Material</collectingCounty>
examined.
</emphasis>
<collectingMunicipality id="A263ACC1CA44E85AFE2EA7D5DCC1FC51" box="[393,483,942,967]" pageId="12" pageNumber="163">Atlantic</collectingMunicipality>
coast of
<collectingCountry id="3AAF762BCA44E85AFDEBA7D5DFE0FC51" box="[588,706,942,967]" name="Venezuela" pageId="12" pageNumber="163">Venezuela</collectingCountry>
,
<location id="47676060CA44E85AFD68A7D5DE3BFC51" LSID="urn:lsid:plazi:treatment:CA1187ADCA44E859FF30A738DEA1FD12:47676060CA44E85AFD68A7D5DE3BFC51" box="[719,793,942,967]" country="Venezuela" county="Material" latitude="10.566112" longLatPrecision="21" longitude="-62.739166" municipality="Atlantic" name="Guiria" pageId="12" pageNumber="163" stateProvince="Sucre">Güiria</location>
,
<geoCoordinate id="278C507CCA44E85AFC82A7D5DE89FC51" box="[805,939,942,967]" direction="north" orientation="latitude" pageId="12" pageNumber="163" precision="15" value="10.566112">10º3358″N</geoCoordinate>
,
<geoCoordinate id="278C507CCA44E85AFC10A7D5D962FC51" box="[951,1088,942,967]" direction="west" orientation="longitude" pageId="12" pageNumber="163" precision="15" value="-62.739166">62º4421″W</geoCoordinate>
, shallow subtidal, soft bottom,
<elevation id="C995D188CA44E85AFF30A7AEDDDAFC7B" box="[151,248,981,1006]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="0.015">
<quantity id="85409B5ECA44E85AFF30A7AEDDDAFC7B" box="[151,248,981,1006]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="0.015">0.015 m</quantity>
</elevation>
2 PVC corer, coll.
<collectorName id="EF4D536DCA44E85AFE7AA7ADDF83FC78" box="[477,673,982,1006]" pageId="12" pageNumber="163">Oscar Díaz-Díaz</collectorName>
,
<collectorName id="EF4D536DCA44E85AFD14A7AEDFDDFC78" box="[691,767,981,1006]" pageId="12" pageNumber="163">March</collectorName>
, 2006,
<emphasis id="70CCEAA9CA44E85AFCFBA7AEDEE1FC78" bold="true" box="[860,963,981,1006]" pageId="12" pageNumber="163">
<typeStatus id="9D038819CA44E85AFCFBA7AEDEE1FC78" box="[860,963,981,1006]" pageId="12" pageNumber="163" type="holotype">holotype</typeStatus>
</emphasis>
(
<specimenCode id="121E9EC0CA44E85AFC70A7ADD9B3FC78" box="[983,1169,982,1006]" collectionCode="ECOSUR" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="12" pageNumber="163">
<collectionCode id="24A9AE7ECA44E85AFC70A7ADD965FC78" box="[983,1095,982,1006]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="12" pageNumber="163">ECOSUR</collectionCode>
0178
</specimenCode>
); same data for seven
<emphasis id="70CCEAA9CA44E85AFF30A782DC2CFB84" bold="true" box="[151,270,1017,1042]" pageId="12" pageNumber="163">
<typeStatus id="9D038819CA44E85AFF30A782DC2CFB84" box="[151,270,1017,1042]" pageId="12" pageNumber="163" type="paratype">paratypes</typeStatus>
</emphasis>
(
<specimenCode id="121E9EC0CA44E85AFE86A781DCF9FB84" box="[289,475,1018,1042]" collectionCode="ECOSUR" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="12" pageNumber="163">
<collectionCode id="24A9AE7ECA44E85AFE86A781DCB3FB84" box="[289,401,1018,1042]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="12" pageNumber="163">ECOSUR</collectionCode>
0179
</specimenCode>
). Non
<typeStatus id="9D038819CA44E85AFD97A781DF42FB84" box="[560,608,1018,1042]" pageId="12" pageNumber="163">type</typeStatus>
material:
<specimenCount id="54BEFD32CA44E85AFD79A781DE5EFB84" box="[734,892,1017,1042]" pageId="12" pageNumber="163" type="generic">12 specimens</specimenCount>
(
<collectionCode id="24A9AE7ECA44E85AFC37A781DEC6FB84" box="[912,996,1018,1042]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15853" name="Estaci<63>n de Investigaciones Marinas de Margarita, Fundaci<63>n La Salle de Ciencias Naturales" pageId="12" pageNumber="163">MOBR</collectionCode>
<date id="3606107BCA44E85AFC56A781D91FFB84" box="[1009,1085,1018,1042]" pageId="12" pageNumber="163" value="1270-01">I-1270</date>
) from
<collectorName id="EF4D536DCA44E85AFB32A782D826FB87" box="[1173,1284,1017,1042]" pageId="12" pageNumber="163">El Peñon</collectorName>
,
<collectingRegion id="807CF859CA44E85AFAB2A781D877FB84" box="[1301,1365,1018,1042]" country="Venezuela" name="Sucre" pageId="12" pageNumber="163">Sucre</collectingRegion>
state
</materialsCitation>
,
<materialsCitation id="F2D03CE6CA44E85AFF30A05BDC00FBC9" ID-GBIF-Occurrence="1324019882" collectorName="Oscar Diaz-Diaz &amp; March" country="Venezuela" elevation="5" latitude="10.450556" location="Venezuela" longLatPrecision="21" longitude="-64.08917" pageId="12" pageNumber="163" specimenCount="1" typeStatus="holotype">
