143 lines
12 KiB
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143 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZBK="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="AA18286887E2D3A015B963EA6471F26A" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus insularum Bistroem, Nilsson & Bergsten, 2015, sp. n." docType="treatment" docUuid="5126E56D-3F25-40E1-99BE-B851F1D84AA6" docUuidSource="ZooBank" docVersion="5" lastPageNumber="240" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="238" updateTime="1668162284477" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bistroem, Olof</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nilsson, Anders N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bergsten, Johannes</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>542</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>379</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-542-1</mods:identifier>
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<mods:identifier type="ZBK">026407877355425BAB10BF1674510F12</mods:identifier>
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<mods:identifier type="ZooBank">026407877355425BAB10BF1674510F12</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127862728" LSID="urn:lsid:zoobank.org:act:5126E56D-3F25-40E1-99BE-B851F1D84AA6" httpUri="http://treatment.plazi.org/id/AA18286887E2D3A015B963EA6471F26A" lastPageId="239" lastPageNumber="240" pageId="237" pageNumber="238">
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<subSubSection pageId="237" pageNumber="238" type="multiple">
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<paragraph pageId="237" pageNumber="238">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection pageId="237" pageNumber="238" type="nomenclature">
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<paragraph pageId="237" pageNumber="238">
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<taxonomicName LSID="http://zoobank.org/5126E56D-3F25-40E1-99BE-B851F1D84AA6" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus insularum" order="Coleoptera" pageId="237" pageNumber="238" phylum="Arthropoda" rank="species" species="insularum">Laccophilus insularum</taxonomicName>
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<taxonomicNameLabel pageId="237" pageNumber="238">sp. n.</taxonomicNameLabel>
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Figs 198-199, 369, 520-521, 578
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</paragraph>
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</subSubSection>
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<subSubSection pageId="237" pageNumber="238" type="type locality">
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<paragraph pageId="237" pageNumber="238">Type locality.</paragraph>
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<paragraph pageId="237" pageNumber="238">
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Madagascar: Ankarafantsika NP, Mahajanga, Boeny (
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-16.30271">S16.30271</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.80995">E046.80995</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="238" lastPageNumber="239" pageId="237" pageNumber="238" type="type material">
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<paragraph pageId="237" pageNumber="238">Type material</paragraph>
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<paragraph lastPageId="238" lastPageNumber="239" pageId="237" pageNumber="238">
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(35 exs.): Holotype: male: "Madagascar Mahajanga: Boeny Ankarafantsika NP, S16.30271, E046.80995, 75 m.a.o. 8.12. 2009 Water net, field# MAD09-39 Bergsten et al." (NHRS). - Paratypes: "Madagascar (95)25.ix. 2001 Ranohira (Fianarantsoa), right affl. of Riv. Ihazofotsy nr Isalo Ranch, stagnant areas/750 m
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<pageBreakToken pageId="238" pageNumber="239" start="start">asl</pageBreakToken>
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, 21.0°C 0.005 mS/cm/Gerecke & Goldschmidt collectors BMNH(E) 2004-46" (BMNH; habitus in Fig. 520); "W-Madagascar (19) Manindaray, W Sakahara 700-800 m asl., 30.01. 1995 G. Dunay & J.
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<normalizedToken originalValue="Janák">Janak</normalizedToken>
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coll." (1 ex. NMW); "S-Madagascar (14) Ambatoveve, Road Betioky-Beneloka 150 m a.s.l., 26.01. 1995 G. Dunay & J.
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<normalizedToken originalValue="Janák">Janak</normalizedToken>
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coll." (1 ex. NMW, 1 ex. MZH); "S-Madagascar (15) Ambialialika, Road Betioky-Beneloka 50 m a.s.l., 27.01. 1995 G. Dunay & J.
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<normalizedToken originalValue="Janák">Janak</normalizedToken>
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coll." (1 ex. NMW; habitus in Fig. 521); "BMNH(E) <745072> DNA voucher/MAD: FIAN: Isalo Source of Piscine Naturelle: Waterhole P41K: N: -22.553: E:45.368, 859 m, 12.V. 2006: Leg. Bergsten et al" (1 ex. MZH); same data but "<794195> DNA voucher" (1 ex. MZH); same data but "<745073> DNA voucher" (1 ex. NHRS); same data but "<745074> DNA voucher" (1 ex. NHRS); "DNA Voucher BMNH(E) <831021>MSL 402:E6/MAD. Isaky-Ivondro Ampasy (E2); rice paddies: P66:
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<geoCoordinate direction="north" orientation="latitude" precision="555" value="24.93">N-24.93</geoCoordinate>
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: E:46.863: 64 m, 08.IV. 2007 Leg. Isambert et al." (1 ex. NHRS); same data but "BMNH(E) <831023>MSL 402:E8" (1 ex. NHRS); Same data but "BMNH(E) <831022>MSL 402:E7" (1 ex. NHRS); "BMNH(E) <742297> DNA voucher/MAD: FIAN: Isalo: River R41D, N:-22.486: E:45.383: 723 m 11.V. 2006 Leg. Bergsten et al" (1 ex. NHRS); "RM: Betsiboka Bas. (PO123) Loc. Ambalambongo aff. de Betsiboka Riv.,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="47.008335">47°00'30"E</geoCoordinate>
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/
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-16.8">16°48'00"S</geoCoordinate>
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, 30.3. 1993 leg. ORSTOM Antananarivo" (1 ex. NHRS/NMW); "RM: Betsiboka Bas.(PO124) Loc. Ambohimanatrika Kamoro Riv.
