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<document id="604457BB2597C7DB9225B9F08E65BD9E" ID-CLB-Dataset="21982" ID-DOI="10.5852/ejt.2014.89" ID-GBIF-Dataset="55dbf55a-f438-463d-b68a-8b48dba366ab" ID-ISSN="2118-9773" ID-Zenodo-Dep="3836768" ID-ZooBank="EAADCD7D-88AB-4F79-8B66-3197FD0BBBC7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590012191492" checkinUser="tatiana" docAuthor="Karner, Michael" docDate="2014" docId="AA291B19B82AFFCFFD837B18FEADFB5A" docLanguage="en" docName="ejt-2014-89.pdf.imf" docOrigin="European Journal of Taxonomy 89" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Psammoecus procerus Karner 2014, sp. nov." docType="treatment" docUuid="C44995A0-A3D4-4C4A-A80A-F45ED4CD4202" docUuidSource="ZooBank" docVersion="8" lastPageNumber="9" masterDocId="56106361B82FFFC7FFE27E1AFFB9FFE6" masterDocTitle="Three new species and new records of African Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="6" updateTime="1698840572497" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="218348750EBE63CDEA84F24FE5580833">Three new species and new records of African Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
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<mods:namePart id="3359241460CBDB4E953E9F10A38BFDC9">Karner, Michael</mods:namePart>
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<mods:title id="7AD9142A78941F78D9AC81D026A29C71">European Journal of Taxonomy</mods:title>
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<mods:part id="0EAB6C18993B6DE44A96B670F818D40F">
<mods:date id="3D424815DC44A31FB49A0DD64FD44EA5">2014</mods:date>
<mods:detail id="73C770F45CC273A5D91BC3AE189089B5" type="pubDate">
<mods:number id="24A6FF022CCBCB017B85030BFF07FFA2">2014-07-04</mods:number>
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<mods:number id="98FA8211BECBD768773C5D914E4218B9">89</mods:number>
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<mods:identifier id="69EE1E58523D156B55391ED4488B4440" type="DOI">10.5852/ejt.2014.89</mods:identifier>
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<treatment id="AA291B19B82AFFCFFD837B18FEADFB5A" ID-DOI="http://doi.org/10.5281/zenodo.3852006" ID-GBIF-Taxon="164243845" ID-Zenodo-Dep="3852006" LSID="urn:lsid:zoobank.org:act:C44995A0-A3D4-4C4A-A80A-F45ED4CD4202" httpUri="http://treatment.plazi.org/id/AA291B19B82AFFCFFD837B18FEADFB5A" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
<subSubSection id="6A9AF984B82AFFC2FD837B18FC6AFAFA" box="[609,979,1282,1308]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="223FAA0FB82AFFC2FD837B18FC6AFAFA" blockId="5.[357,1230,1282,1378]" box="[609,979,1282,1308]" pageId="5" pageNumber="6">
<heading id="79771D63B82AFFC2FD837B18FC6AFAFA" box="[609,979,1282,1308]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="E580D18CB82AFFC2FD837B18FCC8FAFA" authority="Karner, 2014" authorityName="Karner" authorityYear="2014" box="[609,881,1282,1308]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="procerus" status="sp. nov.">
<emphasis id="10F4761DB82AFFC2FD837B18FCC8FAFA" bold="true" box="[609,881,1282,1308]" italics="true" pageId="5" pageNumber="6">Psammoecus procerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BC7CB66B82AFFC2FC9A7B18FC6AFAFA" box="[888,979,1282,1308]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6A9AF984B82AFFC2FD397B3EFCE0FAD8" box="[731,857,1316,1342]" pageId="5" pageNumber="6" type="description">
<paragraph id="223FAA0FB82AFFC2FD397B3EFCE0FAD8" blockId="5.[357,1230,1282,1378]" box="[731,857,1316,1342]" pageId="5" pageNumber="6">