<collectingCountry id="3AAF762BCA44E85AFF30A05BDC28FBAF" box="[151,266,1056,1081]" name="Venezuela" pageId="12" pageNumber="163">Venezuela</collectingCountry>
<geoCoordinate id="278C507CCA44E85AFEB3A05BDCBBFBAF" box="[276,409,1056,1081]" direction="north" orientation="latitude" pageId="12" pageNumber="163" precision="15" value="10.450556">10º2702″N</geoCoordinate>
,
<geoCoordinate id="278C507CCA44E85AFE00A05BDF13FBAF" box="[423,561,1056,1081]" direction="west" orientation="longitude" pageId="12" pageNumber="163" precision="15" value="-64.08917">64º0521″W</geoCoordinate>
, very fine sand
<quantity id="85409B5ECA44E85AFD49A05BDE12FBAE" box="[750,816,1056,1081]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="163" unit="m" value="0.5">
<elevation id="C995D188CA44E85AFD49A05BDE12FBAE" box="[750,816,1056,1081]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="163" unit="m" value="0.5">0.5 m</elevation>
</quantity>
of deep,
<quantity id="85409B5ECA44E85AFC06A05BDEDDFBAE" box="[929,1023,1056,1081]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="0.015">
<elevation id="C995D188CA44E85AFC06A05BDEDDFBAE" box="[929,1023,1056,1081]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="0.015">0.015 m</elevation>
</quantity>
2 PVC corer, coll.
<collectorName id="EF4D536DCA44E85AFB70A05BD8BAFBAE" box="[1239,1432,1056,1080]" pageId="12" pageNumber="163">Oscar Díaz-Díaz</collectorName>
,
<date id="3606107BCA44E85AFF30A03DDC00FBC9" box="[151,290,1094,1119]" pageId="12" pageNumber="163" value="2014-03">
<collectorName id="EF4D536DCA44E85AFF30A03DDDC2FBC9" box="[151,224,1094,1119]" pageId="12" pageNumber="163">March</collectorName>
2014
</date>
</materialsCitation>
; 21 specimens (CB OM-UC 105) from El Peñón, very fine sand
<quantity id="85409B5ECA44E85AFC52A03DD917FBC9" box="[1013,1077,1094,1119]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="163" unit="m" value="0.5">0.5 m</quantity>
deep,
<quantity id="85409B5ECA44E85AFBD9A03DD9FAFBC9" box="[1150,1240,1094,1119]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="0.015">0.015 m</quantity>
2 PVC corer, coll. Oscar Díaz-Díaz,
<date id="3606107BCA44E85AFEC7A010DF2AFB12" box="[352,520,1131,1156]" pageId="12" pageNumber="163" value="2015-02">February 2015</date>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA26530CA44E85BFF60A0F5DE40F9C6" lastPageId="13" lastPageNumber="164" pageId="12" pageNumber="163" type="description">
<paragraph id="420736BBCA44E85AFF60A0F5DC73FAAE" blockId="12.[151,1437,942,2020]" pageId="12" pageNumber="163">
<emphasis id="70CCEAA9CA44E85AFF60A0F5DC73FB31" bold="true" box="[199,337,1166,1191]" pageId="12" pageNumber="163">Description</emphasis>
.
<typeStatus id="9D038819CA44E85AFEC1A0F5DCECFB31" box="[358,462,1166,1191]" pageId="12" pageNumber="163" type="holotype">Holotype</typeStatus>
incomplete, anterior fragment with 77 chaetigers,
<quantity id="85409B5ECA44E85AFB95A0F5D9A5FB31" box="[1074,1159,1166,1191]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="163" unit="mm" value="26.0">26 mm</quantity>
long, 1.0 mm wide at chaetiger 15. Complete
<typeStatus id="9D038819CA44E85AFE17A0CFDF39FB5A" box="[432,539,1204,1228]" pageId="12" pageNumber="163" type="paratype">paratypes</typeStatus>
with 38188 chaetigers (longest specimen in five fragments) plus 13 final segments with no chaetae; 8.0
<quantity id="85409B5ECA44E85AFDAFA0ACDF79FB79" box="[520,603,1239,1263]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="12" pageNumber="163" unit="mm" value="49.0">49 mm</quantity>
long,
<quantity id="85409B5ECA44E85AFD0CA0ACDE15FB79" box="[683,823,1239,1263]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="12" pageNumber="163" unit="mm" value="1.0" valueMax="1.3" valueMin="0.7">0.71.3 mm</quantity>
wide at chaetiger 15; incomplete
<typeStatus id="9D038819CA44E85AFB6AA0A3D81AFB66" box="[1229,1336,1240,1264]" pageId="12" pageNumber="163" type="paratype">paratypes</typeStatus>
with an anterior fragment with 3689 chaetigers, 6.0
<quantity id="85409B5ECA44E85AFD0CA080DFDFFA85" box="[683,765,1275,1300]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="163" unit="mm" value="28.0">28 mm</quantity>
long, 0.71.0 mm wide at chaetiger 15. Color in alcohol yellowish white.