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="47.168335">47°10'06"E</geoCoordinate>
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/
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-16.481945">16°28'55"S</geoCoordinate>
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, 1.4. 1993 leg. ORSTOM Antananarivo" (1 ex. NHRS/NMW); "Mahajanga: Melaky btw. Morafenobe-Ambohijanahary
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.19091">S18.19091</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="45.19986">E045.19986</geoCoordinate>
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, 290 m.a.o., 19.12. 2009 Water net, field#MAD09-74 Bergsten et al / NHRS-JLKB 0000000479" (1 ex. NHRS); same data but not vouchers (6 exs. NHRS); same data but "
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.1988">S18.19880</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="45.15651">E045.15651</geoCoordinate>
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, 313 m.a.o., MAD09-73 / NHRS-JLKB 0000000484" (1 ex. NHRS); same data as holotype (7 exs. NHRS); "Mahajanga: Boeny Ankarafantsika NP,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-16.30271">S16.30271</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.80995">E046.80995</geoCoordinate>
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, 75 m.a.o. 8.12. 2009 Water net, field# MAD09-39 Bergsten et al / NHRS-JLKB 0000000491" (1 ex. NHRS); same data but "29.11. 2009, MAD09-10 / NHRS-J LKB 0000000486 (1 ex. NHRS); Est, Sambava c. 20 m Janac & Moravec leg. / 3.3. 1996 Riv. Anovona env., lux, rizieres / bord de la foret" (1 ex. NMPC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="238" pageNumber="239" type="diagnosis">
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<paragraph pageId="238" pageNumber="239">Diagnosis.</paragraph>
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<paragraph pageId="238" pageNumber="239">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus insularum" order="Coleoptera" pageId="238" pageNumber="239" phylum="Arthropoda" rank="species" species="insularum">Laccophilus insularum</taxonomicName>
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is very closely related to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus livingstoni" order="Coleoptera" pageId="238" pageNumber="239" phylum="Arthropoda" rank="species" species="livingstoni">Laccophilus livingstoni</taxonomicName>
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, occurring in mainland of Africa. Clear differences in appearance of penis apex show that they are not conspecific, but represent good species (vide diagnosis of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus livingstoni" order="Coleoptera" pageId="238" pageNumber="239" phylum="Arthropoda" rank="species" species="livingstoni">Laccophilus livingstoni</taxonomicName>
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on p. 250).
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</paragraph>
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<paragraph pageId="238" pageNumber="239">
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Description (Only diagnostic differences from description of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus livingstoni" order="Coleoptera" pageId="238" pageNumber="239" phylum="Arthropoda" rank="species" species="livingstoni">Laccophilus livingstoni</taxonomicName>
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considered.): Body length 3.0-3.6 mm, width 1.7-2.0 mm. Dorsal, colour pattern uniform; pale spots on elytra may sometimes be in part reduced or expanded (Figs 520-521).
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</paragraph>
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<paragraph pageId="238" pageNumber="239">Head: Uniformly pale; posteriorly not darker.</paragraph>
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<paragraph pageId="238" pageNumber="239">Pronotum: Pale ferrugineous, basally with vague, darker area.</paragraph>
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<paragraph pageId="238" pageNumber="239">Elytra: Basal, pale markings variable in size; sometimes somewhat extended (Figs 520-521).</paragraph>
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<paragraph pageId="238" pageNumber="239">Ventral aspect: Apical ventrite (Fig. 198). Stridulatory apparatus distinct; file of ridges quite broad.</paragraph>
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<paragraph pageId="238" pageNumber="239">Male genitalia: Penis apex distinct; outline rounded and apex expanded both at external and internal edge (Fig. 369).</paragraph>
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<paragraph pageId="238" pageNumber="239">Female: Apical ventrite (Fig. 199).</paragraph>
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</subSubSection>
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<subSubSection pageId="239" pageNumber="240" type="etymology">
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<paragraph pageId="239" pageNumber="240">
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<pageBreakToken pageId="239" pageNumber="240" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="239" pageNumber="240">The name insularum is a Latin noun in genitive plural form meaning "belonging to the island". It refers to the fact that the new species is only known from Madagascar.</paragraph>
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</subSubSection>
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<subSubSection pageId="239" pageNumber="240" type="distribution">
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<paragraph pageId="239" pageNumber="240">Distribution.</paragraph>
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<paragraph pageId="239" pageNumber="240">Madagascar (Fig. 578).</paragraph>
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</subSubSection>
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<subSubSection pageId="239" pageNumber="240" type="collecting circumstances">
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<paragraph pageId="239" pageNumber="240">Collecting circumstances.</paragraph>
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<paragraph pageId="239" pageNumber="240">Insufficiently known. At least once collected in a river, in stagnant sites of the water body.</paragraph>
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</subSubSection>
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</treatment>
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</document> |