<figureCitation id="BABBB68AB82AFFC2FD397B3EFCE0FAD8" box="[731,857,1316,1342]" captionStart="Fig" captionStartId="6.[189,232,1963,1989]" captionTargetBox="[249,1342,271,1923]" captionTargetId="figure@6.[243,1342,265,1923]" captionTargetPageId="6" captionText="Fig. 4. Psammoecus procerus sp. nov., holotype. A. Habitus. B. Head. C. Pronotum. D. Right antenna. EG. Aedeagus (E = ventral, F = lateral, G = dorsal view). Scale lines: A = 1.0 mm, BG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836776" httpUri="https://zenodo.org/record/3836776/files/figure.png" pageId="5" pageNumber="6">Fig. 4AG</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="223FAA0FB82AFFC2FE877B5DFB77FA84" blockId="5.[357,1230,1282,1378]" box="[357,1230,1351,1378]" pageId="5" pageNumber="6">
<uri id="5611A60DB82AFFC2FE877B5DFB77FA84" box="[357,1230,1351,1378]" pageId="5" pageNumber="6">
urn:lsid:zoobank.org:act:
<uuid id="562690DAB82AFFC2FD637B5DFB77FA84" box="[641,1230,1351,1378]" pageId="5" pageNumber="6">C44995A0-A3D4-4C4A-A80A-F45ED4CD4202</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="6A9AF984B82AFFC2FF5F7B93FEA1F9DD" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="223FAA0FB82AFFC2FF5F7B93FE8FFA45" blockId="5.[189,310,1417,1443]" box="[189,310,1417,1443]" pageId="5" pageNumber="6">
<heading id="79771D63B82AFFC2FF5F7B93FE8FFA45" bold="true" box="[189,310,1417,1443]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="10F4761DB82AFFC2FF5F7B93FE8FFA45" bold="true" box="[189,310,1417,1443]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="223FAA0FB82AFFC2FF5F7BADFEA1F9DD" blockId="5.[189,1399,1463,1595]" pageId="5" pageNumber="6">
The following combination of character states is diagnostic for this species: lateral margins of pronotum with very small teeth; antennomeres slender; 1
<superScript id="D5F50747B82AFFC2FD167BC3FD46FA01" attach="left" box="[756,767,1497,1511]" fontSize="6" pageId="5" pageNumber="6">st</superScript>
antennomere 3 times as long as 2
<superScript id="D5F50747B82AFFC2FB417BC3FB0DFA01" attach="none" box="[1187,1204,1497,1511]" fontSize="6" pageId="5" pageNumber="6">nd</superScript>
; microsculpture on vertex and disk of pronotum distinct, reticulate; median lobe of aedeagus slender; parameres long, slender.
</paragraph>
</subSubSection>
<subSubSection id="6A9AF984B82AFFC2FF5F7878FA99F928" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="223FAA0FB82AFFC2FF5F7878FE70F99A" blockId="5.[189,457,1634,1660]" box="[189,457,1634,1660]" pageId="5" pageNumber="6">
<heading id="79771D63B82AFFC2FF5F7878FE70F99A" bold="true" box="[189,457,1634,1660]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="10F4761DB82AFFC2FF5F7878FE70F99A" bold="true" box="[189,457,1634,1660]" pageId="5" pageNumber="6">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="223FAA0FB82AFFC2FF5F788BFA99F928" blockId="5.[189,1399,1681,1743]" pageId="5" pageNumber="6">
This species is most similar to
<taxonomicName id="E580D18CB82AFFC2FDCD7888FCCBF94D" authority="Grouvelle, 1908" authorityName="Grouvelle" authorityYear="1908" box="[559,882,1681,1707]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="10F4761DB82AFFC2FDCD7888FD1EF94D" box="[559,679,1681,1707]" italics="true" pageId="5" pageNumber="6">P. grandis</emphasis>
Grouvelle, 1908
</taxonomicName>
; it can be distinguished by the shape of the antennomeres and male genitalia. It differs from all other African
<taxonomicName id="E580D18CB82AFFC2FC2378AFFBE3F928" authorityName="Latreille" authorityYear="1829" box="[961,1114,1717,1742]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="10F4761DB82AFFC2FC2378AFFBE3F928" box="[961,1114,1717,1742]" italics="true" pageId="5" pageNumber="6">Psammoecus</emphasis>
</taxonomicName>
by its large size.