</paragraph>
<paragraph id="420736BBCA44E85AFF60A138D85AF9CE" blockId="12.[151,1437,942,2020]" pageId="12" pageNumber="163">
Prostomium hourglass-shaped, widest subterminally, bluntly pointed on anterior margin (
<figureCitation id="DA832A3ECA44E85AFAADA138D874FACA" box="[1290,1366,1347,1372]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
AB), posteriorly tapered, with a long, triangular, raised, caruncle (
<figureCitation id="DA832A3ECA44E85AFCE4A11CDEA7FA16" box="[835,901,1383,1408]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
AC) to end on posterior margin of chaetiger 1 (
<figureCitation id="DA832A3ECA44E85AFF38A1F0DDC6FA32" box="[159,228,1419,1444]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
AC). With two pairs of subdermal, black eyes, arranged in a straight line (
<figureCitation id="DA832A3ECA44E85AFBFEA1F0D9BDFA32" box="[1113,1183,1419,1444]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
A): one pair of small, lateral, kidney-shaped eyes and one pair of rounded, medial eyes, or arranged in a trapezoid, first pair rounded, and posterior pair reniform (
<figureCitation id="DA832A3ECA44E85AFE12A1A8DCDFFA7A" box="[437,509,1491,1516]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
B) (eyes not observed in
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). Peristomium long, collar-like, surrounding prostomium, separated from chaetiger 1, forming moderate lateral wings (
<figureCitation id="DA832A3ECA44E85AFC53A18CD91BF986" box="[1012,1081,1527,1552]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
AC). Proboscis not observed. Palps short (
<figureCitation id="DA832A3ECA44E85AFE86A260DC40F9A2" box="[289,354,1563,1588]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
B, C) extending to chaetigers 310 (chaetiger
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); palps with single band of transverse rows of cilia on one side of ventral ciliated groove. Palp sheaths large, smooth, fused to base of palps (
<figureCitation id="DA832A3ECA44E85AFAB4A244D874F9CE" box="[1299,1366,1599,1624]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
B).
</paragraph>
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Notopodial postchaetal lamellae of chaetigers 12 shifted dorsally and deeply serrated. Lamellae of chaetiger 1 bearing 4
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9 (
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) digitiform papillae along distal and middle margins (
<figureCitation id="DA832A3ECA44E85AFBEAA2FCD9B0F936" box="[1101,1170,1671,1696]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
AB, D), basal margin rounded (
<figureCitation id="DA832A3ECA44E85AFEA4A2D0DC64F952" box="[259,326,1707,1732]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
D). Notopodial lamellae of chaetiger 2 with 8
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11 (
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<typeStatus id="9D038819CA44E85AFC1EA2D0D93FF952" box="[953,1053,1707,1732]" pageId="12" pageNumber="163" type="holotype">holotype</typeStatus>
) digitiform papillae along margin (
<figureCitation id="DA832A3ECA44E85AFF38A2B4DDC3F97E" box="[159,225,1743,1768]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
A, E); lamellae of chaetigers 3
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6 with 6
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10 (
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6
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7 in
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) digitiform papillae (
<figureCitation id="DA832A3ECA44E85AFBDBA2B4D99CF97E" box="[1148,1214,1743,1768]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
A, F) along margin; lamellae of chaetiger 7 with 18 (
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) digitiform papillae along margin (
<figureCitation id="DA832A3ECA44E85AFBDFA288D9E2F89A" box="[1144,1216,1779,1804]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
A, G); lamellae of chaetigers 8
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10 with up to 5 digitiform papillae, although some specimens with lamellae on chaetigers 8/11
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37/44 (
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<typeStatus id="9D038819CA44E85AFEA5A340DC44F8C2" box="[258,358,1851,1876]" pageId="12" pageNumber="163" type="holotype">holotype</typeStatus>
) with ruffled margin (
<figureCitation id="DA832A3ECA44E85AFDCCA340DF92F8C2" box="[619,688,1851,1876]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
H); lamellae of chaetigers 37
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44 and subsequent chaetigers with pointed ventral border (
<figureCitation id="DA832A3ECA44E85AFE39A324DCFDF8EE" box="[414,479,1887,1912]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">Fig. 5</figureCitation>
HJ). Ventral and dorsal edges of notopodial and neuropodial lamellae overlapping or touching only on posterior chaetigers (
<figureCitation id="DA832A3ECA44E85AFDC2A3F8DF8EF80A" box="[613,684,1923,1948]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="12" pageNumber="163">
Fig.
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</figureCitation>
IJ, L). Notopodial prechaetal lamellae, small, subtriangular on chaetigers 1
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2 (Fig. DE), thereafter increasing in size, wider, continuing through anterior chaetigers and around chaetiger 43 becoming slender, long and triangular (Fig. I, J), thereafter diminishing in size through posterior chaetigers. Each segment with a pair of dorsal J-shaped double bands of cilia arranged obliquely with a transverse band of cilia between them (
<figureCitation id="DA832A3ECA45E85BFE41A4C7DF0EFF43" box="[486,556,188,213]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
K). Lateral organs between notopodial and neuropodial postchaetal lamellae visible but small on middle chaetigers (
<figureCitation id="DA832A3ECA45E85BFDF7A4A4DFB0FF6E" box="[592,658,223,248]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
L).