</paragraph>
</subSubSection>
<subSubSection id="6A9AF984B82AFFC2FF5F78ECFCA0F8D8" box="[189,793,1782,1854]" pageId="5" pageNumber="6" type="etymology">
<paragraph id="223FAA0FB82AFFC2FF5F78ECFEFDF8F6" blockId="5.[189,324,1782,1808]" box="[189,324,1782,1808]" pageId="5" pageNumber="6">
<heading id="79771D63B82AFFC2FF5F78ECFEFDF8F6" bold="true" box="[189,324,1782,1808]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="10F4761DB82AFFC2FF5F78ECFEFDF8F6" bold="true" box="[189,324,1782,1808]" pageId="5" pageNumber="6">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="223FAA0FB82AFFC2FF5F793EFCA0F8D8" blockId="5.[189,793,1828,1854]" box="[189,793,1828,1854]" pageId="5" pageNumber="6">The specific epithet is the Latin word for “slender”.</paragraph>
</subSubSection>
<caption id="76FFFA87B829FFC1FF5F79B1FAECF80F" ID-DOI="http://doi.org/10.5281/zenodo.3836776" ID-Zenodo-Dep="3836776" httpUri="https://zenodo.org/record/3836776/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1963,1989]" targetBox="[249,1342,271,1923]" targetPageId="6">
<paragraph id="223FAA0FB829FFC1FF5F79B1FAECF80F" blockId="6.[189,1399,1963,2025]" pageId="6" pageNumber="7">
<emphasis id="10F4761DB829FFC1FF5F79B1FEB2F823" bold="true" box="[189,267,1963,1989]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
<taxonomicName id="E580D18CB829FFC1FEF079B6FDA3F823" authority="Karner, 2014" authorityName="Karner" authorityYear="2014" box="[274,538,1964,1989]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="procerus" status="sp. nov.">
<emphasis id="10F4761DB829FFC1FEF079B6FDA3F823" box="[274,538,1964,1989]" italics="true" pageId="6" pageNumber="7">Psammoecus procerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BC7CB66B829FFC1FDC079B1FD39F823" box="[546,640,1963,1989]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="10F4761DB829FFC1FCE079B1FCAEF823" bold="true" box="[770,791,1963,1989]" pageId="6" pageNumber="7">A</emphasis>
. Habitus.
<emphasis id="10F4761DB829FFC1FC7279B1FC1DF823" bold="true" box="[912,932,1963,1989]" pageId="6" pageNumber="7">B</emphasis>
. Head.
<emphasis id="10F4761DB829FFC1FBE279B1FBACF823" bold="true" box="[1024,1045,1963,1989]" pageId="6" pageNumber="7">C</emphasis>
. Pronotum.
<emphasis id="10F4761DB829FFC1FB4579B1FB05F823" bold="true" box="[1191,1212,1963,1989]" pageId="6" pageNumber="7">D</emphasis>
. Right antenna.
<emphasis id="10F4761DB829FFC1FF5F79D4FF4FF80E" bold="true" box="[189,246,1998,2024]" pageId="6" pageNumber="7">EG</emphasis>
. Aedeagus (E = ventral, F = lateral, G = dorsal view). Scale lines: A = 1.0 mm, BG = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="6A9AF984B828FFC0FF5F7F11FCF9FE19" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="223FAA0FB828FFC0FF5F7F11FE13FEC3" blockId="7.[189,426,267,293]" box="[189,426,267,293]" pageId="7" pageNumber="8">
<heading id="79771D63B828FFC0FF5F7F11FE13FEC3" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="10F4761DB828FFC0FF5F7F11FE13FEC3" bold="true" box="[189,426,267,293]" pageId="7" pageNumber="8">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="92E8A052B828FFC0FF087F20FDA5FE7C" ID-GBIF-Occurrence="3761955311" collectingDate="1999-06-07" collectingDateMax="1999-06-13" collectingDateMin="1999-06-07" collectionCode="BMNH" collectorName="A. Polaszek" country="Sao Tome and Principe" location="Bom Successo" municipality="Bom Successo" pageId="7" pageNumber="8" specimenCode="BMNH 2005-138" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="223FAA0FB828FFC0FF087F20FEE5FEB2" blockId="7.[189,1398,314,411]" box="[234,348,314,340]" pageId="7" pageNumber="8">
<emphasis id="10F4761DB828FFC0FF087F20FEE5FEB2" bold="true" box="[234,348,314,340]" pageId="7" pageNumber="8">
<typeStatus id="FD3B14ADB828FFC0FF087F20FEE5FEB2" box="[234,348,314,340]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="223FAA0FB828FFC0FF5F7F47FDA5FE7C" blockId="7.[189,1398,314,411]" pageId="7" pageNumber="8">
<specimenCount id="34866186B828FFC0FF5F7F47FF6AFE91" box="[189,211,349,375]" pageId="7" pageNumber="8" type="male"></specimenCount>
<collectingCountry id="5A97EA9FB828FFC0FF3D7F47FEC1FE91" box="[223,376,347,375]" name="Sao Tome and Principe" pageId="7" pageNumber="8">SAO TOMÉ</collectingCountry>
|
<collectingMunicipality id="C25B3075B828FFC0FE697F47FD80FE91" box="[395,569,349,375]" country="Sao Tome and Principe" name="Bom Successo" pageId="7" pageNumber="8">Bom Successo</collectingMunicipality>
, | ECOFAC compound |
<date id="563E8CCFB828FFC0FCBB7F47FC42FE91" box="[857,1019,349,375]" pageId="7" pageNumber="8" value="1999-06-07" valueMax="1999-06-13" valueMin="1999-06-07">
<collectingDate id="467A7527B828FFC0FCBB7F47FC42FE91" box="[857,1019,349,375]" pageId="7" pageNumber="8" value="1999-06-07" valueMax="1999-06-13" valueMin="1999-06-07">7.13.vi.1999</collectingDate>
</date>
[
<date id="563E8CCFB828FFC0FBE87F47FB70FE91" box="[1034,1225,348,375]" pageId="7" pageNumber="8" value="1999-06-07" valueMax="1999-06-13" valueMin="1999-06-07">713 Jun. 1999</date>
],
<collectorName id="8F75CFD9B828FFC0FB057F47FACFFE91" box="[1255,1398,349,375]" pageId="7" pageNumber="8">A. Polaszek</collectorName>
|
<specimenCode id="72260274B828FFC0FF287F9AFE24FE7D" box="[202,413,384,411]" pageId="7" pageNumber="8">BMNH 2005-138</specimenCode>
(
<collectionCode id="449132CAB828FFC0FE537F9AFDA9FE7C" box="[433,528,384,410]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="7" pageNumber="8">BMNH</collectionCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="92E8A052B828FFC0FF087FDBFCF9FE19" ID-GBIF-Occurrence="3761955322" collectingDate="1999-06-07" collectingDateMax="1999-06-13" collectingDateMin="1999-06-07" collectionCode="BMNH, MKF" collectorName="A. Polaszek" country="Sao Tome and Principe" location="Bom Successo" municipality="Bom Successo" pageId="7" pageNumber="8" specimenCode="BMNH 2005-138" specimenCount="4" specimenCount-female="4" specimenCount-male="1" typeStatus="paratype">