</paragraph>
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Neuropodial postchaetal lamellae of chaetigers 13 serrated and shifted dorsally. Neuropodial lamellae of chaetiger 1 with 1
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5 (
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<typeStatus id="9D038819CA45E85BFE7BA55CDF62FED6" box="[476,576,295,320]" pageId="13" pageNumber="164" type="holotype">holotype</typeStatus>
) digitiform papillae along margin (
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D), lamellae of chaetigers 2
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3 with 1
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5 (
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<typeStatus id="9D038819CA45E85BFE82A537DCABFEF3" box="[293,393,332,357]" pageId="13" pageNumber="164" type="holotype">holotype</typeStatus>
) digitiform papillae (
<figureCitation id="DA832A3ECA45E85BFDD8A537DFE0FEF3" box="[639,706,332,357]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
EF), lamellae of chaetigers 4
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28/39 rounded and smooth (
<figureCitation id="DA832A3ECA45E85BFAC9A537DD86FE1E" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
G) (one specimen with 2 digitiform papillae on right lamellae from chaetiger 4), later becoming wider on chaetigers 29
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40 (
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<typeStatus id="9D038819CA45E85BFE16A5EFDF37FE3B" box="[433,533,404,429]" pageId="13" pageNumber="164" type="holotype">holotype</typeStatus>
) forming a pointed upper border (
<figureCitation id="DA832A3ECA45E85BFC04A5EFDEC3FE3B" box="[931,993,404,429]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig 5</figureCitation>
HI), gradually diminishing in size on subsequent chaetigers (
<figureCitation id="DA832A3ECA45E85BFE39A5CCDCC1FE46" box="[414,483,439,464]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
J). Neuropodial prechaetal lamellae small, rounded and wide (
<figureCitation id="DA832A3ECA45E85BFB02A5CCD9CBFE46" box="[1189,1257,439,464]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
DF); larger in mid region but decreasing gradually in size on posterior chaetigers (
<figureCitation id="DA832A3ECA45E85BFC38A5A7DEC6FE63" box="[927,996,476,501]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
J). All neuropodial prechaetal lamellae not basally fused with neuropodial postchaetal lamellae.
</paragraph>
<paragraph id="420736BBCA45E85BFF60A65FDC00FC82" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Branchiae present from chaetiger 1 (
<figureCitation id="DA832A3ECA45E85BFDC7A65FDF81FDAB" box="[608,675,548,573]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
B, D) to end of body (
<figureCitation id="DA832A3ECA45E85BFC3CA65FDEFCFDAB" box="[923,990,548,573]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
J); branchiae tapered, elongate, smooth; fused along length of notopodial lamellae, branchial tips free, distally pointed on all anterior chaetigers and slightly longer than notopodial lamellae (
<figureCitation id="DA832A3ECA45E85BFDAFA617DF68FD13" box="[520,586,620,645]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
DF). Branchiae on middle and posterior chaetigers smaller than notopodial lamellae (
<figureCitation id="DA832A3ECA45E85BFEADA6F4DC73FD3E" box="[266,337,655,680]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">
Fig.
<date id="3606107BCA45E85BFEE4A6F4DC73FD3E" box="[323,337,655,680]" pageId="13" pageNumber="164">5</date>
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IJ). Each branchia with a band of cilia along inner edge. Accessory branchiae present from chaetigers 13
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30 (
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<typeStatus id="9D038819CA45E85BFE09A6CFDF36FD5B" box="[430,532,692,717]" pageId="13" pageNumber="164" type="holotype">holotype</typeStatus>
), initially with a single, short digitate lobe that arises from dorsolateral side of body behind notopodial base, with number of lobes increasing to seven in posterior chaetigers (
<figureCitation id="DA832A3ECA45E85BFB6AA6ACD82DFD66" box="[1229,1295,727,752]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
M) arranged in two rows.
</paragraph>
<paragraph id="420736BBCA45E85BFF60A764DEE1FB53" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Notochaetae of chaetiger 1 arranged in a dorsal tuft and a ventral fascicle; dorsal tuft with about 30 to 40 very long, slender (
<figureCitation id="DA832A3ECA45E85BFE99A73FDCA6FCCB" box="[318,388,836,861]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
A, C), alimbated, smooth capillaries (
<figureCitation id="DA832A3ECA45E85BFC9CA73FDEA2FCCB" box="[827,896,836,861]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
N) directed upwards (
<figureCitation id="DA832A3ECA45E85BFB27A73FD9E7FCCB" box="[1152,1221,836,861]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
A, C), longer than ventral fascicle; ventral fascicle arranged in two rows; anterior row comprised of stout, moderately reticulated, granulated, unilimbated capillaries (
<figureCitation id="DA832A3ECA45E85BFD93A7F7DF5BFC33" box="[564,633,908,933]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
O), and posterior row of slender, smooth, alimbated capillary chaetae (
<figureCitation id="DA832A3ECA45E85BFF38A7D4DDC0FC5E" box="[159,226,943,968]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
P), longer than chaetae of anterior row, and less numerous than those comprising dorsal tuft. Notochaetae of chaetiger 2 and subsequent chaetigers arranged in three groups; first group with 2 or 3 uppermost wider, smooth, alimbated, capillaries, diminishing gradually in size and number to an anterior row of stout moderate reticulated, granulated, unilimbated capillaries (
<figureCitation id="DA832A3ECA45E85BFD8BA067DF4DFBA3" box="[556,623,1052,1077]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
Q), and a posterior row of slender, very slightly granulated, unilimbated capillaries (
<figureCitation id="DA832A3ECA45E85BFEBCA044DC7CFBCE" box="[283,350,1087,1112]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
R), longer than those of anterior row. Middle chaetigers with same structure of anterior chaetae except for an anterior row of reticulated, long, slightly bilimbate, very pointed capillaries (
<figureCitation id="DA832A3ECA45E85BFB62A01FD82FFBEB" box="[1221,1293,1124,1149]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
S). Posterior chaetigers with dorsal tuft of slender, long, smooth, distally unilimbated capillaries (
<figureCitation id="DA832A3ECA45E85BFBECA0FCD9ACFB36" box="[1099,1166,1159,1184]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
T), and posterior row of slender, smooth, alimbated capillaries. Notopodial hooded hooks absent.