<paragraph id="223FAA0FB828FFC0FF087FDBFEDEFE3D" blockId="7.[189,832,449,511]" box="[234,359,449,475]" pageId="7" pageNumber="8">
<emphasis id="10F4761DB828FFC0FF087FDBFEDEFE3D" bold="true" box="[234,359,449,475]" pageId="7" pageNumber="8">
<typeStatus id="FD3B14ADB828FFC0FF087FDBFEDEFE3D" box="[234,359,449,475]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="223FAA0FB828FFC0FF5F7FFFFCF9FE19" blockId="7.[189,832,449,511]" box="[189,832,484,511]" pageId="7" pageNumber="8">
<specimenCount id="34866186B828FFC0FF5F7FFFFEB9FE19" box="[189,256,485,511]" pageId="7" pageNumber="8" type="female">4 ♀♀</specimenCount>
, with same data as holotype (3
<collectionCode id="449132CAB828FFC0FD907FFFFD74FE19" box="[626,717,485,511]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="7" pageNumber="8">BMNH</collectionCode>
, 1
<collectionCode id="449132CAB828FFC0FD0D7FFFFC8DFE19" box="[751,820,485,511]" pageId="7" pageNumber="8">MKF</collectionCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="6A9AF984B828FFCFFF5F7C3CFAA6FC94" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="223FAA0FB828FFC0FF5F7C3CFE58FDA6" blockId="7.[189,481,550,576]" box="[189,481,550,576]" pageId="7" pageNumber="8">
<heading id="79771D63B828FFC0FF5F7C3CFE58FDA6" bold="true" box="[189,481,550,576]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="10F4761DB828FFC0FF5F7C3CFE58FDA6" bold="true" box="[189,481,550,576]" pageId="7" pageNumber="8">
Description of
<typeStatus id="FD3B14ADB828FFC0FE977C3CFE58FDA6" box="[373,481,550,576]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="223FAA0FB828FFC0FF5F7C4FFA9EFD1D" blockId="7.[189,1398,596,764]" pageId="7" pageNumber="8">
BODY (as in
<figureCitation id="BABBB68AB828FFC0FEB37C4EFE09FD89" box="[337,432,596,623]" captionStart="Fig" captionStartId="6.[189,232,1963,1989]" captionTargetBox="[249,1342,271,1923]" captionTargetId="figure@6.[243,1342,265,1923]" captionTargetPageId="6" captionText="Fig. 4. Psammoecus procerus sp. nov., holotype. A. Habitus. B. Head. C. Pronotum. D. Right antenna. EG. Aedeagus (E = ventral, F = lateral, G = dorsal view). Scale lines: A = 1.0 mm, BG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836776" httpUri="https://zenodo.org/record/3836776/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
). Elongate, slender, total length
<quantity id="E57807EAB828FFC0FCD37C4EFC29FD89" box="[817,912,596,623]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="8" unit="mm" value="3.6">3.6 mm</quantity>
. Elytra testaceous; humeral swelling and lateral margin dark brown, the dark coloration extended to a broad macula near the middle; apical half of suture and apex dark brown. Head and pronotum reddish brown; lateral margins of pronotum a little darker; anterior and posterior margins of pronotum lighter brown. Antennae testaceous, apex of antennomere 6 and antennomeres 7-10 darkened, antennomere 11 yellowish white. Legs yellow.
</paragraph>
<paragraph id="223FAA0FB828FFC0FF5F7D39FC01FBD2" blockId="7.[189,1399,803,1076]" pageId="7" pageNumber="8">
HEAD (as in
<figureCitation id="BABBB68AB828FFC0FEA97D39FE1FFCDB" box="[331,422,803,829]" captionStart="Fig" captionStartId="6.[189,232,1963,1989]" captionTargetBox="[249,1342,271,1923]" captionTargetId="figure@6.[243,1342,265,1923]" captionTargetPageId="6" captionText="Fig. 4. Psammoecus procerus sp. nov., holotype. A. Habitus. B. Head. C. Pronotum. D. Right antenna. EG. Aedeagus (E = ventral, F = lateral, G = dorsal view). Scale lines: A = 1.0 mm, BG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836776" httpUri="https://zenodo.org/record/3836776/files/figure.png" pageId="7" pageNumber="8">Fig. 4B</figureCitation>
). Eyes moderately rounded, temples well defined and curved, moderately narrowed behind eyes, temple angle 44°; head width
<quantity id="E57807EAB828FFC0FD557D5CFC9AFC86" box="[695,803,838,864]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="7" pageNumber="8" unit="mm" value="0.78">0.78 mm</quantity>
, length
<quantity id="E57807EAB828FFC0FC9D7D5CFC52FC86" box="[895,1003,838,864]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="7" pageNumber="8" unit="mm" value="0.46">0.46 mm</quantity>
. Puncturation on vertex irregular; punctures almost as large as an eye facet diameter, sometimes contiguous, sometimes leaving spaces the size of their diameter. Pubescence composed of long, semi-erect setae, directed anteriorly; microsculpture on apical part of vertex distinct, reticulate, on basal part of vertex very shallow. Longitudinal impressions on vertex short, slightly curved, barely attaining eye level. Eyes
<quantity id="E57807EAB828FFC0FC417DC9FBB3FC0B" box="[931,1034,979,1005]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="8" unit="mm" value="0.23">0.23 mm</quantity>
long, distance of inner margins
<quantity id="E57807EAB828FFC0FF5F7DECFE91FBF6" box="[189,296,1014,1041]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="8" unit="mm" value="0.52">0.52 mm</quantity>
. Antennae slender, attaining almost middle of elytra,
<quantity id="E57807EAB828FFC0FC747DECFC46FBF6" box="[918,1023,1014,1040]" metricMagnitude="-3" metricUnit="m" metricValue="1.98" pageId="7" pageNumber="8" unit="mm" value="1.98">1.98 mm</quantity>
long; antennomere proportions: 3.0: 1.0: 1.3: 1.5: 1.7: 1.5: 1.4: 1.3: 1.1: 1.1: 2.1 (
<figureCitation id="BABBB68AB828FFC0FCA77A03FC13FBD2" box="[837,938,1049,1076]" captionStart="Fig" captionStartId="6.[189,232,1963,1989]" captionTargetBox="[249,1342,271,1923]" captionTargetId="figure@6.[243,1342,265,1923]" captionTargetPageId="6" captionText="Fig. 4. Psammoecus procerus sp. nov., holotype. A. Habitus. B. Head. C. Pronotum. D. Right antenna. EG. Aedeagus (E = ventral, F = lateral, G = dorsal view). Scale lines: A = 1.0 mm, BG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836776" httpUri="https://zenodo.org/record/3836776/files/figure.png" pageId="7" pageNumber="8">Fig. 4 D</figureCitation>
).
</paragraph>
<caption id="76FFFA87B828FFC0FF5F79B6FECCF80C" ID-DOI="http://doi.org/10.5281/zenodo.3836778" ID-Zenodo-Dep="3836778" httpUri="https://zenodo.org/record/3836778/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1964,1990]" targetBox="[191,1398,1217,1921]" targetPageId="7">
<paragraph id="223FAA0FB828FFC0FF5F79B6FECCF80C" blockId="7.[189,1399,1964,2026]" pageId="7" pageNumber="8">
<emphasis id="10F4761DB828FFC0FF5F79B6FEB2F821" bold="true" box="[189,267,1964,1991]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
<taxonomicName id="E580D18CB828FFC0FEF679B7FD62F820" authority="Grouvelle, 1908" authorityName="Grouvelle" authorityYear="1908" box="[276,731,1964,1990]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="10F4761DB828FFC0FEF679B7FDB6F820" box="[276,527,1964,1990]" italics="true" pageId="7" pageNumber="8">Psammoecus grandis</emphasis>
Grouvelle, 1908
</taxonomicName>
.
<emphasis id="10F4761DB828FFC0FD0579B6FD45F820" bold="true" box="[743,764,1964,1990]" pageId="7" pageNumber="8">A</emphasis>
. Right antenna.
<emphasis id="10F4761DB828FFC0FC2279B7FC6DF821" bold="true" box="[960,980,1965,1991]" pageId="7" pageNumber="8">B</emphasis>
. Median lobe.
<emphasis id="10F4761DB828FFC0FB6A79B6FB24F820" bold="true" box="[1160,1181,1964,1990]" pageId="7" pageNumber="8">C</emphasis>
. Parameres. Scale lines = 0.5 mm.