</paragraph>
<paragraph id="420736BBCA45E85BFF60A0B4DE5AF9BB" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Neurochaetae of chaetiger 1 similar to notochaetae of chaetiger 1, arranged in two rows: an anterior row of stout, moderate, reticulated, granulated, unilimbate capillaries (
<figureCitation id="DA832A3ECA45E85BFCCDA08FDE8DFA9B" box="[874,943,1268,1293]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
U), and a posterior row of shorter, slender, unilimbate, smooth capillary chaetae (
<figureCitation id="DA832A3ECA45E85BFDEEA16CDFACFAA6" box="[585,654,1303,1328]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">
Fig.
<date id="3606107BCA45E85BFD27A16CDFACFAA6" box="[640,654,1303,1328]" pageId="13" pageNumber="164">5</date>
</figureCitation>
V), capillaries of anterior row shorter than those of posterior row; in addition, a ventral tuft of 4
<emphasis id="70CCEAA9CA45E85BFE6BA147DCF8FAC3" bold="true" box="[460,474,1340,1365]" pageId="13" pageNumber="164"></emphasis>
5 shorter, slender, smooth, alimbated capillaries located in position of sabre chaetae. Chaetigers 29 similar to chaetiger 1 but with long, reticulated, slightly granulated, alimbated sabre chaetae; on chaetigers 10
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15 (
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<typeStatus id="9D038819CA45E85BFE16A1FFDF37FA0B" box="[433,533,1412,1437]" pageId="13" pageNumber="164" type="holotype">holotype</typeStatus>
) sabre chaetae longer, stouter, heavily reticulated (
<figureCitation id="DA832A3ECA45E85BFBC4A1FFD985FA0B" box="[1123,1191,1412,1437]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
W), 2
<emphasis id="70CCEAA9CA45E85BFB4CA1FFD9DBFA0B" bold="true" box="[1259,1273,1412,1437]" pageId="13" pageNumber="164"></emphasis>
3 per fascicle. Unidentate neuropodial hooded hooks (
<figureCitation id="DA832A3ECA45E85BFDF5A1DCDFB1FA56" box="[594,659,1447,1472]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">
Fig.
<date id="3606107BCA45E85BFD20A1DCDFB1FA56" box="[647,659,1447,1472]" pageId="13" pageNumber="164">5</date>
</figureCitation>
XY) (one specimen with one bidentate hook on a posterior fascicle) replacing anterior row of capillary neurochaetae from chaetigers 20
<emphasis id="70CCEAA9CA45E85BFC33A1B7DE80FA73" bold="true" box="[916,930,1484,1509]" pageId="13" pageNumber="164"></emphasis>
27, up to six per neuropodium, accompanied by a row of smooth, slender, unilimbated capillary chaetae (
<figureCitation id="DA832A3ECA45E85BFC91A194DE5AF99E" box="[822,888,1519,1544]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
Y). Hooded hooks distally entire, slightly curved and hood extended distally to, or slightly beyond tip (
<figureCitation id="DA832A3ECA45E85BFD49A26FDE12F9BB" box="[750,816,1556,1581]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">
Fig.
<date id="3606107BCA45E85BFC84A26FDE12F9BB" box="[803,816,1556,1581]" pageId="13" pageNumber="164">5</date>
</figureCitation>
XY).
</paragraph>
<paragraph id="420736BBCA45E85BFF60A24CDE40F9C6" blockId="13.[151,1437,151,2013]" box="[199,866,1591,1616]" pageId="13" pageNumber="164">
Pygidium with midventral flap and two long cirri (
<figureCitation id="DA832A3ECA45E85BFD59A24CDE62F9C6" box="[766,832,1591,1616]" captionStart="FIGURE 5" captionStartId="14.[151,250,1677,1699]" captionTargetBox="[218,1372,193,1660]" captionTargetId="figure@14.[218,1372,193,1660]" captionTargetPageId="14" captionText="FIGURE 5. Dispio bescanzae sp. nov. Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; D I 0.5 mm; K 0.5 mm; L 0.05; M W 0.045 mm; Y 0.05 mm." httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="13" pageNumber="164">Fig. 5</figureCitation>
Z).