</paragraph>
</caption>
<paragraph id="223FAA0FB827FFCFFF5F7F16FB4EFE33" blockId="8.[189,1399,267,470]" pageId="8" pageNumber="9">
PRONOTUM (as in
<figureCitation id="BABBB68AB827FFCFFE697F11FE53FEC3" box="[395,490,267,294]" captionStart="Fig" captionStartId="6.[189,232,1963,1989]" captionTargetBox="[249,1342,271,1923]" captionTargetId="figure@6.[243,1342,265,1923]" captionTargetPageId="6" captionText="Fig. 4. Psammoecus procerus sp. nov., holotype. A. Habitus. B. Head. C. Pronotum. D. Right antenna. EG. Aedeagus (E = ventral, F = lateral, G = dorsal view). Scale lines: A = 1.0 mm, BG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836776" httpUri="https://zenodo.org/record/3836776/files/figure.png" pageId="8" pageNumber="9">Fig. 4C</figureCitation>
). About 1.1 times as wide as head; moderately wider than long; width
<quantity id="E57807EAB827FFCFFAA17F11FF57FEAF" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="9" unit="mm" value="0.86">0.86 mm</quantity>
, length
<quantity id="E57807EAB827FFCFFEAC7F35FE05FEAF" box="[334,444,303,329]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="8" pageNumber="9" unit="mm" value="0.73">0.73 mm</quantity>
. Area close to the posterior angles slightly depressed. Anterior angles separated from the disk by a slightly raised ridge; the small anterolateral space separated by that ridge without microsculpture, bearing 3 short teeth. Left lateral margin with 4, right margin with 5 very short teeth, posterior angle marked by a tooth of same size as lateral teeth. Puncture size on pronotal disk as on vertex, puncturation less dense. Pubescence as on vertex. Microsculpture distinct, reticulate.
</paragraph>
<paragraph id="223FAA0FB827FFCFFF5F7FE7FE8BFD45" blockId="8.[189,1399,509,675]" pageId="8" pageNumber="9">
ELYTRA. Long and slender, 3.3x as long as pronotum, widest just in front of middle, length
<quantity id="E57807EAB827FFCFFAE77FE7FACBFDF1" box="[1285,1394,509,535]" metricMagnitude="-3" metricUnit="m" metricValue="2.44" pageId="8" pageNumber="9" unit="mm" value="2.44">2.44 mm</quantity>
, combined width
<quantity id="E57807EAB827FFCFFE647C3AFE4AFDDC" box="[390,499,544,570]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="9" unit="mm" value="1.4">1.40 mm</quantity>
. Elytral disk with a flat depression between first and second third. Rows of punctures on disk about half as wide as interstices; pubescence composed of long, semierect setae; strial setae about 1/3 shorter than interstrial setae; microsculpture restricted to areas near origin of setae, reticulate.
</paragraph>
<paragraph id="223FAA0FB827FFCFFF5F7CD6FAA6FC94" blockId="8.[189,1399,715,883]" pageId="8" pageNumber="9">
MALE GENITALIA (as in
<figureCitation id="BABBB68AB827FFCFFE5C7CD1FD84FD03" box="[446,573,715,742]" captionStart="Fig" captionStartId="6.[189,232,1963,1989]" captionTargetBox="[249,1342,271,1923]" captionTargetId="figure@6.[243,1342,265,1923]" captionTargetPageId="6" captionText="Fig. 4. Psammoecus procerus sp. nov., holotype. A. Habitus. B. Head. C. Pronotum. D. Right antenna. EG. Aedeagus (E = ventral, F = lateral, G = dorsal view). Scale lines: A = 1.0 mm, BG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836776" httpUri="https://zenodo.org/record/3836776/files/figure.png" pageId="8" pageNumber="9">Fig. 4EG</figureCitation>
). Parameres in ventral view wide, elongate, lateral face parallel, curved medially towards the apex; in lateral view lancet-shaped, almost straight ventrally and curved dorsally, apical third of dorsal face with short setae, tip bent ventrally. Median lobe slender, evenly narrowed toward apex in dorsal view; in lateral view wide to about the middle, narrowed to a parallel-sided section of about 2/5 its length; apical 1/6 narrowed, dorsal face curved in an S-shape, tip bent ventrally.