</paragraph>
</subSubSection>
<subSubSection id="0AA26530CA45E859FF60A227DE14FE46" lastPageId="15" lastPageNumber="166" pageId="13" pageNumber="164" type="discussion">
<paragraph id="420736BBCA45E859FF60A227DC59FEF3" blockId="13.[151,1437,151,2013]" lastBlockId="15.[151,1436,151,645]" lastPageId="15" lastPageNumber="166" pageId="13" pageNumber="164">
<emphasis id="70CCEAA9CA45E85BFF60A227DC19F9E3" bold="true" box="[199,315,1628,1653]" pageId="13" pageNumber="164">Remarks.</emphasis>
<taxonomicName id="85B84D38CA45E85BFEE0A226DF33F9E2" box="[327,529,1628,1652]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="bescanzae" status="sp. nov.">
<emphasis id="70CCEAA9CA45E85BFEE0A226DF33F9E2" box="[327,529,1628,1652]" italics="true" pageId="13" pageNumber="164">Dispio bescanzae</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA45E85BFDB9A227DF5FF9E3" bold="true" box="[542,637,1628,1653]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="6BFF57D2CA45E85BFDB9A227DF5FF9E3" box="[542,637,1628,1653]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to
<taxonomicName id="85B84D38CA45E85BFCC4A226DECEF9E2" box="[867,1004,1629,1652]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="uncinata">
<emphasis id="70CCEAA9CA45E85BFCC4A226DECEF9E2" box="[867,1004,1629,1652]" italics="true" pageId="13" pageNumber="164">D. uncinata</emphasis>
</taxonomicName>
and
<taxonomicName id="85B84D38CA45E85BFB8AA226D9FBF9E2" box="[1069,1241,1629,1652]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="70CCEAA9CA45E85BFB8AA226D9FBF9E2" box="[1069,1241,1629,1652]" italics="true" pageId="13" pageNumber="164">D. panamensis</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA45E85BFB41A227D867F9E3" bold="true" box="[1254,1349,1628,1653]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="6BFF57D2CA45E85BFB41A227D867F9E3" box="[1254,1349,1628,1653]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="85B84D38CA45E85BFAF6A226DC28F90E" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="bescanzae" status="sp. nov.">
<emphasis id="70CCEAA9CA45E85BFAF6A226DC28F90E" italics="true" pageId="13" pageNumber="164">Dispio bescanzae</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA45E85BFEB4A204DC48F90E" bold="true" box="[275,362,1663,1688]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="6BFF57D2CA45E85BFEB4A204DC48F90E" box="[275,362,1663,1688]" pageId="13" pageNumber="164" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="85B84D38CA45E85BFE02A2FADF09F901" box="[421,555,1664,1688]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="uncinata">
<emphasis id="70CCEAA9CA45E85BFE02A2FADF09F901" box="[421,555,1664,1688]" italics="true" pageId="13" pageNumber="164">D. uncinata</emphasis>
</taxonomicName>
have branchiae of the same length and similarly shaped anterior postchaetal notopodial and neuropodial lamellae on the first three chaetigers, all serrated. However,
<taxonomicName id="85B84D38CA45E85BFBD1A2DED819F92A" box="[1142,1339,1700,1724]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="bescanzae" status="sp. nov.">
<emphasis id="70CCEAA9CA45E85BFBD1A2DED819F92A" box="[1142,1339,1700,1724]" italics="true" pageId="13" pageNumber="164">Dispio bescanzae</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA45E85BFAE5A2DFD8BEF92B" bold="true" box="[1346,1436,1700,1725]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="6BFF57D2CA45E85BFAE5A2DFD8BEF92B" box="[1346,1436,1700,1725]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the
<typeStatus id="9D038819CA45E85BFE4AA2B2DF3FF977" box="[493,541,1737,1761]" pageId="13" pageNumber="164">type</typeStatus>
material of
<taxonomicName id="85B84D38CA45E85BFD0FA2B2DE0FF949" box="[680,813,1736,1760]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="uncinata">
<emphasis id="70CCEAA9CA45E85BFD0FA2B2DE0FF949" box="[680,813,1736,1760]" italics="true" pageId="13" pageNumber="164">D. uncinata</emphasis>
</taxonomicName>
, in that the former has a hourglass-shaped prostomium without lateral emarginations, notopodial lamellae on chaetigers 4
<emphasis id="70CCEAA9CA45E85BFC31A297DE86F893" bold="true" box="[918,932,1772,1797]" pageId="13" pageNumber="164"></emphasis>
7/10 with digitiform papillae along margin, lamellae of chaetigers 8/11
<emphasis id="70CCEAA9CA45E85BFE69A374DCFEF8BE" bold="true" box="[462,476,1807,1832]" pageId="13" pageNumber="164"></emphasis>
37/44 with ruffled margin, lamellae of chaetigers 37
<emphasis id="70CCEAA9CA45E85BFB9BA374D968F8BE" bold="true" box="[1084,1098,1807,1832]" pageId="13" pageNumber="164"></emphasis>
44 and subsequent chaetigers with pointed ventral border, ventral and dorsal edges of notopodial and neuropodial lamellae overlapping or touching only on posterior chaetigers, neuropodial lamellae of chaetigers 4
<emphasis id="70CCEAA9CA45E85BFC4AA32CDED9F8E6" bold="true" box="[1005,1019,1879,1904]" pageId="13" pageNumber="164"></emphasis>
28/39 rounded and smooth, a pair of dorsal J-shaped double bands of cilia, anterior notopodial chaetae on posterior row with smooth, alimbated capillary chaetae, anterior neuropodial chaetae on anterior row with unilimbate capillaries, a posterior row of shorter, unilimbate, smooth capillary chaetae, hooded hooks slightly curved and sub-distally entire, the hoods open and extend distally or slightly beyond the tip of the hook.
<taxonomicName id="85B84D38CA47E859FCBDA4E3DEFCFF26" box="[794,990,152,176]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="bescanzae" status="sp. nov.">
<emphasis id="70CCEAA9CA47E859FCBDA4E3DEFCFF26" box="[794,990,152,176]" italics="true" pageId="15" pageNumber="166">Dispio bescanzae</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA47E859FC42A4ECD91CFF26" bold="true" box="[997,1086,151,176]" pageId="15" pageNumber="166">
<taxonomicNameLabel id="6BFF57D2CA47E859FC42A4ECD91CFF26" box="[997,1086,151,176]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="85B84D38CA47E859FB69A4E2D856FF39" box="[1230,1396,152,176]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="70CCEAA9CA47E859FB69A4E2D9C0FF26" box="[1230,1250,153,176]" italics="true" pageId="15" pageNumber="166">D</emphasis>
.
<emphasis id="70CCEAA9CA47E859FB57A4E3D856FF39" box="[1264,1396,152,175]" italics="true" pageId="15" pageNumber="166">panamensis</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA47E859FADDA4ECDDEAFF43" bold="true" pageId="15" pageNumber="166">
<taxonomicNameLabel id="6BFF57D2CA47E859FADDA4ECDDEAFF43" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having deeply serrated anterior notopodial and neuropodial lamellae with papillae, a short caruncle, peristomium of the same length and lateral wings. However,
<taxonomicName id="85B84D38CA47E859FCE5A49ADEF5FF6E" box="[834,983,224,248]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="bescanzae" status="sp. nov.">
<emphasis id="70CCEAA9CA47E859FCE5A49ADEF5FF6E" box="[834,983,224,248]" italics="true" pageId="15" pageNumber="166">D. bescanzae</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA47E859FC78A4A4D91AFF6E" bold="true" box="[991,1080,223,248]" pageId="15" pageNumber="166">
<taxonomicNameLabel id="6BFF57D2CA47E859FC78A4A4D91AFF6E" box="[991,1080,223,248]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="85B84D38CA47E859FB6AA49AD851FF61" box="[1229,1395,224,248]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="70CCEAA9CA47E859FB6AA49AD851FF61" box="[1229,1395,224,248]" italics="true" pageId="15" pageNumber="166">D. panamensis</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA47E859FADDA4A4DDEAFE8B" bold="true" pageId="15" pageNumber="166">
<taxonomicNameLabel id="6BFF57D2CA47E859FADDA4A4DDEAFE8B" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in that the former has a hourglass-shaped prostomium, bluntly pointed anteriorly, a long, triangular caruncle, branchial tips distally pointed on all chaetigers, and the notopodial and neuropodial lamellae overlapping on posterior chaetigers.
</paragraph>
<caption id="16C76633CA46E858FF30A2F6D984F862" httpUri="https://zenodo.org/record/163179/files/figure.png" pageId="14" pageNumber="165" targetBox="[218,1372,193,1660]" targetPageId="14">
<paragraph id="420736BBCA46E858FF30A2F6D984F862" blockId="14.[151,1436,1677,2036]" pageId="14" pageNumber="165">
<emphasis id="70CCEAA9CA46E858FF30A2F6DC36F935" bold="true" box="[151,276,1677,1699]" pageId="14" pageNumber="165">FIGURE 5.</emphasis>
<taxonomicName id="85B84D38CA46E858FEBBA2F5DCEDF935" box="[284,463,1677,1699]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="14" pageNumber="165" phylum="Annelida" rank="species" species="bescanzae" status="sp. nov.">
<emphasis id="70CCEAA9CA46E858FEBBA2F5DCEDF935" box="[284,463,1677,1699]" italics="true" pageId="14" pageNumber="165">Dispio bescanzae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BFF57D2CA46E858FE70A2F6DF0BF935" box="[471,553,1677,1699]" pageId="14" pageNumber="165" rank="species">
<emphasis id="70CCEAA9CA46E858FE70A2F6DF01F935" bold="true" box="[471,547,1677,1699]" pageId="14" pageNumber="165">sp. nov</emphasis>
.
</taxonomicNameLabel>
Holotype (Ecosur- 0178: B-C; paratype (Ecosur- 0179: A, D-Z): (A) anterior region, dorsal view, (B); anterior region, dorsal view; (C) same, lateral view; (D) parapodium of chaetiger 1, anterior view; (E) parapodium of chaetiger 2, anterior view; (F) parapodium of chaetiger 3, anterior view; (G); parapodium of chaetiger 7, anterior view (H) parapodium of chaetiger 35, anterior view; (I) parapodium of chaetiger 58, anterior view; (J) parapodium of chaetiger 90, anterior view; (K) mid-anterior segments, dorsal view; (L) lateral organs from middle chaetigers; (M) accessory branchiae; (N) notopodial chaetae from anterior row of chaetiger 1; (O) notopodial chaetae from ventral fascicle of chaetiger 1, anterior row; (P) notopodial chaetae from ventral fascicle of chaetiger 1, posterior row; (Q) notopodial chaetae from anterior row of chaetiger 2 and subsequent chaetigers; (R) notopodial chaetae from posterior row of chaetiger 2 and subsequent chaetigers; (S) uppermost notopodial chaetae from subsequent chaetigers; (T) lower notopodial chaetae from subsequent chaetigers; (U) notopodial chaetae from anterior row; (V) neuropodial chaetae from posterior row; (W) sabre chaetae; (X) unidentate neuropodial hooded hooks; (Y) neurochaetae bundle from chaetiger 20; (Z) pygidium, dorsal view. Lateral organ (LO). Scale bars: A, B, C, J, Z 0.5 mm; DI 0.5 mm; K 0.5 mm; L 0.05; MW 0.045 mm; Y 0.05 mm.