</paragraph>
</subSubSection>
<subSubSection id="6A9AF984B827FFCFFF5F7D83FCE6FC04" box="[189,863,921,994]" pageId="8" pageNumber="9" type="discussion">
<paragraph id="223FAA0FB827FFCFFF5F7D83FE8AFC55" blockId="8.[189,307,921,947]" box="[189,307,921,947]" pageId="8" pageNumber="9">
<heading id="79771D63B827FFCFFF5F7D83FE8AFC55" bold="true" box="[189,307,921,947]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="10F4761DB827FFCFFF5F7D83FE8AFC55" bold="true" box="[189,307,921,947]" pageId="8" pageNumber="9">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="223FAA0FB827FFCFFF5F7DD2FCE6FC04" blockId="8.[189,863,968,994]" box="[189,863,968,994]" pageId="8" pageNumber="9">
The
<typeStatus id="FD3B14ADB827FFCFFF107DD2FEDBFC04" box="[242,354,968,994]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
vary in body length from
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.
</paragraph>
</subSubSection>
<subSubSection id="6A9AF984B827FFCFFF5F7A13FEADFB5A" pageId="8" pageNumber="9" type="discussion">
<paragraph id="223FAA0FB827FFCFFF5F7A13FE96FBC5" blockId="8.[189,303,1033,1059]" box="[189,303,1033,1059]" pageId="8" pageNumber="9">
<heading id="79771D63B827FFCFFF5F7A13FE96FBC5" bold="true" box="[189,303,1033,1059]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="10F4761DB827FFCFFF5F7A13FE96FBC5" bold="true" box="[189,303,1033,1059]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="223FAA0FB827FFCFFF5F7A22FEADFB5A" blockId="8.[189,1399,1080,1212]" pageId="8" pageNumber="9">
The general appearance and the unusual structure of the male genitalia make this species a very untypical member of the genus
<taxonomicName id="E580D18CB827FFCFFE5B7A46FDEBFB93" authorityName="Latreille" authorityYear="1829" box="[441,594,1116,1141]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="10F4761DB827FFCFFE5B7A46FDEBFB93" box="[441,594,1116,1141]" italics="true" pageId="8" pageNumber="9">Psammoecus</emphasis>
</taxonomicName>
. Its habitus closely resembles that of
<taxonomicName id="E580D18CB827FFCFFBEA7A46FAFCFB93" authority="Grouvelle, 1908" authorityName="Grouvelle" authorityYear="1908" box="[1032,1349,1115,1141]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="10F4761DB827FFCFFBEA7A46FBC7FB93" box="[1032,1150,1115,1141]" italics="true" pageId="8" pageNumber="9">P. grandis</emphasis>
Grouvelle, 1908
</taxonomicName>
, but the antennae as well as the male genitalia of
<taxonomicName id="E580D18CB827FFCFFD247A65FC85FB7E" authorityName="Grouvelle" authorityYear="1908" box="[710,828,1150,1176]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="10F4761DB827FFCFFD247A65FC85FB7E" box="[710,828,1150,1176]" italics="true" pageId="8" pageNumber="9">P. grandis</emphasis>
</taxonomicName>
have an exceedingly different structure (see
<figureCitation id="BABBB68AB827FFCFFAAA7A64FF73FB5A" captionStart="Fig" captionStartId="7.[189,232,1964,1990]" captionTargetBox="[191,1398,1217,1921]" captionTargetId="figure@7.[189,1398,1217,1921]" captionTargetPageId="7" captionText="Fig. 5. Psammoecus grandis Grouvelle, 1908. A. Right antenna. B. Median lobe. C. Parameres. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836778" httpUri="https://zenodo.org/record/3836778/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
AC).
</paragraph>
</subSubSection>
</treatment>
</document>