</paragraph>
</caption>
<paragraph id="420736BBCA47E859FF60A50BDE14FE46" blockId="15.[151,1436,151,645]" pageId="15" pageNumber="166">
In this species we observed one bidentate hooded hook on the posterior chaetiger of one adult specimen. See remarks about bidentate hooks in
<taxonomicName id="85B84D38CA47E859FDB0A5EEDFFBFE3A" box="[535,729,404,428]" class="Polychaeta" family="Spionidae" genus="Dispio" kingdom="Animalia" order="Spionida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="lenislamellata" status="sp. nov.">
<emphasis id="70CCEAA9CA47E859FDB0A5EEDFFBFE3A" box="[535,729,404,428]" italics="true" pageId="15" pageNumber="166">D. lenislamellata</emphasis>
</taxonomicName>
<emphasis id="70CCEAA9CA47E859FD45A5EFDE1EFE3B" bold="true" box="[738,828,404,429]" pageId="15" pageNumber="166">
<taxonomicNameLabel id="6BFF57D2CA47E859FD45A5EFDE1EFE3B" box="[738,828,404,429]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Further differences between this new species and the other species examined are provided in the key and
<tableCitation id="0F3A0300CA47E859FD7BA5CCDE0DFE46" box="[732,815,439,464]" captionStart="TABLE 1" captionStartId="23.[125,210,449,471]" captionText="TABLE 1. Some taxonomic characteristics of Dispio species. NoPoL = Notopođial postchaetal lamellae; AnNePoL = Anterior neuropođial postchaetal lamellae; NePoL = Neuropođial postchaetal lamellae; A. Serrateđ (No. papillae), B. Entire with ruffleđ margin, C. Entire smooth, D. With a notch, E. Appearance bilobeđ, “? ” đenotes absence of information." pageId="15" pageNumber="166">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA26530CA47E859FF60A5A7DE0EFD8E" pageId="15" pageNumber="166" type="etymology">
<paragraph id="420736BBCA47E859FF60A5A7DE0EFD8E" blockId="15.[151,1436,151,645]" pageId="15" pageNumber="166">
<emphasis id="70CCEAA9CA47E859FF60A5A7DC6CFE63" bold="true" box="[199,334,476,501]" pageId="15" pageNumber="166">Etymology.</emphasis>
This species is named after Díaz-Díazs mother-in-law, who recently passed away, and is dedicated to her and all the women in the Bescanza family.
</paragraph>
</subSubSection>
<subSubSection id="0AA26530CA47E859FF60A65FD80CFDF6" pageId="15" pageNumber="166" type="materials_examined">
<paragraph id="420736BBCA47E859FF60A65FDF1EFDAB" blockId="15.[151,1436,151,645]" box="[199,572,548,573]" pageId="15" pageNumber="166">
<materialsCitation id="F2D03CE6CA47E859FF60A65FDF1EFDAB" ID-GBIF-Occurrence="1324019892" box="[199,572,548,573]" country="Venezuela" location="Guiria" pageId="15" pageNumber="166" specimenCount="1" typeStatus="holotype">
<emphasis id="70CCEAA9CA47E859FF60A65FDC47FDAB" bold="true" box="[199,357,548,573]" pageId="15" pageNumber="166">
<typeStatus id="9D038819CA47E859FF60A65FDC20FDAB" box="[199,258,548,573]" pageId="15" pageNumber="166">Type</typeStatus>
locality.
</emphasis>
<location id="47676060CA47E859FECBA65FDC95FDAB" LSID="urn:lsid:plazi:treatment:CA1187ADCA44E859FF30A738DEA1FD12:47676060CA47E859FECBA65FDC95FDAB" box="[364,439,548,573]" country="Venezuela" name="Guiria" pageId="15" pageNumber="166">Güiria</location>
,
<collectingCountry id="3AAF762BCA47E859FE65A65FDF1BFDAB" box="[450,569,548,573]" name="Venezuela" pageId="15" pageNumber="166">Venezuela</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="420736BBCA47E859FF60A63CD80CFDF6" blockId="15.[151,1436,151,645]" box="[199,1326,583,608]" pageId="15" pageNumber="166">
<emphasis id="70CCEAA9CA47E859FF60A63CDC09FDF6" bold="true" box="[199,299,583,608]" pageId="15" pageNumber="166">Ecology.</emphasis>
Specimens were collected in shallow water (
<quantity id="85409B5ECA47E859FC83A63CDE47FDC9" box="[804,869,583,608]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="166" unit="m" value="1.5">1.5 m</quantity>
deep) in fine, medium and coarse sands.
</paragraph>
</subSubSection>
<subSubSection id="0AA26530CA47E859FF60A617DEA1FD12" box="[199,899,620,645]" pageId="15" pageNumber="166" type="description">
<paragraph id="420736BBCA47E859FF60A617DEA1FD12" blockId="15.[151,1436,151,645]" box="[199,899,620,645]" pageId="15" pageNumber="166">
<emphasis id="70CCEAA9CA47E859FF60A617DF26FD13" bold="true" box="[199,516,620,645]" pageId="15" pageNumber="166">Geographical distribution.</emphasis>
Venezuela (Güiria and El Peñón).
</paragraph>
</subSubSection>
</treatment>
</document>